249 lines
38 KiB
XML
249 lines
38 KiB
XML
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<mods:title id="50127B2EC571B2C324D63F35DCB351F2">Three New Species in the Desmopachria striola Species-Group of Desmopachria Babington, 1841 (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Hyphydrini)</mods:title>
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<mods:namePart id="2E8DA0CAFE993A55B6F7136CF6B3F342">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="61A692909CBE75387A7BF01F414B0BC9">Wolfe, G. William</mods:namePart>
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<mods:typeOfResource id="759DB654F47FA2A3DF976187AD3B7A2E">text</mods:typeOfResource>
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<mods:title id="9D5FF8FE154AF9250E6979CF09E434D2">The Coleopterists Bulletin</mods:title>
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<mods:part id="4FBEE3689AA5B62AE944687E137F1D58">
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<mods:date id="6C8EE3295BFA83B8B7436453F0E8B669">2019</mods:date>
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<mods:number id="3A05CFB8B7B0881154702A15F172C2B2">2019-09-22</mods:number>
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<paragraph id="8BE236E7FFB3FFC7CBD6821CFE20FDC2" blockId="3.[183,648,524,652]" pageId="3" pageNumber="623">
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<taxonomicName id="4C5D4D64FFB3FFC7CBD6821CFD90FDDF" authority="Miller and Wolfe" authorityName="Miller and Wolfe" authorityYear="2019" box="[209,618,524,545]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos" status="sp. nov.">
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<emphasis id="B929EAF5FFB3FFC7CBD6821CFE4BFDDF" bold="true" box="[209,433,524,545]" italics="true" pageId="3" pageNumber="623">Desmopachria atropos</emphasis>
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Miller and Wolfe
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,
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<taxonomicNameLabel id="A21A578EFFB3FFC7CA628237FE20FDC2" box="[357,474,551,572]" pageId="3" pageNumber="623" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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Zoobank.org/
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urn:lsid:zoobank.org:act:
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(
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<figureCitation id="13662A62FFB3FFC7CA588266FE41FD75" box="[351,443,630,652]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Figs. 2–5</figureCitation>
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,
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<figureCitation id="13662A62FFB3FFC7CACF8266FE1BFD75" box="[456,481,630,651]" captionStart="Figs" captionStartId="5.[197,236,1461,1480]" captionTargetBox="[177,1188,196,1443]" captionTargetId="figure-137@5.[175,1190,194,1445]" captionTargetPageId="5" captionText="Figs. 25–26. Distribution of new Desmopachria species." figureDoi="http://doi.org/10.5281/zenodo.5402226" httpUri="https://zenodo.org/record/5402226/files/figure.png" pageId="3" pageNumber="623">25</figureCitation>
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)
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<subSubSection id="C347656CFFB3FFC7CBC282B1FEC3FD15" pageId="3" pageNumber="623" type="materials_examined">
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<materialsCitation id="3B353CBAFFB3FFC7CBC282B1FD90FD48" box="[197,618,673,694]" country="Venezuela" location="Barinas State" pageId="3" pageNumber="623" specimenCount="1" stateProvince="Barinas" typeStatus="holotype">
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<typeStatus id="54E68845FFB3FFC7CBC282B1FF0DFD48" box="[197,247,673,694]" pageId="3" pageNumber="623">Type</typeStatus>
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Locality.
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</emphasis>
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<collectingCountry id="F34A7677FFB3FFC7CA6B82B1FE2AFD48" box="[364,464,673,694]" name="Venezuela" pageId="3" pageNumber="623">Venezuela</collectingCountry>
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,
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<collectingRegion id="4999F805FFB3FFC7CAE582B1FD90FD48" box="[482,618,673,694]" country="Venezuela" name="Barinas" pageId="3" pageNumber="623">Barinas State</collectingRegion>
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</materialsCitation>
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,
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<materialsCitation id="3B353CBAFFB3FFC7C97B82B1FE11FD2E" country="Bolivia" location="Ciudad" pageId="3" pageNumber="623" specimenCount="1" typeStatus="holotype">
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<emphasis id="B929EAF5FFB3FFC7C97B82B1FD68FD48" box="[636,658,673,694]" italics="true" pageId="3" pageNumber="623">ca</emphasis>
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<quantity id="4CA59B02FFB3FFC7CBAA82ABFF01FD2E" box="[173,251,699,720]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="623" unit="km" value="13.0">13 km</quantity>
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SE
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<location id="8E82603CFFB3FFC7CA4182ABFE71FD2E" LSID="urn:lsid:plazi:treatment:03F487F1FFB3FFC7CBD6821CFBE7FA63:8E82603CFFB3FFC7CA4182ABFE71FD2E" box="[326,395,699,720]" country="Bolivia" name="Ciudad" pageId="3" pageNumber="623">Ciudad</location>
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<collectingCountry id="F34A7677FFB3FFC7CAA482ABFE11FD2E" box="[419,491,699,720]" name="Bolivia" pageId="3" pageNumber="623">Bolivia</collectingCountry>
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, 8°19.394
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N, 70°28.238
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</emphasis>
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W.
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</paragraph>
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</subSubSection>
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<subSubSection id="C347656CFFB3FFC7CBC282E0FEF2FB51" pageId="3" pageNumber="623" type="diagnosis">
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<paragraph id="8BE236E7FFB3FFC7CBC282E0FEF2FB51" blockId="3.[173,660,673,1730]" pageId="3" pageNumber="623">
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<emphasis id="B929EAF5FFB3FFC7CBC282E0FED6FCFB" bold="true" box="[197,300,752,773]" pageId="3" pageNumber="623">Diagnosis.</emphasis>
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This species differs from others in the group by the shape of the male genitalia. The median lobe is elongate and apically obliquely bent with the apex straight in lateral aspect (
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<figureCitation id="13662A62FFB3FFC7C9288350FD91FCAB" box="[559,619,832,853]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 2</figureCitation>
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). In ventral aspect, the median lobe is broad, laterally broadly rounded, and apically convergent to narrowly produced and pointed (
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<figureCitation id="13662A62FFB3FFC7CAC68380FE03FC5B" box="[449,505,912,933]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 3</figureCitation>
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). Specimens are evenly orange on the dorsum of the head and pronotum and orange-brown on most of the elytra with the region between the suture; the sutural stria darker. In size and coloration,
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<taxonomicName id="4C5D4D64FFB3FFC7CAC883EAFDCFFBF1" authorityName="Miller and Wolfe" authorityYear="2019" box="[463,565,1018,1039]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos">
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<emphasis id="B929EAF5FFB3FFC7CAC883EAFDCFFBF1" box="[463,565,1018,1039]" italics="true" pageId="3" pageNumber="623">D. atropos</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="4C5D4D64FFB3FFC7CBCF8404FDA5FBD4" authority="Miller and Wolfe" authorityName="Miller and Wolfe" authorityYear="2019" box="[200,607,1044,1066]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="lachesis" status="sp. nov.">
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<emphasis id="B929EAF5FFB3FFC7CBCF8404FE50FBD4" box="[200,426,1044,1066]" italics="true" pageId="3" pageNumber="623">Desmopachria lachesis</emphasis>
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Miller and Wolfe
|
||
</taxonomicName>
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,
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<taxonomicNameLabel id="A21A578EFFB3FFC7C96C8405FF0EFBBA" pageId="3" pageNumber="623" rank="species">new species</taxonomicNameLabel>
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, which has the median lobe apically strongly sinuate in lateral aspect and in ventral aspect with the apical rami laterally rounded and apically pointed with a distinct, deep medial emargination (
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<figureCitation id="13662A62FFB3FFC7CBB3848AFF06FB51" box="[180,252,1178,1199]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 10</figureCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C347656CFFB3FFC7CBC284A4FCF5FC5A" pageId="3" pageNumber="623" type="description">
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<paragraph id="8BE236E7FFB3FFC7CBC284A4FCF5FC5A" blockId="3.[173,660,673,1730]" lastBlockId="3.[707,1194,194,1437]" pageId="3" pageNumber="623">
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<emphasis id="B929EAF5FFB3FFC7CBC284A4FE26FB37" bold="true" box="[197,476,1204,1225]" pageId="3" pageNumber="623">Description. Measurements:</emphasis>
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TL =
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, GW = 1.0–
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<quantity id="4CA59B02FFB3FFC7CA2584DFFE88FB1A" box="[290,370,1231,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="623" unit="mm" value="1.1">1.1 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CA59B02FFB3FFC7CACD84DFFDB9FB1A" box="[458,579,1230,1252]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="623" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
|
||
, HW =
|
||
<quantity id="4CA59B02FFB3FFC7CBAA84F9FF0CFB00" box="[173,246,1257,1278]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="623" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CA59B02FFB3FFC7CA3E84F9FE56FB00" box="[313,428,1257,1279]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="623" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
|
||
, TL/GW = 1.4–1.5, HW/ EW = 2.1.
|
||
<emphasis id="B929EAF5FFB3FFC7CA148514FE7FFAE7" bold="true" box="[275,389,1284,1305]" pageId="3" pageNumber="623">Coloration:</emphasis>
|
||
Head and pronotum yelloworange, posterior margin of pronotum medially narrowly brown. Elytron brown-orange, darker medially along sutural stria; surface not iridescent. Head appendages, legs, and ventral surfaces yellow to yellow-orange.
|
||
<emphasis id="B929EAF5FFB3FFC7CA978599FD68FA60" bold="true" box="[400,658,1417,1438]" pageId="3" pageNumber="623">Sculpture and structure:</emphasis>
|
||
Head broad, short; anterior margin of clypeus finely margined with continuous narrow bead; surface of head shiny, very finely and sparsely punctate; eyes large (HW/EW = 2.1); antennae short, antennomeres III–X short, moderately broad, antennomere XI elongate, apically pointed. Pronotum short, broad, laterally distinctly curved with continuous narrow bead; surface shiny, very finely and sparsely punctate, without basal striae. Elytron broad, laterally broadly curved; surface shiny, very finely and sparsely punctate, without basal stria but with distinctive longitudinal sutural stria. Prosternum short, longitudinally compressed; prosternal process short, broad, without medial ventral tubercle, apically broadly triangular. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate; metaventrite wings extremely slender. Metacoxa with medial portion short, less than half length of metaventrite medially, metacoxal lines subsinuate, divergent anteriorly; lateral portion of metacoxa very large, anteriorly strongly expanded; surface shiny, impunctate. Metatrochanter very large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdominal surface shiny and smooth, very finely and sparsely punctate.
|
||
<emphasis id="B929EAF5FFB3FFC7C8738225FBF6FDB4" bold="true" box="[884,1036,565,586]" pageId="3" pageNumber="623">Male genitalia:</emphasis>
|
||
Median lobe in lateral aspect slender but expanded medially, apically narrowed and obliquely bent dorsad, apical portion straight, apex slightly rounded (
|
||
<figureCitation id="13662A62FFB3FFC7C8FE8295FBCDFD64" box="[1017,1079,645,666]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 2</figureCitation>
|
||
); in ventral aspect, narrow basally, widened to medial half, with distinct constriction submedially, apical portion broad, lateral margins broadly rounded to broadly pointed apex, apical portion medially deeply emarginate (
|
||
<figureCitation id="13662A62FFB3FFC7C80C831AFCBCFCE1" box="[779,838,778,799]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 3</figureCitation>
|
||
). Lateral lobe in lateral aspect broad throughout most of length, apically very slender and bent obliquely dorsad, bent portion slender to sharply pointed apex (
|
||
<figureCitation id="13662A62FFB3FFC7C84D834AFC7CFC91" box="[842,902,858,879]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 4</figureCitation>
|
||
); in ventral aspect, moderately broad throughout length, apex broadly rounded (
|
||
<figureCitation id="13662A62FFB3FFC7C9CE839FFCF9FC5A" box="[713,771,911,932]" captionStart="Figs" captionStartId="4.[175,214,1513,1532]" captionTargetBox="[231,1092,195,1497]" captionTargetId="figure-1@4.[230,1093,194,1498]" captionTargetPageId="4" captionText="Figs. 1–24. Desmopachria species. 1) D. atropos, habitus; arrow indicates sutural stria; scale bar = 1.0 mm. Male genitalia: D. atropos: 2–3) Median lobe, right lateral and ventral aspects, respectively; 4–5) Right lateral lobe, right lateral and ventral aspects, respectively. D. clotho: 6) Median lobe, right lateral aspect; 7) Median and right lateral lobes, ventral aspect; 8) Right lateral lobe, right lateral aspect. D. lachesis: 9–10) Median lobe, right lateral and ventral aspects, respectively; 11–12) Right lateral lobe, right lateral and ventral aspects, respectively. D. amyae, redrawn from Miller (2001): 13) Median and left lateral lobes, dorsal aspect; 14) Median lobe, right lateral aspect; 15) Right lateral lobe, right lateral aspect. D. chei, redrawn from Miller (1999): 16) Median and lateral lobes, dorsal aspect; 17) Right lateral lobe, right lateral aspect. D. fossulata, redrawn from Young (1990): 18) Median and lateral lobes, ventral aspect. D. grouvellei, redrawn from Young (1990): 19) Median and lateral lobes, ventral aspect. D. ruginosa, redrawn from Young (1990): 20–21) Median and lateral lobes, ventral and right lateral aspects, respectively. D. striola: 22) Median lobe, right lateral aspect; 23) Median and right lateral lobes, ventral aspect; 24) Right lateral lobe, right lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5402224" httpUri="https://zenodo.org/record/5402224/files/figure.png" pageId="3" pageNumber="623">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347656CFFB3FFC7C9DC83B9FBA0FC27" pageId="3" pageNumber="623" type="etymology">
|
||
<paragraph id="8BE236E7FFB3FFC7C9DC83B9FBA0FC27" blockId="3.[707,1194,194,1437]" pageId="3" pageNumber="623">
|
||
<emphasis id="B929EAF5FFB3FFC7C9DC83B9FCB7FC40" bold="true" box="[731,845,937,958]" pageId="3" pageNumber="623">Etymology.</emphasis>
|
||
This species is named
|
||
<taxonomicName id="4C5D4D64FFB3FFC7CF6683BAFB53FC41" authorityName="Miller and Wolfe" authorityYear="2019" box="[1121,1193,938,959]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos">
|
||
<emphasis id="B929EAF5FFB3FFC7CF6683BAFB53FC41" box="[1121,1193,938,959]" italics="true" pageId="3" pageNumber="623">atropos</emphasis>
|
||
</taxonomicName>
|
||
(unturnable), one of the three Greek fates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347656CFFB3FFC7C9DC83CFFCEDFBA0" pageId="3" pageNumber="623" type="distribution">
|
||
<paragraph id="8BE236E7FFB3FFC7C9DC83CFFC7DFBD7" blockId="3.[707,1194,194,1437]" pageId="3" pageNumber="623">
|
||
<emphasis id="B929EAF5FFB3FFC7C9DC83CFFCA6FC0A" bold="true" box="[731,860,991,1012]" pageId="3" pageNumber="623">Distribution.</emphasis>
|
||
<taxonomicName id="4C5D4D64FFB3FFC7C86183CEFBBEFC0A" authorityName="Miller and Wolfe" authorityYear="2019" box="[870,1092,990,1012]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos">
|
||
<emphasis id="B929EAF5FFB3FFC7C86183CEFBBEFC0A" box="[870,1092,990,1012]" italics="true" pageId="3" pageNumber="623">Desmopachria atropos</emphasis>
|
||
</taxonomicName>
|
||
is known only from one locality in
|
||
<collectingRegion id="4999F805FFB3FFC7C8B983E9FBC7FBF0" box="[958,1085,1017,1038]" country="Venezuela" name="Barinas" pageId="3" pageNumber="623">Barinas State</collectingRegion>
|
||
in western
|
||
<collectingCountry id="F34A7677FFB3FFC7C9C48404FCDEFBD7" box="[707,804,1044,1065]" name="Venezuela" pageId="3" pageNumber="623">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="13662A62FFB3FFC7C8348404FC86FBD7" box="[819,892,1044,1065]" captionStart="Figs" captionStartId="5.[197,236,1461,1480]" captionTargetBox="[177,1188,196,1443]" captionTargetId="figure-137@5.[175,1190,194,1445]" captionTargetPageId="5" captionText="Figs. 25–26. Distribution of new Desmopachria species." figureDoi="http://doi.org/10.5281/zenodo.5402226" httpUri="https://zenodo.org/record/5402226/files/figure.png" pageId="3" pageNumber="623">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE236E7FFB3FFC7C9DC843EFCEDFBA0" blockId="3.[707,1194,194,1437]" pageId="3" pageNumber="623">
|
||
<emphasis id="B929EAF5FFB3FFC7C9DC843EFCD4FBBD" bold="true" box="[731,814,1070,1091]" pageId="3" pageNumber="623">Habitat.</emphasis>
|
||
The
|
||
<typeStatus id="54E68845FFB3FFC7C860843EFC6AFBBD" box="[871,912,1070,1091]" pageId="3" pageNumber="623">type</typeStatus>
|
||
series was collected from a “marsh.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C347656CFFB3FFC7C9DC8473FBE7FA63" pageId="3" pageNumber="623" type="materials_examined">
|
||
<paragraph id="8BE236E7FFB3FFC7C9DC8473FBE7FA63" blockId="3.[707,1194,194,1437]" pageId="3" pageNumber="623">
|
||
<emphasis id="B929EAF5FFB3FFC7C9DC8473FC8CFB86" bold="true" box="[731,886,1123,1144]" pageId="3" pageNumber="623">Type Material.</emphasis>
|
||
<materialsCitation id="3B353CBAFFB3FFC7C8848473FCB9FACD" collectingDate="2012-01-25" collectionCode="MIZA, KUNHM-ENT" collectorName="Short" country="Venezuela" elevation="173" location="Ciudad Bolivia" pageId="3" pageNumber="623" specimenCode="SEMC1030664" specimenCount="2" stateProvince="Barinas" typeStatus="holotype">
|
||
<typeStatus id="54E68845FFB3FFC7C8848473FC21FB86" box="[899,987,1123,1144]" pageId="3" pageNumber="623" type="holotype">Holotype</typeStatus>
|
||
in
|
||
<collectionCode id="ED4CAE22FFB3FFC7CF008473FBBCFB86" box="[1031,1094,1123,1144]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="3" pageNumber="623" type="Museum">MIZA</collectionCode>
|
||
, male labeled “
|
||
<collectingCountry id="F34A7677FFB3FFC7C80C846EFC66FB6D" box="[779,924,1150,1171]" name="Venezuela" pageId="3" pageNumber="623">VENEZUELA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4999F805FFB3FFC7C8AE846EFBD0FB6D" box="[937,1066,1150,1171]" country="Venezuela" name="Barinas" pageId="3" pageNumber="623">Barinas State</collectingRegion>
|
||
8°19.394
|
||
<emphasis id="B929EAF5FFB3FFC7CF8B846AFB6BFB77" box="[1164,1169,1146,1161]" italics="true" pageId="3" pageNumber="623">
|
||
<superScript id="7C289BAFFFB3FFC7CF8B846AFB6BFB77" attach="right" box="[1164,1169,1146,1161]" fontSize="6" pageId="3" pageNumber="623">0</superScript>
|
||
</emphasis>
|
||
N, 70°28.238
|
||
<emphasis id="B929EAF5FFB3FFC7C8218484FCD1FB5D" box="[806,811,1172,1187]" italics="true" pageId="3" pageNumber="623">
|
||
<superScript id="7C289BAFFFB3FFC7C8218484FCD1FB5D" attach="right" box="[806,811,1172,1187]" fontSize="6" pageId="3" pageNumber="623">0</superScript>
|
||
</emphasis>
|
||
W,
|
||
<quantity id="4CA59B02FFB3FFC7C8498489FC7EFB50" box="[846,900,1177,1198]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="623" unit="m" value="173.0">
|
||
<elevation id="0070D1D4FFB3FFC7C8498489FC7EFB50" box="[846,900,1177,1198]" metricMagnitude="2" metricUnit="m" metricValue="1.73" pageId="3" pageNumber="623" unit="m" value="173.0">173m</elevation>
|
||
</quantity>
|
||
ca.
|
||
<quantity id="4CA59B02FFB3FFC7C8A98489FC16FB50" box="[942,1004,1177,1198]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="623" unit="km" value="13.0">13 km</quantity>
|
||
SE
|
||
<location id="8E82603CFFB3FFC7CF108489FB53FB50" LSID="urn:lsid:plazi:treatment:03F487F1FFB3FFC7CBD6821CFBE7FA63:8E82603CFFB3FFC7CF108489FB53FB50" box="[1047,1193,1177,1198]" country="Venezuela" name="Ciudad Bolivia" pageId="3" pageNumber="623" stateProvince="Barinas">Ciudad Bolivia</location>
|
||
leg.
|
||
<collectorName id="26A85331FFB3FFC7C9F784A3FCDCFB36" box="[752,806,1203,1224]" pageId="3" pageNumber="623">Short</collectorName>
|
||
, Arias, & Gustafson marsh;
|
||
<date id="FFE31027FFB3FFC7CF4F84A3FB5EFB36" box="[1096,1188,1203,1224]" pageId="3" pageNumber="623" value="2012-01-25">
|
||
<collectingDate id="EFA7E9CFFFB3FFC7CF4F84A3FB5EFB36" box="[1096,1188,1203,1224]" pageId="3" pageNumber="623" value="2012-01-25">25.i.2012</collectingDate>
|
||
</date>
|
||
; VZ12-0125-02A/
|
||
<specimenCode id="DBFB9E9CFFB3FFC7C87084DEFBF1FB1D" box="[887,1035,1230,1251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="623" type="Museum">SEMC1030664</specimenCode>
|
||
<collectionCode id="ED4CAE22FFB3FFC7CF1E84DEFB53FB1D" box="[1049,1193,1230,1251]" pageId="3" pageNumber="623">KUNHM-ENT</collectionCode>
|
||
[label with barcode]/
|
||
<typeStatus id="54E68845FFB3FFC7C89084F8FBEEFB03" box="[919,1044,1256,1277]" pageId="3" pageNumber="623" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C5D4D64FFB3FFC7CF1A84F7FC12FAE6" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos">
|
||
<emphasis id="B929EAF5FFB3FFC7CF1A84F7FCF1FAE6" italics="true" pageId="3" pageNumber="623">Desmopachria atropos</emphasis>
|
||
Miller and Wolfe, 2017
|
||
</taxonomicName>
|
||
[red label with black line border].”
|
||
</materialsCitation>
|
||
<typeStatus id="54E68845FFB3FFC7C84C850DFC50FACC" box="[843,938,1309,1330]" pageId="3" pageNumber="623" type="paratype">Paratypes</typeStatus>
|
||
: Four labeled same as
|
||
<typeStatus id="54E68845FFB3FFC7CF8E850DFD04FAB3" pageId="3" pageNumber="623" type="holotype">holotype</typeStatus>
|
||
except with different specimen barcode labels (
|
||
<tableCitation id="C6DF035CFFB3FFC7C9F88542FCB5FA99" box="[767,847,1362,1383]" captionStart="Table 1" captionStartId="6.[175,225,195,214]" captionText="Table 1. Snow Entomology Museum Collection, University of Kansas (KUNHM) accession numbers for Desmopachria specimens of new species described herein." pageId="3" pageNumber="623">Table 1</tableCitation>
|
||
) and each with “…
|
||
<typeStatus id="54E68845FFB3FFC7CF348542FB53FA99" box="[1075,1193,1362,1383]" pageId="3" pageNumber="623" type="paratype">PARATYPE</typeStatus>
|
||
<taxonomicName id="4C5D4D64FFB3FFC7C9C4857CFB53FA7C" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" box="[707,1193,1388,1410]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="623" phylum="Arthropoda" rank="species" species="atropos">
|
||
<emphasis id="B929EAF5FFB3FFC7C9C4857CFC5FFA7C" box="[707,933,1388,1410]" italics="true" pageId="3" pageNumber="623">Desmopachria atropos</emphasis>
|
||
Miller and Wolfe, 2017
|
||
</taxonomicName>
|
||
[blue label with black line border].”
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |