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<mods:title id="399A5893001496623D40DB9D20BB8129">Taxonomy and phylogeny of Koinocystididae (Platyhelminthes, Kalyptorhynchia) with the description of three new genera and twelve new species</mods:title>
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Diez, Reygel &amp; Artois
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(
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on live specimens. 
<location id="8EF478C0FFEFFFB30614BA7F798BFEC2" LSID="urn:lsid:plazi:treatment:03829F0DFFEFFFB105DFBBC979DAFD7F:8EF478C0FFEFFFB30614BA7F798BFEC2" box="[860,907,289,315]" country="Cuba" county="Las Sardinas" latitude="19.989166" longLatPrecision="21" longitude="-76.27639" municipality="One" name="One" pageId="33" pageNumber="484" stateProvince="Santiago de Cuba">One</location>
serially-sectioned specimen from 
<location id="8EF478C0FFEFFFB30056BA7E7F9CFEC3" LSID="urn:lsid:plazi:treatment:03829F0DFFEFFFB105DFBBC979DAFD7F:8EF478C0FFEFFFB30056BA7E7F9CFEC3" box="[1310,1436,288,314]" country="Cuba" county="Las Sardinas" latitude="19.989166" longLatPrecision="21" longitude="-76.27639" municipality="One" name="Bueycabon" pageId="33" pageNumber="484" stateProvince="Santiago de Cuba">Bueycabón</location>
(
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; 
<geoCoordinate id="EE1F48DCFFEFFFB30472BA1A7BCAFEA6" box="[314,458,324,351]" degrees="76" direction="west" minutes="57" orientation="longitude" pageId="33" pageNumber="484" precision="15" seconds="28" value="-76.95777">76°57’28”W</geoCoordinate>
) (Type Locality), 
<collectingRegion id="49EFE0F9FFEFFFB307D2BA1A7966FEA7" box="[666,870,324,350]" country="Cuba" name="Santiago de Cuba" pageId="33" pageNumber="484">Santiago de Cuba</collectingRegion>
, 
<collectingCountry id="F33C6E8BFFEFFFB3063ABA1A79ADFEA7" box="[882,941,324,350]" name="Cuba" pageId="33" pageNumber="484">Cuba</collectingCountry>
(
<date id="FF9508DBFFEFFFB306F5BA1A7EA5FEA6" box="[957,1189,324,351]" pageId="33" pageNumber="484" value="2017-11-18">
<collectingDate id="EFD1F133FFEFFFB306F5BA1A7EA5FEA6" box="[957,1189,324,351]" pageId="33" pageNumber="484" value="2017-11-18">November 18, 2017</collectingDate>
</date>
), designated 
<typeStatus id="549090B9FFEFFFB30072BA1A7F9CFEA7" box="[1338,1436,324,350]" pageId="33" pageNumber="484" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3AB6DEFFEFFFB305D7BA377AF2FE7A" box="[159,242,361,387]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="33" pageNumber="484" type="Museum">FMNH</collectionCode>
https://id.luomus.fi/ 
<collectionCode id="ED3AB6DEFFEFFFB3049FBA367801FE7A" box="[471,513,360,387]" pageId="33" pageNumber="484">KV</collectionCode>
.654), intertidal, scraping the upper 
<quantity id="4CD383FEFFEFFFB306C5BA3779D2FE7A" box="[909,978,361,387]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="484" unit="cm" value="10.0">10 cm</quantity>
of the fine sand. 
<collectingMunicipality id="6BF0B461FFEFFFB301DDBA377EC4FE7A" box="[1173,1220,361,387]" pageId="33" pageNumber="484">One</collectingMunicipality>
whole mount from 
<collectingCounty id="62F55697FFEFFFB305DFBAD37B29FE5F" box="[151,297,396,423]" pageId="33" pageNumber="484">Las Sardinas</collectingCounty>
(
<geoCoordinate id="EE1F48DCFFEFFFB30473BAD27BC8FE5E" box="[315,456,396,423]" degrees="19" direction="north" minutes="56" orientation="latitude" pageId="33" pageNumber="484" precision="15" seconds="24" value="19.939999">19°56’24”N</geoCoordinate>
; 
<geoCoordinate id="EE1F48DCFFEFFFB3049DBAD27869FE5E" box="[469,617,396,423]" degrees="76" direction="west" minutes="46" orientation="longitude" pageId="33" pageNumber="484" precision="15" seconds="41" value="-76.77806">76°46’41”W</geoCoordinate>
), 
<collectingRegion id="49EFE0F9FFEFFFB30736BAD2794BFE5F" box="[638,843,396,422]" country="Cuba" name="Santiago de Cuba" pageId="33" pageNumber="484">Santiago de Cuba</collectingRegion>
, 
<collectingCountry id="F33C6E8BFFEFFFB30610BAD27993FE5F" box="[856,915,396,422]" name="Cuba" pageId="33" pageNumber="484">Cuba</collectingCountry>
(
<date id="FF9508DBFFEFFFB306ECBAD37E4AFE5E" box="[932,1098,396,423]" pageId="33" pageNumber="484" value="2016-06-22">
<collectingDate id="EFD1F133FFEFFFB306ECBAD37E4AFE5E" box="[932,1098,396,423]" pageId="33" pageNumber="484" value="2016-06-22">June 22, 2016</collectingDate>
</date>
), fine, silty sand, 
<quantity id="4CD383FEFFEFFFB3005FBAD37F57FE5E" box="[1303,1367,397,423]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="484" unit="m" value="0.1">
<elevation id="0006C928FFEFFFB3005FBAD37F57FE5E" box="[1303,1367,397,423]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="484" unit="m" value="0.1">0.1 m</elevation>
</quantity>
deep, salinity 33 ‰, and two from 
<collectorName id="26DE4BCDFFEFFFB30493BAEE7840FE32" box="[475,576,432,459]" pageId="33" pageNumber="484">El Masío</collectorName>
(
<geoCoordinate id="EE1F48DCFFEFFFB30718BAEE78DDFE32" box="[592,733,432,459]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="33" pageNumber="484" precision="15" seconds="21" value="19.989166">19°59’21”N</geoCoordinate>
; 
<geoCoordinate id="EE1F48DCFFEFFFB307A0BAEE797BFE32" box="[744,891,432,459]" degrees="76" direction="west" minutes="16" orientation="longitude" pageId="33" pageNumber="484" precision="15" seconds="35" value="-76.27639">76°16’35”W</geoCoordinate>
), 
<collectingRegion id="49EFE0F9FFEFFFB306C7BAEE7E57FE33" box="[911,1111,432,458]" country="Cuba" name="Santiago de Cuba" pageId="33" pageNumber="484">Santiago de Cuba</collectingRegion>
, 
<collectingCountry id="F33C6E8BFFEFFFB3012ABAEE7E9DFE33" box="[1122,1181,432,458]" name="Cuba" pageId="33" pageNumber="484">Cuba</collectingCountry>
(
<date id="FF9508DBFFEFFFB301E3BAEF7F4DFE32" box="[1195,1357,432,459]" pageId="33" pageNumber="484" value="2016-06-30">
<collectingDate id="EFD1F133FFEFFFB301E3BAEF7F4DFE32" box="[1195,1357,432,459]" pageId="33" pageNumber="484" value="2016-06-30">June 30, 2016</collectingDate>
</date>
), intertidal, upper 
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of the medium-coarse sand, salinity 33 ‰ (HU 
<collectionCode id="ED3AB6DEFFEFFFB30637BA8B79B1FE16" box="[895,945,469,495]" pageId="33" pageNumber="484">XIII</collectionCode>
.4.28– 
<collectionCode id="ED3AB6DEFFEFFFB306B4BA8B7E2EFE16" box="[1020,1070,469,495]" pageId="33" pageNumber="484">XIII</collectionCode>
.4.30)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFEFFFB3058FBAA77E1EFDEB" box="[199,1054,504,531]" pageId="33" pageNumber="484" type="etymology">
<paragraph id="8B942E1BFFEFFFB3058FBAA77E1EFDEB" blockId="33.[151,1437,288,2043]" box="[199,1054,504,531]" pageId="33" pageNumber="484">
<emphasis id="B95FF209FFEFFFB3058FBAA77B4EFDEA" bold="true" box="[199,334,505,531]" pageId="33" pageNumber="484">Etymology.</emphasis>
The epithet refers to the occurrence of the new species in 
<collectingCountry id="F33C6E8BFFEFFFB30693BAA67E1AFDEB" box="[987,1050,504,530]" name="Cuba" pageId="33" pageNumber="484">Cuba</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFEFFFB3058FB9427B59FD3F" pageId="33" pageNumber="484" type="diagnosis">
<paragraph id="8B942E1BFFEFFFB3058FB9427B59FD3F" blockId="33.[151,1437,288,2043]" pageId="33" pageNumber="484">
<emphasis id="B95FF209FFEFFFB3058FB94278BCFDCE" bold="true" box="[199,700,540,567]" pageId="33" pageNumber="484">
Diagnosis of 
<taxonomicName id="4C2B5598FFEFFFB3042EB9427870FDCE" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[358,624,540,567]" class="Rhabditophora" family="Koinocystididae" genus="Galapagetula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="33" pageNumber="484" phylum="Platyhelminthes" rank="species" species="cubensis" status="sp. nov.">
<emphasis id="B95FF209FFEFFFB3042EB9427870FDCE" bold="true" box="[358,624,540,567]" italics="true" pageId="33" pageNumber="484">Galapagetula cubensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C4F72FFEFFFB30732B94378BCFDCE" box="[634,700,541,567]" pageId="33" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Species of 
<taxonomicName id="4C2B5598FFEFFFB3060FB94279E1FDCF" authorityName="Reygel, Willems &amp; Artois" authorityYear="2011" box="[839,993,540,566]" class="Rhabditophora" family="Koinocystididae" genus="Galapagetula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="33" pageNumber="484" phylum="Platyhelminthes" rank="genus">
<emphasis id="B95FF209FFEFFFB3060FB94279E1FDCF" box="[839,993,540,566]" italics="true" pageId="33" pageNumber="484">Galapagetula</emphasis>
</taxonomicName>
with mouth sphincter. The copulatory bulb encompasses a prostate vesicle, a seminal duct, and the ejaculatory duct, which distally ends in a small and unarmed penis papilla. Prostate glands enter the male duct separately from the seminal duct, forming a divisa-type copulatory organ. Stalk of the resorptive bursa with a strong sphincter. Female duct lacking a sphincter. Common gonopore caudal.
</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFEFFFB1058FB98E79DAFD7F" lastPageId="35" lastPageNumber="486" pageId="33" pageNumber="484" type="description">
<paragraph id="8B942E1BFFEFFFB3058FB98E7EF5FCCB" blockId="33.[151,1437,288,2043]" pageId="33" pageNumber="484">
<emphasis id="B95FF209FFEFFFB3058FB98E7B59FD13" bold="true" box="[199,345,720,746]" pageId="33" pageNumber="484">Description.</emphasis>
Live specimens about 
<quantity id="4CD383FEFFEFFFB3071FB98E78ADFD12" box="[599,685,720,747]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="484" unit="mm" value="1.5">1.5 mm</quantity>
long, translucent, with a pair of small eyes (
<figureCitation id="1310329EFFEFFFB301DDB98E7EFDFD12" box="[1173,1277,720,747]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
: e). Epidermis syncytial, 
<quantity id="4CD383FEFFEFFFB30444B9AA7B63FCF6" box="[268,355,756,783]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="33" pageNumber="484" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 µm</quantity>
thick, fully ciliated. Cilia 
<quantity id="4CD383FEFFEFFFB307DAB9AA78CDFCF6" box="[658,717,756,783]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="484" unit="mm" value="5.0">5 µm</quantity>
long. The epidermis shows many vacuoles, some of which are empty, others are filled with a dark, granular secretion. The dark-stained rhabdites are 
<quantity id="4CD383FEFFEFFFB30116B8477EB5FCCA" box="[1118,1205,793,819]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="33" pageNumber="484" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 µm</quantity>
long.
</paragraph>
<paragraph id="8B942E1BFFEFFFB3058FB86279B7FAFE" blockId="33.[151,1437,288,2043]" pageId="33" pageNumber="484">
The proboscis (
<figureCitation id="1310329EFFEFFFB3043BB8627BD6FCAE" box="[371,470,828,855]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB304B0B863782CFCAE" box="[504,556,829,855]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: pr, 17B) is of the typical koinocystidid construction (see 
<bibRefCitation id="EFBA53EAFFEFFFB301E6B8637F3CFCAE" author="Brunet, M." box="[1198,1340,829,855]" pageId="33" pageNumber="484" pagination="157 - 174" refId="ref37355" refString="Brunet, M. (1972) Koinocystididae de la region de Marseille. Zoologica Scripta, 1, 157 - 174. https: // doi. org / 10.1111 / j. 1463 - 6409.1972. tb 00673. x" type="journal article" year="1972">Brunet 1972</bibRefCitation>
; 
<bibRefCitation id="EFBA53EAFFEFFFB3000FB8627AD3FC82" author="Karling, T. G." pageId="33" pageNumber="484" pagination="241 - 269" refId="ref38008" refString="Karling, T. G. (1980) Revision of Koinocystididae (Turbellaria). Zoologica Scripta, 9, 241 - 269. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00666. x" type="journal article" year="1980">Karling 1980</bibRefCitation>
), without a juncture sphincter. In live specimens it represents about 20% of the body length in live specimens. The exact number of fixators (
<figureCitation id="1310329EFFEFFFB304BDB8DA785AFC66" box="[501,602,900,927]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB307C9B8DB78B5FC66" box="[641,693,901,927]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: pfix) and retractors (
<figureCitation id="1310329EFFEFFFB306E6B8DA7E17FC66" box="[942,1047,900,927]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: pret) muscles could not be determined. The cone retractors (
<figureCitation id="1310329EFFEFFFB30491B8F67842FC3A" box="[473,578,936,963]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: cret) are well developed. A pair of ventral and a pair of dorsal integument retractors (
<figureCitation id="1310329EFFEFFFB3045BB8927B7CFC1E" box="[275,380,972,999]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: iret) are present. Proboscis sheath surrounded by an external layer of longitudinal muscles. Several dilatators (
<figureCitation id="1310329EFFEFFFB30423B8AE7BCFFBF2" box="[363,463,1008,1035]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB304BDB8AF7829FBF2" box="[501,553,1009,1035]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: dil) connect the proboscis sheath with the body wall. Most of these dilatators are very thin, except for some in the proximal half of the proboscis sheath. The proboscis pore (
<figureCitation id="1310329EFFEFFFB30192BF4A7F3EFBD6" box="[1242,1342,1044,1071]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3002CBF4B7F98FBD6" box="[1380,1432,1045,1071]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: prp) is not surrounded by a sphincter. The proboscis sheath is lined by a nucleated and thick epithelium (
<figureCitation id="1310329EFFEFFFB30079BF667F99FBAA" box="[1329,1433,1080,1107]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: pep). This epithelium is continuous with the epithelium covering the proboscis cone. Both epithelia contain oval to circular-shaped glands (reddish stained) (
<figureCitation id="1310329EFFEFFFB3072CBFDE78C7FB62" box="[612,711,1152,1179]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB307A3BFDF791EFB62" box="[747,798,1153,1179]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: epg), albeit the sheath epithelium only in its most proximal part. Several proboscis glands lie just caudal to the proboscis: ventrally and dorsally there are coarse-grained basophilic glands (
<figureCitation id="1310329EFFEFFFB30421BF967BD0FB1A" box="[361,464,1224,1251]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: prg1), and in between them fine-grained eosinophilic (light-pink) (
<figureCitation id="1310329EFFEFFFB30188BF967F27FB1A" box="[1216,1319,1224,1251]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: prg2) and medium-coarse-grained eosinophilic (dark-pink) (
<figureCitation id="1310329EFFEFFFB3078FBFB2792EFAFE" box="[711,814,1260,1287]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: prg3) ones.
</paragraph>
<paragraph id="8B942E1BFFEFFFB3058FBE4E780DF907" blockId="33.[151,1437,288,2043]" pageId="33" pageNumber="484">
The pharynx (
<figureCitation id="1310329EFFEFFFB3042ABE4E7BC4FAD2" box="[354,452,1296,1323]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB304AEBE4F783CFAD2" box="[486,572,1297,1323]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A–D</figureCitation>
: ph, 17B) is located at 30%. It has a diameter of about 15% of the body length. The prepharyngeal cavity is lined by a nucleated epithelium and is surrounded by a layer of longitudinal muscles. Additional circular muscles occur externally from the longitudinal ones in the distal half of the cavity. The mouth (
<figureCitation id="1310329EFFEFFFB305E8BE227B0AFA6E" box="[160,266,1404,1431]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: m) can be closed by a sphincter (
<figureCitation id="1310329EFFEFFFB307D3BE227905FA6E" box="[667,773,1404,1431]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: sph). Two 
<typeStatus id="549090B9FFEFFFB306C6BE2379C9FA6E" box="[910,969,1405,1431]" pageId="33" pageNumber="484">types</typeStatus>
of glands open into the pharynx lumen: coarse-grained eosinophilic ones distally (
<figureCitation id="1310329EFFEFFFB3073ABEFE78DAFA42" box="[626,730,1440,1467]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: phg1), and more to the middle part of the pharynx, additional coarse-grained basophilic glands occur (
<figureCitation id="1310329EFFEFFFB3071EBE9A78BCFA26" box="[598,700,1476,1503]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: phg2).An epithelium lining the pharynx lumen was not observed. The musculature of the pharynx consists of a layer of longitudinal muscles outside of the septum (
<figureCitation id="1310329EFFEFFFB301A5BEB67F55F9FA" box="[1261,1365,1512,1539]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: lm1), which is continuous with the longitudinal muscles surrounding the prepharyngeal cavity, and a thick circular layer just inside (
<figureCitation id="1310329EFFEFFFB30455BD6E7B86F9B2" box="[285,390,1584,1611]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: cm1). The longitudinal muscles at the inside of the septum, observed in several species of 
<taxonomicName id="4C2B5598FFEFFFB305DFBD0B7AF3F997" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[151,243,1621,1646]" class="Rhabditophora" family="Koinocystididae" genus="Itaipusa" kingdom="Animalia" order="Rhabdocoela" pageId="33" pageNumber="484" phylum="Platyhelminthes" rank="genus">
<emphasis id="B95FF209FFEFFFB305DFBD0B7AF3F997" box="[151,243,1621,1646]" italics="true" pageId="33" pageNumber="484">Itaipusa</emphasis>
</taxonomicName>
, are not observed in this species. The pharynx lumen is surrounded by an inner circular muscle layer (
<figureCitation id="1310329EFFEFFFB30027BD0A7ACAF96A" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: cm2) and an outer longitudinal one (
<figureCitation id="1310329EFFEFFFB3072CBD2678CBF96A" box="[612,715,1656,1683]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: lm2). Internal strong longitudinal muscles are present (
<figureCitation id="1310329EFFEFFFB3007BBD267F99F96A" box="[1331,1433,1656,1683]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: ilm). Radial muscles (
<figureCitation id="1310329EFFEFFFB304DABDC27BFAF94E" box="[402,506,1692,1719]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: rm) run between the septum and the wall of the pharynx lumen. The oesophagus (
<figureCitation id="1310329EFFEFFFB305E8BD9E7B03F922" box="[160,259,1728,1755]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB30460BD9F7B5CF922" box="[296,348,1729,1755]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19A</figureCitation>
: oe) is surrounded by the digestive parenchyma (
<figureCitation id="1310329EFFEFFFB306CFBD9E79EFF922" box="[903,1007,1728,1755]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17B</figureCitation>
: dip). This parenchyma shows several nuclei and dark-stained vacuoles.
</paragraph>
<paragraph id="8B942E1BFFEFFFB0058FBC567ACAFE0C" blockId="33.[151,1437,288,2043]" lastBlockId="34.[151,1436,151,501]" lastPageId="34" lastPageNumber="485" pageId="33" pageNumber="484">
The gonads and atrial organs are located in the caudal fourth of the body. The pair of testes (
<figureCitation id="1310329EFFEFFFB301BFBC567F63F8DA" box="[1271,1379,1800,1827]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
: t) is ventral. The vasa deferentia each form a seminal vesicle (
<figureCitation id="1310329EFFEFFFB30650BC72797BF8BE" box="[792,891,1836,1863]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB306D6BC7379BEF8BE" box="[926,958,1837,1863]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">18</figureCitation>
: sv), and these fuse at the point where they enter the copulatory bulb (
<figureCitation id="1310329EFFEFFFB304F5BC0E781FF892" box="[445,543,1872,1899]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3070ABC0F7895F892" box="[578,661,1873,1899]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19B–C</figureCitation>
: cb). At the same point, also the prostate glands (
<figureCitation id="1310329EFFEFFFB301FCBC0E7F1BF892" box="[1204,1307,1872,1899]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
, 
<figureCitation id="1310329EFFEFFFB3006EBC0F7F42F892" box="[1318,1346,1873,1899]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3002CBC0F7F98F892" box="[1380,1432,1873,1899]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19D</figureCitation>
: pg) open into the copulatory bulb. The copulatory bulb encompasses the free prostate vesicle (
<figureCitation id="1310329EFFEFFFB30183BC2A7F1DF876" box="[1227,1309,1908,1935]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3000BBC2B7F99F876" box="[1347,1433,1909,1935]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19B–D</figureCitation>
: pv), the seminal duct (
<figureCitation id="1310329EFFEFFFB304DABCC67BE2F84A" box="[402,482,1944,1971]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3074EBCC7783AF84A" box="[518,570,1945,1971]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19D</figureCitation>
: sd) and the male duct, which ends in an unarmed penis papilla (
<figureCitation id="1310329EFFEFFFB30046BCC67F75F84A" box="[1294,1397,1944,1971]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="33" pageNumber="484">Fig. 17A</figureCitation>
, 
<figureCitation id="1310329EFFEFFFB300C8BCC77F9CF84A" box="[1408,1436,1945,1971]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB305FBBCE37AE5F82E" box="[179,229,1981,2007]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19B</figureCitation>
: pp). The rest of the space of the copulatory bulb is filled with parenchymatic tissue, which appears syncytial (
<figureCitation id="1310329EFFEFFFB305E8BCBE7AF0F802" box="[160,240,2016,2043]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="33" pageNumber="484">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEFFFB3045BBCBF7B45F802" box="[275,325,2017,2043]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="33" pageNumber="484">19B</figureCitation>
: np). The seminal vesicles and the seminal duct are lined by a nucleated epithelium. The copulatory bulb is surrounded by a thick coat of circular muscles, which gradually becomes thinner distally and eventually disappears toward the penis papilla. The prostate vesicle opens proximally into the ejaculatory duct (
<figureCitation id="1310329EFFECFFB00193BBE57F30FF2C" box="[1243,1328,187,213]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
: ed), next to the opening of the seminal duct (
<figureCitation id="1310329EFFECFFB0076EBB81787BFF00" box="[550,635,223,249]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
: sd), hence the copulatory organ is of the divisa 
<typeStatus id="549090B9FFECFFB001D3BB817ECBFF00" box="[1179,1227,223,249]" pageId="34" pageNumber="485">type</typeStatus>
(see 
<bibRefCitation id="EFBA53EAFFECFFB0004FBB817F9CFF00" author="Karling, T. G." box="[1287,1436,222,249]" pageId="34" pageNumber="485" pagination="187 - 279" refId="ref37917" refString="Karling, T. G. (1956) Morphologisch-histologische Untersuchungen an den mannlichen Atrialorganen der Kalyptorhynchia (Turbellaria). Arkiv for Zoologi, Serie 2, 9 (7), 187 - 279." type="journal article" year="1956">Karling 1956</bibRefCitation>
for details). The prostate vesicle consists of a number of a coarse-grained eosinophilic glands. A connection with these prostate glands and the ones entering the copulatory bulb could not be observed, but as nuclei are only found in the extracapsular parts it is most likely that both are part of the same glandular system. The male duct is lined by a nucleated epithelium and surrounded by a layer of longitudinal muscles. Distally, the epithelium is thicker, and the male duct forms the penis papilla, which protrudes into the male atrium (
<figureCitation id="1310329EFFECFFB006A1BACD7E3DFE54" box="[1001,1085,403,429]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
: ma). Only the distal part of the male atrium seems to be surrounded by muscles, which are longitudinal; no muscles were seen around the proximal part.
</paragraph>
<caption id="DF547E93FFECFFB005DFBD8B7821F8E8" ID-DOI="http://doi.org/10.5281/zenodo.4629295" ID-Zenodo-Dep="4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="34" pageNumber="485" startId="34.[151,250,1749,1773]" targetBox="[158,1427,534,1717]" targetPageId="34">
<paragraph id="8B942E1BFFECFFB005DFBD8B7821F8E8" blockId="34.[151,1437,1749,1809]" pageId="34" pageNumber="485">
<emphasis id="B95FF209FFECFFB005DFBD8B7B20F914" bold="true" box="[151,288,1749,1773]" pageId="34" pageNumber="485">FIGURE 17.</emphasis>
<taxonomicName id="4C2B5598FFECFFB0046EBD8B780FF914" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[294,527,1749,1773]" class="Rhabditophora" family="Koinocystididae" genus="Galapagetula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="34" pageNumber="485" phylum="Platyhelminthes" rank="species" species="cubensis" status="sp. nov.">
<emphasis id="B95FF209FFECFFB0046EBD8B780FF914" box="[294,527,1749,1773]" italics="true" pageId="34" pageNumber="485">Galapagetula cubensis</emphasis>
</taxonomicName>
<emphasis id="B95FF209FFECFFB0075DBD8B784EF914" bold="true" box="[533,590,1749,1773]" pageId="34" pageNumber="485">
<taxonomicNameLabel id="A26C4F72FFECFFB0075DBD8B784EF914" box="[533,590,1749,1773]" pageId="34" pageNumber="485" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side.
</paragraph>
</caption>
<paragraph id="8B942E1BFFECFFB1058FBC6179DAFD7F" blockId="34.[151,1437,1855,2026]" lastBlockId="35.[151,1437,151,646]" lastPageId="35" lastPageNumber="486" pageId="34" pageNumber="485">
The vitellaria (
<figureCitation id="1310329EFFECFFB00426BC617BD2F8A0" box="[366,466,1855,1881]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="34" pageNumber="485">Fig. 17A</figureCitation>
&amp; 
<figureCitation id="1310329EFFECFFB004BFBC61784DF8A0" box="[503,589,1855,1881]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="34" pageNumber="485">19A–D</figureCitation>
: vi) extend along both sides of the body from behind the pharynx up to the level of the copulatory bulb. The elongated ovaries (
<figureCitation id="1310329EFFECFFB007A8BC3D7949F884" box="[736,841,1891,1917]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="34" pageNumber="485">Fig. 17A</figureCitation>
: ov) are located on either side of the copulatory bulb. The oocytes are organised in a row, increasing in diameter from the most proximal to the most distal one. The oviducts (
<figureCitation id="1310329EFFECFFB005AABCF57B38F83C" box="[226,312,1963,1989]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
: od) open into the female duct, which distally is a bit swollen (
<figureCitation id="1310329EFFECFFB00148BCF57E52F83C" box="[1024,1106,1963,1989]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="34" pageNumber="485">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFECFFB0013FBCF57ECCF83C" box="[1143,1228,1963,1989]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="34" pageNumber="485">19C–D</figureCitation>
: fd). Both the oviducts and the female duct are lined by a nucleated epithelium. Distally, the female duct is surrounded by a thin lon- gitudinal muscle layer. It enters the female atrium through the latter’s rostro-ventral wall (
<figureCitation id="1310329EFFEDFFB101CEBBC97EEDFF48" box="[1158,1261,151,177]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="35" pageNumber="486">Fig. 17A</figureCitation>
, 
<figureCitation id="1310329EFFEDFFB101B1BBC97F15FF48" box="[1273,1301,151,177]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB10071BBC97F6BFF48" box="[1337,1387,151,177]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19B</figureCitation>
: fa). Two bursae open into the female atrium. A resorptive bursa (
<figureCitation id="1310329EFFEDFFB1060CBBE57995FF2C" box="[836,917,187,213]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB106F2BBE57E0DFF2C" box="[954,1037,187,213]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19B–C</figureCitation>
: b1) opens rostrally into the female atrium and contains degraded material. It is lined by a nucleated epithelium. Its distal, tubiform part (bursal stalk) is surrounded by thin longitudinal muscles. A well-developed sphincter occurs at the place where this bursa opens into the female atrium (
<figureCitation id="1310329EFFEDFFB104D7BA797BF3FEB8" box="[415,499,295,321]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
: sph1, 19B–D: sph). A second, muscular bursa [
<figureCitation id="1310329EFFEDFFB10144BA797E5EFEB8" box="[1036,1118,295,321]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB101CABA797EB6FEB8" box="[1154,1206,295,321]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19D</figureCitation>
: b2; accessory bursa in the terminology of 
<bibRefCitation id="EFBA53EAFFEDFFB104C6BA157872FE9C" author="Reygel, P. C. &amp; Willems, W. R. &amp; Artois, T. J." box="[398,626,330,357]" pageId="35" pageNumber="486" pagination="27 - 40" refId="ref39019" refString="Reygel, P. C., Willems, W. R. &amp; Artois, T. J. (2011) Koinocystididae and Gnathorhynchidae (Platyhelminthes: Rhabdocoela: Kalyptorhynchia) from the Galapagos, with the description of three new species. Zootaxa, 3096 (1), 27 - 40. https: // doi. org / 10.11646 / zootaxa. 3096.1.3" type="journal article" year="2011">
Reygel 
<emphasis id="B95FF209FFEDFFB104AEBA15781FFE9C" box="[486,543,331,357]" italics="true" pageId="35" pageNumber="486">et al.</emphasis>
(2011)
</bibRefCitation>
] opens dorsal-rostrally into the female atrium. It is lined by a nucleated epithelium and surrounded by a layer of longitudinal muscles. The female atrium is lined by a nucleated epithelium and surrounded by an internal layer of longitudinal muscles and an external layer of very strong circular muscles (
<figureCitation id="1310329EFFEDFFB105E8BAE97B08FE28" box="[160,264,439,465]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">Fig. 19C</figureCitation>
: cm). The uterus (
<figureCitation id="1310329EFFEDFFB10493BAE97844FE28" box="[475,580,439,465]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="35" pageNumber="486">Fig. 17A</figureCitation>
, 
<figureCitation id="1310329EFFEDFFB10719BAE9786DFE28" box="[593,621,439,465]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB107DDBAE978E8FE28" box="[661,744,439,465]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19B–C</figureCitation>
: ut) is lined by a nucleated epithelium and by a longitudinal muscle layer. Coarse-grained basophilic (
<figureCitation id="1310329EFFEDFFB10722BA8578BBFE0C" box="[618,699,475,501]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB107A9BA857913FE0C" box="[737,787,475,501]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19B</figureCitation>
: ubg) and fine-grained eosinophilic (
<figureCitation id="1310329EFFEDFFB101FDBA857F06FE0C" box="[1205,1286,475,501]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB10064BA857F5EFE0C" box="[1324,1374,475,501]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19B</figureCitation>
: ueg) glands open into the distal part of the uterus. Proximally and distally from these glands sphincters are present (
<figureCitation id="1310329EFFEDFFB10027BAA17AB7FDC4" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
: sph2 &amp; sph3). The common genital atrium (
<figureCitation id="1310329EFFEDFFB107E3B97D78FFFDC4" box="[683,767,547,573]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
: ca) is lined by a nucleated epithelium and by a longitudinal muscle layer. It opens to the outside through the common gonopore, which is situated exactly terminally (
<figureCitation id="1310329EFFEDFFB10079B9197F98FD98" box="[1329,1432,583,609]" captionStart="FIGURE 17" captionStartId="34.[151,250,1749,1773]" captionTargetBox="[158,1427,534,1717]" captionTargetId="figure-200@34.[151,1436,525,1725]" captionTargetPageId="34" captionText="FIGURE 17. Galapagetula cubensis sp. n.. A, general organisation (from a live animal). B, sagittal reconstruction of the anterior body end from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629295" httpUri="https://zenodo.org/record/4629295/files/figure.png" pageId="35" pageNumber="486">Fig. 17A</figureCitation>
, 
<figureCitation id="1310329EFFEDFFB105DFB9357AB3FD7C" box="[151,179,619,645]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">18</figureCitation>
&amp; 
<figureCitation id="1310329EFFEDFFB1059FB9357B0BFD7C" box="[215,267,619,645]" captionStart="FIGURE 19" captionStartId="36.[151,250,1171,1195]" captionTargetBox="[151,1436,181,1148]" captionTargetId="figure-23@36.[151,1436,181,1148]" captionTargetPageId="36" captionText="FIGURE 19. Galapagetula cubensis sp. n.. A, sagittal section through the anterior body end. B–D, sagittal sections through the genital system." figureDoi="http://doi.org/10.5281/zenodo.4629299" httpUri="https://zenodo.org/record/4629299/files/figure.png" pageId="35" pageNumber="486">19D</figureCitation>
: cg). The gonopore is surrounded by a sphincter (
<figureCitation id="1310329EFFEDFFB1067EB935798AFD7C" box="[822,906,619,645]" captionStart="FIGURE 18" captionStartId="35.[151,250,1934,1958]" captionTargetBox="[157,1426,678,1900]" captionTargetId="figure-278@35.[151,1436,669,1911]" captionTargetPageId="35" captionText="FIGURE 18. Galapagetula cubensis sp. n.. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629297" httpUri="https://zenodo.org/record/4629297/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
: sph4).
</paragraph>
</subSubSection>
</treatment>
</document>