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<document id="BC046402768C16CC7DC5841CAD801A93" ID-CLB-Dataset="6089" ID-DOI="10.11646/zootaxa.4958.1.32" ID-GBIF-Dataset="f8b056ee-69fb-4585-8e6f-ec8a90f937b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692436" ID-ZooBank="4F2516D0-B025-4DA4-9C3D-F4E5093289C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618472120534" checkinUser="plazi" docAuthor="Roca-Cusachs, Marcos, Rider, Id. A. &amp; Jung, Sunghoon" docDate="2021" docId="038420420667C439FF23FC7A0070FB92" docLanguage="en" docName="zootaxa.4958.1.32.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graziasternum graziae Roca-Cusachs &amp; Rider &amp; Jung 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="564" masterDocId="FFBD583A0665C43DFFB4FF890244FFC8" masterDocTitle="New genus and new species of Pentatomidae from Borneo (Hemiptera: Heteroptera)" masterLastPageNumber="569" masterPageNumber="560" pageNumber="562" updateTime="1698936910024" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="926CDA463815337470A746879341AAD9">New genus and new species of Pentatomidae from Borneo (Hemiptera: Heteroptera)</mods:title>
</mods:titleInfo>
<mods:name id="1642CA8BC78A70C493640667A8BE107B" type="personal">
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<mods:namePart id="AF97EC6372C9AB6530AB4F7E3CEF655D">Roca-Cusachs, Marcos</mods:namePart>
<mods:affiliation id="CA77C4D41D59F723E20A2B6233FC52DD">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. marcosrocacusachs @ gmail. com &amp; IRBio. Institut de Recerca a la Biodiversitat, Universitat de Barcelona, Barcelona, Spain.</mods:affiliation>
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<mods:namePart id="E79C1FDCD9CFC965F31479A2AD373A11">Rider, Id. A.</mods:namePart>
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<mods:namePart id="18F57BC4E70672D57164409BF07E9A1F">Jung, Sunghoon</mods:namePart>
<mods:affiliation id="A13C804ADC2B42DCDD6DDB8D82AE6975">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. marcosrocacusachs @ gmail. com &amp; Department of Smart Agriculture Systems, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. Corresponding author. jung @ cnu. ac. kr</mods:affiliation>
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<mods:title id="89E9972476BF93B15212D1821C7C38B2">Zootaxa</mods:title>
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<mods:date id="1A92B0D37D7E4F03B5F338AF7EFCD1F1">2021</mods:date>
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<mods:number id="18D1B533487008BB8EB5CD158CBC4EEA">2021-04-14</mods:number>
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<mods:number id="835941463598411567E9F1B2C2118AAA">4958</mods:number>
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<treatment id="038420420667C439FF23FC7A0070FB92" ID-DOI="http://doi.org/10.5281/zenodo.4692444" ID-GBIF-Taxon="182742133" ID-Zenodo-Dep="4692444" LSID="urn:lsid:plazi:treatment:038420420667C439FF23FC7A0070FB92" httpUri="http://treatment.plazi.org/id/038420420667C439FF23FC7A0070FB92" lastPageId="4" lastPageNumber="564" pageId="2" pageNumber="562">
<subSubSection id="C337C2DF0667C43FFF23FC7A0179FBC5" box="[151,829,1010,1037]" pageId="2" pageNumber="562" type="nomenclature">
<paragraph id="8B9291540667C43FFF23FC7A0179FBC5" blockId="2.[151,829,1010,1073]" box="[151,829,1010,1037]" pageId="2" pageNumber="562">
<heading id="D0DA26380667C43FFF23FC7A0179FBC5" bold="true" box="[151,829,1010,1037]" fontSize="11" level="1" pageId="2" pageNumber="562" reason="1">
<emphasis id="B9594D460667C43FFF23FC7A0179FBC5" bold="true" box="[151,829,1010,1037]" pageId="2" pageNumber="562">
<taxonomicName id="4C2DEAD70667C43FFF23FC7A0097FBC4" authority="Roca-Cusachs &amp; Rider" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[151,723,1010,1037]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="562" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B9594D460667C43FFF23FC7A03EFFBC5" bold="true" box="[151,427,1011,1037]" italics="true" pageId="2" pageNumber="562">Graziasternum graziae</emphasis>
Roca-Cusachs &amp; Rider
</taxonomicName>
,
<taxonomicNameLabel id="A26AF03D0667C43FFD6BFC7A0179FBC5" box="[735,829,1011,1037]" pageId="2" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF23FB9E02AFFBF9" box="[151,235,1047,1073]" pageId="2" pageNumber="562" type="description">
<paragraph id="8B9291540667C43FFF23FB9E02AFFBF9" blockId="2.[151,829,1010,1073]" box="[151,235,1047,1073]" pageId="2" pageNumber="562">
(
<figureCitation id="13168DD10667C43FFF14FB9E02A4FBF9" box="[160,224,1047,1073]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[212,1373,193,1897]" captionTargetId="figure-23@3.[197,1391,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Graziasternum graziae nov. sp.: a, dorsal habitus; b, ventral habitus; c, pygophore (lateral, dorsal, caudal, ventral views, respectively); d, phallus (lateral, dorsal views, respectively); e, right paramere; f, female genital plates, caudoventral view; g, internal female genitalia." figureDoi="http://doi.org/10.5281/zenodo.4692440" httpUri="https://zenodo.org/record/4692440/files/figure.png" pageId="2" pageNumber="562">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF23FBD006CAFB5F" pageId="2" pageNumber="562" type="materials_examined">
<paragraph id="8B9291540667C43FFF23FBD00736FBBB" blockId="2.[151,1437,1113,2040]" box="[151,1394,1113,1140]" pageId="2" pageNumber="562">
<materialsCitation id="3B459B090667C43FFF23FBD00736FBBB" ID-GBIF-Occurrence="3131357301" box="[151,1394,1113,1140]" collectingDate="2011-07" collectionCode="MRC" country="Indonesia" county="Trus Madi mt." location="Male" municipality="Sabah" pageId="2" pageNumber="562" specimenCount="1" typeStatus="holotype">
<emphasis id="B9594D460667C43FFF23FBD0034FFBBB" bold="true" box="[151,267,1113,1139]" pageId="2" pageNumber="562">
<typeStatus id="54962FF60667C43FFF23FBD00342FBBB" box="[151,262,1113,1139]" pageId="2" pageNumber="562" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EF2C78F0667C43FFEA6FBD00308FBBB" LSID="urn:lsid:plazi:treatment:038420420667C439FF23FC7A0070FB92:8EF2C78F0667C43FFEA6FBD00308FBBB" box="[274,332,1113,1139]" country="Indonesia" county="Trus Madi mt." municipality="Sabah" name="Male" pageId="2" pageNumber="562">Male</location>
[
<collectionCode id="ED3C09910667C43FFEEDFBD003E4FBBC" box="[345,416,1113,1140]" country="Turkey" httpUri="http://grbio.org/cool/sdrz-mymz" name="TUBITAK Marmara Research Center Culture Collection" pageId="2" pageNumber="562">MRC</collectionCode>
].
<collectingCountry id="F33AD1C40667C43FFE05FBD0000CFBBB" box="[433,584,1113,1139]" name="Indonesia" pageId="2" pageNumber="562">INDONESIA</collectingCountry>
[Malaysia].
<location id="8EF2C78F0667C43FFD63FBD30168FBBC" LSID="urn:lsid:plazi:treatment:038420420667C439FF23FC7A0070FB92:8EF2C78F0667C43FFD63FBD30168FBBC" box="[727,812,1114,1140]" country="Indonesia" county="Trus Madi mt." municipality="Sabah" name="Borneo" pageId="2" pageNumber="562">Borneo</location>
,
<collectingMunicipality id="6BF60B2E0667C43FFC83FBD0013AFBBB" box="[823,894,1113,1139]" pageId="2" pageNumber="562">Sabah</collectingMunicipality>
,
<collectingCounty id="62F3E9D80667C43FFC3DFBD30668FBBC" box="[905,1068,1113,1140]" pageId="2" pageNumber="562">Trus Madi mt.</collectingCounty>
,
<date id="FF93B7940667C43FFB83FBD006DAFBBB" box="[1079,1182,1113,1139]" pageId="2" pageNumber="562" value="2011-07">
<collectingDate id="EFD74E7C0667C43FFB83FBD006DAFBBB" box="[1079,1182,1113,1139]" pageId="2" pageNumber="562" value="2011-07">VII.2011</collectingDate>
</date>
, local colector leg.
</materialsCitation>
</paragraph>
<paragraph id="8B9291540667C43FFF73FBF406CAFB5F" blockId="2.[151,1437,1113,2040]" box="[199,1166,1149,1176]" pageId="2" pageNumber="562">
<materialsCitation id="3B459B090667C43FFF73FBF403B4FB50" ID-GBIF-Occurrence="3131357302" box="[199,496,1149,1176]" collectionCode="DAR" pageId="2" pageNumber="562" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B9594D460667C43FFF73FBF40302FB5F" bold="true" box="[199,326,1149,1175]" pageId="2" pageNumber="562">
<typeStatus id="54962FF60667C43FFF73FBF40305FB5F" box="[199,321,1149,1175]" pageId="2" pageNumber="562" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D2B5ADD0667C43FFEF9FBF403DCFB5F" box="[333,408,1149,1175]" pageId="2" pageNumber="562" type="male">1 male</specimenCount>
[
<collectionCode id="ED3C09910667C43FFE12FBF403ADFB50" box="[422,489,1149,1176]" country="Australia" lsid="urn:lsid:biocol.org:col:13816" name="Plant Pathology Herbarium" pageId="2" pageNumber="562" type="Herbarium">DAR</collectionCode>
]
</materialsCitation>
,
<materialsCitation id="3B459B090667C43FFE4FFBF406CAFB5F" ID-GBIF-Occurrence="3131357303" box="[507,1166,1149,1176]" collectingDate="2009-05-20" collectionCode="MRC" pageId="2" pageNumber="562" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="9D2B5ADD0667C43FFE4FFBF40018FB5F" box="[507,604,1149,1175]" pageId="2" pageNumber="562" type="female">1 female</specimenCount>
[
<collectionCode id="ED3C09910667C43FFDDEFBF400F4FB50" box="[618,688,1149,1176]" country="Turkey" httpUri="http://grbio.org/cool/sdrz-mymz" name="TUBITAK Marmara Research Center Culture Collection" pageId="2" pageNumber="562">MRC</collectionCode>
], Same data of
<typeStatus id="54962FF60667C43FFCEFFBF401F9FB5F" box="[859,957,1149,1175]" pageId="2" pageNumber="562" type="holotype">holotype</typeStatus>
except:
<date id="FF93B7940667C43FFBA8FBF406CAFB5F" box="[1052,1166,1149,1175]" pageId="2" pageNumber="562" value="2009-05-20">
<collectingDate id="EFD74E7C0667C43FFBA8FBF406CAFB5F" box="[1052,1166,1149,1175]" pageId="2" pageNumber="562" value="2009-05-20">20.V.2009</collectingDate>
</date>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF73FB2807DEFB28" pageId="2" pageNumber="562" type="etymology">
<paragraph id="8B9291540667C43FFF73FB2807DEFB28" blockId="2.[151,1437,1113,2040]" pageId="2" pageNumber="562">
<emphasis id="B9594D460667C43FFF73FB280316FB73" bold="true" box="[199,338,1185,1211]" pageId="2" pageNumber="562">Etymology:</emphasis>
The species
<taxonomicName id="4C2DEAD70667C43FFE52FB2B0019FB73" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[486,605,1186,1212]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="562" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="B9594D460667C43FFE52FB2B0019FB73" box="[486,605,1186,1212]" italics="true" pageId="2" pageNumber="562">G. graziae</emphasis>
</taxonomicName>
is named after our friend and colleague Dr. Jocelia Grazia, to honour her 75th Birthday and in recognition of her extensive contributions in pentatomoid systematics. Noun in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF73FB600712FAA7" pageId="2" pageNumber="562" type="diagnosis">
<paragraph id="8B9291540667C43FFF73FB600712FAA7" blockId="2.[151,1437,1113,2040]" pageId="2" pageNumber="562">
<emphasis id="B9594D460667C43FFF73FB600306FACB" bold="true" box="[199,322,1257,1283]" pageId="2" pageNumber="562">Diagnosis.</emphasis>
This species has all the characters mentioned in the description of the genus. Differs from the other species in the genus by its general colouration brownish (darker than in the other species), head and antennal segments IV and IV concolour with ground colouration, with no black markings. Having the lateral margins of the pronotum ventrally each with a relatively wide, contrastingly dark red which resembles an inverted V-shape.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C439FF73FAF0034DFBFD" lastPageId="4" lastPageNumber="564" pageId="2" pageNumber="562" type="description">
<paragraph id="8B9291540667C43FFF73FAF0000BFA70" blockId="2.[151,1437,1113,2040]" pageId="2" pageNumber="562">
<emphasis id="B9594D460667C43FFF73FAF0031DFA5B" bold="true" box="[199,345,1401,1427]" pageId="2" pageNumber="562">Description.</emphasis>
Overall colouration brown with frontal portion of pronotum and coria reddish, slightly shiny and with coarse evenly scattered punctures.
</paragraph>
<paragraph id="8B9291540667C43FFF73FA4800D6F9A4" blockId="2.[151,1437,1113,2040]" pageId="2" pageNumber="562">Head triangular, dorsal surface concolourous; punctation sparse, slightly reddish; lateral margins of head very weakly concave; juga extending beyond apex of tylus, convergent but not enclosing tylus. Antennae slightly lighter tan in general colouration; segments III to V each with annular reddish ring on apex. Lateral and ventral surfaces of head pale brown with very sparse brownish punctures. Bucculae narrow, widening posteriorly, ridge-like. Rostrum short, reaching between pro- and mesocoxae.</paragraph>
<paragraph id="8B9291540667C439FF73F9FF0646FE89" blockId="2.[151,1437,1113,2040]" lastBlockId="4.[151,1437,151,1114]" lastPageId="4" lastPageNumber="564" pageId="2" pageNumber="562">Pronotum, distinctly bicoloured, brownish posteriorly, gradually becoming dark red anteriorly, evenly punctured; each anterior pronotal angle produced anteriorly as a very small, tooth; anterolateral margins straight, armed with very small granules, slightly deflexed, reflection reaching humeri; humeri small, robust, each produced laterally as an entire, non-toothed process; posterolateral margins sinuous; posterior margin broadly, shallowly concave. Scutellum concolourous with ground colouration, in some specimens anterior medial portion slightly reddish, more or less evenly punctured, punctures slightly more grouped laterally; anterolateral angles of scutellum strongly foveate. Hemelytra brownish-red; hemelytral membranes hyaline, with numerous (910) longitudinal, ramified veins. Sternite lateral margins with a relatively wide, contrastingly dark red which resembles an inverted V-shape mesially pale brownish yellow. sparsely punctate. Ostiolar openings small, rugae long, slender, elongate L-Shaped. Mesosternal carina just reaching procoxae. Metasternum broad, prominent, plate-like, hexagonal, slightly compressed laterally with posterior angles slightly projected caudad towards abdomen; anterior margin of metasternum meeting nearly seamlessly with posterior margin of mesosternum, posterior margin of metasternum bifid, in apposition with basal abdominal protuberance. Legs uniformly yellowish, pilose; superior surfaces of tibiae sulcate; claws black, contrasting with leg colouration. Connexival segments concolourous with general colouration, with no particularities; posterolateral angles slightly but distinctly produced into a minute, sharp denticles. Abdominal venter impunctate, brownish yellow; spiracles concolourous with ventral abdominal surface.</paragraph>
<caption id="DF52C1DC0666C43EFF23F81E03A9F830" ID-DOI="http://doi.org/10.5281/zenodo.4692440" ID-Zenodo-Dep="4692440" httpUri="https://zenodo.org/record/4692440/files/figure.png" pageId="3" pageNumber="563" startId="3.[151,250,1943,1967]" targetBox="[212,1373,193,1897]" targetPageId="3">
<paragraph id="8B9291540666C43EFF23F81E03A9F830" blockId="3.[151,1436,1943,2040]" pageId="3" pageNumber="563">
<emphasis id="B9594D460666C43EFF23F81E0357F867" bold="true" box="[151,275,1943,1967]" pageId="3" pageNumber="563">FIGURE 1.</emphasis>
<taxonomicName id="4C2DEAD70666C43EFEADF8110040F867" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[281,516,1944,1967]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="563" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B9594D460666C43EFEADF8110040F867" box="[281,516,1944,1967]" italics="true" pageId="3" pageNumber="563">Graziasternum graziae</emphasis>
</taxonomicName>
<emphasis id="B9594D460666C43EFDBEF811001FF878" bold="true" box="[522,603,1944,1968]" pageId="3" pageNumber="563">
<taxonomicNameLabel id="A26AF03D0666C43EFDBEF811001FF878" box="[522,603,1944,1968]" pageId="3" pageNumber="563" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
: a, dorsal habitus; b, ventral habitus; c, pygophore (lateral, dorsal, caudal, ventral views, respectively); d, phallus (lateral, dorsal views, respectively); e, right paramere; f, female genital plates, caudoventral view; g, internal female genitalia.
</paragraph>
</caption>
<paragraph id="8B9291540661C439FF73FEC201F0FDD1" blockId="4.[151,1437,151,1114]" pageId="4" pageNumber="564">
Male genitalia (
<figureCitation id="13168DD10661C439FECAFEC203AAFEAD" box="[382,494,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[212,1373,193,1897]" captionTargetId="figure-23@3.[197,1391,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Graziasternum graziae nov. sp.: a, dorsal habitus; b, ventral habitus; c, pygophore (lateral, dorsal, caudal, ventral views, respectively); d, phallus (lateral, dorsal views, respectively); e, right paramere; f, female genital plates, caudoventral view; g, internal female genitalia." figureDoi="http://doi.org/10.5281/zenodo.4692440" httpUri="https://zenodo.org/record/4692440/files/figure.png" pageId="4" pageNumber="564">Fig. 1ce</figureCitation>
): Genital cup well exposed dorsally. Posterolateral angles of pygophore rounded, lobe-like, wide. Ventral rim clearly concave, U-shaped, flat at base. Dorsal rim concave, broadly U-shaped. Parameres somewhat hook-shaped, rounded, not acuminate, each with a small triangular process at apex. Phallotheca dorsoventrally with an inverted piriform shape, laterally somewhat thinner at base, slightly curved. Phallus with two pairs of conjunctival processes. Vesica small, acuminate at apex. Process of vesica laterally lobe-like, rounded, dorsoventrally not divergent at apex, with internal space parallel-sided.
</paragraph>
<paragraph id="8B9291540661C439FF73FDAA01B1FC49" blockId="4.[151,1437,151,1114]" pageId="4" pageNumber="564">
Female genitalia (
<figureCitation id="13168DD10661C439FE3AFDAA03BCFDF6" box="[398,504,547,574]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[212,1373,193,1897]" captionTargetId="figure-23@3.[197,1391,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. Graziasternum graziae nov. sp.: a, dorsal habitus; b, ventral habitus; c, pygophore (lateral, dorsal, caudal, ventral views, respectively); d, phallus (lateral, dorsal views, respectively); e, right paramere; f, female genital plates, caudoventral view; g, internal female genitalia." figureDoi="http://doi.org/10.5281/zenodo.4692440" httpUri="https://zenodo.org/record/4692440/files/figure.png" pageId="4" pageNumber="564">Fig. 1fg</figureCitation>
): Female genital plates with gonocoxae VIII trapezoid, with mesial margins straight, posterior margins sinuous laterally, becoming shallowly concave mesially, each posterolateral angle rounded and divergent in relation to symmetric plate; fused gonocoxae IX trapezoid, nearly triangular; laterotergites IX boomerang shaped, elongate, reaching apices of laterotergites VIII, divergent at apices; sternite X inverted trapezoidal; laterotergites VIII flat, with no special particularities.
<emphasis id="B9594D460661C439FD0CFD3A01C8FD05" box="[696,908,691,717]" italics="true" pageId="4" pageNumber="564">Ductus receptaculi</emphasis>
elongate. Vesicular area membranous; internal rod of vesicular area strongly sclerotized, parallel sided, rectilinear, acuminate at apex and slightly wider posteriorly.
<emphasis id="B9594D460661C439FF23FD720360FCDD" box="[151,292,763,789]" italics="true" pageId="4" pageNumber="564">Pars distalis</emphasis>
nearly two times longer than length of anterior annular flange to
<emphasis id="B9594D460661C439FBB1FD72068AFCDD" box="[1029,1230,763,789]" italics="true" pageId="4" pageNumber="564">capsula seminalis</emphasis>
distance. Anterior annular flange reduced, strongly sclerotized.
<emphasis id="B9594D460661C439FD15FC960130FCF1" box="[673,884,799,825]" italics="true" pageId="4" pageNumber="564">Pars intermedialis</emphasis>
robust, strongly sclerotized, elongate. Posterior annular flange reduced, fused to
<emphasis id="B9594D460661C439FDBEFCCA0097FC95" box="[522,723,835,861]" italics="true" pageId="4" pageNumber="564">capsula seminalis</emphasis>
.
<emphasis id="B9594D460661C439FD55FCCA01F4FC95" box="[737,944,835,861]" italics="true" pageId="4" pageNumber="564">Capsula seminalis</emphasis>
robust, large, adpressed, bearing two short, robust, posteriorly-directed projections which reach posterior annular flange.
</paragraph>
<paragraph id="8B9291540661C439FF73FC02034DFBFD" blockId="4.[151,1437,151,1114]" pageId="4" pageNumber="564">
<emphasis id="B9594D460661C439FF73FC0203A6FC6C" bold="true" box="[199,482,906,933]" pageId="4" pageNumber="564">Measurements (in mm).</emphasis>
Total body length: 10.13/13.41, abdominal width at hemelytral base: 6.38/7.13, maximum abdominal width: 6.71/7.81, length of head from apex to imaginary line between ocelli: 1.37/1.57, width of head (including eyes): 2.21/2.64, width of head before eyes: 1.35/1.66, distance between eyes: 1.2/1.46, pronotal length: 1.89/2.36, maximum pronotal width: 6.55/7.73, scutellar length: 4.51/5.61, scutellar width at base: 3.82/4.56.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0661C439FF73FBB70070FB92" box="[199,564,1086,1114]" pageId="4" pageNumber="564" type="distribution">
<paragraph id="8B9291540661C439FF73FBB70070FB92" blockId="4.[151,1437,151,1114]" box="[199,564,1086,1114]" pageId="4" pageNumber="564">
<emphasis id="B9594D460661C439FF73FBB70325FB90" bold="true" box="[199,353,1086,1112]" pageId="4" pageNumber="564">Distribution.</emphasis>
<collectingCountry id="F33AD1C40661C439FED3FBB6038AFB91" box="[359,462,1087,1113]" name="Malaysia" pageId="4" pageNumber="564">Malaysia</collectingCountry>
(
<collectingRegion id="49E95FB60661C439FE69FBB60063FB92" box="[477,551,1087,1114]" country="Malaysia" name="Sabah" pageId="4" pageNumber="564">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>