599 lines
76 KiB
XML
599 lines
76 KiB
XML
<document id="AFA57D7C1A828CB6455759FD976BBFC8" ID-CLB-Dataset="22207" ID-DOI="10.11646/zootaxa.4778.1.3" ID-GBIF-Dataset="c8f70086-b0f8-4063-8fe1-2150048f14d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3823654" ID-ZooBank="1EFCA09E-E2A7-40D4-A791-235784AFF784" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589345150939" checkinUser="plazi" docAuthor="Derycke, Elizabeth G., Gottscho, Andrew D., Mulcahy, Daniel G. & Queiroz, Kevin De" docDate="2020" docId="0391BC4CFFE1655CC7CDFC28FF30FF49" docLanguage="en" docName="zootaxa.4778.1.3.pdf" docOrigin="Zootaxa 4778 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Uma thurmanae Derycke & Gottscho & Mulcahy & Queiroz 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="87" masterDocId="FFA8C434FFEF6548C75AFF84FFEFFFB3" masterDocTitle="A new cryptic species of fringe-toed lizards from southwestern Arizona with a revised taxonomy of the Uma notata species complex (Squamata: Phrynosomatidae)" masterLastPageNumber="100" masterPageNumber="67" pageNumber="81" updateTime="1698839337530" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D8C1B08325BFE0F5CDF78E95B67FE8E7">A new cryptic species of fringe-toed lizards from southwestern Arizona with a revised taxonomy of the Uma notata species complex (Squamata: Phrynosomatidae)</mods:title>
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<mods:name id="41E079EF69399AFFF14D1C15CFAA7D88" type="personal">
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<mods:roleTerm id="61FD2708D2A470C0D90823A7E3F68AF4">Author</mods:roleTerm>
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<mods:namePart id="FE204B12EEB4828F5163BE0B264CF81C">Derycke, Elizabeth G.</mods:namePart>
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<mods:affiliation id="E2AE9CA40608A616E63463766F54B9A3">Department of Environmental Studies, McDaniel College, 2 College Hill, Westminster, MD 21157</mods:affiliation>
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<mods:namePart id="99017F421584618036ADA467C4BFBF85">Gottscho, Andrew D.</mods:namePart>
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<mods:nameIdentifier id="EB639AC4CF0A461A11529542E8374E6C" type="email">GottschoA@si.edu</mods:nameIdentifier>
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<mods:namePart id="CAE36299C5445A2F774F6E3516D57CA3">Mulcahy, Daniel G.</mods:namePart>
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<mods:affiliation id="97204B798BA31DD3D8C191C874BEAA4A">Global Genome Initiative, National Museum of Natural History, Smithsonian Institution, 10 St and Constitution Ave NW, Washington DC 20560</mods:affiliation>
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<mods:namePart id="CD8DDF8DA41661BD3FAB4D6D2562AF38">Queiroz, Kevin De</mods:namePart>
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<mods:title id="E2CF05985C33F4188B928D3D11FE3A24">Zootaxa</mods:title>
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<mods:part id="89750BE2B22F1A2C5C1EC40AC25CAD2E">
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<mods:date id="72B726D49090EF638DD552EE8FCC524D">2020</mods:date>
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<mods:number id="AD7DE1386289880BE9BBCE59A8578087">2020-05-13</mods:number>
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<mods:number id="FB27463B78BAD9B1BF43F4B086A1AD08">4778</mods:number>
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<treatment id="0391BC4CFFE1655CC7CDFC28FF30FF49" ID-DOI="http://doi.org/10.5281/zenodo.3845815" ID-GBIF-Taxon="164128031" ID-Zenodo-Dep="3845815" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49" httpUri="http://treatment.plazi.org/id/0391BC4CFFE1655CC7CDFC28FF30FF49" lastPageId="20" lastPageNumber="87" pageId="14" pageNumber="81">
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<subSubSection id="C3225ED1FFE16546C7CDFC28FDC1FC59" pageId="14" pageNumber="81" type="nomenclature">
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<paragraph id="8B870D5AFFE16546C7CDFC28FE2BFC75" blockId="14.[151,558,939,1038]" box="[151,452,939,966]" pageId="14" pageNumber="81">
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<heading id="D0CFBA36FFE16546C7CDFC28FE2BFC75" bold="true" box="[151,452,939,966]" fontSize="11" level="1" pageId="14" pageNumber="81" reason="1">
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<emphasis id="B94CD148FFE16546C7CDFC28FE2BFC75" bold="true" box="[151,452,939,966]" pageId="14" pageNumber="81">
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<taxonomicName id="4C3876D9FFE16546C7CDFC28FEB0FC75" authority="Derycke & Gottscho & Mulcahy & Queiroz, 2020" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[151,351,939,966]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="thurmanae" status="sp. nov.">
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<emphasis id="B94CD148FFE16546C7CDFC28FEB0FC75" bold="true" box="[151,351,939,966]" italics="true" pageId="14" pageNumber="81">Uma thurmanae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F6C33FFE16546C63CFC28FE2BFC75" box="[358,452,940,966]" pageId="14" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B870D5AFFE16546C7CDFC54FDC1FC59" blockId="14.[151,558,939,1038]" box="[151,558,976,1002]" pageId="14" pageNumber="81">Mohawk Dunes Fringe-toed Lizards</paragraph>
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</subSubSection>
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<subSubSection id="C3225ED1FFE16546C7CDFC70FF16FBBD" box="[151,249,1011,1038]" pageId="14" pageNumber="81" type="description">
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<paragraph id="8B870D5AFFE16546C7CDFC70FF16FBBD" blockId="14.[151,558,939,1038]" box="[151,249,1011,1038]" pageId="14" pageNumber="81">
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<figureCitation id="130311DFFFE16546C7CDFC70FF16FBBD" box="[151,249,1011,1038]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="15.[151,250,1975,2000]" captionStartId-1="17.[151,250,1825,1850]" captionStartId-2="18.[151,250,1536,1561]" captionStartId-3="19.[151,250,1929,1954]" captionTargetBox-0="[151,1435,417,1950]" captionTargetBox-1="[151,1435,669,1800]" captionTargetBox-2="[151,1435,525,1511]" captionTargetBox-3="[151,1435,273,1904]" captionTargetId-0="figure@15.[151,1435,417,1950]" captionTargetId-1="figure@17.[151,1435,669,1800]" captionTargetId-2="figure@18.[151,1435,525,1511]" captionTargetId-3="figure@19.[151,1435,273,1904]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionText-0="FIGURE 5. (A) Dorsal and (B) ventral view of the adult female holotype (USNM 590033) of Uma thurmanae. Photographs by E. Langan and J. Poindexter." captionText-1="FIGURE 6. Holotype (USNM 590033) of Uma thurmanae showing (A) dorsal and (B) ventral head; (C) dorsal right hindfoot and (D) dorsal right forefoot showing toe fringes. Photographs by E. Langan and J. Poindexter." captionText-2="FIGURE 7. Satellite map of collecting localities for the type series of Uma thurmanae. Locations where specimens were collected, denoted by red stars, lie along the Mohawk Dunes, a long strip running northwest to southeast. The lizards are isolated in this dune system by the parallel Mohawk Mountains to the east, the Gila River and Interstate-8 to the north, and alluvial fans to the west and south, ultimately resulting in speciation. Source: “Mohawk Dunes,” 32° 34’ N, 113° 38’ W, Google Earth. Imagery date: May 29, 2016. Accessed: July 3, 2018." captionText-3="FIGURE 8. (A) An individual of U. thurmanae in its natural habitat. (B) Sand dune habitat of U. thurmanae (camera pointed east with the Mohawk Dunes in the midground and the Mohawk Mountains in the background). Photographs by A. Gottscho." figureDoi-0="http://doi.org/10.5281/zenodo.3823664" figureDoi-1="http://doi.org/10.5281/zenodo.3823666" figureDoi-2="http://doi.org/10.5281/zenodo.3823668" figureDoi-3="http://doi.org/10.5281/zenodo.3823670" httpUri-0="https://zenodo.org/record/3823664/files/figure.png" httpUri-1="https://zenodo.org/record/3823666/files/figure.png" httpUri-2="https://zenodo.org/record/3823668/files/figure.png" httpUri-3="https://zenodo.org/record/3823670/files/figure.png" pageId="14" pageNumber="81">Figs 5–8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3225ED1FFE16546C7CDFBB2FCA7FAC3" pageId="14" pageNumber="81" type="materials_examined">
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<paragraph id="8B870D5AFFE16546C7CDFBB2FCFCFB0F" blockId="14.[151,1437,1077,2041]" pageId="14" pageNumber="81">
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<materialsCitation id="3B500707FFE16546C7CDFBB2FCFCFB0F" ID-GBIF-Occurrence="2612525310" collectingDate="2015-03-29" collectionCode="DC" collectorName="Adult & A. D. Gottscho & Mohawk Dunes" country="United States of America" county="Yuma County" latitude="32.694965" location="Interstate" longLatPrecision="1" longitude="-113.81558" municipality="Smithsonian Institution" pageId="14" pageNumber="81" specimenCode="USNM 590033" specimenCount="1" stateProvince="Arizona" typeStatus="holotype">
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<emphasis id="B94CD148FFE16546C7CDFBB2FEE4FBE3" bold="true" box="[151,267,1078,1104]" pageId="14" pageNumber="81">
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<typeStatus id="5483B3F8FFE16546C7CDFBB2FEE9FBE3" box="[151,262,1078,1104]" pageId="14" pageNumber="81" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectorName id="26CD688CFFE16546C64BFBB2FEBEFBE3" box="[273,337,1078,1104]" pageId="14" pageNumber="81">Adult</collectorName>
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female (
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<specimenCode id="DB9EA521FFE16546C6E8FBB2FD8BFBE3" box="[434,612,1077,1104]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590033</specimenCode>
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;
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<figureCitation id="130311DFFFE16546C535FBB2FD3EFBE3" box="[623,721,1077,1104]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="15.[151,250,1975,2000]" captionStartId-1="17.[151,250,1825,1850]" captionTargetBox-0="[151,1435,417,1950]" captionTargetBox-1="[151,1435,669,1800]" captionTargetId-0="figure@15.[151,1435,417,1950]" captionTargetId-1="figure@17.[151,1435,669,1800]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionText-0="FIGURE 5. (A) Dorsal and (B) ventral view of the adult female holotype (USNM 590033) of Uma thurmanae. Photographs by E. Langan and J. Poindexter." captionText-1="FIGURE 6. Holotype (USNM 590033) of Uma thurmanae showing (A) dorsal and (B) ventral head; (C) dorsal right hindfoot and (D) dorsal right forefoot showing toe fringes. Photographs by E. Langan and J. Poindexter." figureDoi-0="http://doi.org/10.5281/zenodo.3823664" figureDoi-1="http://doi.org/10.5281/zenodo.3823666" httpUri-0="https://zenodo.org/record/3823664/files/figure.png" httpUri-1="https://zenodo.org/record/3823666/files/figure.png" pageId="14" pageNumber="81">Figs 5–6</figureCitation>
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) collected on
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<date id="FF862B9AFFE16546C42AFBB2FBF4FBE3" box="[880,1051,1077,1104]" pageId="14" pageNumber="81" value="2015-03-29">
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<collectingDate id="EFC2D272FFE16546C42AFBB2FBF4FBE3" box="[880,1051,1077,1104]" pageId="14" pageNumber="81" value="2015-03-29">29 March 2015</collectingDate>
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</date>
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by
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<collectorName id="26CD688CFFE16546C31EFBB2FB01FBE3" box="[1092,1262,1078,1105]" pageId="14" pageNumber="81">A. D. Gottscho</collectorName>
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at 17:28 hours. Type locality: the north end of the
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<collectorName id="26CD688CFFE16546C545FBDEFD3CFBC6" box="[543,723,1114,1141]" pageId="14" pageNumber="81">Mohawk Dunes</collectorName>
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(~
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south of
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<location id="8EE75B81FFE16546C4F6FBDFFBFBFBC6" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49:8EE75B81FFE16546C4F6FBDFFBFBFBC6" box="[940,1044,1115,1141]" country="United States of America" county="Yuma County" latitude="32.694965" longLatPrecision="1" longitude="-113.81558" municipality="Smithsonian Institution" name="Interstate" pageId="14" pageNumber="81" stateProvince="Arizona">Interstate</location>
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8),
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<collectingCounty id="62E675D6FFE16546C31AFBDFFB32FBC7" box="[1088,1245,1114,1141]" pageId="14" pageNumber="81">Yuma County</collectingCounty>
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,
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<collectingRegion id="49FCC3B8FFE16546C3BDFBDEFAABFBC7" box="[1255,1348,1114,1140]" country="United States of America" name="Arizona" pageId="14" pageNumber="81">Arizona</collectingRegion>
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,
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<collectingCountry id="F32F4DCAFFE16546C20AFBDEFE8FFB2B" name="United States of America" pageId="14" pageNumber="81">United States of America</collectingCountry>
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(
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<geoCoordinate id="EE0C6B9DFFE16546C62AFBFAFDEAFB2A" box="[368,517,1150,1177]" degrees="32" direction="north" minutes="41.698" orientation="latitude" pageId="14" pageNumber="81" precision="1" value="32.694965">32°41.698 N</geoCoordinate>
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;
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<geoCoordinate id="EE0C6B9DFFE16546C548FBFAFD2FFB2A" box="[530,704,1149,1177]" degrees="113" direction="west" minutes="48.935" orientation="longitude" pageId="14" pageNumber="81" precision="1" value="-113.81558">113° 48.935 W</geoCoordinate>
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; WGS-84 datum).
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<location id="8EE75B81FFE16546C4CCFBFAFBE7FB2B" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49:8EE75B81FFE16546C4CCFBFAFBE7FB2B" box="[918,1032,1150,1176]" country="United States of America" county="Yuma County" latitude="32.694965" longLatPrecision="1" longitude="-113.81558" municipality="Smithsonian Institution" name="Deposited" pageId="14" pageNumber="81" stateProvince="Arizona">Deposited</location>
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at the
|
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<location id="8EE75B81FFE16546C30CFBFAFF03FB0F" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49:8EE75B81FFE16546C30CFBFAFF03FB0F" country="United States of America" county="Yuma County" latitude="32.694965" longLatPrecision="1" longitude="-113.81558" municipality="Smithsonian Institution" name="National Museum of Natural History" pageId="14" pageNumber="81" stateProvince="Arizona">National Museum of Natural History</location>
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,
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<collectingMunicipality id="6BE39720FFE16546C7ADFB26FDEFFB0F" box="[247,512,1186,1212]" pageId="14" pageNumber="81">Smithsonian Institution</collectingMunicipality>
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,
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<collectingRegion id="49FCC3B8FFE16546C551FB26FD7CFB0F" box="[523,659,1186,1212]" country="United States of America" name="Washington" pageId="14" pageNumber="81">Washington</collectingRegion>
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,
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<collectionCode id="ED29959FFFE16546C5C4FB26FD27FB0F" box="[670,712,1186,1212]" pageId="14" pageNumber="81">DC</collectionCode>
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, USA.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B870D5AFFE16546C79DFB42FCA7FAC3" blockId="14.[151,1437,1077,2041]" pageId="14" pageNumber="81">
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<materialsCitation id="3B500707FFE16546C79DFB42FD67FB53" ID-GBIF-Occurrence="2612525304" box="[199,648,1221,1248]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590034" specimenCount="1" typeStatus="paratype">
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<emphasis id="B94CD148FFE16546C79DFB42FED0FB53" bold="true" box="[199,319,1222,1248]" pageId="14" pageNumber="81">
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<typeStatus id="5483B3F8FFE16546C79DFB42FED0FB53" box="[199,319,1222,1248]" pageId="14" pageNumber="81" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.
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||
<specimenCode id="DB9EA521FFE16546C611FB42FE19FB53" box="[331,502,1221,1248]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590034</specimenCode>
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||
adult female
|
||
</materialsCitation>
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,
|
||
<materialsCitation id="3B500707FFE16546C5CBFB42FC57FB53" ID-GBIF-Occurrence="2612525307" box="[657,952,1221,1248]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590035" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C5CBFB42FCD3FB53" box="[657,828,1221,1248]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590035</specimenCode>
|
||
adult male
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C498FB42FB07FB53" ID-GBIF-Occurrence="2612525308" box="[962,1256,1221,1248]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590036" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C498FB42FB83FB53" box="[962,1132,1221,1248]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590036</specimenCode>
|
||
adult male
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C3A8FB42FEA7FAB7" ID-GBIF-Occurrence="2612525303" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590037" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C3A8FB42FA73FB53" box="[1266,1436,1221,1248]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590037</specimenCode>
|
||
juvenile female
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C60EFB6EFD79FAB7" ID-GBIF-Occurrence="2612525309" box="[340,662,1257,1284]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590038" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C60EFB6EFDEFFAB7" box="[340,512,1257,1284]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590038</specimenCode>
|
||
adult female
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C5F8FB6EFBE8FAB7" ID-GBIF-Occurrence="2612525306" box="[674,1031,1257,1284]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590039" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C5F8FB6EFCA0FAB7" box="[674,847,1257,1284]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590039</specimenCode>
|
||
juvenile female
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C349FB6EFAD0FAB7" ID-GBIF-Occurrence="2612525311" box="[1043,1343,1257,1284]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590040" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C349FB6EFB50FAB7" box="[1043,1215,1257,1284]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590040</specimenCode>
|
||
adult male
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C211FB6EFE83FA9B" ID-GBIF-Occurrence="2612525302" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590041" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C211FB6EFF04FA9B" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590041</specimenCode>
|
||
adult male
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C623FA8AFD0FFA9B" ID-GBIF-Occurrence="2612525312" box="[377,736,1293,1320]" collectionCode="USNM" pageId="14" pageNumber="81" specimenCode="USNM 590042" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C623FA8AFDC9FA9B" box="[377,550,1293,1320]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590042</specimenCode>
|
||
juvenile female
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C5B6FA8AFD26FAC3" ID-GBIF-Occurrence="2612525301" collectingDate="2016-03-25" collectingDateMax="2016-03-26" collectingDateMin="2016-03-25" collectionCode="USNM" collectorName="A. D. Gottscho & J. Rabbers & S. Harrington" country="United States of America" county="Yuma County" location="Deposited" municipality="Mohawk Dunes" pageId="14" pageNumber="81" specimenCode="USNM 590043" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
|
||
<specimenCode id="DB9EA521FFE16546C5B6FA8AFC76FA9B" box="[748,921,1293,1320]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="81" type="Museum">USNM 590043</specimenCode>
|
||
adult male. Collected
|
||
<date id="FF862B9AFFE16546C3C0FA89FA73FA9B" box="[1178,1436,1293,1320]" pageId="14" pageNumber="81" value="2016-03-25" valueMax="2016-03-26" valueMin="2016-03-25">
|
||
<collectingDate id="EFC2D272FFE16546C3C0FA89FA73FA9B" box="[1178,1436,1293,1320]" pageId="14" pageNumber="81" value="2016-03-25" valueMax="2016-03-26" valueMin="2016-03-25">25 and 26 March 2016</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26CD688CFFE16546C7E0FAB6FE8FFAFF" box="[186,352,1330,1356]" pageId="14" pageNumber="81">A.D. Gottscho</collectorName>
|
||
,
|
||
<collectorName id="26CD688CFFE16546C636FAB7FE0AFAFF" box="[364,485,1330,1357]" pageId="14" pageNumber="81">J. Rabbers</collectorName>
|
||
, and
|
||
<collectorName id="26CD688CFFE16546C57BFAB6FD54FAFF" box="[545,699,1330,1356]" pageId="14" pageNumber="81">S. Harrington</collectorName>
|
||
from
|
||
<collectingMunicipality id="6BE39720FFE16546C45BFAB6FC59FAFE" box="[769,950,1330,1357]" pageId="14" pageNumber="81">Mohawk Dunes</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62E675D6FFE16546C498FAB7FBB0FAFF" box="[962,1119,1330,1357]" pageId="14" pageNumber="81">Yuma County</collectingCounty>
|
||
, AZ,
|
||
<collectingCountry id="F32F4DCAFFE16546C3C7FAB6FB36FAFF" box="[1181,1241,1330,1356]" name="United States of America" pageId="14" pageNumber="81">USA</collectingCountry>
|
||
.
|
||
<location id="8EE75B81FFE16546C3BEFAB6FAB9FAFF" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49:8EE75B81FFE16546C3BEFAB6FAB9FAFF" box="[1252,1366,1330,1356]" country="United States of America" county="Yuma County" municipality="Mohawk Dunes" name="Deposited" pageId="14" pageNumber="81" stateProvince="Washington">Deposited</location>
|
||
at the
|
||
<location id="8EE75B81FFE16546C7CDFAD2FDD9FAC3" LSID="urn:lsid:plazi:treatment:0391BC4CFFE1655CC7CDFC28FF30FF49:8EE75B81FFE16546C7CDFAD2FDD9FAC3" box="[151,566,1366,1393]" country="United States of America" county="Yuma County" municipality="Mohawk Dunes" name="National Museum of Natural History" pageId="14" pageNumber="81" stateProvince="Washington">National Museum of Natural History</location>
|
||
,
|
||
<collectingRegion id="49FCC3B8FFE16546C51BFAD2FD26FAC3" box="[577,713,1366,1392]" country="United States of America" name="Washington" pageId="14" pageNumber="81">Washington</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B500707FFE16546C58EFAD2FCAAFAC3" ID-GBIF-Occurrence="2612525305" box="[724,837,1366,1392]" collectionCode="DC" country="United States of America" location="USA" pageId="14" pageNumber="81" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED29959FFFE16546C58EFAD2FD11FAC3" box="[724,766,1366,1392]" pageId="14" pageNumber="81">DC</collectionCode>
|
||
,
|
||
<collectingCountry id="F32F4DCAFFE16546C453FAD2FCAAFAC3" box="[777,837,1366,1392]" name="United States of America" pageId="14" pageNumber="81">USA</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3225ED1FFE16546C79DFAFDFEDBF997" pageId="14" pageNumber="81" type="diagnosis">
|
||
<paragraph id="8B870D5AFFE16546C79DFAFDFEDBF997" blockId="14.[151,1437,1077,2041]" pageId="14" pageNumber="81">
|
||
<emphasis id="B94CD148FFE16546C79DFAFDFEADFA27" bold="true" box="[199,322,1401,1428]" pageId="14" pageNumber="81">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C3876D9FFE16546C612FAFFFE11FA27" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[328,510,1402,1428]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFE16546C612FAFFFE11FA27" box="[328,510,1402,1428]" italics="true" pageId="14" pageNumber="81">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
is a morphologically cryptic species that doesn’t exhibit any known fixed morphological differences from other species in the complex, but it can be distinguished from those other species both by statistical differences in morphological characters (
|
||
<tableCitation id="C6BA38E1FFE16546C58BFA46FCC7FA6F" box="[721,808,1473,1500]" captionStart="TABLE 5" captionStartId="8.[151,239,153,177]" captionText="TABLE 5. Post-hoc pairwise comparisons of morphological characters that showed significant p-values in either the ANOVA (Tukey) or Kruskal-Wallis (Nemenyi) tests (see Table 4). Values in bold indicate a significant pairwise difference (p <0.05). See Appendix 2 for character codes." pageId="14" pageNumber="81">Table 5</tableCitation>
|
||
,
|
||
<figureCitation id="130311DFFFE16546C46EFA46FC9AFA6F" box="[820,885,1474,1500]" captionStart="FIGURE 3" captionStartId="12.[151,250,1017,1042]" captionTargetBox="[162,1428,181,988]" captionTargetId="figure@12.[158,1428,181,991]" captionTargetPageId="12" captionText="FIGURE 3. Box-and-whisker plots for discrete morphological characters that exhibited significant pairwise differences between OTUs. See Appendix 2 for details of how the characters were measured. SPLB=Supralabial Scales; SPCIL=Supraciliary Scales; INS=Internasal Scales; FP=Femoral Pores." figureDoi="http://doi.org/10.5281/zenodo.3823660" httpUri="https://zenodo.org/record/3823660/files/figure.png" pageId="14" pageNumber="81">Fig. 3</figureCitation>
|
||
) and in multivariate morphospace (
|
||
<figureCitation id="130311DFFFE16546C25EFA46FAA6FA6F" box="[1284,1353,1474,1500]" captionStart="FIGURE 4" captionStartId="13.[151,250,1852,1877]" captionTargetBox="[321,1269,189,1819]" captionTargetId="figure@13.[312,1276,181,1827]" captionTargetPageId="13" captionText="FIGURE 4. Locations of Uma specimens in the morphospace defined by the first two axes of a Discriminant Function Analysis of morphological characters. Top: when specimens with broken tails are included and TL excluded (n=57), U. inornata can still be distinguished from other OTUs, but U. notata overlaps considerably with U. rufopunctata and U. cowlesi overlaps considerably with U. sp. Bottom: analysis of only the specimens with intact tails such that tail length (TL) could be measured (n=32) allows each OTU to be distinguished." figureDoi="http://doi.org/10.5281/zenodo.3823662" httpUri="https://zenodo.org/record/3823662/files/figure.png" pageId="14" pageNumber="81">Fig. 4</figureCitation>
|
||
). It can also be distinguished from all other members of the
|
||
<taxonomicName id="4C3876D9FFE16546C584FA63FCA7F9B3" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[734,840,1511,1536]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFE16546C584FA63FCA7F9B3" box="[734,840,1511,1536]" italics="true" pageId="14" pageNumber="81">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex by nucleotide substitutions in three mtDNA loci (
|
||
<tableCitation id="C6BA38E1FFE16546C78AF98EFEC9F997" box="[208,294,1546,1572]" captionStart="TABLE 8" captionStartId="16.[151,239,153,177]" captionText="TABLE 8. Alignment positions of 26 fixed single nucleotide polymorphisms in three mitochondrial genes (ATPase 6, cytochrome b, and COI) that diagnose U. sp. from the rest of the U. notata complex. See Appendix 1 for GenBank accession numbers." pageId="14" pageNumber="81">Table 8</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3225ED1FFE16559C79DF9A9FE15FD34" lastPageId="17" lastPageNumber="84" pageId="14" pageNumber="81" type="description">
|
||
<paragraph id="8B870D5AFFE16547C79DF9A9FDCCFE38" blockId="14.[151,1437,1077,2041]" lastBlockId="15.[151,1437,153,395]" lastPageId="15" lastPageNumber="82" pageId="14" pageNumber="81">
|
||
<emphasis id="B94CD148FFE16546C79DF9A9FE32F9FB" bold="true" box="[199,477,1581,1608]" pageId="14" pageNumber="81">
|
||
Description of
|
||
<typeStatus id="5483B3F8FFE16546C62CF9AAFE32F9FB" box="[374,477,1582,1608]" pageId="14" pageNumber="81" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult female; SVL=
|
||
<quantity id="4CC0A0BFFFE16546C595F9AAFCCFF9FA" box="[719,800,1582,1609]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="81" unit="mm" value="68.0">68 mm</quantity>
|
||
; tail length=
|
||
<quantity id="4CC0A0BFFFE16546C4F6F9AAFC12F9FA" box="[940,1021,1582,1609]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="81" unit="mm" value="77.0">77 mm</quantity>
|
||
; head width=
|
||
<quantity id="4CC0A0BFFFE16546C3CFF9AAFB0BF9FA" box="[1173,1252,1582,1609]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="81" unit="mm" value="12.0">12 mm</quantity>
|
||
; head length=
|
||
<quantity id="4CC0A0BFFFE16546C2D9F9A9FF29F9DE" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="81" unit="mm" value="15.0">15 mm</quantity>
|
||
; head height=
|
||
<quantity id="4CC0A0BFFFE16546C631F9D6FE40F9DE" box="[363,431,1618,1645]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="81" unit="mm" value="8.0">8 mm</quantity>
|
||
. Scalation: dorsal head scales smooth, slightly raised; parietal eye present, centrally located in a circular interparietal scale; enlarged post-parietal scales present with three scales separating post-parietal scales from inter-parietal scale; five enlarged anterior auricular scales; four superciliaries; 12 lower festoons; three suboculars, with the third being substantially larger and more elongated than the first two; five enlarged anterior auricular scales; two canthals; four internasal scales; supralabials elongated, and strongly keeled, increasing in size posteriorly; eight from rostral scale to a point below middle of eye; 11 infralabials from mental scale to a point below middle of eye, 15 to corner of mouth; rostral single; five postrostrals, four in broad contact with the rostral scale, one in narrow contact; one mental; one postmental one; gular scales extend between anteriormost sublabial scales to contact postmental scale; gular region covered in small scales varying in shape from hexagonal to rectangular to rhombic to almost round; gular fold present, with enlarged scales; dorsal body scales small, flat, granular, uniform, with no middoral row of enlarged scales; ventral scales larger than dorsal scales, smooth, flat, and uniform; dorsal limb scales slightly larger than dorsal trunk scales; ventral limb scales smaller, more rounded than ventral body scales; anterior scales on front limbs enlarged and strongly keeled; anterior and posterior lateral scales on hind limbs weakly keeled and enlarged; 27 femoral pores on each hind leg in one row; fringes present on all digits, both anteriorly and posteriorly, but appearing smaller and more symmetrical on digits I, II, and V versus larger and more asymmetrical, with the fringes on the posterior edge larger than those on the anterior edge, on digits III and IV; largest fringe scales on the posterior edge of pedal digit IV; 31 fringe scales on left pedal digit IV, 30 on right pedal digit IV. Tail flattened ventrally, with whorls of enlarged scales; four rows of smaller scales separate the eighth and ninth whorls of enlarged scales ventrally.
|
||
</paragraph>
|
||
<caption id="DF475DD2FFE06547C7CDF833FE34F847" ID-DOI="http://doi.org/10.5281/zenodo.3823664" ID-Zenodo-Dep="3823664" httpUri="https://zenodo.org/record/3823664/files/figure.png" pageId="15" pageNumber="82" startId="15.[151,250,1975,2000]" targetBox="[151,1435,417,1950]" targetPageId="15">
|
||
<paragraph id="8B870D5AFFE06547C7CDF833FE34F847" blockId="15.[151,1437,1975,2036]" pageId="15" pageNumber="82">
|
||
<emphasis id="B94CD148FFE06547C7CDF833FEFBF863" bold="true" box="[151,276,1975,2000]" pageId="15" pageNumber="82">FIGURE 5.</emphasis>
|
||
(A) Dorsal and (B) ventral view of the adult female holotype (USNM 590033) of
|
||
<emphasis id="B94CD148FFE06547C333F83DFAF8F863" box="[1129,1303,1976,2000]" italics="true" pageId="15" pageNumber="82">
|
||
<taxonomicName id="4C3876D9FFE06547C333F83DFAFCF863" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[1129,1299,1976,2000]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="82" phylum="Chordata" rank="species" species="thurmanae">Uma thurmanae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Photographs by E. Langan and J. Poindexter.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF475DD2FFFF6558C7CDFF1DFEC1FF49" pageId="16" pageNumber="83">
|
||
<paragraph id="8B870D5AFFFF6558C7CDFF1DFEC1FF49" blockId="16.[151,1437,153,250]" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C7CDFF1DFEEAFF01" bold="true" box="[151,261,153,178]" pageId="16" pageNumber="83">TABLE 8</emphasis>
|
||
. Alignment positions of 26 fixed single nucleotide polymorphisms in three mitochondrial genes (ATPase 6, cytochrome b, and COI) that diagnose
|
||
<emphasis id="B94CD148FFFF6558C568FF3AFDA2FF66" box="[562,589,190,213]" italics="true" pageId="16" pageNumber="83">U.</emphasis>
|
||
sp. from the rest of the
|
||
<taxonomicName id="4C3876D9FFFF6558C412FF3AFC43FF66" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[840,940,190,213]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFFF6558C412FF3AFC43FF66" box="[840,940,190,213]" italics="true" pageId="16" pageNumber="83">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex. See Appendix 1 for GenBank accession numbers.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B870D5AFFFF6558C59AFE8DFC73FE71" pageId="16" pageNumber="83">
|
||
<table id="F938FFFAFFFF9AB7C7ABFE8DFB58FE71" box="[241,1207,265,450]" gridcols="4" gridrows="5" pageId="16" pageNumber="83">
|
||
<tr id="35080F18FFFF9AB7C7ABFE8DFB58FE91" box="[241,1207,265,290]" gridrow="0" pageId="16" pageNumber="83" rowspan-0="1" rowspan-1="1" rowspan-3="1">
|
||
<th id="76D96664FFFF9AB7C5CCFE8DFC73FE91" box="[662,924,265,290]" gridcol="2" gridrow="0" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C59AFE8DFC9DFE91" bold="true" box="[704,882,265,290]" pageId="16" pageNumber="83">ATPase (599 bp)</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFEB5FB58FEF9" box="[241,1207,305,330]" gridrow="1" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFEB5FEA7FEF9" box="[241,328,305,330]" gridcol="0" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C7ABFEB5FEA7FEF9" bold="true" box="[241,328,305,330]" pageId="16" pageNumber="83">Position</emphasis>
|
||
</th>
|
||
<td id="76D96664FFFF9AB7C55CFEB5FDB2FEF9" box="[518,605,305,330]" gridcol="1" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C55CFEB6FDB2FEFA" bold="true" box="[518,605,305,329]" italics="true" pageId="16" pageNumber="83">
|
||
<taxonomicName id="4C3876D9FFFF6558C55CFEB6FDD6FEFA" authorityName=", Norris" authorityYear="1958" box="[518,569,306,329]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="genus">Uma</taxonomicName>
|
||
sp.
|
||
</emphasis>
|
||
</td>
|
||
<td id="76D96664FFFF9AB7C5CCFEB5FB58FEF9" box="[662,1207,305,330]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C4FEFEB5FB58FEFA" bold="true" box="[932,1207,305,330]" pageId="16" pageNumber="83">
|
||
Rest of
|
||
<taxonomicName id="4C3876D9FFFF6558C4ACFEB6FBB8FEFA" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[1014,1111,306,329]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFFF6558C4ACFEB6FBB8FEFA" bold="true" box="[1014,1111,306,329]" italics="true" pageId="16" pageNumber="83">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFEDEFB58FEC1" box="[241,1207,346,370]" gridrow="2" pageId="16" pageNumber="83" rowspan-2="1">
|
||
<th id="76D96664FFFF9AB7C7ABFEDEFEA7FEC1" box="[241,328,346,370]" gridcol="0" gridrow="2" pageId="16" pageNumber="83">73</th>
|
||
<td id="76D96664FFFF9AB7C55CFEDEFDB2FEC1" box="[518,605,346,370]" gridcol="1" gridrow="2" pageId="16" pageNumber="83">C</td>
|
||
<td id="76D96664FFFF9AB7C37FFEDEFB58FEC1" box="[1061,1207,346,370]" gridcol="3" gridrow="2" pageId="16" pageNumber="83">T</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFE06FB58FE29" box="[241,1207,386,410]" gridrow="3" pageId="16" pageNumber="83" rowspan-2="1">
|
||
<th id="76D96664FFFF9AB7C7ABFE06FEA7FE29" box="[241,328,386,410]" gridcol="0" gridrow="3" pageId="16" pageNumber="83">298</th>
|
||
<td id="76D96664FFFF9AB7C55CFE06FDB2FE29" box="[518,605,386,410]" gridcol="1" gridrow="3" pageId="16" pageNumber="83">C</td>
|
||
<td id="76D96664FFFF9AB7C37FFE06FB58FE29" box="[1061,1207,386,410]" gridcol="3" gridrow="3" pageId="16" pageNumber="83">A</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFE2DFB58FE71" box="[241,1207,425,450]" gridrow="4" pageId="16" pageNumber="83" rowspan-0="1" rowspan-1="1" rowspan-3="1">
|
||
<td id="76D96664FFFF9AB7C5CCFE2DFC73FE71" box="[662,924,425,450]" gridcol="2" gridrow="4" pageId="16" pageNumber="83">Cytochrome b (1031 bp)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C7ABFE55FB58FE59" blockId="16.[241,1207,465,1093]" box="[241,1207,465,490]" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C7ABFE55FB58FE59" bold="true" box="[241,1207,465,490]" pageId="16" pageNumber="83">
|
||
Position
|
||
<taxonomicName id="4C3876D9FFFF6558C55CFE56FDD6FE5A" authorityName=", Norris" authorityYear="1958" box="[518,569,466,489]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94CD148FFFF6558C55CFE56FDD6FE5A" bold="true" box="[518,569,466,489]" italics="true" pageId="16" pageNumber="83">Uma</emphasis>
|
||
</taxonomicName>
|
||
sp. Rest of
|
||
<taxonomicName id="4C3876D9FFFF6558C4ACFE56FBB8FE5A" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[1014,1111,466,489]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFFF6558C4ACFE56FBB8FE5A" bold="true" box="[1014,1111,466,489]" italics="true" pageId="16" pageNumber="83">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C64AFE7EFBD9FDA1" blockId="16.[241,1207,465,1093]" box="[272,1078,506,530]" pageId="16" pageNumber="83">20 T C</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C64AFDA6FBDAFD8A" blockId="16.[241,1207,465,1093]" box="[272,1077,546,570]" pageId="16" pageNumber="83">23 C T</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C64AFDCEFBD9FDD1" blockId="16.[241,1207,465,1093]" box="[272,1078,586,610]" pageId="16" pageNumber="83">89 A C</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FDF6FBABFD39" blockId="16.[241,1207,465,1093]" box="[266,1092,626,650]" pageId="16" pageNumber="83">122 C A/G</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FD1FFBD9FD00" blockId="16.[241,1207,465,1093]" box="[266,1078,667,691]" pageId="16" pageNumber="83">203 T C</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FD47FBD8FD68" blockId="16.[241,1207,465,1093]" box="[266,1079,707,731]" pageId="16" pageNumber="83">218 A G</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FD6FFBD8FCB0" blockId="16.[241,1207,465,1093]" box="[266,1079,747,771]" pageId="16" pageNumber="83">383 G A</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FC97FBD9FC98" blockId="16.[241,1207,465,1093]" box="[266,1078,787,812]" pageId="16" pageNumber="83">455 T C</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FCBFFBD8FCE7" blockId="16.[241,1207,465,1093]" box="[266,1079,827,852]" pageId="16" pageNumber="83">485 G A</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FCE7FBDAFCC8" blockId="16.[241,1207,465,1093]" box="[266,1077,867,892]" pageId="16" pageNumber="83">615 C T</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FC08FBD8FC17" blockId="16.[241,1207,465,1093]" box="[266,1079,908,932]" pageId="16" pageNumber="83">773 C A</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FC30FBD8FC7F" blockId="16.[241,1207,465,1093]" box="[266,1079,948,972]" pageId="16" pageNumber="83">920 G A</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FC58FBDAFC47" blockId="16.[241,1207,465,1093]" box="[266,1077,988,1013]" pageId="16" pageNumber="83">962 C T</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C650FB80FBD9FBAE" blockId="16.[241,1207,465,1093]" box="[266,1078,1028,1053]" pageId="16" pageNumber="83">975 T C</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C659FBA8FBDAFBF7" blockId="16.[241,1207,465,1093]" box="[259,1077,1068,1093]" pageId="16" pageNumber="83">1016 C T</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C58AFBD0FBD9FA4C" pageId="16" pageNumber="83">
|
||
<table id="F938FFFAFFFF9AB7C7ABFBD0FB58FA4C" box="[241,1207,1108,1535]" gridcols="3" gridrows="11" pageId="16" pageNumber="83">
|
||
<tr id="35080F18FFFF9AB7C7ABFBD0FB58FBDE" box="[241,1207,1108,1133]" gridrow="0" pageId="16" pageNumber="83" rowspan-0="1">
|
||
<th id="76D96664FFFF9AB7C55CFBD0FB58FBDE" box="[518,1207,1108,1133]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C58AFBD0FC8DFBDE" bold="true" box="[720,866,1108,1133]" pageId="16" pageNumber="83">CO1 (657 bp)</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFBF8FB58FB26" box="[241,1207,1148,1173]" gridrow="1" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFBF8FEA7FB26" box="[241,328,1148,1173]" gridcol="0" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C7ABFBF8FEA7FB26" bold="true" box="[241,328,1148,1173]" pageId="16" pageNumber="83">Position</emphasis>
|
||
</th>
|
||
<td id="76D96664FFFF9AB7C55CFBF8FDB2FB26" box="[518,605,1148,1173]" gridcol="1" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C55CFBF9FDB2FB26" bold="true" box="[518,605,1149,1173]" italics="true" pageId="16" pageNumber="83">
|
||
<taxonomicName id="4C3876D9FFFF6558C55CFBF9FDD6FB27" authorityName=", Norris" authorityYear="1958" box="[518,569,1149,1172]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="genus">Uma</taxonomicName>
|
||
sp.
|
||
</emphasis>
|
||
</td>
|
||
<td id="76D96664FFFF9AB7C4FEFBF8FB58FB26" box="[932,1207,1148,1173]" gridcol="2" gridrow="1" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C4FEFBF9FB58FB26" bold="true" box="[932,1207,1148,1173]" pageId="16" pageNumber="83">
|
||
Rest of
|
||
<taxonomicName id="4C3876D9FFFF6558C4ACFBF9FBB8FB27" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[1014,1111,1149,1172]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="83" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFFF6558C4ACFBF9FBB8FB27" bold="true" box="[1014,1111,1149,1172]" italics="true" pageId="16" pageNumber="83">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFB21FB58FB0E" box="[241,1207,1189,1213]" gridrow="2" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFB21FEA7FB0E" box="[241,328,1189,1213]" gridcol="0" gridrow="2" pageId="16" pageNumber="83">42</th>
|
||
<td id="76D96664FFFF9AB7C55CFB21FDB2FB0E" box="[518,605,1189,1213]" gridcol="1" gridrow="2" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFB21FB58FB0E" box="[932,1207,1189,1213]" gridcol="2" gridrow="2" pageId="16" pageNumber="83">A</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFB49FB58FB56" box="[241,1207,1229,1253]" gridrow="3" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFB49FEA7FB56" box="[241,328,1229,1253]" gridcol="0" gridrow="3" pageId="16" pageNumber="83">84</th>
|
||
<td id="76D96664FFFF9AB7C55CFB49FDB2FB56" box="[518,605,1229,1253]" gridcol="1" gridrow="3" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFB49FB58FB56" box="[932,1207,1229,1253]" gridcol="2" gridrow="3" pageId="16" pageNumber="83">C</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFB71FB58FABD" box="[241,1207,1269,1294]" gridrow="4" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFB71FEA7FABD" box="[241,328,1269,1294]" gridcol="0" gridrow="4" pageId="16" pageNumber="83">258</th>
|
||
<td id="76D96664FFFF9AB7C55CFB71FDB2FABD" box="[518,605,1269,1294]" gridcol="1" gridrow="4" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFB71FB58FABD" box="[932,1207,1269,1294]" gridcol="2" gridrow="4" pageId="16" pageNumber="83">C</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFA9AFB58FA85" box="[241,1207,1310,1334]" gridrow="5" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFA9AFEA7FA85" box="[241,328,1310,1334]" gridcol="0" gridrow="5" pageId="16" pageNumber="83">417</th>
|
||
<td id="76D96664FFFF9AB7C55CFA9AFDB2FA85" box="[518,605,1310,1334]" gridcol="1" gridrow="5" pageId="16" pageNumber="83">G</td>
|
||
<td id="76D96664FFFF9AB7C4FEFA9AFB58FA85" box="[932,1207,1310,1334]" gridcol="2" gridrow="5" pageId="16" pageNumber="83">A</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFAC2FB58FAED" box="[241,1207,1350,1374]" gridrow="6" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFAC2FEA7FAED" box="[241,328,1350,1374]" gridcol="0" gridrow="6" pageId="16" pageNumber="83">420</th>
|
||
<td id="76D96664FFFF9AB7C55CFAC2FDB2FAED" box="[518,605,1350,1374]" gridcol="1" gridrow="6" pageId="16" pageNumber="83">G</td>
|
||
<td id="76D96664FFFF9AB7C4FEFAC2FB58FAED" box="[932,1207,1350,1374]" gridcol="2" gridrow="6" pageId="16" pageNumber="83">A</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFAE9FB58FA35" box="[241,1207,1389,1414]" gridrow="7" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFAE9FEA7FA35" box="[241,328,1389,1414]" gridcol="0" gridrow="7" pageId="16" pageNumber="83">456</th>
|
||
<td id="76D96664FFFF9AB7C55CFAE9FDB2FA35" box="[518,605,1389,1414]" gridcol="1" gridrow="7" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFAE9FB58FA35" box="[932,1207,1389,1414]" gridcol="2" gridrow="7" pageId="16" pageNumber="83">C</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFA12FB58FA1C" box="[241,1207,1430,1455]" gridrow="8" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFA12FEA7FA1C" box="[241,328,1430,1455]" gridcol="0" gridrow="8" pageId="16" pageNumber="83">516</th>
|
||
<td id="76D96664FFFF9AB7C55CFA12FDB2FA1C" box="[518,605,1430,1455]" gridcol="1" gridrow="8" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFA12FB58FA1C" box="[932,1207,1430,1455]" gridcol="2" gridrow="8" pageId="16" pageNumber="83">C</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFA3AFB58FA64" box="[241,1207,1470,1495]" gridrow="9" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFA3AFEA7FA64" box="[241,328,1470,1495]" gridcol="0" gridrow="9" pageId="16" pageNumber="83">543</th>
|
||
<td id="76D96664FFFF9AB7C55CFA3AFDB2FA64" box="[518,605,1470,1495]" gridcol="1" gridrow="9" pageId="16" pageNumber="83">A</td>
|
||
<td id="76D96664FFFF9AB7C4FEFA3AFB58FA64" box="[932,1207,1470,1495]" gridcol="2" gridrow="9" pageId="16" pageNumber="83">G</td>
|
||
</tr>
|
||
<tr id="35080F18FFFF9AB7C7ABFA62FB58FA4C" box="[241,1207,1510,1535]" gridrow="10" pageId="16" pageNumber="83">
|
||
<th id="76D96664FFFF9AB7C7ABFA62FEA7FA4C" box="[241,328,1510,1535]" gridcol="0" gridrow="10" pageId="16" pageNumber="83">567</th>
|
||
<td id="76D96664FFFF9AB7C55CFA62FDB2FA4C" box="[518,605,1510,1535]" gridcol="1" gridrow="10" pageId="16" pageNumber="83">T</td>
|
||
<td id="76D96664FFFF9AB7C4FEFA62FB58FA4C" box="[932,1207,1510,1535]" gridcol="2" gridrow="10" pageId="16" pageNumber="83">C</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B870D5AFFFF6558C79DF9AAFD0DF8DA" blockId="16.[151,1437,1582,2041]" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C79DF9AAFDADF9FA" bold="true" box="[199,578,1582,1609]" pageId="16" pageNumber="83">Coloration and pattern (in life).</emphasis>
|
||
Dorsum with an ocellated pattern. The individual ocelli are roughly circular beige markings, each with a rusty central spot and radiating lines, surrounded by black. The lateral ocelli are aligned in rows so that their black borders form broken black longitudinal lines between the rows of ocelli. Underside of torso cream-colored and unblemished except for two black ventrolateral blotches slightly anterior to the midpoint between axilla and groin. Gular region marked by five black diagonal lines oriented anterolateral to posteromedial. Limbs lighter in coloration, consisting of a light-colored background with dark brown and gray spots. Eight black markings present ventrally on the tail, beginning
|
||
<quantity id="4CC0A0BFFFFF6558C5E3F882FCC5F892" box="[697,810,1798,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.765" pageId="16" pageNumber="83" unit="mm" value="17.65">17.65 mm</quantity>
|
||
posterior to the cloaca as a small cluster of black dots in the center of the tail, and increasing in length and width posteriorly. In preservative, dorsal coloration is yellowed, with muted contrast between ocelli and background.
|
||
</paragraph>
|
||
<paragraph id="8B870D5AFFFF6559C79DF8F6FE15FD34" blockId="16.[151,1437,1582,2041]" lastBlockId="17.[151,1437,152,647]" lastPageId="17" lastPageNumber="84" pageId="16" pageNumber="83">
|
||
<emphasis id="B94CD148FFFF6558C79DF8F6FED7F83E" bold="true" box="[199,312,1906,1933]" pageId="16" pageNumber="83">Variation</emphasis>
|
||
.
|
||
<typeStatus id="5483B3F8FFFF6558C61FF8F7FE5DF83E" box="[325,434,1907,1933]" pageId="16" pageNumber="83" type="paratype">Paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="5483B3F8FFFF6558C520F8F7FD33F83E" box="[634,732,1907,1933]" pageId="16" pageNumber="83" type="holotype">holotype</typeStatus>
|
||
in coloration and pattern. Adult males were identified as specimens with SVL ≥
|
||
<quantity id="4CC0A0BFFFFF6558C63FF812FE5BF802" box="[357,436,1942,1969]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="83" unit="mm" value="65.0">65 mm</quantity>
|
||
and the presence of enlarged postanal scales; adult females were identified as specimens with SVL ≥
|
||
<quantity id="4CC0A0BFFFFF6558C67AF83EFE80F866" box="[288,367,1978,2005]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="83" unit="mm" value="60.0">60 mm</quantity>
|
||
and no enlarged postanal scales (
|
||
<bibRefCitation id="EFA970ABFFFF6558C5B8F83FFC81F866" author="Norris, K." box="[738,878,1978,2005]" pageId="16" pageNumber="83" pagination="251 - 326" refId="ref14358" refString="Norris, K. (1958) The evolution and systematics of the iguanid genus Uma and its relation to the evolution of other North American desert reptiles. Bulletin of the American Museum of Natural History, 114, 251 - 326." type="journal article" year="1958">Norris 1958</bibRefCitation>
|
||
). For adults of both sexes combined, rostral width averaged
|
||
<quantity id="4CC0A0BFFFFF6558C659F85AFE87F84A" box="[259,360,2014,2041]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" pageId="16" pageNumber="83" unit="mm" value="1.69">1.69 mm</quantity>
|
||
(range 1.34 to 1.89); mental width
|
||
<quantity id="4CC0A0BFFFFF6558C5ADF85AFCB4F84A" box="[759,859,2014,2041]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="83" unit="mm" value="0.55">0.55 mm</quantity>
|
||
(0.35–0.73); head length
|
||
<quantity id="4CC0A0BFFFFF6558C327F85AFB00F84A" box="[1149,1263,2014,2041]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="16" pageNumber="83" unit="mm" value="16.9">16.90 mm</quantity>
|
||
(14.66–18.54); head width
|
||
<quantity id="4CC0A0BFFFFE6559C646FF1CFE60FF00" box="[284,399,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.351" pageId="17" pageNumber="84" unit="mm" value="13.51">13.51 mm</quantity>
|
||
(12.00–15.53); head depth
|
||
<quantity id="4CC0A0BFFFFE6559C59DFF1CFCC3FF00" box="[711,812,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" pageId="17" pageNumber="84" unit="mm" value="9.75">9.75 mm</quantity>
|
||
(7.89–12.11). Body width averaged
|
||
<quantity id="4CC0A0BFFFFE6559C396FF1DFAD0FF00" box="[1228,1343,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.677" pageId="17" pageNumber="84" unit="mm" value="26.77">26.77 mm</quantity>
|
||
(23.16– 31.32); ventrolateral blotch width
|
||
<quantity id="4CC0A0BFFFFE6559C57BFF39FD67FF64" box="[545,648,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.28" pageId="17" pageNumber="84" unit="mm" value="4.28">4.28 mm</quantity>
|
||
(3.27–5.59); femur length
|
||
<quantity id="4CC0A0BFFFFE6559C49BFF38FBD9FF64" box="[961,1078,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.857" pageId="17" pageNumber="84" unit="mm" value="18.57">18.57 mm</quantity>
|
||
(16.00–21.05); length of pedal digit IV 16.00 mm (13.80–18.19); tail length
|
||
<quantity id="4CC0A0BFFFFE6559C5FAFF65FCFCFF48" box="[672,787,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="8.102" pageId="17" pageNumber="84" unit="mm" value="81.02">81.02 mm</quantity>
|
||
(67.65–93.16); tail base width
|
||
<quantity id="4CC0A0BFFFFE6559C320FF65FB01FF48" box="[1146,1262,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.472" pageId="17" pageNumber="84" unit="mm" value="14.72">14.72 mm</quantity>
|
||
(12.96–17.39). Postrostral scales had a median of four (range 4–5); one mental scale; one postmental scale; postmental scale came in contact with gular scales in all but two specimens (USNM 590036 and USNM 590040). Infralabial scales had a median of 11 scales (range 10–11); eight supralabial scales per side except in USNM 590043 (seven scales) and USNM 590041 (nine scales); four internasal scales (3–6); four supercilliaries (4–5); 12 festoons on lower eyelid (11–14); five enlarged anterior auricular scales (5–6); 27 femoral pores (25–30); 30 fringe scales on posterior edge of pedal digit IV (25–33). USNM 590035 had one enlarged post-anal scale on the right. Dorsum ocelli had black borders and rusty central spots, with the background coloration ranging from off-white to slate gray. Limbs usually lighter in coloration than dorsum, with grey or light brown spotting. Ventral coloration ranged from cream to tawny. Ventrolateral markings ranged from a clustering of small black dots to solid black blotches. Gular markings ranged from five to six pairs of black diagonal lines. Black ventral tail markings ranged from six to eight. Juveniles duller than adults in dorsal coloration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF475DD2FFFE6559C7CDF8A5FBB4F8ED" ID-DOI="http://doi.org/10.5281/zenodo.3823666" ID-Zenodo-Dep="3823666" httpUri="https://zenodo.org/record/3823666/files/figure.png" pageId="17" pageNumber="84" startId="17.[151,250,1825,1850]" targetBox="[151,1435,669,1800]" targetPageId="17">
|
||
<paragraph id="8B870D5AFFFE6559C7CDF8A5FBB4F8ED" blockId="17.[151,1436,1825,1886]" pageId="17" pageNumber="84">
|
||
<emphasis id="B94CD148FFFE6559C7CDF8A5FEE2F889" bold="true" box="[151,269,1825,1850]" pageId="17" pageNumber="84">FIGURE 6</emphasis>
|
||
. Holotype (USNM 590033) of
|
||
<taxonomicName id="4C3876D9FFFE6559C514F8A6FD1BF88A" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[590,756,1825,1849]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFE6559C514F8A6FD1BF88A" box="[590,756,1825,1849]" italics="true" pageId="17" pageNumber="84">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
showing (A) dorsal and (B) ventral head; (C) dorsal right hindfoot and (D) dorsal right forefoot showing toe fringes. Photographs by E. Langan and J. Poindexter.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3225ED1FFFE655BC79DF80FFC01FF48" lastPageId="19" lastPageNumber="86" pageId="17" pageNumber="84" type="distribution">
|
||
<paragraph id="8B870D5AFFFE655AC79DF80FFEB9FE44" blockId="17.[151,1437,1931,2030]" lastBlockId="18.[151,1436,153,503]" lastPageId="18" lastPageNumber="85" pageId="17" pageNumber="84">
|
||
<emphasis id="B94CD148FFFE6559C79DF80FFE8EF815" bold="true" box="[199,353,1931,1958]" pageId="17" pageNumber="84">Distribution.</emphasis>
|
||
<taxonomicName id="4C3876D9FFFE6559C630F809FDCDF815" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[362,546,1932,1958]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFE6559C630F809FDCDF815" box="[362,546,1932,1958]" italics="true" pageId="17" pageNumber="84">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Mohawk Dunes, Yuma County,
|
||
<collectingRegion id="49FCC3B8FFFE6559C3EBF808FAE1F815" box="[1201,1294,1932,1958]" country="United States of America" name="Arizona" pageId="17" pageNumber="84">Arizona</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32F4DCAFFFE6559C246F808FABDF815" box="[1308,1362,1932,1958]" name="United States of America" pageId="17" pageNumber="84">USA</collectingCountry>
|
||
(Figs. 1,7). The type material comes from the northern, central, and southern parts of the main dune system. The distance between our northernmost and southernmost samples is ~
|
||
<quantity id="4CC0A0BFFFFE6559C447F850FC87F85D" box="[797,872,2004,2030]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="84" unit="km" value="24.0">24 km</quantity>
|
||
. Nearly all of the probable range of
|
||
<taxonomicName id="4C3876D9FFFE6559C25AF851FA73F85D" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[1280,1436,2004,2030]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="84" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFE6559C25AF851FA73F85D" box="[1280,1436,2004,2030]" italics="true" pageId="17" pageNumber="84">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
falls within the boundaries of the Barry M. Goldwater Range operated by the
|
||
<collectingCountry id="F32F4DCAFFFD655AC35BFF1DFBDCFF00" box="[1025,1075,153,179]" name="United States of America" pageId="18" pageNumber="85">U.S.</collectingCountry>
|
||
Marine Corps, although the
|
||
<typeStatus id="5483B3F8FFFD655AC22DFF1DFF32FF64" pageId="18" pageNumber="85" type="holotype">holotype</typeStatus>
|
||
was collected on land administered by the Bureau of Land Management on the northernmost extent of the Mohawk Dunes abutting Interstate 8 (north of the Range). While this manuscript was in preparation additional individuals of what are likely members of
|
||
<taxonomicName id="4C3876D9FFFD655AC512FE82FD0CFEAC" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[584,739,261,287]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC512FE82FD0CFEAC" box="[584,739,261,287]" italics="true" pageId="18" pageNumber="85">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
were observed east of the Mohawk Mountains near Dateland, Yuma County,
|
||
<collectingRegion id="49FCC3B8FFFD655AC664FEADFE77FEF0" box="[318,408,297,323]" country="United States of America" name="Arizona" pageId="18" pageNumber="85">Arizona</collectingRegion>
|
||
(D. Leavitt,
|
||
<collectingRegion id="49FCC3B8FFFD655AC573FEADFD6CFEF0" box="[553,643,297,323]" country="United States of America" name="Arizona" pageId="18" pageNumber="85">Arizona</collectingRegion>
|
||
Dept. Game and Fish, pers. comm.). Examination of satellite imagery reveals that likely habitat for this species may extend east of the mountains through a pass in the southern end of the range (~
|
||
<geoCoordinate id="EE0C6B9DFFFD655AC644FEF5FE7FFE38" box="[286,400,369,395]" degrees="32" direction="north" minutes="31" orientation="latitude" pageId="18" pageNumber="85" precision="925" value="32.516666">32° 31’ N</geoCoordinate>
|
||
, ~
|
||
<geoCoordinate id="EE0C6B9DFFFD655AC6F1FEF5FDDFFE38" box="[427,560,368,395]" degrees="113" direction="west" minutes="35" orientation="longitude" pageId="18" pageNumber="85" precision="925" value="-113.583336">113° 35’ W</geoCoordinate>
|
||
). Further work is needed to assess the distributional limits of this species. For this to occur, researchers would need to secure special use permits from the
|
||
<collectingCountry id="F32F4DCAFFFD655AC4B1FE11FBF2FE1C" box="[1003,1053,405,431]" name="United States of America" pageId="18" pageNumber="85">U.S.</collectingCountry>
|
||
Air Force and/or Marine Corps to access restricted areas on this active range. Even if the range includes Dateland,
|
||
<taxonomicName id="4C3876D9FFFD655AC347FE3EFB57FE60" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[1053,1208,441,467]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC347FE3EFB57FE60" box="[1053,1208,441,467]" italics="true" pageId="18" pageNumber="85">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Yuma County,
|
||
<collectingRegion id="49FCC3B8FFFD655AC7AEFE59FEBDFE44" box="[244,338,477,503]" country="United States of America" name="Arizona" pageId="18" pageNumber="85">Arizona</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF475DD2FFFD655AC7CDF984FDB7F91A" ID-DOI="http://doi.org/10.5281/zenodo.3823668" ID-Zenodo-Dep="3823668" httpUri="https://zenodo.org/record/3823668/files/figure.png" pageId="18" pageNumber="85" startId="18.[151,250,1536,1561]" targetBox="[151,1435,525,1511]" targetPageId="18">
|
||
<paragraph id="8B870D5AFFFD655AC7CDF984FDB7F91A" blockId="18.[151,1437,1536,1705]" pageId="18" pageNumber="85">
|
||
<emphasis id="B94CD148FFFD655AC7CDF984FEFBF9AA" bold="true" box="[151,276,1536,1561]" pageId="18" pageNumber="85">FIGURE 7.</emphasis>
|
||
Satellite map of collecting localities for the type series of
|
||
<taxonomicName id="4C3876D9FFFD655AC430F986FBFFF9AA" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[874,1040,1537,1561]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC430F986FBFFF9AA" box="[874,1040,1537,1561]" italics="true" pageId="18" pageNumber="85">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
. Locations where specimens were collected, denoted by red stars, lie along the Mohawk Dunes, a long strip running northwest to southeast. The lizards are isolated in this dune system by the parallel Mohawk Mountains to the east, the Gila River and Interstate-8 to the north, and alluvial fans to the west and south, ultimately resulting in speciation. Source: “Mohawk Dunes,” 32° 34’ N, 113° 38’ W, Google Earth. Imagery date: May 29, 2016. Accessed: July 3, 2018.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B870D5AFFFD655BC79DF953FC01FF48" blockId="18.[151,1437,1750,2030]" lastBlockId="19.[151,1436,152,251]" lastPageId="19" lastPageNumber="86" pageId="18" pageNumber="85">
|
||
<emphasis id="B94CD148FFFD655AC79DF953FE63F942" bold="true" box="[199,396,1750,1777]" pageId="18" pageNumber="85">Natural History.</emphasis>
|
||
<taxonomicName id="4C3876D9FFFD655AC6C9F95CFDA5F942" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[403,586,1751,1777]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC6C9F95CFDA5F942" box="[403,586,1751,1777]" italics="true" pageId="18" pageNumber="85">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
is a lowland, arenicolous, diurnal, insectivorous species specialized for life in windblown sand (
|
||
<figureCitation id="130311DFFFFD655AC625F97FFE2CF8A6" box="[383,451,1787,1813]" captionStart="FIGURE 8" captionStartId="19.[151,250,1929,1954]" captionTargetBox="[151,1435,273,1904]" captionTargetId="figure@19.[151,1435,273,1904]" captionTargetPageId="19" captionText="FIGURE 8. (A) An individual of U. thurmanae in its natural habitat. (B) Sand dune habitat of U. thurmanae (camera pointed east with the Mohawk Dunes in the midground and the Mohawk Mountains in the background). Photographs by A. Gottscho." figureDoi="http://doi.org/10.5281/zenodo.3823670" httpUri="https://zenodo.org/record/3823670/files/figure.png" pageId="18" pageNumber="85">Fig. 8</figureCitation>
|
||
) much like other members of the
|
||
<taxonomicName id="4C3876D9FFFD655AC412F978FC91F8A6" authorityName=", Norris" authorityYear="1958" box="[840,894,1788,1813]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94CD148FFFD655AC412F978FC91F8A6" box="[840,894,1788,1813]" italics="true" pageId="18" pageNumber="85">Uma</emphasis>
|
||
</taxonomicName>
|
||
clade. Its natural history is presumed to be like that of other members of the
|
||
<taxonomicName id="4C3876D9FFFD655AC6BFF8A4FDBEF88A" baseAuthorityName="Schmidt" baseAuthorityYear="1953" box="[485,593,1824,1849]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="notata">
|
||
<emphasis id="B94CD148FFFD655AC6BFF8A4FDBEF88A" box="[485,593,1824,1849]" italics="true" pageId="18" pageNumber="85">U. notata</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<bibRefCitation id="EFA970ABFFFD655AC593F89BFC80F88A" author="Stebbins, R." box="[713,879,1823,1849]" pageId="18" pageNumber="85" pagination="311 - 322" refId="ref14793" refString="Stebbins, R. (1944) Some aspects of the ecology of the iguanid genus Uma. Ecological Monographs, 14 (3), 311 - 322. https: // doi. org / 10.2307 / 1948446" type="journal article" year="1944">Stebbins 1944</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA970ABFFFD655AC426F89BFBE8F88A" author="Norris, K." box="[892,1031,1822,1849]" pageId="18" pageNumber="85" pagination="251 - 326" refId="ref14358" refString="Norris, K. (1958) The evolution and systematics of the iguanid genus Uma and its relation to the evolution of other North American desert reptiles. Bulletin of the American Museum of Natural History, 114, 251 - 326." type="journal article" year="1958">Norris 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA970ABFFFD655AC34FF89BFB55F88A" author="Mayhew, W." box="[1045,1210,1823,1849]" pageId="18" pageNumber="85" pagination="9 - 18" refId="ref14240" refString="Mayhew, W. (1966) Reproduction in the arenicolous lizard Uma notata. Ecology, 47 (1), 9 - 18. https: // doi. org / 10.2307 / 1935740" type="journal article" year="1966">Mayhew 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA970ABFFFD655AC39DF89BFA82F88A" author="Stebbins, R." box="[1223,1389,1823,1849]" pageId="18" pageNumber="85" refId="ref14833" refString="Stebbins, R. (2003) A Field Guide to Western Reptiles and Amphibians. Third edition. Houghton Mifflin Company, Boston and New York, 533 pp." type="book" year="2003">Stebbins 2003</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3876D9FFFD655AC2DBF8A4FEFFF8EE" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC2DBF8A4FEFFF8EE" italics="true" pageId="18" pageNumber="85">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
(under the name
|
||
<taxonomicName id="4C3876D9FFFD655AC695F8C0FD6EF8EE" authorityName="Cope" authorityYear="1895" box="[463,641,1859,1885]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="rufopunctata">
|
||
<emphasis id="B94CD148FFFD655AC695F8C0FD6EF8EE" box="[463,641,1859,1885]" italics="true" pageId="18" pageNumber="85">U. rufopunctata</emphasis>
|
||
</taxonomicName>
|
||
) was the subject of an ecological study conducted at the Barry M. Goldwater Range (
|
||
<bibRefCitation id="EFA970ABFFFD655AC66BF8E3FDCFF832" author="Bagne, K. E. & Finch, D. M." box="[305,544,1895,1921]" pageId="18" pageNumber="85" refId="ref12293" refString="Bagne, K. E. & Finch, D. M. (2012) Vulnerability of species to climate change in the Southwest: threatened, endangered, and at-risk species at the Barry M. Goldwater Range, Arizon a. U. S. Department of Agriculture Forest Service, Fort Collins, 139 pp. https: // doi. org / 10.2737 / RMRS-GTR- 284" type="book" year="2012">Bagne & Finch 2012</bibRefCitation>
|
||
). The study used
|
||
<taxonomicName id="4C3876D9FFFD655AC585F8ECFCABF832" box="[735,836,1895,1922]" pageId="18" pageNumber="88" rank="species" species="system">a System</taxonomicName>
|
||
for Assessing Vulnerability of Species (SAVS;
|
||
<bibRefCitation id="EFA970ABFFFD655AC20FF8ECFEFEF816" author="Bagne, K. E. & Friggens, M. M. & Finch, D. M." pageId="18" pageNumber="85" refId="ref12229" refString="Bagne, K. E., Friggens, M. M. & Finch, D. M. (2011) A system for assessing vulnerability of species (SAVS) to climate change. U. S. Department of Agriculture Forest Service, Fort Collins, 28 pp. https: // doi. org / 10.2737 / RMRS-GTR- 257" type="book" year="2011">
|
||
Bagne
|
||
<emphasis id="B94CD148FFFD655AC7CDF808FF26F816" box="[151,201,1931,1957]" italics="true" pageId="18" pageNumber="85">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
) to evaluate the various wildlife species found within the bombing range in terms of their resilience or vulnerability to both climate change and military activity in order to propose efficient management strategies. Of the 23 categories for which a species could possibly be impacted by through climate change,
|
||
<taxonomicName id="4C3876D9FFFD655AC3EFF850FA83F85E" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[1205,1388,2003,2029]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="85" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFD655AC3EFF850FA83F85E" box="[1205,1388,2003,2029]" italics="true" pageId="18" pageNumber="85">Uma thurmanae</emphasis>
|
||
</taxonomicName>
|
||
had an overall vulnerability score of 5.2, the fourth highest score of the 15 species evaluated. The most threatening impacts of both climate change and military activity are burrows collapsing from military vehicular activity, decrease in vegetative shade, and inhibited egg development from lack of moist sand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF475DD2FFFC655BC7CDF80DFA61F875" ID-DOI="http://doi.org/10.5281/zenodo.3823670" ID-Zenodo-Dep="3823670" httpUri="https://zenodo.org/record/3823670/files/figure.png" pageId="19" pageNumber="86" startId="19.[151,250,1929,1954]" targetBox="[151,1435,273,1904]" targetPageId="19">
|
||
<paragraph id="8B870D5AFFFC655BC7CDF80DFA61F875" blockId="19.[151,1436,1929,1990]" pageId="19" pageNumber="86">
|
||
<emphasis id="B94CD148FFFC655BC7CDF80DFEFCF811" bold="true" box="[151,275,1929,1954]" pageId="19" pageNumber="86">FIGURE 8.</emphasis>
|
||
(A) An individual of
|
||
<taxonomicName id="4C3876D9FFFC655BC6A9F80EFD6EF812" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[499,641,1929,1953]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="86" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFC655BC6A9F80EFD6EF812" box="[499,641,1929,1953]" italics="true" pageId="19" pageNumber="86">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
in its natural habitat. (B) Sand dune habitat of
|
||
<taxonomicName id="4C3876D9FFFC655BC339F80EFB1EF812" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[1123,1265,1929,1953]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="86" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFC655BC339F80EFB1EF812" box="[1123,1265,1929,1953]" italics="true" pageId="19" pageNumber="86">U. thurmanae</emphasis>
|
||
</taxonomicName>
|
||
(camera pointed east with the Mohawk Dunes in the midground and the Mohawk Mountains in the background). Photographs by A. Gottscho.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3225ED1FFFB655CC79DFF1DFF30FF49" pageId="20" pageNumber="87" type="etymology">
|
||
<paragraph id="8B870D5AFFFB655CC79DFF1DFF30FF49" blockId="20.[151,1436,152,250]" pageId="20" pageNumber="87">
|
||
<emphasis id="B94CD148FFFB655CC79DFF1DFEA1FF00" bold="true" box="[199,334,153,179]" pageId="20" pageNumber="87">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C3876D9FFFB655CC574FF1CFD48FF01" authorityName="Derycke & Gottscho & Mulcahy & Queiroz" authorityYear="2020" box="[558,679,152,178]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="87" phylum="Chordata" rank="species" species="thurmanae">
|
||
<emphasis id="B94CD148FFFB655CC574FF1CFD48FF01" box="[558,679,152,178]" italics="true" pageId="20" pageNumber="87">thurmanae</emphasis>
|
||
</taxonomicName>
|
||
honors
|
||
<taxonomicName id="4C3876D9FFFB655CC459FF1DFBECFF01" authority="Karuna Thurman" authorityName="Karuna Thurman" box="[771,1027,152,179]" class="Reptilia" family="Phrynosomatidae" genus="Uma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="87" phylum="Chordata" rank="genus">Uma Karuna Thurman</taxonomicName>
|
||
(born
|
||
<date id="FF862B9AFFFB655CC314FF1CFB17FF00" box="[1102,1272,152,179]" pageId="20" pageNumber="87" value="1970-04-29">April 29, 1970</date>
|
||
), an American actress and pop icon, for her philanthropic contributions and outreach promoting wildlife conservation and human rights.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |