305 lines
36 KiB
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305 lines
36 KiB
XML
<document id="7BAB26031C5AC69AD00983FB4B5D5D2D" ID-CLB-Dataset="132960" ID-DOI="10.11646/phytotaxa.559.2.7" ID-GBIF-Dataset="8d205305-4c48-47dd-9d00-3d1b0b660aa1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7021601" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661410718254" checkinUser="plazi" docAuthor="Geethakumary, M. P., Deepu, S., Kissling, J. & Pandurangan, A. G." docDate="2022" docId="0397E93AFF89F16607EBFEA6DCF3F4A6" docLanguage="en" docName="phytotaxa.559.2.7.pdf" docOrigin="Phytotaxa 559 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Exacum Linnaeus 1753" docType="key" docVersion="9" lastPageNumber="191" masterDocId="FFAE9142FF8FF1600763FFADD959F006" masterDocTitle="Revisiting the taxonomy of Exacum courtallense (Gentianaceae) and recognizing E. courtallense var. laxiflorum at the species rank" masterLastPageNumber="192" masterPageNumber="185" pageNumber="191" updateTime="1699373709092" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="BF4FF86B3E034242959713F2F3A9EB1D">
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<mods:title id="4E1AB1BBB0044409DC0D5B37F9E084CB">Revisiting the taxonomy of Exacum courtallense (Gentianaceae) and recognizing E. courtallense var. laxiflorum at the species rank</mods:title>
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<mods:name id="792F774D573D809653611ED066D29D9C" type="personal">
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<mods:namePart id="BFA642885DC207A96BF5A77DCCAD1C80">Geethakumary, M. P.</mods:namePart>
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<mods:nameIdentifier id="25923D1826051FD37C20A9B4FF747F7D" type="ORCID">0000-0002-2136-5987</mods:nameIdentifier>
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<mods:affiliation id="10CEFC28839941803928814B3C5879A8">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India & geethatbgt @ gmail. com; https: // orcid. org / 0000 - 0002 - 2136 - 5987</mods:affiliation>
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<mods:nameIdentifier id="831D3CD4280355A7EF41F9B660C9B80A" type="email">geethatbgt@gmail.com</mods:nameIdentifier>
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<mods:namePart id="09C18611D7BC60E6FBBA7AAC72736558">Deepu, S.</mods:namePart>
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<mods:affiliation id="705A19DBB4573E6E1A11658C8DC4F0DB">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India & Forest Ecology Department, Kerala Forest Research Institute, Thrissur, Kerala 680653, India</mods:affiliation>
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<mods:name id="2EF45E6A172ACC7A276B868DE6C08694" type="personal">
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<mods:role id="084CFD1A5BD3F741059E80D19DC4A7FD">
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<mods:namePart id="6E642F1F24D89221DD29DA9C0960DDE4">Kissling, J.</mods:namePart>
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<mods:nameIdentifier id="31CF2555427733098A24F4DFAFB64536" type="ORCID">0000-0002-1078-4000</mods:nameIdentifier>
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<mods:affiliation id="3500D7E51B47A390ACE8BCE9D75D2F13">Institute of biology, University of Neuchâtel, Rue Emile-Argand 11, 2000 Neuchâtel, Switzerland & jonathan. kissling @ unine. ch; https: // orcid. org / 0000 - 0002 - 1078 - 4000</mods:affiliation>
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<mods:nameIdentifier id="9D3CCCB53F586E303ABF59A9E944BFC3" type="email">jonathan.kissling@unine.ch</mods:nameIdentifier>
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<mods:namePart id="72595D06BE3ABDC65D51943A3CB424FD">Pandurangan, A. G.</mods:namePart>
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<mods:affiliation id="3C816E382EAD9BC6803EF9785A20176E">Plant Systematic and Evolutionary Science Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute, Karimancode P. O., Thiruvananthapuram, Kerala 695562, India & Centre for Innovation in Science and Social Action, Thiruvananthapuram, Kerala 695010, India</mods:affiliation>
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<mods:typeOfResource id="9BAA00666FF3DBA18E38B77A5811D0D5">text</mods:typeOfResource>
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<mods:title id="C6FF0555186FA1C2D0E630451594D54A">Phytotaxa</mods:title>
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<mods:part id="1AE34851FE6375780C89422242FBEB52">
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<mods:date id="DB56B94D098CF0D9B29388A0EDC6299E">2022</mods:date>
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<mods:number id="19B2C5451CDEC207D68B24095BE624DB">2022-08-23</mods:number>
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<mods:number id="71CF580812A95D57D94DF34E17F9D260">559</mods:number>
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<mods:detail id="231F8B0056E1F5F87ED2B2EA6AD393FB" type="issue">
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<mods:number id="9B66DCF4F6481693538C59B1512EBF9C">2</mods:number>
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<mods:start id="46A9D72AD9B83136DC63567C54F6A628">185</mods:start>
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<mods:identifier id="B439C6C4E720FC19719EB044D1BA1C5F" type="CLB-Dataset">132960</mods:identifier>
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<mods:identifier id="534A2F5C6F78DB2CA1FE7BF1AF1B71FD" type="DOI">10.11646/phytotaxa.559.2.7</mods:identifier>
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<treatment id="0397E93AFF89F16607EBFEA6DCF3F4A6" ID-DOI="http://doi.org/10.5281/zenodo.7041579" ID-GBIF-Taxon="199891830" ID-Zenodo-Dep="7041579" LSID="urn:lsid:plazi:treatment:0397E93AFF89F16607EBFEA6DCF3F4A6" httpUri="http://treatment.plazi.org/id/0397E93AFF89F16607EBFEA6DCF3F4A6" lastPageNumber="191" pageId="6" pageNumber="191">
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<subSubSection id="C3240BA7FF89F16607EBFEA6DBDEF123" box="[136,647,266,293]" pageId="6" pageNumber="191" type="nomenclature">
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<paragraph id="8B81582CFF89F16607EBFEA6DBDEF123" blockId="6.[136,647,266,293]" box="[136,647,266,293]" pageId="6" pageNumber="191">
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<heading id="D0C9EF40FF89F16607EBFEA6DBDEF123" bold="true" box="[136,647,266,293]" fontSize="11" level="1" pageId="6" pageNumber="191" reason="1">
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<emphasis id="B94A843EFF89F16607EBFEA6DBDEF123" bold="true" box="[136,647,266,293]" italics="true" pageId="6" pageNumber="191">
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Key to
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<taxonomicName id="4C3E23AFFF89F16607B4FEA6D86EF123" ID-CoL="4HPR" authority="Linnaeus (1753: 112)" authorityName="Linnaeus" authorityPageNumber="112" authorityYear="1753" box="[215,311,267,293]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="genus">Exacum</taxonomicName>
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species in the Western Ghats
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3240BA7FF89F16607EBFEE5DCF3F4A6" pageId="6" pageNumber="191" type="key">
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<keyStep id="30CA4089FF89F16607EBFEE5DCF2F17F" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFEE5DCF2F15B" blockId="6.[136,1451,327,1185]" box="[136,1451,327,349]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFEE5DCF2F15B" box="[136,1451,327,349]" pageId="6" pageNumber="191">1. Flowers 4-merous ...............................................................................................................................................................................2</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFEC8DCF2F17F" blockId="6.[136,1451,327,1185]" box="[136,1451,356,378]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFEC8DCF2F17F" box="[136,1451,356,378]" pageId="6" pageNumber="191">- Flowers 5-merous ...............................................................................................................................................................................8</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CA4089FF89F16607EBFE2CDCF2F1B5" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFE2CDCF2F190" blockId="6.[136,1451,327,1185]" box="[136,1451,385,407]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFE2CDCF2F190" box="[136,1451,385,407]" pageId="6" pageNumber="191">2. Calyx lobes without wings .................................................................................................................................................................3</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFE33DCF2F1B5" blockId="6.[136,1451,327,1185]" box="[136,1451,414,436]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFE33DCF2F1B5" box="[136,1451,414,436]" pageId="6" pageNumber="191">- Calyx lobes with prominent wings .....................................................................................................................................................4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CA4089FF89F16607EBFE16DCF2F1EB" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFE16DCF2F1D6" blockId="6.[136,1451,327,1185]" box="[136,1451,442,465]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFE16DCF2F1D6" box="[136,1451,442,465]" pageId="6" pageNumber="191">
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3. Flowers subsessile; calyx wings without prominent veins................................................................
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<taxonomicName id="4C3E23AFFF89F166031FFE16DCF2F1D6" ID-CoL="6HLNS" authority="Linnaeus (1753: 112)" authorityName="Linnaeus" authorityPageNumber="112" authorityYear="1753" box="[1148,1451,442,465]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="sessile">
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<emphasis id="B94A843EFF89F166031FFE16DD8EF1D6" bold="true" box="[1148,1239,442,464]" italics="true" pageId="6" pageNumber="191">E. sessile</emphasis>
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<bibRefCitation id="EFAF25DDFF89F16603BEFE16DCF2F1D6" author="Linnaeus, C." box="[1245,1451,442,465]" pageId="6" pageNumber="191" refId="ref4821" refString="Linnaeus, C. (1753) Species Plantarum. Vol. 1. Impensis Laurentii Salvii, Stockholm, 560 pp." type="book" year="1753">Linnaeus (1753: 112)</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFE75DCF2F1EB" blockId="6.[136,1451,327,1185]" box="[136,1451,471,493]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFE75DCF2F1EB" box="[136,1451,471,493]" pageId="6" pageNumber="191">
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- Flowers pedicellate; calyx wings with prominent veins...................................................................
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<taxonomicName id="4C3E23AFFF89F166031FFE75DCF2F1EB" ID-CoL="3DJCY" authority="C. B. Clarke (1885: 98)" authorityName="C. B. Clarke" authorityPageNumber="98" authorityYear="1885" box="[1148,1451,471,493]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="lawii">
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<emphasis id="B94A843EFF89F166031FFE75DD91F1EB" bold="true" box="[1148,1224,471,493]" italics="true" pageId="6" pageNumber="191">E. lawii</emphasis>
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C.B.
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<bibRefCitation id="EFAF25DDFF89F1660263FE7ADCF2F1EB" author="Clarke, C. B." box="[1280,1451,471,493]" pageId="6" pageNumber="191" pagination="98" refId="ref4276" refString="Clarke, C. B. (1885) [Exacum lawii]. In: Hooker, J. D., Flora of British India, vol. 4. L. Reeve & Co., London, pp. 98." type="book chapter" year="1885">Clarke (1885: 98)</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CA4089FF89F16607EBFE59DCF2F221" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFE59DCF2F20F" blockId="6.[136,1451,327,1185]" box="[136,1451,500,522]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFE59DCF2F20F" box="[136,1451,500,522]" pageId="6" pageNumber="191">
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4. Leaves 3-veined (midvein with one secondary vein on each side) ......................................
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<taxonomicName id="4C3E23AFFF89F166034FFE58DCF2F20F" ID-CoL="3DJDS" authority="Linnaeus (1753: 112)" authorityName="Linnaeus" authorityPageNumber="112" authorityYear="1753" box="[1068,1451,500,522]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="pedunculatum">
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<emphasis id="B94A843EFF89F166034FFE58DD8EF20C" bold="true" box="[1068,1239,500,522]" italics="true" pageId="6" pageNumber="191">E. pedunculatum</emphasis>
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<bibRefCitation id="EFAF25DDFF89F16603BEFE59DCF2F20F" author="Linnaeus, C." box="[1245,1451,500,522]" pageId="6" pageNumber="191" refId="ref4821" refString="Linnaeus, C. (1753) Species Plantarum. Vol. 1. Impensis Laurentii Salvii, Stockholm, 560 pp." type="book" year="1753">Linnaeus (1753: 112)</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFDBFDCF2F221" blockId="6.[136,1451,327,1185]" box="[136,1451,529,551]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFDBFDCF2F221" box="[136,1451,529,551]" pageId="6" pageNumber="191">- Leaves 5-veined (midvein with two secondary veins on each side) ..................................................................................................5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CA4089FF89F16607EBFD83DCF2F266" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFD83DCF2F242" blockId="6.[136,1451,327,1185]" box="[136,1451,558,580]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFD83DCF2F242" box="[136,1451,558,580]" pageId="6" pageNumber="191">5. Anthers oblong ...................................................................................................................................................................................6</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFDE6DCF2F266" blockId="6.[136,1451,327,1185]" box="[136,1451,587,609]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFDE6DCF2F266" box="[136,1451,587,609]" pageId="6" pageNumber="191">- Anthers linear .....................................................................................................................................................................................7</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30CA4089FF89F16607EBFDCADCF2F29C" pageId="6" pageNumber="191">
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<paragraph id="8B81582CFF89F16607EBFDCADCF2F27B" blockId="6.[136,1451,327,1185]" box="[136,1451,615,637]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFDCADCF2F27B" box="[136,1451,615,637]" pageId="6" pageNumber="191">
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6. Leaves with a distinct petiole; calyx wings truncate ......................................................................
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<taxonomicName id="4C3E23AFFF89F1660305FDC5DCF2F27B" ID-CoL="3DJDW" authority="Grisebach (1845: 46)" authorityName="Grisebach" authorityPageNumber="46" authorityYear="1845" box="[1126,1451,615,637]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="petiolare">
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<emphasis id="B94A843EFF89F1660305FDC5DD80F27B" bold="true" box="[1126,1241,615,637]" italics="true" pageId="6" pageNumber="191">E. petiolare</emphasis>
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<bibRefCitation id="EFAF25DDFF89F16603BCFDCADCF2F27B" author="Grisebach, A. H. R." box="[1247,1451,615,637]" pageId="6" pageNumber="191" pagination="46" refId="ref4352" refString="Grisebach, A. H. R. (1845) [Exacum]. In: A. P. de Candolle, Prodromus Systematis Naturalis Regni Vegetabilis, vol. 9. Sumptibus Sociorum Treuttel et Wu ¨ rtz, Paris, pp. 46. https: // doi. org / 10.5962 / bhl. title. 286" type="book chapter" year="1845">Grisebach (1845: 46)</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B81582CFF89F16607EBFD28DCF2F29C" blockId="6.[136,1451,327,1185]" box="[136,1451,644,666]" pageId="6" pageNumber="191">
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<keyLead id="30CFFB19FF89F16607EBFD28DCF2F29C" box="[136,1451,644,666]" pageId="6" pageNumber="191">
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- Leaves sessile; calyx wings cordate ..............................................................................................
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<taxonomicName id="4C3E23AFFF89F1660300FD28DCF2F29C" ID-CoL="3DJDZ" authority="Grisebach (1845: 46)" authorityName="Grisebach" authorityPageNumber="46" authorityYear="1845" box="[1123,1451,644,666]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="pumilum">
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<emphasis id="B94A843EFF89F1660300FD28DD81F29C" bold="true" box="[1123,1240,644,666]" italics="true" pageId="6" pageNumber="191">E. pumilum</emphasis>
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<bibRefCitation id="EFAF25DDFF89F16603BCFD29DCF2F29C" author="Grisebach, A. H. R." box="[1247,1451,644,666]" pageId="6" pageNumber="191" pagination="46" refId="ref4352" refString="Grisebach, A. H. R. (1845) [Exacum]. In: A. P. de Candolle, Prodromus Systematis Naturalis Regni Vegetabilis, vol. 9. Sumptibus Sociorum Treuttel et Wu ¨ rtz, Paris, pp. 46. https: // doi. org / 10.5962 / bhl. title. 286" type="book chapter" year="1845">Grisebach (1845: 46)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFD0FDCF2F2D5" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFD0FDCF2F2B0" blockId="6.[136,1451,327,1185]" box="[136,1451,673,695]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFD0FDCF2F2B0" box="[136,1451,673,695]" pageId="6" pageNumber="191">
|
|
7. Calyx lobes cuspidate with broad rounded dorsal wing; anthers curved ................................
|
|
<taxonomicName id="4C3E23AFFF89F1660323FD0CDCF2F2B0" ID-CoL="3DJF6" authority="Roxburgh (1832: 398)" authorityName="Roxburgh" authorityPageNumber="398" authorityYear="1832" box="[1088,1451,673,695]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="tetragonum">
|
|
<emphasis id="B94A843EFF89F1660323FD0CDD96F2B0" bold="true" box="[1088,1231,673,694]" italics="true" pageId="6" pageNumber="191">E. tetragonum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F16603B6FD0CDCF2F2B0" author="Roxburgh, W." box="[1237,1451,673,695]" pageId="6" pageNumber="191" pagination="398" refId="ref4987" refString="Roxburgh, W. (1832) [Exacum tetragonum]. In: Roxbourgh, W & Carey, W., Flora indica, vol. 1. Serampore, pp. 398. https: // doi. org / 10.5962 / bhl. title. 590" type="book chapter" year="1832">Roxburgh (1832: 398)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFD13DCF2F2D5" blockId="6.[136,1451,327,1185]" box="[136,1451,701,724]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFD13DCF2F2D5" box="[136,1451,701,724]" pageId="6" pageNumber="191">
|
|
- Calyx lobes acuminate with a narrow dorsal wing; anthers straight ............................................
|
|
<taxonomicName id="4C3E23AFFF89F166033CFD13DCF2F2D5" ID-CoL="3DJCG" authority="Klackenberg (1985: 43)" authorityName="Klackenberg" authorityPageNumber="43" authorityYear="1985" box="[1119,1451,701,724]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="grande">
|
|
<emphasis id="B94A843EFF89F166033CFD13DD99F2D5" bold="true" box="[1119,1216,701,723]" italics="true" pageId="6" pageNumber="191">E. grande</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F16603A5FD13DCF2F2D5" author="Klackenberg, J." box="[1222,1451,702,724]" pageId="6" pageNumber="191" pagination="1 - 144" refId="ref4799" refString="Klackenberg, J. (1985) The genus Exacum (Gentianaceae). Opera Botanica 84: 1 - 144." type="journal article" year="1985">Klackenberg (1985: 43)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFD76DCF2F30B" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFD76DCF2F2F6" blockId="6.[136,1451,327,1185]" box="[136,1451,731,753]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFD76DCF2F2F6" box="[136,1451,731,753]" pageId="6" pageNumber="191">8. Dorsal side of the anther with a papilla present near the apex ...........................................................................................................9</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFD55DCF2F30B" blockId="6.[136,1451,327,1185]" box="[136,1451,759,781]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFD55DCF2F30B" box="[136,1451,759,781]" pageId="6" pageNumber="191">- Dorsal side of the anther without a papilla.......................................................................................................................................14</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFCB9DCF2F340" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFCB9DCF2F32F" blockId="6.[136,1451,327,1185]" box="[136,1451,788,810]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFCB9DCF2F32F" box="[136,1451,788,810]" pageId="6" pageNumber="191">9. Leaves petiolate or ± sessile.............................................................................................................................................................10</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFC9FDCF2F340" blockId="6.[136,1451,327,1185]" box="[136,1451,817,839]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC9FDCF2F340" box="[136,1451,817,839]" pageId="6" pageNumber="191">- Leaves amplexicaul or semiamplexicaul..........................................................................................................................................12</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFCE3DCF2F386" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFCE3DCF2F362" blockId="6.[136,1451,327,1185]" box="[136,1451,845,868]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFCE3DCF2F362" box="[136,1451,845,868]" pageId="6" pageNumber="191">
|
|
10. Perennial herbs; anthers oblong............................................................................
|
|
<taxonomicName id="4C3E23AFFF89F16604AFFCE3DCF2F362" ID-CoL="3DJCS" authority="Geethakumary et al. (2016 a: 226)" authorityName="Geethakumary" authorityPageNumber="226" authorityYear="2016" box="[972,1451,845,868]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="idukkianum">
|
|
<emphasis id="B94A843EFF89F16604AFFCE3DD39F365" bold="true" box="[972,1120,845,867]" italics="true" pageId="6" pageNumber="191">E. idukkianum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660305FCE3DCF2F362" author="Geethakumary, M. P. & Pandurangan, A. G. & Ravichandran, P. & Deepu, S." box="[1126,1451,846,868]" pageId="6" pageNumber="191" pagination="225 - 229" refId="ref4414" refString="Geethakumary, M. P., Pandurangan, A. G., Ravichandran, P. & Deepu, S. (2016 a) A new species of Exacum (Gentianaceae) from Western Ghats, India. Phytotaxa 282 (3): 225 - 229. https: // doi. org / 10.11646 / phytotaxa. 282.3.6" type="journal article" year="2016">
|
|
Geethakumary
|
|
<emphasis id="B94A843EFF89F166039AFCE2DC73F365" box="[1273,1322,846,868]" italics="true" pageId="6" pageNumber="191">et al.</emphasis>
|
|
(2016a: 226)
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFCC6DCF2F386" blockId="6.[136,1451,327,1185]" box="[136,1451,875,897]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFCC6DCF2F386" box="[136,1451,875,897]" pageId="6" pageNumber="191">- Annual herbs; anthers linear.............................................................................................................................................................11</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFC25DCF2F3BF" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFC25DCF2F39B" blockId="6.[136,1451,327,1185]" box="[136,1451,903,925]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC25DCF2F39B" box="[136,1451,903,925]" pageId="6" pageNumber="191">
|
|
11. Stem 4-angled, winged; fruiting pedicel drooping.......................................................................
|
|
<taxonomicName id="4C3E23AFFF89F166033DFC25DCF2F39B" ID-CoL="3DJFL" authority="Arnott (1839: 176)" authorityName="Arnott" authorityPageNumber="176" authorityYear="1839" box="[1118,1451,903,925]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="wightianum">
|
|
<emphasis id="B94A843EFF89F166033DFC25DDA8F39B" bold="true" box="[1118,1265,903,925]" italics="true" pageId="6" pageNumber="191">E. wightianum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660396FC25DCF2F39B" author="Arnott, G. A. W." box="[1269,1451,903,925]" pageId="6" pageNumber="191" pagination="175 - 176" refId="ref4158" refString="Arnott, G. A. W. (1839) Exaci species ex peninsula Indica ac ex insula Ceylano. Annales de Sciences Naturelles; Botanique, Ser. 2, 11: 175 - 176." type="journal article" year="1839">Arnott (1839: 176)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFC08DCF2F3BF" blockId="6.[136,1451,327,1185]" box="[136,1451,932,954]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC08DCF2F3BF" box="[136,1451,932,954]" pageId="6" pageNumber="191">
|
|
- Stem 4-lineolate, not winged; fruiting pedicel not drooping..................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660359FC08DCF2F3BF" ID-CoL="3DJCW" authority="Gopalan (2002: 271)" authorityName="Gopalan" authorityPageNumber="271" authorityYear="2002" box="[1082,1451,932,954]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="klackenbergii">
|
|
<emphasis id="B94A843EFF89F1660359FC08DD84F3BC" bold="true" box="[1082,1245,932,954]" italics="true" pageId="6" pageNumber="191">E. klackenbergii</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660380FC09DCF2F3BF" author="Gopalan, R." box="[1251,1451,932,954]" pageId="6" pageNumber="191" pagination="271 - 273" refId="ref4544" refString="Gopalan, R. (2002) A new species of Exacum L. Gentianaceae from Agasthiyamalai (Pothigai), southern Western Ghats India. Journal of Bombay Natural History Society 99 (2): 271 - 273." type="journal article" year="2002">Gopalan (2002: 271)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFC6CDCF2F3F5" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFC6CDCF2F3D0" blockId="6.[136,1451,327,1185]" box="[136,1451,961,983]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC6CDCF2F3D0" box="[136,1451,961,983]" pageId="6" pageNumber="191">
|
|
12. Style shorter than anthers .........................................................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660487FC6CDCF2F3D0" ID-CoL="3DJCV" authority="Geethakumary et al. (2016 b: 429)" authorityName="Geethakumary" authorityPageNumber="429" authorityYear="2016" box="[996,1451,961,983]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="keralense">
|
|
<emphasis id="B94A843EFF89F1660487FC6CDD06F3D1" bold="true" box="[996,1119,961,983]" italics="true" pageId="6" pageNumber="191">E. keralense</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660306FC6CDCF2F3D0" author="Geethakumary, M. P. & K. M. P. Kumar & Pandurangan, A. G. & Deepu, S." box="[1125,1451,961,983]" pageId="6" pageNumber="191" pagination="429 - 433" refId="ref4477" refString="Geethakumary, M. P., K. M. P. Kumar, Pandurangan, A. G. & Deepu, S. (2016 b) Exacum keralense (Gentianaceae), a new species from the Western Ghats, India. Botany Letters 163 (4): 429 - 433. https: // doi. org / 10.1080 / 23818107.2016.1225267" type="journal article" year="2016">
|
|
Geethakumary
|
|
<emphasis id="B94A843EFF89F166039BFC6FDC70F3D0" box="[1272,1321,961,983]" italics="true" pageId="6" pageNumber="191">et al.</emphasis>
|
|
(2016b: 429)
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFC73DCF2F3F5" blockId="6.[136,1451,327,1185]" box="[136,1451,990,1012]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC73DCF2F3F5" box="[136,1451,990,1012]" pageId="6" pageNumber="191">- Style longer than anther....................................................................................................................................................................13</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFC56DCF2F42B" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFC56DCF2F416" blockId="6.[136,1451,327,1185]" box="[136,1451,1019,1041]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFC56DCF2F416" box="[136,1451,1019,1041]" pageId="6" pageNumber="191">
|
|
13. Leaves ovate, papilla in the dorsal side of anther distinct....................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660349FC56DCF2F416" ID-CoL="3DJB8" authority="Beddome (1874: 24)" authorityName="Beddome" authorityPageNumber="24" authorityYear="1874" box="[1066,1451,1019,1041]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="atropurpureum">
|
|
<emphasis id="B94A843EFF89F1660349FC56DD87F416" bold="true" box="[1066,1246,1019,1040]" italics="true" pageId="6" pageNumber="191">E. atropurpureum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660387FC56DCF2F416" author="Beddome, R. H." box="[1252,1451,1019,1041]" pageId="6" pageNumber="191" pagination="118" refId="ref4196" refString="Beddome, R. H. (1874) Icones Plantarum Indiae Orientalis. J. B. Pharoah, Madras, pp. 24, t. 118. https: // doi. org / 10.5962 / bhl. title. 92" type="journal article" year="1874">Beddome (1874: 24)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFBB5DCF2F42B" blockId="6.[136,1451,327,1185]" box="[136,1451,1047,1069]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFBB5DCF2F42B" box="[136,1451,1047,1069]" pageId="6" pageNumber="191">
|
|
- Leaves elliptic, papilla in the dorsal side of anther indistinct ............................................
|
|
<taxonomicName id="4C3E23AFFF89F1660342FBB5DCF2F42B" ID-CoL="3DJB2" authority="Beddome (1874: 33)" authorityName="Beddome" authorityPageNumber="33" authorityYear="1874" box="[1057,1451,1047,1069]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="anamallayanum">
|
|
<emphasis id="B94A843EFF89F1660342FBB5DD87F42B" bold="true" box="[1057,1246,1047,1069]" italics="true" pageId="6" pageNumber="191">E. anamallayanum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660387FBBADCF2F42B" author="Beddome, R. H." box="[1252,1451,1047,1069]" pageId="6" pageNumber="191" pagination="118" refId="ref4196" refString="Beddome, R. H. (1874) Icones Plantarum Indiae Orientalis. J. B. Pharoah, Madras, pp. 24, t. 118. https: // doi. org / 10.5962 / bhl. title. 92" type="journal article" year="1874">Beddome (1874: 33)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFB99DCF3F461" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFB99DCF2F44C" blockId="6.[136,1451,327,1185]" box="[136,1451,1076,1098]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFB99DCF2F44C" box="[136,1451,1076,1098]" pageId="6" pageNumber="191">14. Succulent herbs with a pincushion habit ..........................................................................................................................................15</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFBFFDCF3F461" blockId="6.[136,1451,327,1185]" box="[136,1450,1105,1127]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFBFFDCF3F461" box="[136,1450,1105,1127]" pageId="6" pageNumber="191">
|
|
- Slightly to much-branched ascending herbs..................................................................................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660246FBFCDCF3F461" ID-CoL="9VGDR" authorityName="Geethakumary, Deepu, Kissling & Pandurangan A. Habit" authorityYear="2022" baseAuthorityName="Geethakumary, Deepu, Kissling & Pandurangan A. Habit" baseAuthorityYear="2022" box="[1317,1450,1105,1127]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="laxiflorum">
|
|
<emphasis id="B94A843EFF89F1660246FBFCDCF3F461" bold="true" box="[1317,1450,1105,1127]" italics="true" pageId="6" pageNumber="191">E. laxiflorum</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="30CA4089FF89F16607EBFBC3DCF3F4A6" pageId="6" pageNumber="191">
|
|
<paragraph id="8B81582CFF89F16607EBFBC3DCF2F485" blockId="6.[136,1451,327,1185]" box="[136,1451,1134,1156]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFBC3DCF2F485" box="[136,1451,1134,1156]" pageId="6" pageNumber="191">
|
|
15 Flowers solitary, long-pedicellate..........................................................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660352FBC3DCF2F485" ID-CoL="3DJF8" authority="Beddome (1874: 24)" authorityName="Beddome" authorityPageNumber="24" authorityYear="1874" box="[1073,1451,1134,1156]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="travancoricum">
|
|
<emphasis id="B94A843EFF89F1660352FBC3DD87F485" bold="true" box="[1073,1246,1134,1155]" italics="true" pageId="6" pageNumber="191">E. travancoricum</emphasis>
|
|
<bibRefCitation id="EFAF25DDFF89F1660387FBC3DCF2F485" author="Beddome, R. H." box="[1252,1451,1134,1156]" pageId="6" pageNumber="191" pagination="118" refId="ref4196" refString="Beddome, R. H. (1874) Icones Plantarum Indiae Orientalis. J. B. Pharoah, Madras, pp. 24, t. 118. https: // doi. org / 10.5962 / bhl. title. 92" type="journal article" year="1874">Beddome (1874: 24)</bibRefCitation>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8B81582CFF89F16607EBFB26DCF3F4A6" blockId="6.[136,1451,327,1185]" box="[136,1450,1162,1185]" pageId="6" pageNumber="191">
|
|
<keyLead id="30CFFB19FF89F16607EBFB26DCF3F4A6" box="[136,1450,1162,1185]" pageId="6" pageNumber="191">
|
|
- Flowers in multiflorous cymes, short-pedicellate .......................................................................................................
|
|
<taxonomicName id="4C3E23AFFF89F1660276FB26DCF3F4A6" ID-CoL="3DJBT" authorityName="Arnott. A. Habit" authorityYear="1839" box="[1301,1450,1162,1184]" class="Magnoliopsida" family="Gentianaceae" genus="Exacum" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="191" phylum="Tracheophyta" rank="species" species="courtallense">
|
|
<emphasis id="B94A843EFF89F1660276FB26DCF3F4A6" bold="true" box="[1301,1450,1162,1184]" italics="true" pageId="6" pageNumber="191">E. courtallense</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |