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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1152.98258" ID-Pensoft-Pub="1313-2970-1152-45" ID-Pensoft-UUID="7CD89C59351C5B439C82F9AEEA86CC51" ID-ZooBank="D20AFC88096345FEA8B074D00728424F" ModsDocID="1313-2970-1152-45" checkinTime="1678295859973" checkinUser="pensoft" docAuthor="Innabi, Julie, Stout, Carla C. &amp; Valdes, Angel" docDate="2023" docId="035E119B8A7557F085A85A99EBACFD40" docLanguage="en" docName="ZooKeys 1152: 45-95" docOrigin="ZooKeys 1152" docPubDate="2023-03-07" docSource="http://dx.doi.org/10.3897/zookeys.1152.98258" docTitle="Sclerodoris tuberculata Eliot 1904" docType="treatment" docVersion="1" id="7CD89C59351C5B439C82F9AEEA86CC51" lastPageNumber="45" masterDocId="7CD89C59351C5B439C82F9AEEA86CC51" masterDocTitle="Seven new &quot; cryptic &quot; species of Discodorididae (Mollusca, Gastropoda, Nudibranchia) from New Caledonia" masterLastPageNumber="95" masterPageNumber="45" pageNumber="45" updateTime="1678295859973" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new &quot; cryptic &quot; species of Discodorididae (Mollusca, Gastropoda, Nudibranchia) from New Caledonia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Innabi, Julie</mods:namePart>
<mods:affiliation>Department of Biological Sciences, California State Polytechnic University Pomona, 3801 West Temple Avenue, Pomona, California 91768, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stout, Carla C.</mods:namePart>
<mods:affiliation>Department of Biological Sciences, California State Polytechnic University Pomona, 3801 West Temple Avenue, Pomona, California 91768, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Valdes, Angel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2347-4896</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, California State Polytechnic University Pomona, 3801 West Temple Avenue, Pomona, California 91768, USA</mods:affiliation>
<mods:nameIdentifier type="email">aavaldes@cpp.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1152</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>95</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1152.98258</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1152.98258</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1152-45</mods:identifier>
<mods:identifier type="ZooBank">D20AFC88096345FEA8B074D00728424F</mods:identifier>
<mods:identifier type="Pensoft-UUID">7CD89C59351C5B439C82F9AEEA86CC51</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:035E119B8A7557F085A85A99EBACFD40" httpUri="http://treatment.plazi.org/id/035E119B8A7557F085A85A99EBACFD40" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName ID-CoL="4VPRY" ID-ENA="2049550" LSID="035E119B-8A75-57F0-85A8-5A99EBACFD40" authority="Eliot, 1904" authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">Sclerodoris tuberculata Eliot, 1904</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Photographs of live animals of the genus Sclerodoris Eliot, 1904 A Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197 on black background B-D &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., Holotype (MNHN IM- 2013 - 86193) on black background (B), MNHN IM- 2013 - 86195 on black background (C), MNHN IM- 2013 - 86194 on black background (D) E Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198) on black background." figureDoi="10.3897/zookeys.1152.98258.figure14" httpUri="https://binary.pensoft.net/fig/820842" pageId="0" pageNumber="45">Figs 14A</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawing of the reproductive system of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, general view (A), detail of the bursa copulatrix and seminal receptable (B), penial spines (C) D-F Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), general view (D), detail of the bursa copulatrix and seminal receptable (E), penial spines (F) G &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86193. Abbreviations: ag, accessory gland; am, ampulla; bc, bursa copulatrix; dd, deferent duct; fg, female gland complex; pr, prostate; sr, seminal receptacle; vg, vagina." figureDoi="10.3897/zookeys.1152.98258.figure15" httpUri="https://binary.pensoft.net/fig/820843" pageId="0" pageNumber="45">, 15A-C</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. SEM of the radula of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, innermost teeth (A), mid-lateral teeth (B), outer lateral teeth (C) D-F &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86195, innermost teeth (D), mid-lateral teeth (E), outer lateral teeth (F) G-I Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), innermost teeth (G), mid-lateral teeth (H), outer lateral teeth (I)." figureDoi="10.3897/zookeys.1152.98258.figure16" httpUri="https://binary.pensoft.net/fig/820844" pageId="0" pageNumber="45">, 16A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Kelaart" authorityYear="1858" class="Gastropoda" family="Discodorididae" genus="Doris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Doris castanea" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="castanea">Doris castanea</taxonomicName>
?
<taxonomicName authorityName="Kelaart" authorityYear="1858" class="Gastropoda" family="Discodorididae" genus="Doris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Doris castanea" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="castanea">Doris castanea</taxonomicName>
Kelaart, 1858: 110. Type locality: Sober Island, Tricomalie [= Trincomalee] harbor, Ceylon [= Sri Lanka].
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">Sclerodoris tuberculata</taxonomicName>
Eliot, 1904: 381-382. Type locality: Prison Island [= Changuu], Zanzibar harbor, Tanzania.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris minor" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="minor">Sclerodoris minor</taxonomicName>
Eliot, 1904: 381. Type locality: Chuaka [= Chwaka], Zanzibar, Tanzania.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">Sclerodoris rubra</taxonomicName>
Eliot, 1904: 382-383. Type locality: reef off the east coast of Zanzibar, Tanzania.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Gastropoda" family="Discodorididae" genus="Halgerda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halgerda rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">Halgerda rubra</taxonomicName>
Bergh, 1905: 126-127, pl. 4 fig. 2, pl. 15 figs 34-36. Type locality: Bandas [= Banda Islands], Indonesia, 36 m depth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation collectingDate="2018-09-26" country="New Caledonia" elevation="7" latitude="-20.581667" location="Pointe de Pandop" longLatPrecision="126" longitude="164.275" municipality="Koumac" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:035E119B8A7557F085A85A99EBACFD40:FAD91CF5E83365A37E00BA2958D49A03" country="New Caledonia" latitude="-20.581667" longLatPrecision="126" longitude="164.275" municipality="Koumac" name="Pointe de Pandop">Pointe de Pandop</location>
,
<collectingMunicipality>Koumac</collectingMunicipality>
,
<collectingCountry name="New Caledonia">New Caledonia</collectingCountry>
(
<geoCoordinate degrees="20" direction="south" minutes="34.9" orientation="latitude" precision="92" value="-20.581667">20°34.9'S</geoCoordinate>
,
<geoCoordinate degrees="164" direction="east" minutes="16.5" orientation="longitude" precision="92" value="164.275">164°16.5'E</geoCoordinate>
),
<elevation metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">
<quantity metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">7 m</quantity>
depth
</elevation>
[Koumac 2.1 stn. KR868],
<collectingDate value="2018-09-26">26 Sep 2018</collectingDate>
,
<specimenCount type="generic">1 specimen</specimenCount>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.4" unit="mm" value="44.0">44 mm</quantity>
long, dissected (MNHN IM-2013-86197, isolate JI10)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
Body oval, flattened, with an irregular, coriaceous texture (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Photographs of live animals of the genus Sclerodoris Eliot, 1904 A Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197 on black background B-D &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., Holotype (MNHN IM- 2013 - 86193) on black background (B), MNHN IM- 2013 - 86195 on black background (C), MNHN IM- 2013 - 86194 on black background (D) E Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198) on black background." figureDoi="10.3897/zookeys.1152.98258.figure14" httpUri="https://binary.pensoft.net/fig/820842" pageId="0" pageNumber="45">14A</figureCitation>
). Branchial and rhinophoral sheaths somewhat elevated, simple, circular. Gill composed of eight short, tripinnate branchial leaves, arranged upright. Rhinophores short, lamellated, with 18 lamellae. Visceral hump clearly elevated over the rest of the mantle, with several lateral protuberances and a conspicuous depression mid-length. Dorsum completely covered with small caryophyllidia. Body color red, with several large, irregularly opaque white patches, mainly on the mantle margin and some white pigment irregularly scattered all over. Rhinophores and branchial leaves are the same color as the dorsum.
</paragraph>
<caption doi="10.3897/zookeys.1152.98258.figure14" httpUri="https://binary.pensoft.net/fig/820842" pageId="0" pageNumber="45" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 14.</emphasis>
Photographs of live animals of the genus
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
Eliot, 1904
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904, MNHN IM-2013-86197 on black background
<emphasis bold="true" pageId="0" pageNumber="45">B-D</emphasis>
&quot;
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
&quot;
<taxonomicName authorityName="&quot;" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris dutertrei" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="dutertrei">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris dutertrei</emphasis>
</taxonomicName>
sp. nov., Holotype (MNHN IM-2013-86193) on black background (
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
), MNHN IM-2013-86195 on black background (
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
), MNHN IM-2013-86194 on black background (
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="45">E</emphasis>
<taxonomicName authorityName="Innabi &amp; Stout &amp; Valdés" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris faninozi" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="faninozi">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris faninozi</emphasis>
</taxonomicName>
sp. nov., Holotype (MNHN IM-2013-86198) on black background.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
Reproductive system (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawing of the reproductive system of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, general view (A), detail of the bursa copulatrix and seminal receptable (B), penial spines (C) D-F Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), general view (D), detail of the bursa copulatrix and seminal receptable (E), penial spines (F) G &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86193. Abbreviations: ag, accessory gland; am, ampulla; bc, bursa copulatrix; dd, deferent duct; fg, female gland complex; pr, prostate; sr, seminal receptacle; vg, vagina." figureDoi="10.3897/zookeys.1152.98258.figure15" httpUri="https://binary.pensoft.net/fig/820843" pageId="0" pageNumber="45">15A, B</figureCitation>
) with a long, wide, convoluted ampulla with several folds, which connects with the female gland complex and the oval, flattened prostate. The prostate narrows substantially into a long, straight duct, before expanding into the short, wide deferent duct. The penis is armed with triangular spines, varying in size (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawing of the reproductive system of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, general view (A), detail of the bursa copulatrix and seminal receptable (B), penial spines (C) D-F Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), general view (D), detail of the bursa copulatrix and seminal receptable (E), penial spines (F) G &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86193. Abbreviations: ag, accessory gland; am, ampulla; bc, bursa copulatrix; dd, deferent duct; fg, female gland complex; pr, prostate; sr, seminal receptacle; vg, vagina." figureDoi="10.3897/zookeys.1152.98258.figure15" httpUri="https://binary.pensoft.net/fig/820843" pageId="0" pageNumber="45">15C</figureCitation>
) with thickened bases and sharp cusps. The vagina is elongate, narrow, as wide as the deferent duct, connecting directly to the large, oval bursa copulatrix. The elongate seminal receptacle also connects to the bursa copulatrix next to the vaginal connection, and the short uterine duct that enters the female gland complex (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drawing of the reproductive system of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, general view (A), detail of the bursa copulatrix and seminal receptable (B), penial spines (C) D-F Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), general view (D), detail of the bursa copulatrix and seminal receptable (E), penial spines (F) G &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86193. Abbreviations: ag, accessory gland; am, ampulla; bc, bursa copulatrix; dd, deferent duct; fg, female gland complex; pr, prostate; sr, seminal receptacle; vg, vagina." figureDoi="10.3897/zookeys.1152.98258.figure15" httpUri="https://binary.pensoft.net/fig/820843" pageId="0" pageNumber="45">15B</figureCitation>
). The bursa copulatrix is ~ 4
<normalizedToken originalValue="×">x</normalizedToken>
as large as the seminal receptable. An accessory gland connects to the genial atrium where the deferent duct and the vagina meet. The accessory gland is granular in texture and approximately as large as the seminal receptable.
</paragraph>
<caption doi="10.3897/zookeys.1152.98258.figure15" httpUri="https://binary.pensoft.net/fig/820843" pageId="0" pageNumber="45" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 15.</emphasis>
Drawing of the reproductive system of specimens of the genus
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
Eliot, 1904
<emphasis bold="true" pageId="0" pageNumber="45">A-C</emphasis>
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904, MNHN IM-2013-86197, general view (
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
), detail of the bursa copulatrix and seminal receptable (
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
), penial spines (
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="45">D-F</emphasis>
<taxonomicName authorityName="Innabi &amp; Stout &amp; Valdés" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris faninozi" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="faninozi">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris faninozi</emphasis>
</taxonomicName>
sp. nov., Holotype (MNHN IM-2013-86198), general view (
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
), detail of the bursa copulatrix and seminal receptable (
<emphasis bold="true" pageId="0" pageNumber="45">E</emphasis>
), penial spines (
<emphasis bold="true" pageId="0" pageNumber="45">F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="45">G</emphasis>
&quot;
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
&quot;
<taxonomicName authorityName="&quot;" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris dutertrei" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="dutertrei">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris dutertrei</emphasis>
</taxonomicName>
sp. nov., MNHN IM-2013-86193. Abbreviations: ag, accessory gland; am, ampulla; bc, bursa copulatrix; dd, deferent duct; fg, female gland complex; pr, prostate; sr, seminal receptacle; vg, vagina.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
Radular formula 38
<normalizedToken originalValue="×">x</normalizedToken>
49.0.49 in a 44-mm long specimen (MNHN IM-2013-86197). Rachidian teeth absent. Inner and mid-lateral teeth hamate, having an elongate cusp and lacking denticles (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. SEM of the radula of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, innermost teeth (A), mid-lateral teeth (B), outer lateral teeth (C) D-F &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86195, innermost teeth (D), mid-lateral teeth (E), outer lateral teeth (F) G-I Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), innermost teeth (G), mid-lateral teeth (H), outer lateral teeth (I)." figureDoi="10.3897/zookeys.1152.98258.figure16" httpUri="https://binary.pensoft.net/fig/820844" pageId="0" pageNumber="45">16A, B</figureCitation>
). Innermost teeth very small in comparison to mid-laterals (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. SEM of the radula of specimens of the genus Sclerodoris Eliot, 1904 A-C Sclerodoris tuberculata Eliot, 1904, MNHN IM- 2013 - 86197, innermost teeth (A), mid-lateral teeth (B), outer lateral teeth (C) D-F &quot; Sclerodoris &quot; Sclerodoris dutertrei sp. nov., MNHN IM- 2013 - 86195, innermost teeth (D), mid-lateral teeth (E), outer lateral teeth (F) G-I Sclerodoris faninozi sp. nov., Holotype (MNHN IM- 2013 - 86198), innermost teeth (G), mid-lateral teeth (H), outer lateral teeth (I)." figureDoi="10.3897/zookeys.1152.98258.figure16" httpUri="https://binary.pensoft.net/fig/820844" pageId="0" pageNumber="45">16A</figureCitation>
). The teeth increase in size gradually towards the medial portion of the half-row. Outermost teeth small, decreasing in size gradually, composed of a short, blunt cusp with numerous small denticles (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEM of the radula of specimens of Rostanga poddubetskaiae sp. nov., MNHN IM- 2013 - 86205 A innermost teeth B mid-lateral teeth C outer lateral teeth." figureDoi="10.3897/zookeys.1152.98258.figure13" httpUri="https://binary.pensoft.net/fig/820841" pageId="0" pageNumber="45">13C</figureCitation>
). No jaw was observed, labial cuticle smooth.
</paragraph>
<caption doi="10.3897/zookeys.1152.98258.figure16" httpUri="https://binary.pensoft.net/fig/820844" pageId="0" pageNumber="45" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 16.</emphasis>
SEM of the radula of specimens of the genus
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
Eliot, 1904
<emphasis bold="true" pageId="0" pageNumber="45">A-C</emphasis>
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904, MNHN IM-2013-86197, innermost teeth (
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
), mid-lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
), outer lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="45">D-F</emphasis>
&quot;
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris</emphasis>
</taxonomicName>
&quot;
<taxonomicName authorityName="&quot;" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris dutertrei" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="dutertrei">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris dutertrei</emphasis>
</taxonomicName>
sp. nov., MNHN IM-2013-86195, innermost teeth (
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
), mid-lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">E</emphasis>
), outer lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="45">G-I</emphasis>
<taxonomicName authorityName="Innabi &amp; Stout &amp; Valdés" authorityYear="2023" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris faninozi" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="faninozi">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris faninozi</emphasis>
</taxonomicName>
sp. nov., Holotype (MNHN IM-2013-86198), innermost teeth (
<emphasis bold="true" pageId="0" pageNumber="45">G</emphasis>
), mid-lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">H</emphasis>
), outer lateral teeth (
<emphasis bold="true" pageId="0" pageNumber="45">I</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="biology_ecology">
<paragraph pageId="0" pageNumber="45">Biology.</paragraph>
<paragraph pageId="0" pageNumber="45">Rare, found under rocks at 7 m depth. Widespread in the Indo-Pacific region. The single specimen was found under a rock while SCUBA diving where it was highly cryptic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="remarks">
<paragraph pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation author="Eliot, CNE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="45" pagination="354 - 385" refId="B28" refString="Eliot, CNE, 1904. On some nudibranchs from East Africa and Zanzibar part. III. Proceedings of the Zoological Society of London 1903: 354 - 385" title="On some nudibranchs from East Africa and Zanzibar part. III." volume="1903" year="1904">Eliot (1904)</bibRefCitation>
described
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
based on one specimen collected in Zanzibar as follows: &quot;Dark brown with sandy spots, exactly like a sponge splashed with sand. Underside clear bright brownish red. Branchial pocket crenulate. The middle part of back covered with conical warts, which form an irregular keel; smaller warts on mantle-edge. Rhinophores red; branchiae eight, voluminous; axes red, tips white. Animal alters shape, sometimes rather high, sometimes quite flat like
<taxonomicName authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Platydoris" kingdom="Animalia" lsidName="Platydoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Platydoris</emphasis>
</taxonomicName>
. Consistency quite hard and rather rough. Two depressions with deep black markings as in some species of
<taxonomicName authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Trippa" kingdom="Animalia" lsidName="Trippa" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Trippa</emphasis>
</taxonomicName>
.&quot; In the same paper
<bibRefCitation author="Eliot, CNE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="45" pagination="354 - 385" refId="B28" refString="Eliot, CNE, 1904. On some nudibranchs from East Africa and Zanzibar part. III. Proceedings of the Zoological Society of London 1903: 354 - 385" title="On some nudibranchs from East Africa and Zanzibar part. III." volume="1903" year="1904">Eliot (1904)</bibRefCitation>
introduced two additional species also resembling sponges,
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris minor" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris minor</emphasis>
</taxonomicName>
Eliot, 1904, and
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubra</emphasis>
</taxonomicName>
Eliot, 1904, both synonyms of
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
is considered a widespread species in the Indo-Pacific region and is well documented in the literature (
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.2001.tb00689.x" author="Valdes, A" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="103 - 198" refId="B84" refString="Valdes, A, Gosliner, TM, 2001. Systematics and phylogeny of the caryophyllidia-bearing dorids (Mollusca, Nudibranchia), with the description of a new genus and four new species from Indo-Pacific deep waters. Zoological Journal of the Linnean Society 133 (2): 103 - 198, DOI: https://doi.org/10.1111/j.1096-3642.2001.tb00689.x" title="Systematics and phylogeny of the caryophyllidia-bearing dorids (Mollusca, Nudibranchia), with the description of a new genus and four new species from Indo-Pacific deep waters." url="https://doi.org/10.1111/j.1096-3642.2001.tb00689.x" volume="133" year="2001">
<normalizedToken originalValue="Valdés">Valdes</normalizedToken>
and Gosliner 2001
</bibRefCitation>
;
<bibRefCitation author="Yonow, N" journalOrPublisher="Pensoft, Sofia" pageId="0" pageNumber="45" refId="B88" refString="Yonow, N, 2008. Sea Slugs of the Red Sea. Pensoft, Sofia" title="Sea Slugs of the Red Sea." year="2008">Yonow 2008</bibRefCitation>
;
<bibRefCitation author="Gosliner, TM" journalOrPublisher="New World Publications, Jacksonville, Florida" pageId="0" pageNumber="45" refId="B38" refString="Gosliner, TM, Valdes, A, Behrens, DW, 2018. Nudibranch and Sea Slug Identification: Indo-Pacific (2nd Ed.). New World Publications, Jacksonville, Florida" title="Nudibranch and Sea Slug Identification: Indo-Pacific (2 nd Ed.)." year="2018">Gosliner et al. 2018</bibRefCitation>
;
<bibRefCitation author="Nakano, R" journalOrPublisher="Bun-ichi Sogo, Shizuoka, Japan" pageId="0" pageNumber="45" refId="B60" refString="Nakano, R, 2018. Field Guide to Sea Slugs and Nudibranchs of Japan. Bun-ichi Sogo, Shizuoka, Japan" title="Field Guide to Sea Slugs and Nudibranchs of Japan." year="2018">Nakano 2018</bibRefCitation>
). The material here examined is consistent with the original description of
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
and subsequent records; however, a record of this species from New Caledonia (
<bibRefCitation author="Herve, J-F" journalOrPublisher="Catherine Ledru, Noumea, New Caledonia" pageId="0" pageNumber="45" refId="B42" refString="Herve, J-F, 2010. Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches. Catherine Ledru, Noumea, New Caledonia" title="Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches." year="2010">
<normalizedToken originalValue="Hervé">Herve</normalizedToken>
2010
</bibRefCitation>
) is probably the closely related species
<taxonomicName baseAuthorityName="Baba" baseAuthorityYear="1949" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubicunda" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubicunda">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubicunda</emphasis>
</taxonomicName>
(Baba, 1949).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation author="Eliot, CNE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="45" pagination="636 - 691" refId="B29" refString="Eliot, CNE, 1906. On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne. Proceedings of the Zoological Society of London 1906: 636 - 691" title="On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne." volume="1906" year="1906">Eliot (1906)</bibRefCitation>
suggested that
<taxonomicName authorityName="Kelaart" authorityYear="1858" class="Gastropoda" family="Discodorididae" genus="Doris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Doris castanea" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="45">Doris castanea</emphasis>
</taxonomicName>
Kelaart, 1858 was possibly the same species as
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904, but indicated the identity of the latter could not be established with certainty based on the type material.
<bibRefCitation author="Eliot, CNE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="45" pagination="636 - 691" refId="B29" refString="Eliot, CNE, 1906. On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne. Proceedings of the Zoological Society of London 1906: 636 - 691" title="On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne." volume="1906" year="1906">Eliot (1906</bibRefCitation>
: pl. 42, figs 6, 7) reproduced the original drawing by Kelaart, which clearly resembles a dark specimen of
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
. Later,
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" author="Eliot, CNE" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="86 - 122" refId="B30" refString="Eliot, CNE, 1908. Reports on the Marine Biology of the Sudanese Red Sea.-XI. Notes of a Collection of Nudibranchs from the Red Sea. Zoological Journal of the Linnean Society 31 (204): 86 - 122, DOI: https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" title="Reports on the Marine Biology of the Sudanese Red Sea. - XI. Notes of a Collection of Nudibranchs from the Red Sea." url="https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" volume="31" year="1908">Eliot (1908)</bibRefCitation>
regarded
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubra</emphasis>
</taxonomicName>
Eliot, 1904 as a senior synonym of
<taxonomicName class="Gastropoda" family="Discodorididae" genus="Halgerda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halgerda rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Halgerda rubra</emphasis>
</taxonomicName>
Bergh, 1905.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.21.1947.561" author="Allan, JK" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="45" pagination="433 - 463" refId="B2" refString="Allan, JK, 1947. Nudibranchia from the Clarence River Heads, north coast, New South Wales. Records of the Australian Museum 21: 433 - 463, DOI: https://doi.org/10.3853/j.0067-1975.21.1947.561" title="Nudibranchia from the Clarence River Heads, north coast, New South Wales." url="https://doi.org/10.3853/j.0067-1975.21.1947.561" volume="21" year="1947">Allan (1947)</bibRefCitation>
reported
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
from New South Wales, Australia, under the genus name
<taxonomicName authorityName="Bergh" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Peronodoris" kingdom="Animalia" lsidName="Peronodoris" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Peronodoris</emphasis>
</taxonomicName>
Bergh, 1904 and commented on
<bibRefCitation author="Eliot, CNE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="45" pagination="636 - 691" refId="B29" refString="Eliot, CNE, 1906. On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne. Proceedings of the Zoological Society of London 1906: 636 - 691" title="On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder and Hancock preserved in the Hancock Museum at Newcastle-on-Tyne." volume="1906" year="1906">
<normalizedToken originalValue="Eliot’s">Eliot's</normalizedToken>
(1906)
</bibRefCitation>
proposed synonymy between this species and
<taxonomicName lsidName="D. castanea" pageId="0" pageNumber="45" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="45">D. castanea</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.21.1947.561" author="Allan, JK" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="45" pagination="433 - 463" refId="B2" refString="Allan, JK, 1947. Nudibranchia from the Clarence River Heads, north coast, New South Wales. Records of the Australian Museum 21: 433 - 463, DOI: https://doi.org/10.3853/j.0067-1975.21.1947.561" title="Nudibranchia from the Clarence River Heads, north coast, New South Wales." url="https://doi.org/10.3853/j.0067-1975.21.1947.561" volume="21" year="1947">Allan (1947)</bibRefCitation>
indicated that &quot;although the colour sketch of the upper surface of
<normalizedToken originalValue="Kelaart’s">Kelaart's</normalizedToken>
specimen resembles that of our specimen to a very slight degree,&quot; the undersurface is exactly like the color sketch of the New South Wales material of
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.21.1947.561" author="Allan, JK" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="45" pagination="433 - 463" refId="B2" refString="Allan, JK, 1947. Nudibranchia from the Clarence River Heads, north coast, New South Wales. Records of the Australian Museum 21: 433 - 463, DOI: https://doi.org/10.3853/j.0067-1975.21.1947.561" title="Nudibranchia from the Clarence River Heads, north coast, New South Wales." url="https://doi.org/10.3853/j.0067-1975.21.1947.561" volume="21" year="1947">Allan (1947)</bibRefCitation>
concluded that whether
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
was eventually to become a synonym of
<taxonomicName lsidName="D. castanea" pageId="0" pageNumber="45" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="45">D. castanea</emphasis>
</taxonomicName>
remained to be seen, as fresh material from the two type localities needs to be examined before this can be determined.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" author="Rudman, WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="59 - 88" refId="B75" refString="Rudman, WB, 1978. The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific. Zoological Journal of the Linnean Society 62 (1): 59 - 88, DOI: https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" title="The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific." url="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" volume="62" year="1978">Rudman (1978)</bibRefCitation>
endorsed
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" author="Eliot, CNE" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="86 - 122" refId="B30" refString="Eliot, CNE, 1908. Reports on the Marine Biology of the Sudanese Red Sea.-XI. Notes of a Collection of Nudibranchs from the Red Sea. Zoological Journal of the Linnean Society 31 (204): 86 - 122, DOI: https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" title="Reports on the Marine Biology of the Sudanese Red Sea. - XI. Notes of a Collection of Nudibranchs from the Red Sea." url="https://doi.org/10.1111/j.1096-3642.1908.tb00457.x" volume="31" year="1908">
<normalizedToken originalValue="Eliot’s">Eliot's</normalizedToken>
(1908)
</bibRefCitation>
decision to synonymize
<taxonomicName class="Gastropoda" family="Discodorididae" genus="Halgerda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halgerda rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Halgerda rubra</emphasis>
</taxonomicName>
Bergh, 1905 with
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubra</emphasis>
</taxonomicName>
Eliot, 1904. At the same time
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" author="Rudman, WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="59 - 88" refId="B75" refString="Rudman, WB, 1978. The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific. Zoological Journal of the Linnean Society 62 (1): 59 - 88, DOI: https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" title="The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific." url="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" volume="62" year="1978">Rudman (1978)</bibRefCitation>
regarded
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubra" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubra">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubra</emphasis>
</taxonomicName>
Eliot, 1904 and
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris minor" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris minor</emphasis>
</taxonomicName>
Eliot, 1904 as synonyms of
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904, and based on the Principle of First Reviser (
<bibRefCitation author="Herve, J-F" journalOrPublisher="Catherine Ledru, Noumea, New Caledonia" pageId="0" pageNumber="45" refId="B43" refString="ICZN (1999) International Code of Zoological Nomenclature (4th edn.).International Trust for Zoological Nomenclature, London, 306 pp." year="2010">ICZN 1999</bibRefCitation>
: Article 24),
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" author="Rudman, WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="59 - 88" refId="B75" refString="Rudman, WB, 1978. The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific. Zoological Journal of the Linnean Society 62 (1): 59 - 88, DOI: https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" title="The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific." url="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" volume="62" year="1978">Rudman (1978)</bibRefCitation>
established
<taxonomicName lsidName="S. tuberculata" pageId="0" pageNumber="45" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">S. tuberculata</emphasis>
</taxonomicName>
as the valid name for this species.
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" author="Rudman, WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="59 - 88" refId="B75" refString="Rudman, WB, 1978. The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific. Zoological Journal of the Linnean Society 62 (1): 59 - 88, DOI: https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" title="The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific." url="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" volume="62" year="1978">Rudman (1978)</bibRefCitation>
also commented that the original description of
<taxonomicName lsidName="D. castanea" pageId="0" pageNumber="45" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="45">D. castanea</emphasis>
</taxonomicName>
by
<bibRefCitation author="Kelaart, EF" journalOrPublisher="Journal of the Ceylon Branch of the Royal Asiatic Society" pageId="0" pageNumber="45" pagination="76 - 124" refId="B50" refString="Kelaart, EF, 1858. New and little known species of Ceylon nudibranchiate molluscs, and zoophytes. Journal of the Ceylon Branch of the Royal Asiatic Society 3: 76 - 124" title="New and little known species of Ceylon nudibranchiate molluscs, and zoophytes." volume="3" year="1858">Kelaart (1858)</bibRefCitation>
was most inadequate and therefore best to ignore it. In this paper we follow
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" author="Rudman, WB" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="45" pagination="59 - 88" refId="B75" refString="Rudman, WB, 1978. The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific. Zoological Journal of the Linnean Society 62 (1): 59 - 88, DOI: https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" title="The dorid opisthobranch genera Halgerda Bergh and Sclerodoris Eliot from the Indo-West Pacific." url="https://doi.org/10.1111/j.1096-3642.1978.tb00523.x" volume="62" year="1978">
<normalizedToken originalValue="Rudman’s">Rudman's</normalizedToken>
(1978)
</bibRefCitation>
conclusion and regard
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
Eliot, 1904 as the valid name for this species with the synonymies established above. We also leave the question of the identity of
<taxonomicName lsidName="D. castanea" pageId="0" pageNumber="45" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="45">D. castanea</emphasis>
</taxonomicName>
as unresolved.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<bibRefCitation author="Herve, J-F" journalOrPublisher="Catherine Ledru, Noumea, New Caledonia" pageId="0" pageNumber="45" refId="B42" refString="Herve, J-F, 2010. Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches. Catherine Ledru, Noumea, New Caledonia" title="Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches." year="2010">
<normalizedToken originalValue="Hervé">Herve</normalizedToken>
(2010)
</bibRefCitation>
reported
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
from New Caledonia but based on the photographs published (
<bibRefCitation author="Herve, J-F" journalOrPublisher="Catherine Ledru, Noumea, New Caledonia" pageId="0" pageNumber="45" refId="B42" refString="Herve, J-F, 2010. Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches. Catherine Ledru, Noumea, New Caledonia" title="Guide des Nudibranchs de Nouvelle-Caledonie et Autres Opisthobranches." year="2010">
<normalizedToken originalValue="Hervé">Herve</normalizedToken>
2010
</bibRefCitation>
: 214), it seems that these records correspond to
<taxonomicName baseAuthorityName="Baba" baseAuthorityYear="1949" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris rubicunda" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="rubicunda">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris rubicunda</emphasis>
</taxonomicName>
(Baba, 1949). The present study is the first confirmed record of
<taxonomicName authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Sclerodoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sclerodoris tuberculata" order="Nudibranchia" pageId="0" pageNumber="45" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="45">Sclerodoris tuberculata</emphasis>
</taxonomicName>
from New Caledonia.
</paragraph>
</subSubSection>
</treatment>
</document>
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