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<document id="A87275F1CC6F403B5A83B09B72D5895C" ID-CLB-Dataset="302266" ID-DOI="10.11646/zootaxa.5497.1.4" ID-GBIF-Dataset="75139c5b-5d59-48fa-9a6f-06e501f6a8ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="13617904" ID-ZooBank="7F924C76-CE6D-4F31-9020-400800FAE279" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1725025343624" checkinUser="plazi" docAuthor="Cifuentes, Julio, Robla, Jairo &amp; Garcia, Lluc" docDate="2024" docId="038E862E4479FF99FF0D4D2FFB980BC3" docLanguage="en" docName="zootaxa.5497.1.4.pdf" docOrigin="Zootaxa 5497 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Armadillidium galiciense , Schmolzer 1955" docType="treatment" docVersion="2" lastPageNumber="93" masterDocId="FFB7FE564470FF93FF9A4C31FFDB0F5A" masterDocTitle="Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae)" masterLastPageNumber="99" masterPageNumber="83" pageNumber="92" updateTime="1725476738656" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E9AF0EE11AF48DD630CCEA9175FAC64">Description of Armadillidium boalense sp. nov. from northern Spain, with remarks and a key of the genus in the Iberian Peninsula and the Balearic Islands (Isopoda: Oniscidea: Armadillidiidae)</mods:title>
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<mods:namePart id="1767C980D5E94C1DB6EB5F8269118743">Cifuentes, Julio</mods:namePart>
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<mods:namePart id="28BA5BB578EC8AECA2BB41E893929722">Robla, Jairo</mods:namePart>
<mods:affiliation id="8CB83A5655D91C6B7BD25D47CEE290E5">Department of Conservation Biology and Global Change, Doñana Biological Station - CSIC, Sevilla, Spain</mods:affiliation>
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<mods:namePart id="10115FC1217728FE58A3966B00348F81">Garcia, Lluc</mods:namePart>
<mods:nameIdentifier id="47EA7B502A37DCF49AED5FDA8B05DB59" type="ORCID">0000-0002-6043-288X</mods:nameIdentifier>
<mods:nameIdentifier id="551FCA6A8C26A0D0C2C37D6550A963FC" type="email">llucgarciaisopoda@gmail.com</mods:nameIdentifier>
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<mods:title id="64583591BEE349C0B490844B2B596C0D">Zootaxa</mods:title>
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<mods:date id="8565C76DC788D0ED6BFDF5BD2AFEFC6A">2024</mods:date>
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<mods:number id="D9712F274278F45DD4961479649AE4B9">2024-08-22</mods:number>
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<mods:number id="9EA1593FE58067E3A38AB59FBC341651">5497</mods:number>
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<subSubSection id="C33D64B34479FF9AFF0D4D2FFD400E62" box="[151,667,286,313]" pageId="9" pageNumber="92" type="nomenclature">
<paragraph id="8B9837384479FF9AFF0D4D2FFD400E62" blockId="9.[151,667,286,349]" box="[151,667,286,313]" pageId="9" pageNumber="92">
<heading id="D0D080544479FF9AFF0D4D2FFD400E62" bold="true" box="[151,667,286,313]" fontSize="11" level="1" pageId="9" pageNumber="92" reason="1">
<taxonomicName id="4C274CBB4479FF9AFF0D4D2FFD400E62" ID-CoL="684QS" authority="Schmolzer, 1955" authorityName=", Schmolzer" authorityYear="1955" box="[151,667,286,313]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFF0D4D2FFD400E62" bold="true" box="[151,667,286,313]" pageId="9" pageNumber="92">
<emphasis id="B953EB2A4479FF9AFF0D4D2FFE120E63" bold="true" box="[151,457,286,313]" italics="true" pageId="9" pageNumber="92">Armadillidium galiciense</emphasis>
<bibRefCitation id="EFB64AC94479FF9AFE4A4D2FFD400E62" author="Schmolzer, K." box="[464,667,286,312]" pageId="9" pageNumber="92" pagination="155 - 215" refId="ref12406" refString="Schmolzer, K. (1955) Isopoda terrarum mediterranearum. 1. Mitteilung: Uber neue und bekannte Landasseln der Pyrenaenhalbinsel. Eos, 31, 155 - 215." type="journal article" year="1955">Schmölzer, 1955</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C33D64B34479FF9AFF0D4D72FF020E07" box="[151,217,323,349]" pageId="9" pageNumber="92" type="description">
<paragraph id="8B9837384479FF9AFF0D4D72FF020E07" blockId="9.[151,667,286,349]" box="[151,217,323,349]" pageId="9" pageNumber="92">
<figureCitation id="131C2BBD4479FF9AFF0D4D72FF020E07" box="[151,217,323,349]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="9" pageNumber="92">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D64B34479FF9AFF0D4DB6FCCB0EC5" box="[151,784,391,415]" pageId="9" pageNumber="92" type="reference_group">
<paragraph id="8B9837384479FF9AFF0D4DB6FCCB0EC5" blockId="9.[151,784,391,415]" box="[151,784,391,415]" pageId="9" pageNumber="92">
<treatmentCitationGroup id="AB3710164479FF9AFF0D4DB6FCCB0EC5" box="[151,784,391,415]" pageId="9" pageNumber="92">
<treatmentCitation id="0A8611294479FF9AFF0D4DB6FCCB0EC5" author="Schmolzer, K." box="[151,784,391,415]" page="189" pageId="9" pageNumber="92" year="1955">
<taxonomicName id="4C274CBB4479FF9AFF0D4DB6FD890EC5" ID-CoL="684QS" authority=", Schmolzer, 1955, p. 189" authorityName=", Schmolzer" authorityPageNumber="189" authorityYear="1955" box="[151,594,391,415]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFF0D4DB6FE430EC5" box="[151,408,391,415]" italics="true" pageId="9" pageNumber="92">Armadillidium galiciense</emphasis>
,
<bibRefCitation id="EFB64AC94479FF9AFE3F4DB6FD890EC5" author="Schmolzer, K." box="[421,594,391,415]" pageId="9" pageNumber="92" pagination="155 - 215" refId="ref12406" refString="Schmolzer, K. (1955) Isopoda terrarum mediterranearum. 1. Mitteilung: Uber neue und bekannte Landasseln der Pyrenaenhalbinsel. Eos, 31, 155 - 215." type="journal article" year="1955">Schmölzer, 1955</bibRefCitation>
</taxonomicName>
, p. 189–192, fig. 9
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</paragraph>
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<caption id="DF5867B04479FF9AFF02496FFAEA0AC0" ID-DOI="http://doi.org/10.5281/zenodo.13617918" ID-Zenodo-Dep="13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="9" pageNumber="92" startId="9.[152,255,1374,1398]" targetBox="[151,1436,439,1350]" targetPageId="9" targetType="figure">
<paragraph id="8B9837384479FF9AFF02496FFAEA0AC0" blockId="9.[151,1437,1374,1434]" pageId="9" pageNumber="92">
<emphasis id="B953EB2A4479FF9AFF02496FFECC0A2C" bold="true" box="[152,279,1374,1398]" pageId="9" pageNumber="92">FIGURE 7.</emphasis>
<taxonomicName id="4C274CBB4479FF9AFE87496FFDF00A2C" authorityName=", Schmolzer" authorityYear="1955" box="[285,555,1374,1398]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFE87496FFDF00A2C" box="[285,555,1374,1398]" italics="true" pageId="9" pageNumber="92">Armadillidium galiciense</emphasis>
</taxonomicName>
. Slide of the Holotype.
<emphasis id="B953EB2A4479FF9AFCB0496FFCE70A2C" bold="true" box="[810,828,1374,1398]" pageId="9" pageNumber="92">A</emphasis>
. Pleopods 1 ♂ and Genital papilla (one exopod missing).
<emphasis id="B953EB2A4479FF9AFF0D49B3FF730AC0" bold="true" box="[151,168,1410,1434]" pageId="9" pageNumber="92">B</emphasis>
. Pleopods 2 ♂.
<emphasis id="B953EB2A4479FF9AFED149B3FE860AC0" bold="true" box="[331,349,1410,1434]" pageId="9" pageNumber="92">C</emphasis>
. Pereopod 1 ♂.
<emphasis id="B953EB2A4479FF9AFD9849B3FDCF0AC0" bold="true" box="[514,532,1410,1434]" pageId="9" pageNumber="92">D</emphasis>
. Pereopod 7 ♂.
<emphasis id="B953EB2A4479FF9AFD2049B3FD100AC0" bold="true" box="[698,715,1410,1434]" pageId="9" pageNumber="92">E</emphasis>
. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm).
</paragraph>
</caption>
<subSubSection id="C33D64B34479FF9AFF0D49D5FAEB084C" pageId="9" pageNumber="92" type="materials_examined">
<paragraph id="8B9837384479FF9AFF0D49D5FE5E0AA4" blockId="9.[151,389,1508,1534]" box="[151,389,1508,1534]" pageId="9" pageNumber="92">
<emphasis id="B953EB2A4479FF9AFF0D49D5FE5E0AA4" bold="true" box="[151,389,1508,1534]" pageId="9" pageNumber="92">Material examined</emphasis>
</paragraph>
<paragraph id="8B9837384479FF9AFF0D4A1DFD790930" blockId="9.[151,1436,1580,1643]" pageId="9" pageNumber="92">
<materialsCitation id="3B4F3D654479FF9AFF0D4A1DFD790930" country="Spain" location="Spain" pageId="9" pageNumber="92" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F33077A84479FF9AFF0D4A1DFF0C091C" box="[151,215,1580,1606]" name="Spain" pageId="9" pageNumber="92">Spain</collectingCountry>
•
<specimenCount id="9D21FCB14479FF9AFF754A1CFEDF091D" box="[239,260,1581,1607]" count="1" pageId="9" pageNumber="92" type="male">♂</specimenCount>
<emphasis id="B953EB2A4479FF9AFE904A1DFEAC091C" bold="true" box="[266,375,1580,1606]" pageId="9" pageNumber="92">
<typeStatus id="549C899A4479FF9AFE904A1DFEAC091C" box="[266,375,1580,1606]" pageId="9" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
, male; high-definition photographs of the slide containing male pleopods 1–2 and pereopods 1 and 7 (see comments and taxonomic remarks).
</materialsCitation>
</paragraph>
<paragraph id="8B9837384479FF9AFF0D4A85FE550994" blockId="9.[151,398,1716,1742]" box="[151,398,1716,1742]" pageId="9" pageNumber="92">
<heading id="D0D080544479FF9AFF0D4A85FE550994" bold="true" box="[151,398,1716,1742]" fontSize="11" level="1" pageId="9" pageNumber="92" reason="1">
<emphasis id="B953EB2A4479FF9AFF0D4A85FE550994" bold="true" box="[151,398,1716,1742]" pageId="9" pageNumber="92">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9837384479FF9AFF0D4ACDFAEB084C" blockId="9.[151,1328,1788,1815]" box="[151,1328,1788,1815]" pageId="9" pageNumber="92">
<materialsCitation id="3B4F3D654479FF9AFF0D4ACDFAEB084C" box="[151,1328,1788,1815]" pageId="9" pageNumber="92">
<collectingCountry id="F33077A84479FF9AFF0D4ACDFF0C084C" box="[151,215,1788,1814]" name="Spain" pageId="9" pageNumber="92">Spain</collectingCountry>
•
<specimenCount id="9D21FCB14479FF9AFF754ACDFEC8084C" box="[239,275,1788,1814]" count="1" pageId="9" pageNumber="92" type="female">1♀</specimenCount>
;
<collectingRegion id="49E3F9DA4479FF9AFEBB4ACDFEAE084C" box="[289,373,1788,1814]" country="Spain" name="Galicia" pageId="9" pageNumber="92">Galicia</collectingRegion>
, Pontevedra, Cerdedo;
<date id="FF9911F84479FF9AFDE34ACDFCD6084D" box="[633,781,1788,1815]" pageId="9" pageNumber="92" value="1952-04-31">31 Apr. 1952</date>
; Franz leg.; Schmölzer det.;
<collectionCode id="ED36AFFD4479FF9AFBD54ACDFB78084C" box="[1103,1187,1788,1814]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="9" pageNumber="92" type="Museum">MNCN</collectionCode>
20.04/9634.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D64B34479FF99FF0D4B51FBCB0EF7" lastPageId="10" lastPageNumber="93" pageId="9" pageNumber="92" type="diagnosis">
<paragraph id="8B9837384479FF9AFF0D4B51FE730820" blockId="9.[151,424,1888,1914]" box="[151,424,1888,1914]" pageId="9" pageNumber="92">
<heading id="D0D080544479FF9AFF0D4B51FE730820" bold="true" box="[151,424,1888,1914]" fontSize="11" level="1" pageId="9" pageNumber="92" reason="1">
<emphasis id="B953EB2A4479FF9AFF0D4B51FE730820" bold="true" box="[151,424,1888,1914]" pageId="9" pageNumber="92">Taxonomical remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9837384479FF99FF0D4B99FBCB0EF7" blockId="9.[151,1436,1960,2023]" lastBlockId="10.[151,1437,151,429]" lastPageId="10" lastPageNumber="93" pageId="9" pageNumber="92">
<bibRefCitation id="EFB64AC94479FF9AFF0D4B99FEEC0898" author="Vandel, A." box="[151,311,1960,1986]" pageId="9" pageNumber="92" pagination="417 - 931" refId="ref12668" refString="Vandel, A. (1962) Isopodes terrestres (deuxieme partie). Faune de France, Paris, 66, 417 - 931." type="journal article" year="1962">Vandel (1962)</bibRefCitation>
considered
<taxonomicName id="4C274CBB4479FF9AFE254B98FD8B0898" authorityName=", Schmolzer" authorityYear="1955" box="[447,592,1960,1987]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFE254B98FD8B0898" box="[447,592,1960,1987]" italics="true" pageId="9" pageNumber="92">A. galiciense</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="4C274CBB4479FF9AFCDB4B98FC770898" authorityName="Brandt" authorityYear="1833" box="[833,940,1961,1987]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B953EB2A4479FF9AFCDB4B98FC770898" box="[833,940,1961,1987]" italics="true" pageId="9" pageNumber="92">A. pictum</emphasis>
</taxonomicName>
, a criterion followed by other authors (
<bibRefCitation id="EFB64AC94479FF9AFAFE4B99FF0908BC" author="Cruz, A." pageId="9" pageNumber="92" pagination="155 - 181" refId="ref11258" refString="Cruz, A. (1993) El genero Armadillidium Brandt, 1833 en la Peninsula Iberica y Baleares (Isopoda, Oniscidea, Armadillidiidae). Boletin Asociacion espanola Entomologia, 17 (1) 155 - 181." type="journal article" year="1993">Cruz 1993</bibRefCitation>
;
<bibRefCitation id="EFB64AC94479FF9AFF454BFDFE5408BC" author="Cifuentes, J." box="[223,399,1996,2022]" pageId="9" pageNumber="92" refId="ref10875" refString="Cifuentes, J. (2019) Los isopodos terrestres de Galicia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia, 75 (2), e 098. https: // doi. org / 10.3989 / graellsia. 2019. v 75.243" type="journal volume" year="2019">Cifuentes 2019</bibRefCitation>
). However,
<taxonomicName id="4C274CBB4479FF9AFD8C4BFCFD7308BC" authorityName=", Schmolzer" authorityYear="1955" box="[534,680,1996,2023]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFD8C4BFCFD7308BC" box="[534,680,1996,2023]" italics="true" pageId="9" pageNumber="92">A. galiciense</emphasis>
</taxonomicName>
is a valid and distinct species.
<taxonomicName id="4C274CBB4479FF9AFB924BFCFB4108BC" authorityName=", Schmolzer" authorityYear="1955" box="[1032,1178,1996,2023]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A4479FF9AFB924BFCFB4108BC" box="[1032,1178,1996,2023]" italics="true" pageId="9" pageNumber="92">A. galiciense</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C274CBB4479FF9AFAAA4BFCFA4708BC" authorityName="Brandt" authorityYear="1833" box="[1328,1436,1997,2023]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B953EB2A4479FF9AFAAA4BFCFA4708BC" box="[1328,1436,1997,2023]" italics="true" pageId="9" pageNumber="92">A. pictum</emphasis>
</taxonomicName>
mainly in the
<emphasis id="B953EB2A447AFF99FEAF4CA6FE4E0FEB" box="[309,405,151,177]" italics="true" pageId="10" pageNumber="93">endopod</emphasis>
of the
<emphasis id="B953EB2A447AFF99FE7C4CA6FD4E0FEB" box="[486,661,151,177]" italics="true" pageId="10" pageNumber="93">male pleopod 1</emphasis>
(
<figureCitation id="131C2BBD447AFF99FD3F4CA6FD240FEB" box="[677,767,151,177]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="10" pageNumber="93">Fig. 7A</figureCitation>
), which is straight, progressively tapered, and only slightly bent and pointed distally; and in the exopod, which is longitudinally elongated, without posterior lobe and twice as wide as long. It also differs in the
<emphasis id="B953EB2A447AFF99FDDB4CEEFD2B0FA3" box="[577,752,223,249]" italics="true" pageId="10" pageNumber="93">male pleopod 2</emphasis>
(
<figureCitation id="131C2BBD447AFF99FC9B4CEEFC830FA3" box="[769,856,223,249]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="10" pageNumber="93">Fig. 7B</figureCitation>
) (not illustrated in the original description), by the
<emphasis id="B953EB2A447AFF99FF0D4D32FF330E47" box="[151,232,259,285]" italics="true" pageId="10" pageNumber="93">exopod</emphasis>
with a triangular posterior lobe, not bent nor narrowed as in
<taxonomicName id="4C274CBB447AFF99FC1C4D32FC2B0E46" authorityName="Brandt" authorityYear="1833" box="[902,1008,259,285]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B953EB2A447AFF99FC1C4D32FC2B0E46" box="[902,1008,259,285]" italics="true" pageId="10" pageNumber="93">A. pictum</emphasis>
</taxonomicName>
. According to the original description, the coloration of the body is also different. For all these reasons, it does not seem justified that
<taxonomicName id="4C274CBB447AFF99FB774D16FA5B0E1B" authorityName=", Schmolzer" authorityYear="1955" box="[1261,1408,295,321]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A447AFF99FB774D16FA5B0E1B" box="[1261,1408,295,321]" italics="true" pageId="10" pageNumber="93">A. galiciense</emphasis>
</taxonomicName>
is considered a junior subjective synonym of
<taxonomicName id="4C274CBB447AFF99FD144D7AFD270E3E" authorityName="Brandt" authorityYear="1833" box="[654,764,331,357]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B953EB2A447AFF99FD144D7AFD270E3E" box="[654,764,331,357]" italics="true" pageId="10" pageNumber="93">A. pictum</emphasis>
</taxonomicName>
and we can confirm the validity of this species. From
<taxonomicName id="4C274CBB447AFF99FA1E4D7AFF200ED3" authorityName="Cifuentes &amp; Robla &amp; Garcia" authorityYear="2024" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="boalense" status="sp. nov.">
<emphasis id="B953EB2A447AFF99FA1E4D7AFF200ED3" italics="true" pageId="10" pageNumber="93">A. boalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2605651447AFF99FE984D5EFE850ED3" box="[258,350,367,393]" pageId="10" pageNumber="93" rank="species">
<emphasis id="B953EB2A447AFF99FE984D5EFE8C0ED3" bold="true" box="[258,343,367,393]" pageId="10" pageNumber="93">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs mainly in the exopods of the male pleopods 1–2 and in having the male pereopods 1 and 7 (
<figureCitation id="131C2BBD447AFF99FF2E4DA2FEF60EF7" box="[180,301,403,429]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="10" pageNumber="93">Fig. 7C–D</figureCitation>
) without evident modifications, instead of strongly differentiated.
</paragraph>
</subSubSection>
<subSubSection id="C33D64B3447AFF99FF0D4DCFFB3C0DF3" pageId="10" pageNumber="93" type="distribution">
<paragraph id="8B983738447AFF99FF0D4DCFFE7A0D43" blockId="10.[151,417,510,537]" box="[151,417,510,537]" pageId="10" pageNumber="93">
<emphasis id="B953EB2A447AFF99FF0D4DCFFE7A0D43" bold="true" box="[151,417,510,537]" pageId="10" pageNumber="93">Distribution remarks</emphasis>
</paragraph>
<paragraph id="8B983738447AFF99FF0D4E76FB3C0DF3" blockId="10.[151,1436,582,681]" pageId="10" pageNumber="93">
<bibRefCitation id="EFB64AC9447AFF99FF0D4E76FEBA0D3B" author="Schmolzer, K." box="[151,353,582,609]" pageId="10" pageNumber="93" pagination="155 - 215" refId="ref12406" refString="Schmolzer, K. (1955) Isopoda terrarum mediterranearum. 1. Mitteilung: Uber neue und bekannte Landasseln der Pyrenaenhalbinsel. Eos, 31, 155 - 215." type="journal article" year="1955">Schmölzer (1955)</bibRefCitation>
described
<taxonomicName id="4C274CBB447AFF99FE464E76FDB50D3B" authorityName=", Schmolzer" authorityYear="1955" box="[476,622,583,609]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A447AFF99FE464E76FDB50D3B" box="[476,622,583,609]" italics="true" pageId="10" pageNumber="93">A. galiciense</emphasis>
</taxonomicName>
as an endemic species of northwestern
<collectingCountry id="F33077A8447AFF99FBB54E76FBA90D3B" box="[1071,1138,583,609]" name="Spain" pageId="10" pageNumber="93">Spain</collectingCountry>
, based on specimens from
<collectingRegion id="49E3F9DA447AFF99FF0D4E5AFF330DDF" box="[151,232,619,645]" country="Spain" name="Galicia" pageId="10" pageNumber="93">Galicia</collectingRegion>
(
<figureCitation id="131C2BBD447AFF99FF6D4E5AFEA40DDF" box="[247,383,619,645]" captionStart="FIGURE 1" captionStartId="2.[152,255,1303,1327]" captionTargetBox="[158,1430,492,1271]" captionTargetId="figure-103@2.[151,1436,486,1276]" captionTargetPageId="2" captionText="FIGURE 1. A. Distribution of Armadillidium species in Asturias (red, within Spain seen in darker blue). B. Habitat of Armadillidium boalense sp. nov. in Os Chaos (Boal, Asturias). C. A more detailed photograph of the rocky wall where A. boalense sp. nov. specimens were found. D. Distribution of Armadillidium species in the group ‘pictum’ in northern Iberian Peninsula with the provinces with available data of northern Spain in gray. In both maps: New records (★) and bibliographic records (●). A. album (blue), A. assimile (orange), A. boalense sp. nov. (red), A. galiciense (green), A. nasatum (pink), A. pictum (purple), cf. A. pictum (purple?) and A. vulgare (black)." figureDoi="http://doi.org/10.5281/zenodo.13685712" httpUri="https://zenodo.org/record/13685712/files/figure.png" pageId="10" pageNumber="93">Fig. 1A, 1D</figureCitation>
). The same author reported this species in other localities of the Cantabric region:
<collectingRegion id="49E3F9DA447AFF99FA8A4E5AFAB60DDF" box="[1296,1389,619,645]" country="Spain" name="Asturias" pageId="10" pageNumber="93">Asturias</collectingRegion>
and
<collectingRegion id="49E3F9DA447AFF99FF0D4EBEFEDF0DF3" box="[151,260,655,681]" country="Spain" name="Cantabria" pageId="10" pageNumber="93">Cantabria</collectingRegion>
(
<bibRefCitation id="EFB64AC9447AFF99FE894EBEFE170DF3" author="Schmolzer, K." box="[275,460,655,681]" pageId="10" pageNumber="93" pagination="1 - 161" refId="ref12467" refString="Schmolzer, K. (1971) Die Landisopoden der Iberischen Halbinsel. Monografias de Ciencia moderna, 80, 1 - 161." type="journal article" year="1971">Schmölzer 1971</bibRefCitation>
) (
<figureCitation id="131C2BBD447AFF99FE7E4EBEFDB60DF3" box="[484,621,655,681]" captionStart="FIGURE 1" captionStartId="2.[152,255,1303,1327]" captionTargetBox="[158,1430,492,1271]" captionTargetId="figure-103@2.[151,1436,486,1276]" captionTargetPageId="2" captionText="FIGURE 1. A. Distribution of Armadillidium species in Asturias (red, within Spain seen in darker blue). B. Habitat of Armadillidium boalense sp. nov. in Os Chaos (Boal, Asturias). C. A more detailed photograph of the rocky wall where A. boalense sp. nov. specimens were found. D. Distribution of Armadillidium species in the group ‘pictum’ in northern Iberian Peninsula with the provinces with available data of northern Spain in gray. In both maps: New records (★) and bibliographic records (●). A. album (blue), A. assimile (orange), A. boalense sp. nov. (red), A. galiciense (green), A. nasatum (pink), A. pictum (purple), cf. A. pictum (purple?) and A. vulgare (black)." figureDoi="http://doi.org/10.5281/zenodo.13685712" httpUri="https://zenodo.org/record/13685712/files/figure.png" pageId="10" pageNumber="93">Fig. 1A, 1D</figureCitation>
). This species has never been reported again since 1971.
</paragraph>
</subSubSection>
<subSubSection id="C33D64B3447AFF99FF0D4EC3FB980BC3" pageId="10" pageNumber="93" type="discussion">
<paragraph id="8B983738447AFF99FF0D4EC3FE630C57" blockId="10.[151,440,754,781]" box="[151,440,754,781]" pageId="10" pageNumber="93">
<heading id="D0D08054447AFF99FF0D4EC3FE630C57" bold="true" box="[151,440,754,781]" fontSize="11" level="1" pageId="10" pageNumber="93" reason="1">
<emphasis id="B953EB2A447AFF99FF0D4EC3FE630C57" bold="true" box="[151,440,754,781]" pageId="10" pageNumber="93">
Comments on
<typeStatus id="549C899A447AFF99FED64EC2FE630C57" box="[332,440,755,781]" pageId="10" pageNumber="93" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B983738447AFF99FF0D4F0AFB980BC3" blockId="10.[151,1437,827,1177]" pageId="10" pageNumber="93">
According to the original description of
<taxonomicName id="4C274CBB447AFF99FDF74F0AFCDA0C0F" authorityName=", Schmolzer" authorityYear="1955" box="[621,769,827,853]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A447AFF99FDF74F0AFCDA0C0F" box="[621,769,827,853]" italics="true" pageId="10" pageNumber="93">A. galiciense</emphasis>
</taxonomicName>
, it is a clearly different species from
<taxonomicName id="4C274CBB447AFF99FB214F0AFAF20C0E" authorityName="Brandt" authorityYear="1833" box="[1211,1321,827,853]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B953EB2A447AFF99FB214F0AFAF20C0E" box="[1211,1321,827,853]" italics="true" pageId="10" pageNumber="93">A. pictum</emphasis>
</taxonomicName>
and from
<taxonomicName id="4C274CBB447AFF99FF0D4F6EFEC60C23" authorityName="Cifuentes &amp; Robla &amp; Garcia" authorityYear="2024" box="[151,285,863,889]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="boalense" status="sp. nov.">
<emphasis id="B953EB2A447AFF99FF0D4F6EFEC60C23" box="[151,285,863,889]" italics="true" pageId="10" pageNumber="93">A. boalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2605651447AFF99FEBC4F6EFE5F0C23" box="[294,388,863,889]" pageId="10" pageNumber="93" rank="species">
<emphasis id="B953EB2A447AFF99FEBC4F6EFEA60C23" bold="true" box="[294,381,863,889]" pageId="10" pageNumber="93">sp. nov</emphasis>
.
</taxonomicNameLabel>
The
<typeStatus id="549C899A447AFF99FE584F6EFDFF0C23" box="[450,548,863,889]" pageId="10" pageNumber="93" type="holotype">holotype</typeStatus>
slide (
<figureCitation id="131C2BBD447AFF99FDEE4F6EFD350C23" box="[628,750,863,889]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="10" pageNumber="93">Fig. 7A–E</figureCitation>
) contained the male pleopods 1 and 2 (the left exopod 1 is missing), the genital papilla and the pereopods 1 and 7. It is hand-labelled (
<figureCitation id="131C2BBD447AFF99FC704FB2FB9B0CC7" box="[1002,1088,899,925]" captionStart="FIGURE 7" captionStartId="9.[152,255,1374,1398]" captionTargetBox="[151,1436,439,1350]" captionTargetId="figure-67@9.[151,1436,439,1350]" captionTargetPageId="9" captionText="FIGURE 7. Armadillidium galiciense. Slide of the Holotype. A. Pleopods 1 ♂ and Genital papilla (one exopod missing). B. Pleopods 2 ♂. C. Pereopod 1 ♂. D. Pereopod 7 ♂. E. Detail of label (scale bars: A–B, 0.65 mm; C–D, 0.75 mm)." figureDoi="http://doi.org/10.5281/zenodo.13617918" httpUri="https://zenodo.org/record/13617918/files/figure.png" pageId="10" pageNumber="93">Fig. 7E</figureCitation>
) as following: «
<taxonomicName id="4C274CBB447AFF99FB634FB2FA470CC7" authorityName="Brandt" authorityYear="1831" box="[1273,1436,899,925]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B953EB2A447AFF99FB634FB2FA470CC7" box="[1273,1436,899,925]" italics="true" pageId="10" pageNumber="93">Armadillidium</emphasis>
</taxonomicName>
(A.)
<emphasis id="B953EB2A447AFF99FF574F96FE9E0C9B" box="[205,325,935,961]" italics="true" pageId="10" pageNumber="93">galiciensis</emphasis>
(sic) Schmlzr (sic)» inked-written and «
<typeStatus id="549C899A447AFF99FC8B4F96FC520C9B" box="[785,905,935,961]" pageId="10" pageNumber="93" type="holotype">Holotypus</typeStatus>
» pencil-written. Additional information inked-written included «Sp 169/1F Leg. Franz» and «K. SCHMÖLZER INNSBRUCK». It is important to comment that in its original description,
<bibRefCitation id="EFB64AC9447AFF99FE504FDEFD4C0B53" author="Schmolzer, K." box="[458,663,1006,1033]" pageId="10" pageNumber="93" pagination="155 - 215" refId="ref12406" refString="Schmolzer, K. (1955) Isopoda terrarum mediterranearum. 1. Mitteilung: Uber neue und bekannte Landasseln der Pyrenaenhalbinsel. Eos, 31, 155 - 215." type="journal article" year="1955">Schmölzer (1955)</bibRefCitation>
did not expressly designated an
<typeStatus id="549C899A447AFF99FB8D4FDEFBA20B53" box="[1047,1145,1007,1033]" pageId="10" pageNumber="93" type="holotype">holotype</typeStatus>
but indicated «Cerdedo» (Pontevedra province,
<collectingRegion id="49E3F9DA447AFF99FE014822FE340B77" box="[411,495,1043,1069]" country="Spain" name="Galicia" pageId="10" pageNumber="93">Galicia</collectingRegion>
,
<collectingCountry id="F33077A8447AFF99FE664822FDE60B77" box="[508,573,1043,1069]" name="Spain" pageId="10" pageNumber="93">Spain</collectingCountry>
) as its type locality, corresponding to the locality «(Sp 179, loc. typ.), leg. F». Otherwise, the slide annotated as «
<typeStatus id="549C899A447AFF99FDC84806FD1F0B0B" box="[594,708,1079,1105]" pageId="10" pageNumber="93" type="holotype">holotypus</typeStatus>
» corresponded to the locality of Pyrnedo (sic), (i.e.,., Piornedo, province of Lugo,
<collectingRegion id="49E3F9DA447AFF99FEF6486AFE1B0B2F" box="[364,448,1115,1141]" country="Spain" name="Galicia" pageId="10" pageNumber="93">Galicia</collectingRegion>
,
<collectingCountry id="F33077A8447AFF99FE56486AFDCB0B2F" box="[460,528,1115,1141]" name="Spain" pageId="10" pageNumber="93">Spain</collectingCountry>
), recorded in the original description as «(loc. Sp 169), leg. F». So, a complete review of the type series of
<taxonomicName id="4C274CBB447AFF99FE54484EFD840BC3" authorityName=", Schmolzer" authorityYear="1955" box="[462,607,1151,1177]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="galiciense">
<emphasis id="B953EB2A447AFF99FE54484EFD840BC3" box="[462,607,1151,1177]" italics="true" pageId="10" pageNumber="93">A. galiciense</emphasis>
</taxonomicName>
should be addressed in a subsequent work.
</paragraph>
</subSubSection>
</treatment>
</document>
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