plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/F4/87/03F487FF47704D169A71FB9BFC66FAB0.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

207 lines
32 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D2A68E8FC993D80A912D28CB08278322" ID-CLB-Dataset="72867" ID-DOI="10.1080/00222933.2022.2046887" ID-GBIF-Dataset="93716e8b-7637-4345-8eaf-ecb3984baa19" ID-ISSN="1464-5262" ID-Zenodo-Dep="6530252" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652081793889" checkinUser="plazi" docAuthor="Valois, Marcely C., Silva, Fernando A. B. &amp; Vaz-de-Mello, Fernando Z." docDate="2022" docId="03F487FF47704D169A71FB9BFC66FAB0" docLanguage="en" docName="JNATHIST.56.1-4.119-147.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2022.2046887" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Dichotomius (Selenocopris) favi Kohlmann and Solis 1997" docType="treatment" docVersion="7" lastPageNumber="133" masterDocId="FFCDFF87477D4D189B42FF9DFF91FFE2" masterDocTitle="A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="147" masterPageNumber="119" pageNumber="132" updateTime="1699329890315" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7ED8E3E706FA25FB5AC3A18F5A7D617F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4D1065CF90F216E733824D0598636F9">
<mods:title id="826B83E2159E0BA16CEA545C0A8C59B7">A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="8B137C3DBCB27F4320269B467BA6C5C7" type="personal">
<mods:role id="D7E678DE794947EBD95DE6A08E1FD571">
<mods:roleTerm id="F6051DF5967AE896D2BCDEB04BB5F5E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3828127CDAEDD3A83C69C203A3A5D757">Valois, Marcely C.</mods:namePart>
</mods:name>
<mods:name id="4EB29E66E4A37A1098AFF92858ADAEC2" type="personal">
<mods:role id="CFCA87410F72A27EBBD2D7021D83DEF8">
<mods:roleTerm id="1734A4EC08BFECCE5155BEF95965C5AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BF6813B3348011B62E31773D2C2789D">Silva, Fernando A. B.</mods:namePart>
</mods:name>
<mods:name id="D8001696C611AD22FF3F04578A24F5F9" type="personal">
<mods:role id="64606B0521CA1DD6C9CDE2A163CDAAD4">
<mods:roleTerm id="FAC9562C3032765BEDAFE320DAC03929">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="235F3A1AC16140C9A97AF000DC2616A8">Vaz-de-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7417D80DC2F0986C6AFF3DB71636F479">text</mods:typeOfResource>
<mods:relatedItem id="ED2EB375DD6878593CBC856B7A2ADF97" type="host">
<mods:titleInfo id="2C70A6E07C7A4C9818DBEBF26CD0B07D">
<mods:title id="2753679686D9BF388FE1060025D3B032">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="5A34B61A12CCBE0A136E3F1E09C65004">
<mods:date id="2123E2C934BBE0D91FAF26895F807074">2022</mods:date>
<mods:detail id="3D151F539D5FAB61706BCA0F9D6CFB27" type="pubDate">
<mods:number id="5E70CECA1D52A15468C0547EAFC282D5">2022-04-13</mods:number>
</mods:detail>
<mods:detail id="EC37A7774C1FC631D6FE9A5104DD3396" type="volume">
<mods:number id="255A02C0811AEDC348401C99E1259D03">56</mods:number>
</mods:detail>
<mods:detail id="27ED28C404A0357B319B4F61782EC708" type="issue">
<mods:number id="6040BE82E07E1A134F169DFB8D0F915B">1 - 4</mods:number>
</mods:detail>
<mods:extent id="6024BEA1FEA0E015BABABD00E82194F4" unit="page">
<mods:start id="24DB8E3C55577144686D56EBC48B9E0E">119</mods:start>
<mods:end id="606ED045B8C16403E56F059974615C6C">147</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="717A935C89319D24749ED822DE301701">
<mods:url id="52AA5733DE96DFD2E2E859BB23400735">http://dx.doi.org/10.1080/00222933.2022.2046887</mods:url>
</mods:location>
<mods:classification id="F1602FDC014FAA77B9822BB3B1F67F59">journal article</mods:classification>
<mods:identifier id="095BFE90F8E1F2D2C5D4C9A843F82713" type="CLB-Dataset">72867</mods:identifier>
<mods:identifier id="D94A50DDB976AE98492AFC83CC2367C5" type="DOI">10.1080/00222933.2022.2046887</mods:identifier>
<mods:identifier id="C29DF840012A995984A6D5F2246C9A48" type="GBIF-Dataset">93716e8b-7637-4345-8eaf-ecb3984baa19</mods:identifier>
<mods:identifier id="2D14D594467F538CBFF5CB4F602BA671" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D4BFA4E53188C9EA5BC4AC38037A8C41" type="Zenodo-Dep">6530252</mods:identifier>
</mods:mods>
<treatment id="03F487FF47704D169A71FB9BFC66FAB0" ID-DOI="http://doi.org/10.5281/zenodo.6645864" ID-GBIF-Taxon="195913547" ID-Zenodo-Dep="6645864" LSID="urn:lsid:plazi:treatment:03F487FF47704D169A71FB9BFC66FAB0" httpUri="http://treatment.plazi.org/id/03F487FF47704D169A71FB9BFC66FAB0" lastPageId="14" lastPageNumber="133" pageId="13" pageNumber="132">
<subSubSection id="C347656247704D159A71FB9BFC61FBC2" box="[307,1008,1030,1056]" pageId="13" pageNumber="132" type="nomenclature">
<paragraph id="8BE236E947704D159A71FB9BFC61FBC2" blockId="13.[307,1008,1030,1056]" box="[307,1008,1030,1056]" pageId="13" pageNumber="132">
<heading id="D0AA818547704D159A71FB9BFC61FBC2" box="[307,1008,1030,1056]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="132" reason="5">
<taxonomicName id="4C5D4D6A47704D159A71FB9BFC61FBC2" ID-CoL="35NBM" authority="Kohlmann and Solis, 1997" authorityName="Kohlmann and Solis" authorityYear="1997" box="[307,1008,1030,1056]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="favi" subGenus="Selenocopris">
<emphasis id="B929EAFB47704D159A71FB9BFD27FBC2" bold="true" box="[307,694,1030,1056]" italics="true" pageId="13" pageNumber="132">Dichotomius (Selenocopris) favi</emphasis>
<bibRefCitation id="EFCC4B1847704D1599FCFB9BFC61FBC2" author="Kohlmann B &amp; Solis A" box="[702,1008,1030,1056]" pageId="13" pageNumber="132" pagination="343 - 382" refId="ref14937" refString="Kohlmann B, Solis A. 1997. El genero Dichotomius en Costa Rica. Giorn. Ital. Entomol. 8: 343 - 382." type="journal article" year="1997">Kohlmann and Solís, 1997</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347656247704D15996EFBD9FD66FBBC" box="[556,759,1092,1118]" pageId="13" pageNumber="132" type="description">
<paragraph id="8BE236E947704D15996EFBD9FD66FBBC" blockId="13.[556,759,1092,1118]" box="[556,759,1092,1118]" pageId="13" pageNumber="132">
(
<figureCitation id="13662A6C47704D159976FBD9FD33FBBC" box="[564,674,1092,1118]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figures 2</figureCitation>
(g–l), 6)
</paragraph>
</subSubSection>
<subSubSection id="C347656247704D159BA4FBE5FD62FB35" pageId="13" pageNumber="132" type="reference_group">
<paragraph id="8BE236E947704D159BA4FBE5FD62FB35" blockId="13.[228,1117,1144,1239]" pageId="13" pageNumber="132">
<treatmentCitationGroup id="AB4D11C747704D159BA4FBE5FD62FB35" pageId="13" pageNumber="132">
<treatmentCitation id="0AFC10F847704D159BA4FBE5FD6DFB73" author="Kohlmann B &amp; Solis A" box="[230,764,1144,1169]" page="345" pageId="13" pageNumber="132" year="1997">
<taxonomicName id="4C5D4D6A47704D159BA4FBE5FD6DFB73" ID-CoL="35NBM" authority="Kohlmann and Solis 1997: 345" authorityName="Kohlmann and Solis" authorityPageNumber="345" authorityYear="1997" box="[230,764,1144,1169]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47704D159BA4FBE5FE0FFB73" box="[230,414,1144,1169]" italics="true" pageId="13" pageNumber="132">Dichotomius favi</emphasis>
<bibRefCitation id="EFCC4B1847704D159AE7FBE5FD6DFB73" author="Kohlmann B &amp; Solis A" box="[421,764,1144,1169]" pageId="13" pageNumber="132" pagination="343 - 382" refId="ref14937" refString="Kohlmann B, Solis A. 1997. El genero Dichotomius en Costa Rica. Giorn. Ital. Entomol. 8: 343 - 382." type="journal article" year="1997">Kohlmann and Solís 1997: 345</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 363;
<treatmentCitation id="0AFC10F847704D15987EFBE5FC7FFB73" author="Krajcik M" box="[828,1006,1144,1169]" page="53" pageId="13" pageNumber="132" year="2006">
<bibRefCitation id="EFCC4B1847704D15987EFBE5FC7FFB73" author="Krajcik M" box="[828,1006,1144,1169]" pageId="13" pageNumber="132" pagination="1 - 189" refId="ref14964" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae. Animma. X. Supplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 53</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFC10F847704D1598BBFBE5FBC8FB73" author="Krajcik M" box="[1017,1113,1144,1169]" page="91" pageId="13" pageNumber="132" year="2012">
<bibRefCitation id="EFCC4B1847704D1598BBFBE5FBC8FB73" author="Krajcik M" box="[1017,1113,1144,1169]" pageId="13" pageNumber="132" pagination="278" refId="ref14991" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Anima. X Supplement 5: 278." type="journal article" year="2012">2012: 91</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFC10F847704D159BA6FB06FDBAFB56" author="Arias-Buritica JA &amp; Delgado-Gomez P &amp; Gonzalez FA &amp; Vaz-de-Mello FZ" box="[228,555,1179,1204]" page="876" pageId="13" pageNumber="132" year="2011">
<bibRefCitation id="EFCC4B1847704D159BA6FB06FDBAFB56" author="Arias-Buritica JA &amp; Delgado-Gomez P &amp; Gonzalez FA &amp; Vaz-de-Mello FZ" box="[228,555,1179,1204]" pageId="13" pageNumber="132" pagination="875 - 878" refId="ref14408" refString="Arias-Buritica JA, Delgado-Gomez P, Gonzalez FA, Vaz-de-Mello FZ. 2011. Nuevos registros de escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) para el departamento de Choco (Colombia). Zool Mex. 27 (3): 875 - 878." type="journal article" year="2011">Ariás-Buriticá et al. 2011: 876</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFC10F847704D159974FB06FCFEFB56" author="Solis A &amp; Kohlmann B" box="[566,879,1179,1204]" page="6" pageId="13" pageNumber="132" year="2012">
<bibRefCitation id="EFCC4B1847704D159974FB06FCFEFB56" author="Solis A &amp; Kohlmann B" box="[566,879,1179,1204]" pageId="13" pageNumber="132" pagination="1 - 32" refId="ref15591" refString="Solis A, Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa. 3482 (1): 1 - 32." type="journal article" year="2012">Solis and Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFC10F847704D159839FB06FD7EFB35" author="Vaz-de-Mello FZ &amp; Nunes RV" page="299" pageId="13" pageNumber="132" year="2016">
<bibRefCitation id="EFCC4B1847704D159839FB06FD7EFB35" author="Vaz-de-Mello FZ &amp; Nunes RV" pageId="13" pageNumber="132" pagination="294 - 300" refId="ref15886" refString="Vaz-de-Mello FZ, Nunes RV. 2016. New species, redescription and taxonomic notes in the Dichotomius (Selenocopris) batesi (Harold) species-group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 4139 (2): 294 - 300." type="journal article" year="2016">Vaz-de-Mello and Nunes 2016: 299</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C347656247704D159BE2FA99FEA8F9F3" pageId="13" pageNumber="132" type="diagnosis">
<paragraph id="8BE236E947704D159BE2FA99FE85FAFC" blockId="13.[160,1156,1284,1553]" box="[160,276,1284,1310]" pageId="13" pageNumber="132">
<emphasis id="B929EAFB47704D159BE2FA99FE85FAFC" bold="true" box="[160,276,1284,1310]" italics="true" pageId="13" pageNumber="132">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE236E947704D159BE2FAB5FEA8F9F3" blockId="13.[160,1156,1284,1553]" pageId="13" pageNumber="132">
<taxonomicName id="4C5D4D6A47704D159BE2FAB5FECAFAA3" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,347,1320,1345]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47704D159BE2FAB5FECAFAA3" box="[160,347,1320,1345]" italics="true" pageId="13" pageNumber="132">Dichotomius favi</emphasis>
</taxonomicName>
is suggested to be closely related to
<taxonomicName id="4C5D4D6A47704D15985EFAB5FC17FAA3" authorityName="Kohlmann and Solis" authorityYear="1997" box="[796,902,1320,1345]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47704D15985EFAB5FC17FAA3" box="[796,902,1320,1345]" italics="true" pageId="13" pageNumber="132">D. danieli</emphasis>
</taxonomicName>
(see the diagnosis of
<taxonomicName id="4C5D4D6A47704D159BE2FAD7FE96FA81" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,263,1354,1379]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47704D159BE2FAD7FE96FA81" box="[160,263,1354,1379]" italics="true" pageId="13" pageNumber="132">D. danieli</emphasis>
</taxonomicName>
). However, the following combination of characters can be used to distinguish
<taxonomicName id="4C5D4D6A47704D159BE2FAF0FF77FA64" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,230,1389,1414]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47704D159BE2FAF0FF77FA64" box="[160,230,1389,1414]" italics="true" pageId="13" pageNumber="132">D. favi</emphasis>
</taxonomicName>
from its relatives: surface of head with different-sized ocellate punctures, forming transverse rows at clypeal surface; elytral striae thin and deeply punctate, punctures spaced by their diameter; ventral surface of the mesofemora with large coarse punctures at apical third; and paramera rectangular, obliquely truncate at apex in dorsal view (
<figureCitation id="13662A6C47704D159BEAFA65FE94F9F3" box="[168,261,1528,1553]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(j,k)).
</paragraph>
</subSubSection>
<subSubSection id="C347656247704D169BE2F9A3FC7DFB90" lastPageId="14" lastPageNumber="133" pageId="13" pageNumber="132" type="description">
<paragraph id="8BE236E947704D159BE2F9A3FE03F9BA" blockId="13.[160,1155,1598,1728]" box="[160,402,1598,1624]" pageId="13" pageNumber="132">
<emphasis id="B929EAFB47704D159BE2F9A3FE03F9BA" bold="true" box="[160,402,1598,1624]" italics="true" pageId="13" pageNumber="132">Redescription (male)</emphasis>
</paragraph>
<paragraph id="8BE236E947704D169BE2F9FFFC7DFB90" blockId="13.[160,1155,1598,1728]" lastBlockId="14.[160,1156,142,1138]" lastPageId="14" lastPageNumber="133" pageId="13" pageNumber="132">
<emphasis id="B929EAFB47704D159BE2F9FFFEA0F999" bold="true" box="[160,305,1634,1659]" pageId="13" pageNumber="132">Body length</emphasis>
:
<quantity id="4CA59B0C47704D159A7FF9FFFE31F999" box="[317,416,1634,1659]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" pageId="13" pageNumber="132" unit="mm" value="8.55">8.55 mm</quantity>
. Dorsum black, barely shiny on head and pronotum, elytra opaque (
<figureCitation id="13662A6C47704D159BEAF918FE97F97C" box="[168,262,1669,1694]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(g)).
<emphasis id="B929EAFB47704D159A7AF918FEE7F97C" bold="true" box="[312,374,1669,1694]" pageId="13" pageNumber="132">Head</emphasis>
: clypeal edge bidentate, teeth acute, separated by a deep V-shaped emargination. Clypeogenal junction rounded. Cephalic process produced into a small central tubercle (
<figureCitation id="13662A6C47734D169A1DFF13FE2AFF45" box="[351,443,142,167]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(g)). Antenal club with light brown tumescence.
<emphasis id="B929EAFB47734D169895FF13FBBAFF45" bold="true" box="[983,1067,142,167]" pageId="14" pageNumber="133">Thorax</emphasis>
: pronotum approximately twice wider than long (
<figureCitation id="13662A6C47734D1699C7FF2CFD73FF28" box="[645,738,177,202]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(g)). Anteromedian region of the disc lacking knobs, horns or excavations. Surface with dense ocellate punctures, equally spaced. Mid-longitudinal sulcus feebly marked medially. Surface of hypomera covered with dense ocellate punctures, spaced by less than their diameter; some punctures bearing orange setae. Prosternum covered with ocellate punctures. Mesoventrite covered with ocellate punctures, usually bearing short orange setae. Metaventrite surface with ocellate punctures, except on disc; punctures on anterior region usually bearing short setae. Meso-metaventral suture distinct. Mid-longitudinal sulcus of metaventrite feebly marked posteriorly. Lateral lobes of metaventrite with ocellate punctures larger than those on anterior region.
<emphasis id="B929EAFB47734D169A8FFE74FD82FDE0" bold="true" box="[461,531,489,514]" pageId="14" pageNumber="133">Elytra</emphasis>
: slightly convex in lateral view (
<figureCitation id="13662A6C47734D1698CCFE74FC7EFDE0" box="[910,1007,489,514]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(g)). Humeral callus distinct. Interstriae I–VII slight convex, barely shiny, with shagreened microsculpture; medial region densely covered with smooth dots.
<emphasis id="B929EAFB47734D169869FDB3FCF3FDA5" bold="true" box="[811,866,558,583]" pageId="14" pageNumber="133">Legs</emphasis>
: protibial spur curved at apical fifth. Metatibial spur deeply emarginate apically. Ventral surface of the profemora covered with coarse setiferous punctures along its entire extension. Ventral surface of the metafemora with ocellate punctures concentrated on apical half; basal half smooth. Meso and metafemora bearing a thin longitudinal groove along its entire extension, wider on apical half.
<emphasis id="B929EAFB47734D169A5DFD41FE05FD17" bold="true" box="[287,404,732,757]" pageId="14" pageNumber="133">Abdomen</emphasis>
: ventrites covered with ocellate punctures in lateral regions; punctures in medial region in a single row along anterior edge. Sixth ventrite 3 times longer than fifth. Pygidium twice wider than long; disc strongly convex apically. Surface of pygidial disc with ocellate punctures concentrated along anterior half, and spaced by half their diameter. Apex of the pygidium margined.
<emphasis id="B929EAFB47734D1699BFFCFBFCCFFC9D" bold="true" box="[765,862,870,895]" pageId="14" pageNumber="133">Tegmen</emphasis>
: in dorsal view, paramera rectangular, one-third longer than its width medially (
<figureCitation id="13662A6C47734D1699B6FC14FCC1FC40" box="[756,848,905,930]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(j,k)); apex slightly obliquely truncate; inner edges close to each other at basal fifth (
<figureCitation id="13662A6C47734D169864FC31FC17FC27" box="[806,902,940,965]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(j,k)). Dorsal surface of paramera smooth (
<figureCitation id="13662A6C47734D169A3AFC53FE44FC05" box="[376,469,974,999]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(j)). Apical half of paramera noticeably narrower than base in lateral view (
<figureCitation id="13662A6C47734D169A76FC6CFE04FBE8" box="[308,405,1009,1034]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(l)). Ventral sclerotised membrane between paramera thinner and curved at basal half; angulated laterally (
<figureCitation id="13662A6C47734D169937FB89FD45FBCF" box="[629,724,1044,1069]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="14" pageNumber="133">Figure 2</figureCitation>
(k)).
<emphasis id="B929EAFB47734D169847FB89FCCDFBCF" bold="true" box="[773,860,1044,1069]" pageId="14" pageNumber="133">Female</emphasis>
: differs from male by the clypeal teeth larger and acute. Clypeal edge with well defined transverse ridges. Sixth abdominal ventrite 3 times longer than fifth. Disc of the pygidium swollen.
</paragraph>
</subSubSection>
<subSubSection id="C347656247734D169BE2FB30FC66FAB0" pageId="14" pageNumber="133" type="materials_examined">
<paragraph id="8BE236E947734D169BE2FB30FE2EFB25" blockId="14.[160,1155,1197,1362]" box="[160,447,1197,1223]" pageId="14" pageNumber="133">
<emphasis id="B929EAFB47734D169BE2FB30FE2EFB25" bold="true" box="[160,447,1197,1223]" italics="true" pageId="14" pageNumber="133">Type material examined</emphasis>
</paragraph>
<paragraph id="8BE236E947734D169BE2FB4CFC66FAB0" blockId="14.[160,1155,1197,1362]" pageId="14" pageNumber="133">
<typeStatus id="54E6884B47734D169BE2FB4CFE96FB08" box="[160,263,1233,1258]" pageId="14" pageNumber="133" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C5D4D6A47734D169A70FB4CFE7FFB08" authorityName="Kohlmann and Solis" authorityYear="1997" box="[306,494,1233,1258]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47734D169A70FB4CFE7FFB08" box="[306,494,1233,1258]" italics="true" pageId="14" pageNumber="133">Dichotomius favi</emphasis>
</taxonomicName>
<specimenCount id="9D5BFD6047734D169ABAFB4CFD9DFB0A" box="[504,524,1233,1256]" pageId="14" pageNumber="133" type="male">♂</specimenCount>
[
<collectionCode id="ED4CAE2C47734D169959FB4CFDF9FB08" box="[539,616,1233,1258]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="14" pageNumber="133">MNCR</collectionCode>
]: ‘Sector Cerro Cocori. Fca./de E. Rojas,
<specimenCount id="9D5BFD6047734D169F76FB4CFB12FB08" box="[1076,1155,1233,1258]" pageId="14" pageNumber="133" type="generic">
<quantity id="4CA59B0C47734D169F76FB4CFBEEFB08" box="[1076,1151,1233,1258]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="133" unit="m" value="150.0">150 m</quantity>
.
</specimenCount>
Prov./
<collectingRegion id="4999F80B47734D169BA0FB69FEBBFAEF" box="[226,298,1268,1293]" country="Costa Rica" name="Limon" pageId="14" pageNumber="133">Limon</collectingRegion>
, Costa Rica. Oct/1992, E. Rojas/L–N 286000. 567,500’ // ‘
<taxonomicName id="4C5D4D6A47734D1698F4FB69FBEAFAEF" authorityName="Kohlmann and Solis" authorityYear="1997" box="[950,1147,1268,1293]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="favi">Dichotomius favi</taxonomicName>
/ Kohlmann and Solís’ // ‘
<collectingCountry id="F34A767947734D169AF8FA8BFDDDFACD" box="[442,588,1302,1327]" name="Costa Rica" pageId="14" pageNumber="133">COSTA RICA</collectingCountry>
/INBIO [barcode] CRI000 931,796’ // ‘HOLOTYPO/
<taxonomicName id="4C5D4D6A47734D169BE2FAA4FECFFAB0" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,350,1337,1362]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="favi">Dichotomius favi</taxonomicName>
/Kohlmann y Solís’. Examined via photographs (
<figureCitation id="13662A6C47734D16983AFAA4FC43FAB0" box="[888,978,1337,1362]" captionStart="Figure 1" captionStartId="4.[160,225,1665,1688]" captionTargetBox="[195,1140,661,1606]" captionTargetId="figure-144@4.[163,1153,646,1634]" captionTargetPageId="4" captionText="Figure 1. Dorsal habitus of male. (a) Dichotomius benesi; (b) D. berthalutzae sp. nov.; (c) D. femoratus; (d) D. fortepunctatus; (e) D. gamboaensis." figureDoi="http://doi.org/10.5281/zenodo.6530254" httpUri="https://zenodo.org/record/6530254/files/figure.png" pageId="14" pageNumber="133">Figure 1</figureCitation>
(i)).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink