246 lines
30 KiB
XML
246 lines
30 KiB
XML
<document id="525DCF17FA089C8DA3817915AE04F462" ID-DOI="10.11646/zootaxa.5109.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6336940" ID-ZooBank="7B6C0651-0935-4C77-B157-3ABF53E3AD81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646727632931" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Ślipiński, Adam" docDate="2022" docId="038B11419C78701BD4D8FD86ECACFDC1" docLanguage="en" docName="zootaxa.5109.1.1.pdf.imf" docOrigin="Zootaxa 5109 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5109.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Murmidius okinawanus Jałoszyński & Ślipiński 2022, sp. n." docType="treatment" docVersion="5" lastPageNumber="56" masterDocId="FFB269399C4D702CD44FFFD9E928FFB2" masterDocTitle="Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="54" updateTime="1699303175474" updateUser="plazi">
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<mods:mods id="343A18F712F82F5F559622702E51318A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3757CE83CD1348196C031FAB2ED33025">
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<mods:title id="0A6996E9881DBA4449545AA251847E31">Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="592224AAB22666BDF6E0457EDD9EA1EC" type="personal">
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<mods:roleTerm id="3303DD04ED37D0B1DC3F241740EBCE11">Author</mods:roleTerm>
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<mods:namePart id="06BEC4CB3DE051382BD805A583D0B7B2">Jałoszyński, Paweł</mods:namePart>
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<mods:nameIdentifier id="C3A13459C98E6416F6DABE1998AFC3AE" type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
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<mods:nameIdentifier id="1AE1566E22FF6C596E28FE717CAC1415" type="email">scydmaenus@yahoo.com</mods:nameIdentifier>
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<mods:name id="5CD0290CCD17CC81AA73FEE4AA3E5347" type="personal">
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<mods:roleTerm id="3E81E1AEA5767AF103BB41C69E39892B">Author</mods:roleTerm>
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<mods:namePart id="F28483D2A4DA9714C86A8C8887AA0AD1">Ślipiński, Adam</mods:namePart>
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<mods:nameIdentifier id="0DD826E9C7E7D343D2954DC6797FC08A" type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
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<mods:nameIdentifier id="9B13C8840D0FB24A6F0AF9021EA63867" type="email">scydmaenus@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="795F6DC0FCD5D4111FBF12FCFA27E43E">text</mods:typeOfResource>
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<mods:titleInfo id="9B6A2F215046BE0D9EE0B686E17CB241">
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<mods:title id="09B0764E35288975FC84C8A329178C6D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EB8F89CA1FF87FCCFDF7BA04F3FB16E6">
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<mods:date id="4A124D40B4E302CEE950FDE2C6251325">2022</mods:date>
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<mods:number id="0253B9E9877246495A585BB7EC51C56F">2022-03-08</mods:number>
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<mods:detail id="21DEBEC47110F8382A230C323E83EE10" type="volume">
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<mods:number id="0BCC7C5AC4409F6058F0407FFA060F30">5109</mods:number>
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</mods:detail>
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<mods:detail id="2F0D59639AB79DF47168D68277B7056B" type="issue">
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<mods:number id="1C87A2A94CD5C776F83485C9FDAFCB24">1</mods:number>
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<mods:url id="0B058EBDA9D96029753936E28E858FFD">http://dx.doi.org/10.11646/zootaxa.5109.1.1</mods:url>
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<mods:classification id="4AA6C586EECCC0F2E2ADB511E182C1D2">journal article</mods:classification>
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<mods:identifier id="1750FF250179309DF2797F5150578045" type="DOI">10.11646/zootaxa.5109.1.1</mods:identifier>
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<mods:identifier id="3759572EEAE4271B4DB270B17B8E3173" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="038B11419C78701BD4D8FD86ECACFDC1" LSID="urn:lsid:plazi:treatment:038B11419C78701BD4D8FD86ECACFDC1" httpUri="http://treatment.plazi.org/id/038B11419C78701BD4D8FD86ECACFDC1" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
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<subSubSection id="C338F3DC9C787019D4D8FD86EB2AFDC8" box="[151,514,607,634]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="8B9DA0579C787019D4D8FD86EB2AFDC8" blockId="53.[151,514,607,671]" box="[151,514,607,634]" pageId="53" pageNumber="54">
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<heading id="D0D5173B9C787019D4D8FD86EB2AFDC8" bold="true" box="[151,514,607,634]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
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<emphasis id="B9567C459C787019D4D8FD86EB2AFDC8" bold="true" box="[151,514,607,634]" pageId="53" pageNumber="54">
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<taxonomicName id="4C22DBD49C787019D4D8FD86E891FDC8" authority="Jałoszyński & Ślipiński, 2022" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[151,441,607,634]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="okinawanus" status="sp. nov.">
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<emphasis id="B9567C459C787019D4D8FD86E891FDC8" bold="true" box="[151,441,607,634]" italics="true" pageId="53" pageNumber="54">Murmidius okinawanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A265C13E9C787019D58FFDB9EB2AFDC8" box="[448,514,608,634]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338F3DC9C787019D4D8FD5DE86BFD2D" box="[151,323,644,671]" pageId="53" pageNumber="54" type="description">
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<paragraph id="8B9DA0579C787019D4D8FD5DE86BFD2D" blockId="53.[151,514,607,671]" box="[151,323,644,671]" pageId="53" pageNumber="54">
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||
(
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<figureCitation id="1319BCD29C787019D4D0FD5DE813FD2D" box="[159,315,644,671]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Figs 156–162</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C338F3DC9C787019D4D8FD15EB4FFC9C" pageId="53" pageNumber="54" type="materials_examined">
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<paragraph id="8B9DA0579C787019D4D8FD15EB4FFC9C" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
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<emphasis id="B9567C459C787019D4D8FD15E897FD54" bold="true" box="[151,447,716,742]" pageId="53" pageNumber="54">
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<collectingCounty id="62FCD8DB9C787019D4D8FD15E9FAFD54" box="[151,210,716,742]" pageId="53" pageNumber="54">Type</collectingCounty>
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material examined.
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</emphasis>
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<materialsCitation id="3B4AAA0A9C787019D589FD15ED2CFCB8" collectingDate="2019-02-15" collectionCode="NSMT" country="Japan" county="Japan" elevation="100" location="Ryukyus" municipality="Japan" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
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<emphasis id="B9567C459C787019D589FD15EA26FD57" bold="true" box="[454,782,715,742]" pageId="53" pageNumber="54">
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<typeStatus id="54991EF59C787019D589FD15EB1BFD54" box="[454,563,716,742]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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(
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<collectingCounty id="62FCD8DB9C787019D60DFD12EBB9FD57" box="[578,657,715,741]" pageId="53" pageNumber="54">
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||
<collectingMunicipality id="6BF93A2D9C787019D60DFD12EBB9FD57" box="[578,657,715,741]" pageId="53" pageNumber="54">Japan</collectingMunicipality>
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||
</collectingCounty>
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||
:
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||
<collectingCounty id="62FCD8DB9C787019D6D2FD12EA20FD57" box="[669,776,715,741]" pageId="53" pageNumber="54">
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||
<location id="8EFDF68C9C787019D6D2FD12EA20FD57" LSID="urn:lsid:plazi:treatment:038B11419C78701BD4D8FD86ECACFDC1:8EFDF68C9C787019D6D2FD12EA20FD57" box="[669,776,715,741]" country="Japan" county="Japan" municipality="Japan" name="Ryukyus" pageId="53" pageNumber="54" stateProvince="Okinawa">Ryukyus</location>
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</collectingCounty>
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)
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</emphasis>
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:
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<specimenCount id="9D246BDE9C787019D752FD15EA1CFD54" box="[797,820,716,742]" pageId="53" pageNumber="54" type="male">♂</specimenCount>
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, “
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<collectingCountry id="F335E0C79C787019D705FD15EA8DFD54" box="[842,933,716,742]" name="Japan" pageId="53" pageNumber="54">JAPAN</collectingCountry>
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,
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<collectingRegion id="49E66EB59C787019D7E0FD15ED19FD54" box="[943,1073,716,742]" country="Japan" name="Okinawa" pageId="53" pageNumber="54">OKINAWA</collectingRegion>
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Island / Nago City, Nago Castle Site / ~
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<quantity id="4CDA0DB29C787019D4A5FD29E81DFCB8" box="[234,309,752,778]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="m" value="100.0">
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||
<elevation id="000F47649C787019D4A5FD29E81DFCB8" box="[234,309,752,778]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="m" value="100.0">100 m</elevation>
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</quantity>
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||
,
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||
<date id="FF9C86979C787019D50CFD29E8E9FCB9" box="[323,449,752,779]" pageId="53" pageNumber="54" value="2019-02-15">
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||
<collectingDate id="EFD87F7F9C787019D50CFD29E8E9FCB9" box="[323,449,752,779]" pageId="53" pageNumber="54" value="2019-02-15">15.02.2019</collectingDate>
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||
</date>
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||
/ humid forest / leg. P. JAŁOSZYŃSKI” (
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<collectionCode id="ED3338929C787019D7E7FD29EAD4FCB8" box="[936,1020,752,778]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="53" pageNumber="54">NSMT</collectionCode>
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)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B4AAA0A9C787019D05FFD29EB4CFC9C" collectingDate="2019-02-15" collectionCode="MNHW" country="Japan" county="Japan" elevation="100" location="Ryukyus" municipality="Japan" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
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||
<emphasis id="B9567C459C787019D05FFD29EDA0FCB8" bold="true" box="[1040,1160,752,778]" pageId="53" pageNumber="54">
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||
<typeStatus id="54991EF59C787019D05FFD29EDA0FCB8" box="[1040,1160,752,778]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
: 3 exx of unknown sex, same data as for holotype (cPJ,
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<collectionCode id="ED3338929C787019D5B4FCCDEB74FC9C" box="[507,604,788,814]" pageId="53" pageNumber="54">MNHW</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338F3DC9C787019D488FCE1EC0CFBC0" pageId="53" pageNumber="54" type="diagnosis">
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||
<paragraph id="8B9DA0579C787019D488FCE1EC0CFBC0" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
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<emphasis id="B9567C459C787019D488FCE1E86AFCE0" bold="true" box="[199,322,824,850]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
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Body elongate oval and weakly convex, pitchy black; antennal club only slightly elongate; anterior clypeal margin smooth; pronotum lacking lateral and sublateral longitudinal impressions; clypeus with extremely fine microreticulation, frons and vertex with indistinct, superficial traces of microreticulation, elytra lacking microsculpture; elytra with strong and dense punctures not arranged in rows; prosternal carinae only as long as about fourth of prosternum; mesoventral plate with evenly rounded anterior margin and complete but fine submarginal carina, lacking lateral carinae; elytral epipleura strongly narrowed at the level of metaventrite where they are only as wide as half width of mesofemur; metaventrite with large, circular, unevenly distributed punctures on sides; penis stout, in lateral view distinctly curved; in ventral view tapering from near middle to subtriangular and blunt apex; tegmen short and broad, with rounded apex bearing one pair of moderately long and several pairs of short setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C338F3DC9C78701BD488FBA5EA41FEED" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
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<paragraph id="8B9DA0579C787019D488FBA5EA6CFA94" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
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<emphasis id="B9567C459C787019D488FBA5E871FB24" bold="true" box="[199,345,1148,1174]" pageId="53" pageNumber="54">Description.</emphasis>
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BL
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<quantity id="4CDA0DB29C787019D5C1FBA5EB1FFB24" box="[398,567,1148,1175]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.5" metricValueMin="1.28" pageId="53" pageNumber="54" unit="mm" value="1.39" valueMax="1.5" valueMin="1.28">1.28–1.50 mm</quantity>
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; BL/EW 1.45–1.62. Body elongate oval (
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<figureCitation id="1319BCD29C787019D05AFBA5ED9CFB25" box="[1045,1204,1148,1175]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Figs 156–157</figureCitation>
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) and in lateral view weakly convex (
|
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<figureCitation id="1319BCD29C787019D51DFB79E89EFB09" box="[338,438,1184,1211]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Fig. 159</figureCitation>
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); pigmentation nearly black, in some specimens head and pronotum indistinctly lighter, legs only slightly lighter than; dorsum covered with extremely short whitish setae (discernible under magnification 80 ×); cuticle glossy, on clypeus distinctly but very finely microreticulate, superficial traces of reticulation can also be seen on frons, vertex and, even less distinct, on pronotum.
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</paragraph>
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<paragraph id="8B9DA0579C787019D488FAE9EB85FA21" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
|
||
Head
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<quantity id="4CDA0DB29C787019D547FAE9E882FAF8" box="[264,426,1328,1355]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="53" pageNumber="54" unit="mm" value="0.37" valueMax="0.39" valueMin="0.35">0.35–0.39 mm</quantity>
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wide; frons and vertex weakly convex, with fine but distinct and dense punctures separated by spaces subequal to their diameters; clypeus less glossy than frons; eyes large, strongly convex, coarsely faceted. Antennal club only slightly elongate (
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<figureCitation id="1319BCD29C787019D671FAA1EB88FA21" box="[574,672,1400,1427]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Fig. 157</figureCitation>
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||
).
|
||
</paragraph>
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<paragraph id="8B9DA0579C787019D488FA45ED08F9F4" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
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Pronotum (
|
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<figureCitation id="1319BCD29C787019D509FA45E8CCFA05" box="[326,484,1436,1463]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Figs 156–157</figureCitation>
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) strongly transverse, widest at base; PL
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<quantity id="4CDA0DB29C787019D7FEFA45ED70FA04" box="[945,1112,1436,1463]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="53" pageNumber="54" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 mm</quantity>
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||
, PW
|
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<quantity id="4CDA0DB29C787019D0D8FA45EC15FA04" box="[1175,1341,1436,1462]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.0" metricValueMin="6.3" pageId="53" pageNumber="54" unit="mm" value="0.665" valueMax="0.7" valueMin="0.63">0.63–0.70 mm</quantity>
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||
, PL/PW 0.37–0.43; lateral margins weakly rounded and moderately strongly convergent anterad, with narrow lateral carinae; mesal corners of antennal cavities weakly projecting anterad; lateral and sublateral longitudinal impressions lacking; anterior pronotal margin with complete marginal line. Punctures on disc similar to those on frons and vertex but indistinctly larger and sparser, nearly evenly distributed, large punctures lacking.
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</paragraph>
|
||
<paragraph id="8B9DA0579C787019D488F989EDAEF900" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
|
||
Prosternum (
|
||
<figureCitation id="1319BCD29C787019D518F989E89DF9D9" box="[343,437,1616,1643]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Fig. 158</figureCitation>
|
||
) with barely discernible transverse microreticulation and with fine, inconspicuous setiferous punctures; prosternal carinae short, about as long as 1/4 of prosternum; notosternal carinae nearly straight and nearly reaching anterior prosternal margin, the latter with distinct narrow marginal carina.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C787019D488F965EC58F8D4" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
|
||
Elytra (
|
||
<figureCitation id="1319BCD29C787019D552F965E894F965" box="[285,444,1724,1751]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Figs 156–157</figureCitation>
|
||
) together oval, with strongly rounded sides, broadest distinctly in front of middle, EL
|
||
<quantity id="4CDA0DB29C787019D4D8F939E814F948" box="[151,316,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" metricValueMax="1.13" metricValueMin="0.95" pageId="53" pageNumber="54" unit="mm" value="1.04" valueMax="1.13" valueMin="0.95">0.95–1.13 mm</quantity>
|
||
, EW
|
||
<quantity id="4CDA0DB29C787019D537F939EB35F948" box="[376,541,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="9.5" metricValueMin="8.8" pageId="53" pageNumber="54" unit="mm" value="0.915" valueMax="0.95" valueMin="0.88">0.88–0.95 mm</quantity>
|
||
, EL/EW 1.09–1.22. Humerus with distinct callus; elytra densely punctate, punc- tures small but distinct, not arranged in rows, those on anteromedian region of each elytron almost coarse, separated by spaces slightly shorter than their diameters, punctures reducing in diameter and depth toward sides and apices. Epipleura conspicuously narrowed near middle (
|
||
<figureCitation id="1319BCD29C787019D6F8F895EA30F8D4" box="[695,792,1868,1894]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Fig. 160</figureCitation>
|
||
, arrowhead), so that lateral elytral margin is concave.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C787019D488F8A9EB2FF838" blockId="53.[151,1437,715,2002]" box="[199,519,1904,1930]" pageId="53" pageNumber="54">Hind wings fully developed.</paragraph>
|
||
<paragraph id="8B9DA0579C787019D488F84DEB33F860" blockId="53.[151,1437,715,2002]" pageId="53" pageNumber="54">
|
||
Mesoventral plate (
|
||
<figureCitation id="1319BCD29C787019D5D1F84DE8D5F81D" box="[414,509,1940,1967]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="53" pageNumber="54">Fig. 160</figureCitation>
|
||
) lacking lateral longitudinal carinae; anterior margin evenly, broadly rounded, with fine, complete submarginal carina.
|
||
</paragraph>
|
||
<caption id="DF5DF0DF9C7B701AD4D8F8ECEA97F8C3" ID-DOI="http://doi.org/10.5281/zenodo.6336993" ID-Zenodo-Dep="6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,264,1845,1869]" targetBox="[151,1436,181,1821]" targetPageId="54">
|
||
<paragraph id="8B9DA0579C7B701AD4D8F8ECEA97F8C3" blockId="54.[151,1437,1845,1905]" pageId="54" pageNumber="55">
|
||
<emphasis id="B9567C459C7B701AD4D8F8ECE847F8FF" bold="true" box="[151,367,1845,1869]" pageId="54" pageNumber="55">FIGURES 156–162.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C22DBD49C7B701AD6D4F8ECEAA3F8FF" authority="Jałoszyński & Ślipiński, 2022" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[667,907,1845,1869]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="okinawanus" status="sp. nov.">
|
||
<emphasis id="B9567C459C7B701AD6D4F8ECEAA3F8FF" box="[667,907,1845,1869]" italics="true" pageId="54" pageNumber="55">Murmidius okinawanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9567C459C7B701AD7DDF8ECEAE5F8FF" bold="true" box="[914,973,1845,1869]" pageId="54" pageNumber="55">
|
||
<taxonomicNameLabel id="A265C13E9C7B701AD7DDF8ECEAE5F8FF" box="[914,973,1845,1869]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9DA0579C7A701BD488FF4EED0FFF61" blockId="55.[151,1437,151,628]" pageId="55" pageNumber="56">
|
||
Metaventrite (
|
||
<figureCitation id="1319BCD29C7A701BD52AFF4EE8ECFF00" box="[357,452,151,178]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="55" pageNumber="56">Fig. 160</figureCitation>
|
||
) with large, circular and distinct punctures unevenly distributed on sides; median region with fine, inconspicuous setiferous punctures. Discrimen externally well-marked.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C7A701BD488FF05EA17FF44" blockId="55.[151,1437,151,628]" box="[199,831,220,247]" pageId="55" pageNumber="56">Tarsi with short, inconspicuous setae on tarsomeres 1–3.</paragraph>
|
||
<paragraph id="8B9DA0579C7A701BD488FF26EA41FEED" blockId="55.[151,1437,151,628]" pageId="55" pageNumber="56">
|
||
Aedeagus (
|
||
<figureCitation id="1319BCD29C7A701BD50AFF26E8CAFEA8" box="[325,482,255,282]" captionStart="FIGURES 156–162" captionStartId="54.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-19@54.[151,1436,181,1821]" captionTargetPageId="54" captionText="FIGURES 156–162. Morphological structures of Murmidius okinawanus sp. n. Body in dorsal (156–157), ventral (158) and lateral (159) views; pterothorax in ventral view (160); and aedeagus (161–162)." figureDoi="http://doi.org/10.5281/zenodo.6336993" httpUri="https://zenodo.org/record/6336993/files/figure.png" pageId="55" pageNumber="56">Figs 161–162</figureCitation>
|
||
)
|
||
<quantity id="4CDA0DB29C7A701BD5BDFF27EB70FEAB" box="[498,600,254,281]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="55" pageNumber="56" unit="mm" value="0.35">0.35 mm</quantity>
|
||
long, stout; penis in ventral view gradually tapering from near middle toward subtriangular, blunt apex, in lateral view distinctly curved; tegmen stout, with rounded apex bearing one pair of moderately long setae and several pairs of much shorter ones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C7A701BD488FEBFEB92FE30" box="[199,698,358,386]" pageId="55" pageNumber="56" type="distribution">
|
||
<paragraph id="8B9DA0579C7A701BD488FEBFEB92FE30" blockId="55.[151,1437,151,628]" box="[199,698,358,386]" pageId="55" pageNumber="56">
|
||
<emphasis id="B9567C459C7A701BD488FEBFE849FE32" bold="true" box="[199,353,358,384]" pageId="55" pageNumber="56">Distribution.</emphasis>
|
||
Sub-Oriental (Okinawa-jima).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C7A701BD488FE50EBCFFE11" box="[199,743,393,420]" pageId="55" pageNumber="56" type="etymology">
|
||
<paragraph id="8B9DA0579C7A701BD488FE50EBCFFE11" blockId="55.[151,1437,151,628]" box="[199,743,393,420]" pageId="55" pageNumber="56">
|
||
<emphasis id="B9567C459C7A701BD488FE50E860FE11" bold="true" box="[199,328,393,419]" pageId="55" pageNumber="56">Etymology</emphasis>
|
||
. Named after the island of
|
||
<collectingRegion id="49E66EB59C7A701BD635FE50EBCBFE11" box="[634,739,393,419]" country="Japan" name="Okinawa" pageId="55" pageNumber="56">Okinawa</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338F3DC9C7A701BD488FE75ECACFDC1" pageId="55" pageNumber="56" type="discussion">
|
||
<paragraph id="8B9DA0579C7A701BD488FE75EBFFFDE3" blockId="55.[151,1437,151,628]" pageId="55" pageNumber="56">
|
||
<emphasis id="B9567C459C7A701BD488FE75E813FE74" bold="true" box="[199,315,428,454]" pageId="55" pageNumber="56">Remarks.</emphasis>
|
||
This nearly black species is unique among all members of
|
||
<taxonomicName id="4C22DBD49C7A701BD782FE75ED60FE74" box="[973,1096,428,454]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9567C459C7A701BD782FE75ED60FE74" box="[973,1096,428,454]" italics="true" pageId="55" pageNumber="56">Murmidius</emphasis>
|
||
</taxonomicName>
|
||
in having the elytral epipleura strongly narrowed near middle, so that the lateral elytral edge is concave between meso- and metacoxae and the epipleuron in its narrowest site in this region is only half as wide as mesofemur. The dark pigmentation and body shape make
|
||
<taxonomicName id="4C22DBD49C7A701BD56CFDCCE8E7FD9C" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[291,463,532,558]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="okinawanus">
|
||
<emphasis id="B9567C459C7A701BD56CFDCCE8E7FD9C" box="[291,463,532,558]" italics="true" pageId="55" pageNumber="56">M. okinawanus</emphasis>
|
||
</taxonomicName>
|
||
similar to
|
||
<taxonomicName id="4C22DBD49C7A701BD606FDCCEB93FD9C" box="[585,699,532,558]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="hebrus">
|
||
<emphasis id="B9567C459C7A701BD606FDCCEB93FD9C" box="[585,699,532,558]" italics="true" pageId="55" pageNumber="56">M. hebrus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C22DBD49C7A701BD6BCFDCCEA49FD9C" authorityName="Jałoszyński & Ślipiński" authorityYear="2022" box="[755,865,532,558]" class="Insecta" family="Murmidiidae" genus="Murmidius" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="merkli">
|
||
<emphasis id="B9567C459C7A701BD6BCFDCCEA49FD9C" box="[755,865,532,558]" italics="true" pageId="55" pageNumber="56">M. merkli</emphasis>
|
||
</taxonomicName>
|
||
; but the aedeagi, and especially the apical setation of the tegmen, are clearly different in each of them.
|
||
</paragraph>
|
||
<paragraph id="8B9DA0579C7A701BD488FD80ECACFDC1" blockId="55.[151,1437,151,628]" box="[199,1412,601,628]" pageId="55" pageNumber="56">
|
||
The
|
||
<typeStatus id="54991EF59C7A701BD4B5FD83E802FDC6" box="[250,298,602,628]" pageId="55" pageNumber="56">type</typeStatus>
|
||
series of this species was collected by sifting moist leaf litter and rotten wood in subtropical forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |