treatments-xml/data/03/A6/87/03A687B1630526566CCE73D2FE7865EF.xml

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<mods:title id="2484F45C0A43FC24D8777F4E7C1C1DAF">Alaskacladius gen. nov., (Diptera: Chironomidae), a unique new orthoclad from Alaska</mods:title>
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<mods:affiliation id="B6F9E94C57E1DD0721069EA5A42B5FF8">Department of Environmental and Life Sciences, Trent University, 1600 West Bank Drive, Peterborough, ON, Canada a. namayan @ taxanama. com, arminnamayandeh @ trentu. ca; https: // orcid. org / 0000 - 0003 - 2136 - 0497 &amp; Department of Physiology, School of Medicine, Wayne State University, Detroit, MI 48201, USA</mods:affiliation>
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. 
<materialsCitation id="3B673CFA63052652697570EFFA2E63CA" collectorName="B. Titaro" country="Canada" location="British Columbia" pageId="3" pageNumber="574" specimenCount="1" specimenCount-female="1" stateProvince="British Columbia" typeStatus="paratype">
<emphasis id="B97BEAB563052652697570EFFA2B63AE" box="[1324,1436,1695,1720]" italics="true" pageId="3" pageNumber="574">
<typeStatus id="54B4880563052652697570EFFA2163AE" box="[1324,1430,1695,1720]" pageId="3" pageNumber="574">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D09FD2E630526526CCE70B2FF4B63CA" box="[151,252,1730,1756]" pageId="3" pageNumber="574" type="female">1 female</specimenCount>
, 
<collectingCountry id="F3187637630526526D5070B2FED763CA" box="[265,352,1730,1756]" name="Canada" pageId="3" pageNumber="574">Canada</collectingCountry>
, 
<collectingRegion id="49CBF845630526526D3570B2FD8563CA" box="[364,562,1730,1756]" country="Canada" name="British Columbia" pageId="3" pageNumber="574">British Columbia</collectingRegion>
, Vancouver, Stanley Park, 49.301, -123.14; leg. 
<collectorName id="26FA537163052652680E70B3FB0863CA" box="[1111,1215,1730,1757]" pageId="3" pageNumber="574">B. Titaro</collectorName>
; deposited at CBG
</materialsCitation>
. 
<materialsCitation id="3B673CFA630526526CCE7097FDBE6232" collectorName="S. Querengesser" country="Canada" latitude="53.257" location="Haida Gwaii" longLatPrecision="64" longitude="-132.089" municipality="Queen Charlotte Islands" pageId="3" pageNumber="574" specimenCount="1" specimenCount-female="1" stateProvince="British Columbia" typeStatus="paratype">
<emphasis id="B97BEAB5630526526CCE7097FEB06216" box="[151,263,1767,1792]" italics="true" pageId="3" pageNumber="574">
<typeStatus id="54B48805630526526CCE7097FEB66216" box="[151,257,1767,1792]" pageId="3" pageNumber="574">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D09FD2E630526526D497096FEC06216" box="[272,375,1766,1792]" pageId="3" pageNumber="574" type="female">1 female</specimenCount>
, 
<collectingCountry id="F3187637630526526DDD7096FE6C6216" box="[388,475,1766,1792]" name="Canada" pageId="3" pageNumber="574">Canada</collectingCountry>
, 
<collectingRegion id="49CBF845630526526DB17096FD196216" box="[488,686,1766,1792]" country="Canada" name="British Columbia" pageId="3" pageNumber="574">British Columbia</collectingRegion>
, 
<collectingMunicipality id="6BD4ACDD630526526EE57096FC656216" box="[700,978,1766,1792]" pageId="3" pageNumber="574">Queen Charlotte Islands</collectingMunicipality>
, 
<location id="8ED0607C630526526F867096FBC46216" LSID="urn:lsid:plazi:treatment:03A687B1630526566CCE73D2FE7865EF:8ED0607C630526526F867096FBC46216" box="[991,1139,1766,1792]" country="Canada" latitude="53.257" longLatPrecision="64" longitude="-132.089" municipality="Queen Charlotte Islands" name="Haida Gwaii" pageId="3" pageNumber="574" stateProvince="British Columbia">Haida Gwaii</location>
; 
<geoCoordinate id="EE3B50606305265268D87096FB666217" box="[1153,1233,1766,1793]" degrees="53.257" direction="north" orientation="latitude" pageId="3" pageNumber="574" precision="55" value="53.257">53.257</geoCoordinate>
, 
<geoCoordinate id="EE3B50606305265268877096FAF16216" box="[1246,1350,1766,1792]" degrees="132.089" direction="west" orientation="longitude" pageId="3" pageNumber="574" precision="55" value="-132.089">-132.089</geoCoordinate>
; leg. 
<collectorName id="26FA53716305265269DF7096FE846232" pageId="3" pageNumber="574">S. Querengesser</collectorName>
; deposited at CBG
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315652C630526526C9E715EFD196287" pageId="3" pageNumber="574" type="etymology">
<paragraph id="8BB036A7630526526C9E715EFDAA627A" blockId="3.[151,1437,1586,1972]" pageId="3" pageNumber="574">
<emphasis id="B97BEAB5630526526C9E715EFEF9625E" bold="true" box="[199,334,1838,1864]" pageId="3" pageNumber="574">Etymology.</emphasis>
The new species is named after John Hudson, who has enormously contributed to our 
<taxonomicName id="4C0F4D246305265268A5715EFA2B625E" box="[1276,1436,1838,1864]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="574" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
collection from 
<collectingRegion id="49CBF845630526526D117122FE21627A" box="[328,406,1874,1900]" country="United States of America" name="Alaska" pageId="3" pageNumber="574">Alaska</collectingRegion>
since 1994.
</paragraph>
<paragraph id="8BB036A7630526526C9E7106FD196287" blockId="3.[151,1437,1586,1972]" box="[199,686,1910,1937]" pageId="3" pageNumber="574">
<emphasis id="B97BEAB5630526526C9E7106FE666286" bold="true" box="[199,465,1910,1936]" pageId="3" pageNumber="574">Diagnostic characters.</emphasis>
Same as the genus.
</paragraph>
</subSubSection>
<subSubSection id="C315652C630526566C9E71EAFE7865EF" lastPageId="7" lastPageNumber="578" pageId="3" pageNumber="574" type="description">
<paragraph id="8BB036A7630526526C9E71EAFEEE62A2" blockId="3.[151,1437,1586,1972]" box="[199,345,1946,1972]" pageId="3" pageNumber="574">
<emphasis id="B97BEAB5630526526C9E71EAFEEE62A2" bold="true" box="[199,345,1946,1972]" pageId="3" pageNumber="574">Description.</emphasis>
</paragraph>
<caption id="DF70662F630226556CCE7055FF6B6377" ID-DOI="http://doi.org/10.5281/zenodo.8243765" ID-Zenodo-Dep="8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="4" pageNumber="575" startId="4.[151,250,1573,1597]" targetBox="[161,1427,191,1536]" targetPageId="4" targetType="figure">
<paragraph id="8BB036A7630226556CCE7055FF6B6377" blockId="4.[151,1436,1573,1633]" pageId="4" pageNumber="575">
<emphasis id="B97BEAB5630226556CCE7055FEA5632B" bold="true" box="[151,274,1573,1597]" pageId="4" pageNumber="575">FIGURE 2.</emphasis>
<taxonomicName id="4C0F4D24630226556D417055FD9F632B" authority="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram, 2023" authorityName="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram" authorityYear="2023" box="[280,552,1573,1597]" class="Insecta" family="Chironomidae" genus="Alaskacladius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="575" phylum="Arthropoda" rank="species" species="johnhudsoni" status="gen. nov., sp. nov.">
<emphasis id="B97BEAB5630226556D417055FD9F632B" box="[280,552,1573,1597]" italics="true" pageId="4" pageNumber="575">Alaskacladius johnhudsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857CE630226556E777056FD52632B" box="[558,741,1574,1598]" pageId="4" pageNumber="575" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal.
</paragraph>
</caption>
<paragraph id="8BB036A7630226556C9E70C4FED763DB" blockId="4.[151,1436,1715,2030]" box="[199,352,1715,1742]" pageId="4" pageNumber="575">
<emphasis id="B97BEAB5630226556C9E70C4FED763DB" bold="true" box="[199,352,1715,1742]" pageId="4" pageNumber="575">Male (n = 7).</emphasis>
</paragraph>
<paragraph id="8BB036A7630226556C9E70A7FB1C63E7" blockId="4.[151,1436,1715,2030]" box="[199,1195,1751,1778]" pageId="4" pageNumber="575">
Total length 3.0–3.8, 
<quantity id="4CF79B42630226556DEF70A8FDB863E4" box="[438,527,1752,1778]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="575" unit="mm" value="3.3">3.3 mm</quantity>
. Wing 2.4–2.8, 
<quantity id="4CF79B42630226556EE670A7FCA263E4" box="[703,789,1751,1778]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="575" unit="mm" value="2.6">2.6 mm</quantity>
long and 0.58–0.66, 
<quantity id="4CF79B4263022655685B70A7FBD163E4" box="[1026,1126,1751,1778]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="575" unit="mm" value="0.62">0.62 mm</quantity>
wide.
</paragraph>
<paragraph id="8BB036A7630226556C9E708BFE6E622F" blockId="4.[151,1436,1715,2030]" pageId="4" pageNumber="575">
<emphasis id="B97BEAB5630226556C9E708BFD7C6203" box="[199,715,1787,1814]" italics="true" pageId="4" pageNumber="575">General coloration of the mounted specimens</emphasis>
. Head, thorax, halter, legs and tergites brown. Sternites slightly lighter. Wing greyish brown.
</paragraph>
<paragraph id="8BB036A7630226546C9E7133FD4165C3" blockId="4.[151,1436,1715,2030]" lastBlockId="5.[151,1436,150,501]" lastPageId="5" lastPageNumber="576" pageId="4" pageNumber="575">
<emphasis id="B97BEAB5630226556C9E7133FEB3624B" box="[199,260,1859,1885]" italics="true" pageId="4" pageNumber="575">Head</emphasis>
(
<figureCitation id="13342A22630226556D4B7133FEDE6248" box="[274,361,1859,1886]" captionStart="FIGURE 1" captionStartId="3.[151,250,1336,1360]" captionTargetBox="[155,1430,187,1303]" captionTargetId="figure-17@3.[151,1437,181,1312]" captionTargetPageId="3" captionText="FIGURE 1. Alaskacladius johnhudsoni gen. nov., sp. nov., male. A. head; B. thorax; C. wing; D. hypopygium." figureDoi="http://doi.org/10.5281/zenodo.8243763" httpUri="https://zenodo.org/record/8243763/files/figure.png" pageId="4" pageNumber="575">Fig. 1A</figureCitation>
). Antenna with 13 flagellomere, last flagellomere with 10 sensilla chaetica, 2 
<superScript id="7C7A9BEF6302265568E57133FB7B6247" attach="none" box="[1212,1228,1859,1873]" fontSize="6" pageId="4" pageNumber="575">nd</superScript>
–3 
<superScript id="7C7A9BEF6302265568B17133FB416247" attach="left" box="[1256,1270,1859,1873]" fontSize="6" pageId="4" pageNumber="575">rd</superScript>
segments each with 2 sensilla chaetica, groove starts at 4 
<superScript id="7C7A9BEF630226556E3C7117FDC56263" attach="left" box="[613,626,1895,1909]" fontSize="6" pageId="4" pageNumber="575">th</superScript>
segment, AR 1.5–1.6. Eyes bare, with very short dorsomedial extension, temporal setae 7–8 with 1–2 postoculars, 6 uniserial orbitals, and 4–5 outer verticals (
<figureCitation id="13342A2263022655681471FBFB1262B0" box="[1101,1189,1931,1958]" captionStart="FIGURE 1" captionStartId="3.[151,250,1336,1360]" captionTargetBox="[155,1430,187,1303]" captionTargetId="figure-17@3.[151,1437,181,1312]" captionTargetPageId="3" captionText="FIGURE 1. Alaskacladius johnhudsoni gen. nov., sp. nov., male. A. head; B. thorax; C. wing; D. hypopygium." figureDoi="http://doi.org/10.5281/zenodo.8243763" httpUri="https://zenodo.org/record/8243763/files/figure.png" pageId="4" pageNumber="575">Fig. 1A</figureCitation>
). Tentorium 187–202, 196 μm long, narrow and long apically, base slightly expanded, large tentorial pit close to apex (
<figureCitation id="13342A2263022655688B71DFFA9A62DC" box="[1234,1325,1967,1994]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="4" pageNumber="575">Fig. 2A</figureCitation>
). Clypeus nearly squared, 67–100, 88 μm long and 98–131, 112 μm wide, bearing 8 setae, setae 254–355 μm long. Palpal segments lengths (in μm): 73–99, 82; 52–79, 65; 114–169, 145; 109–135,121; 145–193, 163. Third palpomere with 1 sensilla clavata medially, 4 laterally, and 3 ventrally.
</paragraph>
<paragraph id="8BB036A7630326546C9E76AFFC37640B" blockId="5.[151,1436,150,501]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E76AFFEAF65EF" box="[199,280,223,249]" italics="true" pageId="5" pageNumber="576">Thorax</emphasis>
(
<figureCitation id="13342A22630326546D7076AFFE3565EF" box="[297,386,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1336,1360]" captionTargetBox="[155,1430,187,1303]" captionTargetId="figure-17@3.[151,1437,181,1312]" captionTargetPageId="3" captionText="FIGURE 1. Alaskacladius johnhudsoni gen. nov., sp. nov., male. A. head; B. thorax; C. wing; D. hypopygium." figureDoi="http://doi.org/10.5281/zenodo.8243763" httpUri="https://zenodo.org/record/8243763/files/figure.png" pageId="5" pageNumber="576">Fig. 1B</figureCitation>
). Acrostichals 14–16; dorsocentrals 12; prealars 9–13, 
<quantity id="4CF79B42630326546FAE76AFFB9465EF" box="[1015,1059,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="576" unit="in" value="8.0">8 in</quantity>
a semicircular pattern; scutellars 
<quantity id="4CF79B42630326546CCE7773FF4D640B" box="[151,250,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="5" pageNumber="576" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
two rows. Antepronotal lobes bearing 8–10 setae basally.
</paragraph>
<paragraph id="8BB036A7630326546C9E7757FC67649F" blockId="5.[151,1436,150,501]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E7757FEB66456" box="[199,257,295,320]" italics="true" pageId="5" pageNumber="576">Wing</emphasis>
(
<figureCitation id="13342A22630326546D497757FEDF6457" box="[272,360,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1336,1360]" captionTargetBox="[155,1430,187,1303]" captionTargetId="figure-17@3.[151,1437,181,1312]" captionTargetPageId="3" captionText="FIGURE 1. Alaskacladius johnhudsoni gen. nov., sp. nov., male. A. head; B. thorax; C. wing; D. hypopygium." figureDoi="http://doi.org/10.5281/zenodo.8243763" httpUri="https://zenodo.org/record/8243763/files/figure.png" pageId="5" pageNumber="576">Fig. 1C</figureCitation>
). Wing bare with fine punctation visible at 40 x magnification. Brachiolum with 1 seta. Squama with 19 setae. R with 19 setae, and R 
<subScript id="178B34E2630326546E69772DFD8F647D" attach="left" box="[560,568,349,363]" fontSize="6" pageId="5" pageNumber="576">1</subScript>
with 6 setae. R 
<subScript id="178B34E2630326546EBB772DFD4B647D" attach="left" box="[738,764,349,363]" fontSize="6" pageId="5" pageNumber="576">4+5</subScript>
with 11 setae, starting on apical 2/3 
<superScript id="7C7A9BEF6303265468D3773AFB2F644E" attach="left" box="[1162,1176,330,344]" fontSize="6" pageId="5" pageNumber="576">rd</superScript>
of the vein; other veins bare. Costa extension 71 μm. Cu1 moderately curved. Anal lobe rounded.
</paragraph>
<paragraph id="8BB036A7630326546C9E77E3FABA64E3" blockId="5.[151,1436,150,501]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E77E3FEB364BA" box="[199,260,403,428]" italics="true" pageId="5" pageNumber="576">Legs.</emphasis>
Femur, tibia and tarsus of all legs with long and strong beard. Pulvilli well-developed. Fore tibia spur 78–83, 80 μm long, mid tibia spurs 37–38 μm, and 29–37 μm long, hind tibia spurs 72–77, 74 and 28 μm long, hind tibia comb with around 17 bristles. Pseudospurs absent. Lengths and proportions of legs as in 
<tableCitation id="C68D031C6303265468EA77ABFAB064E3" box="[1203,1287,475,501]" captionStart="TABLE 1" captionStartId="5.[152,244,548,572]" captionText="TABLE 1. Male leg lengths (μm) and proportions of Alaskacladius johnhudsoni gen. nov., sp. nov.." pageId="5" pageNumber="576">Table 1</tableCitation>
.
</paragraph>
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<paragraph id="8BB036A7630326546CC17454FB7D672A" blockId="5.[152,1226,548,573]" box="[152,1226,548,573]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546CC17454FEB8672A" bold="true" box="[152,271,548,572]" pageId="5" pageNumber="576">TABLE 1.</emphasis>
Male leg lengths (μm) and proportions of 
<taxonomicName id="4C0F4D24630326546E8C7454FC42672A" authority="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram, 2023" authorityName="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram" authorityYear="2023" box="[725,1013,548,572]" class="Insecta" family="Chironomidae" genus="Alaskacladius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="johnhudsoni" status="gen. nov., sp. nov.">
<emphasis id="B97BEAB5630326546E8C7454FC42672A" box="[725,1013,548,572]" italics="true" pageId="5" pageNumber="576">Alaskacladius johnhudsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857CE630326546FA57455FB74672B" box="[1020,1219,549,573]" pageId="5" pageNumber="576" rank="species">
<emphasis id="B97BEAB5630326546FA57455FBD7672B" bold="true" box="[1020,1120,549,573]" pageId="5" pageNumber="576">gen. nov.</emphasis>
, 
<emphasis id="B97BEAB56303265468347455FB74672B" bold="true" box="[1133,1219,549,573]" pageId="5" pageNumber="576">sp. nov.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BB036A7630326546D76743CFAC367FF" pageId="5" pageNumber="576">
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<emphasis id="B97BEAB5630326546D76743CFEF46772" bold="true" box="[303,323,588,612]" pageId="5" pageNumber="576">fe</emphasis>
</th>
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<emphasis id="B97BEAB5630326546DFE743CFE016772" bold="true" box="[423,438,588,612]" pageId="5" pageNumber="576">ti</emphasis>
</th>
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<emphasis id="B97BEAB5630326546E4D743CFD9E6772" bold="true" box="[532,553,588,612]" pageId="5" pageNumber="576">ta</emphasis>
<emphasis id="B97BEAB5630326546E70742CFD87677C" bold="true" box="[553,560,604,618]" pageId="5" pageNumber="576">1</emphasis>
</th>
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<emphasis id="B97BEAB5630326546ED0743CFD296772" bold="true" box="[649,670,588,612]" pageId="5" pageNumber="576">ta</emphasis>
<emphasis id="B97BEAB5630326546EC7742CFD12677C" bold="true" box="[670,677,604,618]" pageId="5" pageNumber="576">2</emphasis>
</th>
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<emphasis id="B97BEAB5630326546EA4743CFCA56772" bold="true" box="[765,786,588,612]" pageId="5" pageNumber="576">ta</emphasis>
<emphasis id="B97BEAB5630326546F4B742CFCAE677C" bold="true" box="[786,793,604,618]" pageId="5" pageNumber="576">3</emphasis>
</th>
<th id="76EE5D996303D9AE6F35743CFC25677C" box="[876,914,588,618]" gridcol="6" gridrow="0" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546F28743CFC316772" bold="true" box="[881,902,588,612]" pageId="5" pageNumber="576">ta</emphasis>
<emphasis id="B97BEAB5630326546FDF742CFC3A677C" bold="true" box="[902,909,604,618]" pageId="5" pageNumber="576">4</emphasis>
</th>
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<emphasis id="B97BEAB5630326546FB2743CFBB76772" bold="true" box="[1003,1024,588,612]" pageId="5" pageNumber="576">ta</emphasis>
<emphasis id="B97BEAB5630326546859742CFBB0677C" bold="true" box="[1024,1031,604,618]" pageId="5" pageNumber="576">5</emphasis>
</th>
<th id="76EE5D996303D9AE6807743CFB3D677C" box="[1118,1162,588,618]" gridcol="8" gridrow="0" pageId="5" pageNumber="576">
<emphasis id="B97BEAB563032654683B743CFB326772" bold="true" box="[1122,1157,588,612]" pageId="5" pageNumber="576">LR</emphasis>
</th>
<th id="76EE5D996303D9AE6885743CFB48677C" box="[1244,1279,588,618]" gridcol="9" gridrow="0" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546885743CFB486772" bold="true" box="[1244,1279,588,612]" pageId="5" pageNumber="576">BV</emphasis>
</th>
<th id="76EE5D996303D9AE690D743CFAC3677C" box="[1364,1396,588,618]" gridcol="10" gridrow="0" pageId="5" pageNumber="576">
<emphasis id="B97BEAB563032654690D743CFAC36772" bold="true" box="[1364,1396,588,612]" pageId="5" pageNumber="576">SV</emphasis>
</th>
</tr>
<tr id="353F34E56303D9AE6CE7740CFAC3678C" box="[190,1396,636,666]" gridrow="1" pageId="5" pageNumber="576">
<th id="76EE5D996303D9AE6CE7740CFF62678C" box="[190,213,636,666]" gridcol="0" gridrow="1" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546CE7740CFF7A6782" bold="true" box="[190,205,636,660]" pageId="5" pageNumber="576">P</emphasis>
<emphasis id="B97BEAB5630326546C9774FCFF62678C" bold="true" box="[206,213,652,666]" pageId="5" pageNumber="576">1</emphasis>
</th>
<td id="76EE5D996303D9AE6D7F740CFEFB678C" box="[294,332,636,666]" gridcol="1" gridrow="1" pageId="5" pageNumber="576">838</td>
<td id="76EE5D996303D9AE6DCC740CFE7F678C" box="[405,456,636,666]" gridcol="2" gridrow="1" pageId="5" pageNumber="576">911</td>
<td id="76EE5D996303D9AE6E56740CFD82678C" box="[527,565,636,666]" gridcol="3" gridrow="1" pageId="5" pageNumber="576">575</td>
<td id="76EE5D996303D9AE6EDD740CFD1D678C" box="[644,682,636,666]" gridcol="4" gridrow="1" pageId="5" pageNumber="576">359</td>
<td id="76EE5D996303D9AE6EA1740CFCA9678C" box="[760,798,636,666]" gridcol="5" gridrow="1" pageId="5" pageNumber="576">237</td>
<td id="76EE5D996303D9AE6F35740CFC25678C" box="[876,914,636,666]" gridcol="6" gridrow="1" pageId="5" pageNumber="576">157</td>
<td id="76EE5D996303D9AE6FBF740CFBBB678C" box="[998,1036,636,666]" gridcol="7" gridrow="1" pageId="5" pageNumber="576">117</td>
<td id="76EE5D996303D9AE6807740CFB3D678C" box="[1118,1162,636,666]" gridcol="8" gridrow="1" pageId="5" pageNumber="576">0.63</td>
<td id="76EE5D996303D9AE6885740CFB48678C" box="[1244,1279,636,666]" gridcol="9" gridrow="1" pageId="5" pageNumber="576">2.7</td>
<td id="76EE5D996303D9AE690D740CFAC3678C" box="[1364,1396,636,666]" gridcol="10" gridrow="1" pageId="5" pageNumber="576">3.0</td>
</tr>
<tr id="353F34E56303D9AE6CE774D6FAC367D2" box="[190,1396,678,708]" gridrow="2" pageId="5" pageNumber="576">
<th id="76EE5D996303D9AE6CE774D6FF6267D2" box="[190,213,678,708]" gridcol="0" gridrow="2" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546CE774D6FF7A67A8" bold="true" box="[190,205,678,702]" pageId="5" pageNumber="576">P</emphasis>
<emphasis id="B97BEAB5630326546C9774C6FF6267D2" bold="true" box="[206,213,694,708]" pageId="5" pageNumber="576">2</emphasis>
</th>
<td id="76EE5D996303D9AE6D7F74D6FEFB67D2" box="[294,332,678,708]" gridcol="1" gridrow="2" pageId="5" pageNumber="576">879</td>
<td id="76EE5D996303D9AE6DCC74D6FE7F67D2" box="[405,456,678,708]" gridcol="2" gridrow="2" pageId="5" pageNumber="576">881</td>
<td id="76EE5D996303D9AE6E5674D6FD8267D2" box="[527,565,678,708]" gridcol="3" gridrow="2" pageId="5" pageNumber="576">396</td>
<td id="76EE5D996303D9AE6EDD74D6FD1D67D2" box="[644,682,678,708]" gridcol="4" gridrow="2" pageId="5" pageNumber="576">221</td>
<td id="76EE5D996303D9AE6EA174D6FCA967D2" box="[760,798,678,708]" gridcol="5" gridrow="2" pageId="5" pageNumber="576">180</td>
<td id="76EE5D996303D9AE6F3574D6FC2567D2" box="[876,914,678,708]" gridcol="6" gridrow="2" pageId="5" pageNumber="576">109</td>
<td id="76EE5D996303D9AE6FBF74D6FBBB67D2" box="[998,1036,678,708]" gridcol="7" gridrow="2" pageId="5" pageNumber="576">110</td>
<td id="76EE5D996303D9AE680774D6FB3D67D2" box="[1118,1162,678,708]" gridcol="8" gridrow="2" pageId="5" pageNumber="576">0.45</td>
<td id="76EE5D996303D9AE688574D6FB4867D2" box="[1244,1279,678,708]" gridcol="9" gridrow="2" pageId="5" pageNumber="576">3.5</td>
<td id="76EE5D996303D9AE690D74D6FAC367D2" box="[1364,1396,678,708]" gridcol="10" gridrow="2" pageId="5" pageNumber="576">4.4</td>
</tr>
<tr id="353F34E56303D9AE6CE774A0FAC367F8" box="[190,1396,720,750]" gridrow="3" pageId="5" pageNumber="576">
<th id="76EE5D996303D9AE6CE774A0FF6267F8" box="[190,213,720,750]" gridcol="0" gridrow="3" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546CE774A0FF7A67FE" bold="true" box="[190,205,720,744]" pageId="5" pageNumber="576">P</emphasis>
<emphasis id="B97BEAB5630326546C977490FF6267F8" bold="true" box="[206,213,736,750]" pageId="5" pageNumber="576">3</emphasis>
</th>
<td id="76EE5D996303D9AE6D7F74A0FEFB67F8" box="[294,332,720,750]" gridcol="1" gridrow="3" pageId="5" pageNumber="576">940</td>
<td id="76EE5D996303D9AE6DCC74A0FE7F67F8" box="[405,456,720,750]" gridcol="2" gridrow="3" pageId="5" pageNumber="576">1071</td>
<td id="76EE5D996303D9AE6E5674A0FD8267F8" box="[527,565,720,750]" gridcol="3" gridrow="3" pageId="5" pageNumber="576">639</td>
<td id="76EE5D996303D9AE6EDD74A0FD1D67F8" box="[644,682,720,750]" gridcol="4" gridrow="3" pageId="5" pageNumber="576">350</td>
<td id="76EE5D996303D9AE6EA174A0FCA967F8" box="[760,798,720,750]" gridcol="5" gridrow="3" pageId="5" pageNumber="576">231</td>
<td id="76EE5D996303D9AE6F3574A0FC2567F8" box="[876,914,720,750]" gridcol="6" gridrow="3" pageId="5" pageNumber="576">127</td>
<td id="76EE5D996303D9AE6FBF74A0FBBB67F8" box="[998,1036,720,750]" gridcol="7" gridrow="3" pageId="5" pageNumber="576">104</td>
<td id="76EE5D996303D9AE680774A0FB3D67F8" box="[1118,1162,720,750]" gridcol="8" gridrow="3" pageId="5" pageNumber="576">0.60</td>
<td id="76EE5D996303D9AE688574A0FB4867F8" box="[1244,1279,720,750]" gridcol="9" gridrow="3" pageId="5" pageNumber="576">3.3</td>
<td id="76EE5D996303D9AE690D74A0FAC367F8" box="[1364,1396,720,750]" gridcol="10" gridrow="3" pageId="5" pageNumber="576">3.1</td>
</tr>
</table>
</paragraph>
<caption id="DF70662F630326546CCE73A7FB7C6305" ID-DOI="http://doi.org/10.5281/zenodo.8243767" ID-Zenodo-Dep="8243767" httpUri="https://zenodo.org/record/8243767/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,250,1495,1519]" targetBox="[151,1436,808,1471]" targetPageId="5" targetType="figure">
<paragraph id="8BB036A7630326546CCE73A7FB7C6305" blockId="5.[151,1436,1495,1555]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546CCE73A7FEA460F9" bold="true" box="[151,275,1495,1519]" pageId="5" pageNumber="576">FIGURE 3.</emphasis>
<taxonomicName id="4C0F4D24630326546D4173A7FD9E60F9" authority="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram, 2023" authorityName="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram" authorityYear="2023" box="[280,553,1495,1519]" class="Insecta" family="Chironomidae" genus="Alaskacladius" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="johnhudsoni" status="gen. nov., sp. nov.">
<emphasis id="B97BEAB5630326546D4173A7FD9E60F9" box="[280,553,1495,1519]" italics="true" pageId="5" pageNumber="576">Alaskacladius johnhudsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857CE630326546E6973A7FD5C60F9" box="[560,747,1495,1519]" pageId="5" pageNumber="576" rank="species">
<emphasis id="B97BEAB5630326546E6973A7FD3960F9" bold="true" box="[560,654,1495,1519]" pageId="5" pageNumber="576">gen. nov.</emphasis>
, 
<emphasis id="B97BEAB5630326546EC273A7FD5C60F9" bold="true" box="[667,747,1495,1519]" pageId="5" pageNumber="576">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. A. hypopygium, arrow indicates the virga; B. variation in the shape of turned gonostylus C. variation in the shape of turned gonostylus. The scale bars represent 50 µm.
</paragraph>
</caption>
<paragraph id="8BB036A7630326546C9E7016FC506241" blockId="5.[151,1437,1637,2023]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E7016FEE16369" box="[199,342,1638,1663]" italics="true" pageId="5" pageNumber="576">Hypopygium</emphasis>
(
<figureCitation id="13342A22630326546D307015FE786369" box="[361,463,1637,1663]" captionStart="FIGURE 1" captionStartId="3.[151,250,1336,1360]" captionTargetBox="[155,1430,187,1303]" captionTargetId="figure-17@3.[151,1437,181,1312]" captionTargetPageId="3" captionText="FIGURE 1. Alaskacladius johnhudsoni gen. nov., sp. nov., male. A. head; B. thorax; C. wing; D. hypopygium." figureDoi="http://doi.org/10.5281/zenodo.8243763" httpUri="https://zenodo.org/record/8243763/files/figure.png" pageId="5" pageNumber="576">Figs. 1D</figureCitation>
&amp; 
<figureCitation id="13342A22630326546DA37015FD976369" box="[506,544,1637,1663]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="5" pageNumber="576">2A</figureCitation>
). Anal point prominent, cone-shaped, broad basally, 32–42, 36 μm long and 78–111, 90 μm wide, bearing around 24 simple moderately thick setae. Virga with 4 thin and compacted long spines placed vertically close to sternapodeme, 76–92, 84 μm long, and two hyaline lamellae adjacent to them, lamella with striation. Sternapodeme 117–148, 124 μm long, slightly arched, oral projections developed. Phallapodeme 42–56, 48 μm long. Gonocoxite 163–202, 184 μm long. Inferior volsella trapezoid shape, located halfway along the gonocoxite. Gonostylus prominent, 117–143, 133 μm long, expanded from basal 1/3 
<superScript id="7C7A9BEF63032654682C7169FB346231" attach="left" box="[1141,1155,1817,1831]" fontSize="6" pageId="5" pageNumber="576">rd</superScript>
to the apex, outer corner (projection) well-developed, megaseta 15–19, 16 μm long. HR 1.4, HV 2.4.
</paragraph>
<paragraph id="8BB036A7630326546C9E7111FECC626D" blockId="5.[151,1437,1637,2023]" box="[199,379,1889,1915]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E7111FECC626D" bold="true" box="[199,379,1889,1915]" pageId="5" pageNumber="576">Female (n = 2).</emphasis>
</paragraph>
<paragraph id="8BB036A7630326546C9E71F5FCDE6289" blockId="5.[151,1437,1637,2023]" box="[199,873,1925,1952]" pageId="5" pageNumber="576">
Total length 
<quantity id="4CF79B42630326546D0D71F5FE1A62B6" box="[340,429,1925,1952]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="576" unit="mm" value="2.8">2.8 mm</quantity>
. Wing 
<quantity id="4CF79B42630326546DA271F5FDE662B6" box="[507,593,1925,1952]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="576" unit="mm" value="1.9">1.9 mm</quantity>
long and 
<quantity id="4CF79B42630326546E9971F5FC9362B6" box="[704,804,1925,1952]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="576" unit="mm" value="0.55">0.55 mm</quantity>
wide.
</paragraph>
<paragraph id="8BB036A7630326546C9E71D9FE6E62F1" blockId="5.[151,1437,1637,2023]" pageId="5" pageNumber="576">
<emphasis id="B97BEAB5630326546C9E71D9FD7C62D5" box="[199,715,1961,1987]" italics="true" pageId="5" pageNumber="576">General coloration of the mounted specimens</emphasis>
. Head, thorax, halter, legs and tergites brown. Sternites slightly lighter. Wing greyish brown.
</paragraph>
<caption id="DF70662F630026576CCE7531FB9566D3" ID-DOI="http://doi.org/10.5281/zenodo.8243769" ID-Zenodo-Dep="8243769" httpUri="https://zenodo.org/record/8243769/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,833,857]" targetBox="[345,1239,184,805]" targetPageId="6" targetType="figure">
<paragraph id="8BB036A7630026576CCE7531FB9566D3" blockId="6.[151,1436,833,965]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576CCE7531FEA1664F" bold="true" box="[151,278,833,857]" pageId="6" pageNumber="577">FIGURE 4.</emphasis>
Neighbor-Joining (NJ) and Maximum Likelihood (ML) trees of the 
<taxonomicName id="4C0F4D24630026576FB57531FB34664F" authorityName="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram" authorityYear="2023" box="[1004,1155,833,857]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="571" phylum="Arthropoda" rank="subFamily" subFamily="Orthocladiinae">Orthocladiinae</taxonomicName>
genera and two outgroups 
<taxonomicName id="4C0F4D24630026576CCE7515FDA8666B" authority="Sublette, 1964" authorityName="Sublette" authorityYear="1964" box="[151,543,869,893]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B97BEAB5630026576CCE7515FE30666B" box="[151,391,869,893]" italics="true" pageId="6" pageNumber="577">Procladius denticulatus</emphasis>
Sublette, 1964
</taxonomicName>
and 
<taxonomicName id="4C0F4D24630026576E167516FC50666B" authority="Sublette, 1964" authorityName="Sublette" authorityYear="1964" box="[591,999,869,893]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="neopunctipennis">
<emphasis id="B97BEAB5630026576E167516FCF8666B" box="[591,847,870,893]" italics="true" pageId="6" pageNumber="577">Tanypus neopunctipennis</emphasis>
Sublette, 1964
</taxonomicName>
inferred from the COI nucleotide sequence data (567 bp). Numbers on branches represent the bootstrap value for Neighbor-Joining (NJ) and Maximum Likelihood (ML) (10000 replicates). Clades without numbers are supported with bootstrap support of ≤ 95.
</paragraph>
</caption>
<paragraph id="8BB036A7630026576C9E7268FA2A6124" blockId="6.[151,1437,1048,1552]" box="[199,1437,1048,1075]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9E7268FEB36124" box="[199,260,1048,1074]" italics="true" pageId="6" pageNumber="577">Head</emphasis>
(
<figureCitation id="13342A22630026576D4B7268FEDC6124" box="[274,363,1048,1074]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2A</figureCitation>
). Antenna segment one 77–87, 82 μm long; second segment 51 μm long, other segments missing.
</paragraph>
<paragraph id="8BB036A7630026576CCE724DFA2B6141" blockId="6.[151,1437,1048,1552]" box="[151,1436,1085,1111]" pageId="6" pageNumber="577">Eyes bare, with no dorsomedial extension, temporal setae 8–10 with 6–7 uniserial orbitals, and 2–3 outer verticals</paragraph>
<paragraph id="8BB036A7630026576CCE7211FA2A61B6" blockId="6.[151,1437,1048,1552]" pageId="6" pageNumber="577">
(
<figureCitation id="13342A22630026576CC67211FF4C616A" box="[159,251,1121,1148]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2A</figureCitation>
). Tentorium 192–197, 195 μm long, narrow and long apically, base slightly expanded, large tentorial pit close to apex (
<figureCitation id="13342A22630026576D6772F6FE2D61B6" box="[318,410,1158,1184]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2A</figureCitation>
). Clypeus 57–66, 62 μm long and 97 μm wide, bearing 8 setae, setae 54–58, 56 μm long.
</paragraph>
<paragraph id="8BB036A7630026576CCE72DBFA2B61D3" blockId="6.[151,1437,1048,1552]" box="[151,1436,1195,1222]" pageId="6" pageNumber="577">Palpal segments lengths (in μm; n = 1): 67, 45, 98, 103, 139. Third palpomere with 1 sensilla clavata medially, 4</paragraph>
<paragraph id="8BB036A7630026576CCE72A0FE1A61FC" blockId="6.[151,1437,1048,1552]" box="[151,429,1232,1258]" pageId="6" pageNumber="577">laterally, and 3 ventrally.</paragraph>
<paragraph id="8BB036A7630026576C9E7285FE2C6025" blockId="6.[151,1437,1048,1552]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9E7285FEAF6019" box="[199,280,1269,1295]" italics="true" pageId="6" pageNumber="577">Thorax</emphasis>
(
<figureCitation id="13342A22630026576D717285FE366019" box="[296,385,1269,1295]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2B</figureCitation>
). Acrostichals 14; dorsocentrals 18; prealars 10; scutellars 
<quantity id="4CF79B426300265768457285FBFF6019" box="[1052,1096,1269,1295]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="577" unit="in" value="8.0">8 in</quantity>
two rows. Antepronotal lobes bearing 5 setae basally.
</paragraph>
<paragraph id="8BB036A7630026576C9E734FFA2A604F" blockId="6.[151,1437,1048,1552]" box="[199,1437,1342,1369]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9E734FFEB6604E" box="[199,257,1343,1368]" italics="true" pageId="6" pageNumber="577">Wing</emphasis>
(
<figureCitation id="13342A22630026576D4D734EFEC7604E" box="[276,368,1342,1368]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2C</figureCitation>
). Wing bare with fine punctation visible at 40 x magnification. Brachiolum with 1–2 setae.
</paragraph>
<paragraph id="8BB036A7630026576CCE7313FBBE60B4" blockId="6.[151,1437,1048,1552]" pageId="6" pageNumber="577">
Squama with 3–6, 5 setae. R with 17–21, 19 setae, and R 
<subScript id="178B34E2630026576F7F7306FC996092" attach="left" box="[806,814,1398,1412]" fontSize="6" pageId="6" pageNumber="577">1</subScript>
with 12–14, 13 setae. R 
<subScript id="178B34E263002657681D7305FBE96095" attach="left" box="[1092,1118,1397,1411]" fontSize="6" pageId="6" pageNumber="577">4+5</subScript>
with 25–31, 28 setae; other veins bare. Costa extension 86 μm. Cu 
<subScript id="178B34E2630026576E1E73EAFDF860BE" attach="left" box="[583,591,1434,1448]" fontSize="6" pageId="6" pageNumber="577">1</subScript>
moderately curved. Anal lobe reduced.
</paragraph>
<paragraph id="8BB036A7630026576C9E73DDFA2B60D1" blockId="6.[151,1437,1048,1552]" box="[199,1436,1453,1479]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9E73DDFEB360D0" box="[199,260,1453,1478]" italics="true" pageId="6" pageNumber="577">Legs.</emphasis>
Pulvilli well-developed. Fore tibia spur 32 μm long, mid tibia spurs 24 μm, hind tibia spurs 62–64, 63
</paragraph>
<paragraph id="8BB036A7630026576CCE73A1FEBA6306" blockId="6.[151,1437,1048,1552]" pageId="6" pageNumber="577">
and 21 μm long, hind tibia comb with around 12 bristles. Pseudospurs absent. Lengths and proportions of legs as in 
<tableCitation id="C68D031C630026576CEA7386FEBE6306" box="[179,265,1526,1552]" captionStart="TABLE 2" captionStartId="6.[152,244,1600,1624]" captionText="TABLE 2. Female leg lengths (μm) and proportions of Alaskacladius johnhudsoni gen. nov., sp. nov.." pageId="6" pageNumber="577">Table 2</tableCitation>
.
</paragraph>
<caption id="DF70662F630026576CC17030FB56634E" box="[152,1249,1600,1624]" pageId="6" pageNumber="577" startId="6.[152,244,1600,1624]" targetType="table">
<paragraph id="8BB036A7630026576CC17030FB56634E" blockId="6.[152,1249,1600,1624]" box="[152,1249,1600,1624]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576CC17030FEB8634E" bold="true" box="[152,271,1600,1624]" pageId="6" pageNumber="577">TABLE 2.</emphasis>
Female leg lengths (μm) and proportions of 
<taxonomicName id="4C0F4D24630026576EB47030FBBA634E" authority="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram, 2023" authorityName="Namayandeh &amp; Hudson &amp; Ghaderi &amp; Vasquez &amp; Ram" authorityYear="2023" box="[749,1037,1600,1624]" class="Insecta" family="Chironomidae" genus="Alaskacladius" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="johnhudsoni" status="gen. nov., sp. nov.">
<emphasis id="B97BEAB5630026576EB47030FBBA634E" box="[749,1037,1600,1624]" italics="true" pageId="6" pageNumber="577">Alaskacladius johnhudsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857CE63002657684D7030FB6C634E" box="[1044,1243,1600,1624]" pageId="6" pageNumber="577" rank="species">
<emphasis id="B97BEAB563002657684D7030FBCF634E" bold="true" box="[1044,1144,1600,1624]" pageId="6" pageNumber="577">gen. nov.</emphasis>
, 
<emphasis id="B97BEAB56300265768DC7030FB6C634E" bold="true" box="[1157,1243,1600,1624]" pageId="6" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BB036A7630026576D667018FAC36212" pageId="6" pageNumber="577">
<table id="F90FC4076300D9AE6C917018FAC3621F" box="[200,1396,1640,1801]" gridcols="11" gridrows="4" pageId="6" pageNumber="577">
<tr id="353F34E56300D9AE6C917018FAC36390" box="[200,1396,1640,1670]" gridrow="0" pageId="6" pageNumber="577" rowspan-0="1">
<th id="76EE5D996300D9AE6D6F7018FEEB6390" box="[310,348,1640,1670]" gridcol="1" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576D667018FEE46396" bold="true" box="[319,339,1640,1664]" pageId="6" pageNumber="577">fe</emphasis>
</th>
<th id="76EE5D996300D9AE6DFA7018FE7E6390" box="[419,457,1640,1670]" gridcol="2" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576DF67018FE096396" bold="true" box="[431,446,1640,1664]" pageId="6" pageNumber="577">ti</emphasis>
</th>
<th id="76EE5D996300D9AE6E497018FD816390" box="[528,566,1640,1670]" gridcol="3" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576E4C7018FD9D6396" bold="true" box="[533,554,1640,1664]" pageId="6" pageNumber="577">ta</emphasis>
<emphasis id="B97BEAB5630026576E737007FD866393" bold="true" box="[554,561,1655,1669]" pageId="6" pageNumber="577">1</emphasis>
</th>
<th id="76EE5D996300D9AE6EDF7018FD1B6390" box="[646,684,1640,1670]" gridcol="4" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576ED27018FD176396" bold="true" box="[651,672,1640,1664]" pageId="6" pageNumber="577">ta</emphasis>
<emphasis id="B97BEAB5630026576EF97007FD106393" bold="true" box="[672,679,1655,1669]" pageId="6" pageNumber="577">2</emphasis>
</th>
<th id="76EE5D996300D9AE6EA27018FC966390" box="[763,801,1640,1670]" gridcol="5" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576F597018FCA26396" bold="true" box="[768,789,1640,1664]" pageId="6" pageNumber="577">ta</emphasis>
<emphasis id="B97BEAB5630026576F4C7007FCAB6393" bold="true" box="[789,796,1655,1669]" pageId="6" pageNumber="577">3</emphasis>
</th>
<th id="76EE5D996300D9AE6F2F7018FC256390" box="[886,914,1640,1670]" gridcol="6" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576F2F7018FC3C6396" bold="true" box="[886,907,1640,1664]" pageId="6" pageNumber="577">ta</emphasis>
<emphasis id="B97BEAB5630026576FD27007FC256393" bold="true" box="[907,914,1655,1669]" pageId="6" pageNumber="577">4</emphasis>
</th>
<th id="76EE5D996300D9AE6FB67018FBBC6390" box="[1007,1035,1640,1670]" gridcol="7" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576FB67018FBB36396" bold="true" box="[1007,1028,1640,1664]" pageId="6" pageNumber="577">ta</emphasis>
<emphasis id="B97BEAB563002657685D7008FBBC6390" bold="true" box="[1028,1035,1656,1670]" pageId="6" pageNumber="577">5</emphasis>
</th>
<th id="76EE5D996300D9AE68387018FB3A6390" box="[1121,1165,1640,1670]" gridcol="8" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB563002657683C7018FB3F6396" bold="true" box="[1125,1160,1640,1664]" pageId="6" pageNumber="577">LR</emphasis>
</th>
<th id="76EE5D996300D9AE68877018FAB66390" box="[1246,1281,1640,1670]" gridcol="9" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB56300265768877018FAB66396" bold="true" box="[1246,1281,1640,1664]" pageId="6" pageNumber="577">BV</emphasis>
</th>
<th id="76EE5D996300D9AE690D7018FAC36390" box="[1364,1396,1640,1670]" gridcol="10" gridrow="0" pageId="6" pageNumber="577">
<emphasis id="B97BEAB563002657690D7018FAC36396" bold="true" box="[1364,1396,1640,1664]" pageId="6" pageNumber="577">SV</emphasis>
</th>
</tr>
<tr id="353F34E56300D9AE6C9170E8FAC363A3" box="[200,1396,1688,1717]" gridrow="1" pageId="6" pageNumber="577">
<th id="76EE5D996300D9AE6C9170E8FF6963A3" box="[200,222,1688,1717]" gridcol="0" gridrow="1" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9170E8FF6063A6" bold="true" box="[200,215,1688,1712]" pageId="6" pageNumber="577">P</emphasis>
<emphasis id="B97BEAB5630026576C8E70D7FF6963A3" bold="true" box="[215,222,1703,1717]" pageId="6" pageNumber="577">1</emphasis>
</th>
<td id="76EE5D996300D9AE6D6F70E8FEEB63A3" box="[310,348,1688,1717]" gridcol="1" gridrow="1" pageId="6" pageNumber="577">596</td>
<td id="76EE5D996300D9AE6DFA70E8FE7E63A3" box="[419,457,1688,1717]" gridcol="2" gridrow="1" pageId="6" pageNumber="577">685</td>
<td id="76EE5D996300D9AE6E4970E8FD8163A3" box="[528,566,1688,1717]" gridcol="3" gridrow="1" pageId="6" pageNumber="577">354</td>
<td id="76EE5D996300D9AE6EDF70E8FD1B63A3" box="[646,684,1688,1717]" gridcol="4" gridrow="1" pageId="6" pageNumber="577">197</td>
<td id="76EE5D996300D9AE6EA270E8FC9663A3" box="[763,801,1688,1717]" gridcol="5" gridrow="1" pageId="6" pageNumber="577">126</td>
<td id="76EE5D996300D9AE6F2F70E8FC2563A3" box="[886,914,1688,1717]" gridcol="6" gridrow="1" pageId="6" pageNumber="577">94</td>
<td id="76EE5D996300D9AE6FB670E8FBBC63A3" box="[1007,1035,1688,1717]" gridcol="7" gridrow="1" pageId="6" pageNumber="577">94</td>
<td id="76EE5D996300D9AE683870E8FB3A63A3" box="[1121,1165,1688,1717]" gridcol="8" gridrow="1" pageId="6" pageNumber="577">0.52</td>
<td id="76EE5D996300D9AE688770E8FAB663A3" box="[1246,1281,1688,1717]" gridcol="9" gridrow="1" pageId="6" pageNumber="577">3.2</td>
<td id="76EE5D996300D9AE690D70E8FAC363A3" box="[1364,1396,1688,1717]" gridcol="10" gridrow="1" pageId="6" pageNumber="577">3.6</td>
</tr>
<tr id="353F34E56300D9AE6C9170B2FAC363C9" box="[200,1396,1730,1759]" gridrow="2" pageId="6" pageNumber="577">
<th id="76EE5D996300D9AE6C9170B2FF6963C9" box="[200,222,1730,1759]" gridcol="0" gridrow="2" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9170B2FF6063CC" bold="true" box="[200,215,1730,1754]" pageId="6" pageNumber="577">P</emphasis>
<emphasis id="B97BEAB5630026576C8E70A1FF6963C9" bold="true" box="[215,222,1745,1759]" pageId="6" pageNumber="577">2</emphasis>
</th>
<td id="76EE5D996300D9AE6D6F70B2FEEB63C9" box="[310,348,1730,1759]" gridcol="1" gridrow="2" pageId="6" pageNumber="577">627</td>
<td id="76EE5D996300D9AE6DFA70B2FE7E63C9" box="[419,457,1730,1759]" gridcol="2" gridrow="2" pageId="6" pageNumber="577">589</td>
<td id="76EE5D996300D9AE6E4970B2FD8163C9" box="[528,566,1730,1759]" gridcol="3" gridrow="2" pageId="6" pageNumber="577">269</td>
<td id="76EE5D996300D9AE6EDF70B2FD1B63C9" box="[646,684,1730,1759]" gridcol="4" gridrow="2" pageId="6" pageNumber="577">137</td>
<td id="76EE5D996300D9AE6EA270B2FC9663C9" box="[763,801,1730,1759]" gridcol="5" gridrow="2" pageId="6" pageNumber="577">87</td>
<td id="76EE5D996300D9AE6F2F70B2FC2563C9" box="[886,914,1730,1759]" gridcol="6" gridrow="2" pageId="6" pageNumber="577">54</td>
<td id="76EE5D996300D9AE6FB670B2FBBC63C9" box="[1007,1035,1730,1759]" gridcol="7" gridrow="2" pageId="6" pageNumber="577">48</td>
<td id="76EE5D996300D9AE683870B2FB3A63C9" box="[1121,1165,1730,1759]" gridcol="8" gridrow="2" pageId="6" pageNumber="577">0.46</td>
<td id="76EE5D996300D9AE688770B2FAB663C9" box="[1246,1281,1730,1759]" gridcol="9" gridrow="2" pageId="6" pageNumber="577">4.6</td>
<td id="76EE5D996300D9AE690D70B2FAC363C9" box="[1364,1396,1730,1759]" gridcol="10" gridrow="2" pageId="6" pageNumber="577">4.5</td>
</tr>
<tr id="353F34E56300D9AE6C91709CFAC3621F" box="[200,1396,1772,1801]" gridrow="3" pageId="6" pageNumber="577">
<th id="76EE5D996300D9AE6C91709CFF69621F" box="[200,222,1772,1801]" gridcol="0" gridrow="3" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C91709CFF606212" bold="true" box="[200,215,1772,1796]" pageId="6" pageNumber="577">P</emphasis>
<emphasis id="B97BEAB5630026576C8E708BFF69621F" bold="true" box="[215,222,1787,1801]" pageId="6" pageNumber="577">3</emphasis>
</th>
<td id="76EE5D996300D9AE6D6F709CFEEB621F" box="[310,348,1772,1801]" gridcol="1" gridrow="3" pageId="6" pageNumber="577">701</td>
<td id="76EE5D996300D9AE6DFA709CFE7E621F" box="[419,457,1772,1801]" gridcol="2" gridrow="3" pageId="6" pageNumber="577">750</td>
<td id="76EE5D996300D9AE6E49709CFD81621F" box="[528,566,1772,1801]" gridcol="3" gridrow="3" pageId="6" pageNumber="577">424</td>
<td id="76EE5D996300D9AE6EDF709CFD1B621F" box="[646,684,1772,1801]" gridcol="4" gridrow="3" pageId="6" pageNumber="577">229</td>
<td id="76EE5D996300D9AE6EA2709CFC96621F" box="[763,801,1772,1801]" gridcol="5" gridrow="3" pageId="6" pageNumber="577">184</td>
<td id="76EE5D996300D9AE6F2F709CFC25621F" box="[886,914,1772,1801]" gridcol="6" gridrow="3" pageId="6" pageNumber="577">98</td>
<td id="76EE5D996300D9AE6FB6709CFBBC621F" box="[1007,1035,1772,1801]" gridcol="7" gridrow="3" pageId="6" pageNumber="577">95</td>
<td id="76EE5D996300D9AE6838709CFB3A621F" box="[1121,1165,1772,1801]" gridcol="8" gridrow="3" pageId="6" pageNumber="577">0.56</td>
<td id="76EE5D996300D9AE6887709CFAB6621F" box="[1246,1281,1772,1801]" gridcol="9" gridrow="3" pageId="6" pageNumber="577">3.1</td>
<td id="76EE5D996300D9AE690D709CFAC3621F" box="[1364,1396,1772,1801]" gridcol="10" gridrow="3" pageId="6" pageNumber="577">3.4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB036A7630026576C9E712CFA2B62F4" blockId="6.[151,1437,1884,2018]" pageId="6" pageNumber="577">
<emphasis id="B97BEAB5630026576C9E712CFE866260" box="[199,305,1884,1910]" italics="true" pageId="6" pageNumber="577">Genitalia</emphasis>
(
<figureCitation id="13342A22630026576D1C712CFE776260" box="[325,448,1884,1910]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2D–E</figureCitation>
). Gonocoxite well-reduced, not extending beyond the anterior of tergite IX, bearing around 8 setae. Seminal capsules semi-circular, 71–79, 76 µm long, and 91–103, 96 µm wide. Spermathecal ducts with loop (
<figureCitation id="13342A22630026576D4D71D4FEC762A8" box="[276,368,1956,1982]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="6" pageNumber="577">Fig. 2D</figureCitation>
). Notum 100 µm long, ramus 91 µm long. Gonapophysis VIII divided into large ventrolateral lobe well separated basally and apically from dorsomesal lobe; apodeme lobe distinct between these lobes. Tergite
</paragraph>
<paragraph id="8BB036A7630126566CCE76E7FD8665C3" blockId="7.[151,1436,150,249]" pageId="7" pageNumber="578">
IX not divided, large, and crescent-shaped, bearing around 25 setae (
<figureCitation id="13342A22630126566FD376E7FC5765A7" box="[906,992,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="7" pageNumber="578">Fig. 2E</figureCitation>
). Cercus semi-circular (
<figureCitation id="13342A226301265668BE76E7FA8B65A7" box="[1255,1340,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1573,1597]" captionTargetBox="[161,1427,191,1536]" captionTargetId="figure-23@4.[151,1437,181,1549]" captionTargetPageId="4" captionText="FIGURE 2. Alaskacladius johnhudsoni gen. nov., sp. nov., female.A. head; B. thorax; C. wing; D. genitalia ventral; E. genitalia dorsal." figureDoi="http://doi.org/10.5281/zenodo.8243765" httpUri="https://zenodo.org/record/8243765/files/figure.png" pageId="7" pageNumber="578">Fig. 2E</figureCitation>
), 63–75, 69 µm long and 73–75, 74 µm wide.
</paragraph>
<paragraph id="8BB036A7630126566C9E76AFFE7865EF" blockId="7.[151,1436,150,249]" box="[199,463,223,249]" pageId="7" pageNumber="578">
<emphasis id="B97BEAB5630126566C9E76AFFEE565EF" bold="true" box="[199,338,223,249]" pageId="7" pageNumber="578">Immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>