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<document id="27B0BF3945474FAE364F3F5A2E7C4804" ID-DOI="10.11646/zootaxa.5351.5.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8392291" ID-ZooBank="C6C7769C-94F6-496A-9293-2F0E1E201330" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695993006982" checkinUser="plazi" docAuthor="Davranoglou, Leonidas-Romanos, Baňař, Petr, Kakiopoulos, George, Balázs, Attila, Avtzis, Dimitrios N. &amp; Hlaváč, Peter" docDate="2023" docId="03F487CEE47EFFC8FF235286FA4245A4" docLanguage="en" docName="zootaxa.5351.5.4.pdf" docOrigin="Zootaxa 5351 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.5.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afropselaphus euboicus Davranoglou, Hlavac &amp; Banar 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="567" masterDocId="FFCDFFB6E478FFC0FFB4541CFFC84257" masterDocTitle="Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)" masterLastPageNumber="570" masterPageNumber="559" pageNumber="565" updateTime="1695993641248" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="02F091401F57333B4C76F5755C75F09C">Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)</mods:title>
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<mods:name id="32061234159211F9096FFE71AE854858" type="personal">
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<mods:namePart id="6D7428F32D9523C5CA2ED101AC335DCA">Davranoglou, Leonidas-Romanos</mods:namePart>
<mods:affiliation id="99D7BAE7BF4532426F57C899671C63C8">Oxford University Museum of Natural History, University of Oxford, Parks Road, Oxford OX 1 3 PW, UK;</mods:affiliation>
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<mods:namePart id="3924AAC55FA68E39EFF3B40033044F1C">Baňař, Petr</mods:namePart>
<mods:affiliation id="7135C2004E10E3DEFA9C33F95AFEC045">Department of Entomology, Moravian Museum, Zelný trh 6, 659 37 Brno, Czech Republic;</mods:affiliation>
</mods:name>
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<mods:namePart id="E3F353926C76436F7CB252F5F709B80B">Kakiopoulos, George</mods:namePart>
<mods:nameIdentifier id="926A46E23EAF0EA8C14886A81BCE9ECF" type="ORCID">0000-0003-3486-5099</mods:nameIdentifier>
<mods:affiliation id="9D596950DFF8F059CD8D23F1B7D53A46">Ilidos 60 - 62 street, 115 27, Athens, Greece; https: // orcid. org / 0000 - 0003 - 3486 - 5099</mods:affiliation>
</mods:name>
<mods:name id="C4922CC55E7CC60612F69C8E799CEB40" type="personal">
<mods:role id="FBE9EEBDD718FA68ABD4D28AFF198DA5">
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</mods:role>
<mods:namePart id="F347BA67D0AA4D6CCCE6D8A7790F67CB">Balázs, Attila</mods:namePart>
<mods:affiliation id="E7706C37664A2769DBF1E7ABC4D36047">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Mendel University in Brno, Zemědělská 1, 613 00 Brno, Czech Republic;</mods:affiliation>
</mods:name>
<mods:name id="DE2F85124C2E9E0F760836760F8FE69E" type="personal">
<mods:role id="B6F968AE5ECBC50D8A244F3B55059F23">
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<mods:namePart id="8C8B9B69205C50AAFA51A927578A2BF5">Avtzis, Dimitrios N.</mods:namePart>
<mods:nameIdentifier id="6C69E807A4C5EEC4A4AB4DA890B8C46F" type="ORCID">0000-0002-7772-6892</mods:nameIdentifier>
<mods:affiliation id="FF6FC12D63AEEBFDFA7A0CACD55CA9FD">Forest Research Institute, Hellenic Agricultural Organization Demeter, 570 06 Vassilika-Thessaloniki, Greece; dimitrios. avtzis @ gmail. com; https: // orcid. org / 0000 - 0002 - 7772 - 6892</mods:affiliation>
<mods:nameIdentifier id="491EA1B21F51DF6764EB80C27633FB36" type="email">dimitrios.avtzis@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CFF5FA371C3E7DF04EE04FBB444EA27F">Hlaváč, Peter</mods:namePart>
<mods:nameIdentifier id="EEB8DDA2B2A2928F97AE252716225841" type="ORCID">0000-0001-5060-0811</mods:nameIdentifier>
<mods:affiliation id="8126746A5907E1268567DEC526EC20AE">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice; peterclaviger @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
<mods:nameIdentifier id="B1C05F7A7782B727A5B5050EC1E67EBC" type="email">peterclaviger@gmail.com</mods:nameIdentifier>
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<mods:title id="772565CEAD6A680137DA29281DB72A77">Zootaxa</mods:title>
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<mods:date id="0269DD60F0000E7F5EC3897FE512143B">2023</mods:date>
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<treatment id="03F487CEE47EFFC8FF235286FA4245A4" ID-DOI="http://doi.org/10.5281/zenodo.8392349" ID-Zenodo-Dep="8392349" LSID="urn:lsid:plazi:treatment:03F487CEE47EFFC8FF235286FA4245A4" httpUri="http://treatment.plazi.org/id/03F487CEE47EFFC8FF235286FA4245A4" lastPageId="8" lastPageNumber="567" pageId="6" pageNumber="565">
<subSubSection id="C3476553E47EFFC6FF235286FC5944E2" box="[151,913,1689,1717]" pageId="6" pageNumber="565" type="nomenclature">
<paragraph id="8BE236D8E47EFFC6FF235286FC5944E2" blockId="6.[151,913,1689,1753]" box="[151,913,1689,1717]" pageId="6" pageNumber="565">
<heading id="D0AA81B4E47EFFC6FF235286FC5944E2" bold="true" box="[151,913,1689,1717]" fontSize="11" level="1" pageId="6" pageNumber="565" reason="1">
<emphasis id="B929EACAE47EFFC6FF235286FC5944E2" bold="true" box="[151,913,1689,1717]" pageId="6" pageNumber="565">
<taxonomicName id="4C5D4D5BE47EFFC6FF235286FC8A44E3" authority="Davranoglou, Hlavac &amp; Banar" authorityName="Davranoglou, Hlavac &amp; Banar" authorityYear="2023" box="[151,834,1689,1717]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE47EFFC6FF235286FE7344E2" bold="true" box="[151,443,1690,1717]" italics="true" pageId="6" pageNumber="565">Afropselaphus euboicus</emphasis>
Davranoglou, Hlaváč &amp; Baňař
</taxonomicName>
,
<taxonomicNameLabel id="A21A57B1E47EFFC6FCFB5287FC5944E2" box="[847,913,1691,1717]" pageId="6" pageNumber="565" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3476553E47EFFC6FF2352A2FEB3448E" box="[151,379,1726,1753]" pageId="6" pageNumber="565" type="description">
<paragraph id="8BE236D8E47EFFC6FF2352A2FEB3448E" blockId="6.[151,913,1689,1753]" box="[151,379,1726,1753]" pageId="6" pageNumber="565">
(
<figureCitation id="13662A5DE47EFFC6FF2B52A2FF31448F" box="[159,249,1726,1752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="6" pageNumber="565">Figs 2C</figureCitation>
<figureCitation id="13662A5DE47EFFC6FEBD52A2FEE2448F" box="[265,298,1726,1752]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="6" pageNumber="565">6C</figureCitation>
,
<figureCitation id="13662A5DE47EFFC6FE8152A3FEBB448E" box="[309,371,1727,1753]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="6" pageNumber="565">7E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47EFFC6FF23531AFCF945E6" pageId="6" pageNumber="565" type="materials_examined">
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<emphasis id="B929EACAE47EFFC6FF23531AFE194576" bold="true" box="[151,465,1798,1825]" pageId="6" pageNumber="565">
Material studied.
<typeStatus id="54E6887AE47EFFC6FED0531BFE194576" box="[356,465,1799,1825]" pageId="6" pageNumber="565">Holotype</typeStatus>
</emphasis>
,♁:
<emphasis id="B929EACAE47EFFC6FE48531AFDBA4577" bold="true" box="[508,626,1798,1824]" pageId="6" pageNumber="565">
<collectingCountry id="F34A7648E47EFFC6FE48531AFDBA4577" box="[508,626,1798,1824]" name="Greece" pageId="6" pageNumber="565">GREECE</collectingCountry>
</emphasis>
,with one label “GREECE,Euboea, Steni,
<geoCoordinate id="EE69501FE47EFFC6FB89531AFB204576" box="[1085,1256,1798,1825]" degrees="38" direction="north" minutes="35" orientation="latitude" pageId="6" pageNumber="565" precision="1" seconds="18.45" value="38.588455">38°3518.45”N</geoCoordinate>
<geoCoordinate id="EE69501FE47EFFC6FB58531AFA504576" box="[1260,1432,1798,1825]" degrees="23" direction="east" minutes="50" orientation="longitude" pageId="6" pageNumber="565" precision="1" seconds="52.38" value="23.847883">23°5052.38”E</geoCoordinate>
,
<quantity id="4CA59B3DE47EFFC6FF235336FF294512" box="[151,225,1834,1861]" metricMagnitude="2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="565" unit="m" value="508.0">508 m</quantity>
,
<date id="FFE31018E47EFFC6FF595337FE974512" box="[237,351,1835,1861]" pageId="6" pageNumber="565" value="2018-10-21">21.x.2018</date>
,
<taxonomicName id="4C5D4D5BE47EFFC6FED85336FE184513" box="[364,464,1834,1860]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="6" pageNumber="565" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EACAE47EFFC6FED85336FE184513" box="[364,464,1834,1860]" italics="true" pageId="6" pageNumber="565">Platanus</emphasis>
</taxonomicName>
sifting litter, L. Davranoglou lgt.” [white, printed] (
<collectionCode id="ED4CAE1DE47EFFC6FBAC5337FBA64512" box="[1048,1134,1835,1861]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="565">NMPC</collectionCode>
).
<emphasis id="B929EACAE47EFFC6FB355337FB314512" bold="true" box="[1153,1273,1835,1861]" pageId="6" pageNumber="565">
<typeStatus id="54E6887AE47EFFC6FB355337FB314512" box="[1153,1273,1835,1861]" pageId="6" pageNumber="565">Paratypes</typeStatus>
</emphasis>
, 3 ♁♁,
<specimenCount id="9D5BFD51E47EFFC6FAE25337FA5E4512" box="[1366,1430,1835,1861]" pageId="6" pageNumber="565" type="female">4 ♀♀</specimenCount>
, 2 exx, same data as for holotype (
<collectionCode id="ED4CAE1DE47EFFC6FD925353FD49453E" box="[550,641,1871,1897]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="6" pageNumber="565">MMBC</collectionCode>
,
<collectionCode id="ED4CAE1DE47EFFC6FD245353FD13453E" box="[656,731,1871,1897]" pageId="6" pageNumber="565">PCPH</collectionCode>
).
<emphasis id="B929EACAE47EFFC6FD455352FC0B453F" bold="true" box="[753,963,1870,1896]" pageId="6" pageNumber="565">Other specimens.</emphasis>
1 ♁,
<specimenCount id="9D5BFD51E47EFFC6FBBF5353FB87453E" box="[1035,1103,1871,1897]" pageId="6" pageNumber="565" type="female">2 ♀♀</specimenCount>
: Greece, Euboea, Stropones,
<taxonomicName id="4C5D4D5BE47EFFC6FF23536FFEC945DA" box="[151,257,1907,1933]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="565" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EACAE47EFFC6FF23536FFEC945DA" box="[151,257,1907,1933]" italics="true" pageId="6" pageNumber="565">Castanea</emphasis>
</taxonomicName>
forest leaf litter,
<date id="FFE31018E47EFFC6FE77536EFDEE45DB" box="[451,550,1906,1932]" pageId="6" pageNumber="565" value="2017-05-06">6.v.2017</date>
, G. Kakiopoulos lgt.; 1 ♁: Greece, Euboea, Glyfada, evergreen
<taxonomicName id="4C5D4D5BE47EFFC6FB43536FFA9D45DA" box="[1271,1365,1907,1933]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="565" phylum="Tracheophyta" rank="genus">
<emphasis id="B929EACAE47EFFC6FB43536FFA9D45DA" box="[1271,1365,1907,1933]" italics="true" pageId="6" pageNumber="565">Quercus</emphasis>
</taxonomicName>
forest leaf-litter,
<quantity id="4CA59B3DE47EFFC6FEB8538BFE9D45E6" box="[268,341,1943,1969]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="565" unit="m" value="400.0">400 m</quantity>
,
<date id="FFE31018E47EFFC6FED5538AFE1545E7" box="[353,477,1942,1968]" pageId="6" pageNumber="565" value="2017-03-19">19.iii.2017</date>
, G. Kakiopoulos lgt. (
<collectionCode id="ED4CAE1DE47EFFC6FD60538BFCED45E6" box="[724,805,1943,1969]" pageId="6" pageNumber="565">PCGK</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47EFFC7FF7353A6FAAA4282" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565" type="diagnosis">
<paragraph id="8BE236D8E47EFFC7FF7353A6FAAA4282" blockId="6.[151,1437,1798,2041]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF7353A6FE8A4583" bold="true" box="[199,322,1978,2004]" pageId="6" pageNumber="565">Diagnosis.</emphasis>
Head 1.5 times longer than wide, scape 2.3 times longer than wide, tergite 1 (IV) 1.4 times broader than long, sternite 2 (IV) 2.3 times wider than long; metaventrite of male produced into acute spine (
<figureCitation id="13662A5DE47EFFC6FB5253C2FAF545AF" box="[1254,1341,2014,2040]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="6" pageNumber="565">Fig. 6C</figureCitation>
); central spine of internal sac of aedeagus with a membranous branch that overlays second hook-like branch (
<figureCitation id="13662A5DE47FFFC7FAA2548BFA4742E6" box="[1302,1423,151,177]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="7" pageNumber="566">Fig. 7E, F</figureCitation>
); second spine of aedeagus simple, located on top of membrane bearing small number of denticles (
<figureCitation id="13662A5DE47FFFC7FB5454A7FA9D4282" box="[1248,1365,187,213]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="7" pageNumber="566">Fig. 7E, F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF226650E47FFFC7FF2350C4FF77476F" ID-DOI="http://doi.org/10.5281/zenodo.8392309" ID-Zenodo-Dep="8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="7" pageNumber="566" startId="7.[151,250,1240,1264]" targetBox="[325,1263,237,1217]" targetPageId="7" targetType="figure">
<paragraph id="8BE236D8E47FFFC7FF2350C4FF77476F" blockId="7.[151,1436,1240,1336]" pageId="7" pageNumber="566">
<emphasis id="B929EACAE47FFFC7FF2350C4FEDA46A7" bold="true" box="[151,274,1240,1264]" pageId="7" pageNumber="566">FIGURE 6.</emphasis>
Lateral view of metaventrite in the examined
<taxonomicName id="4C5D4D5BE47FFFC7FD5750C4FC8746A7" authorityName="Jeannel" authorityYear="1950" box="[739,847,1240,1264]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="tribe" tribe="Pselaphini">Pselaphini</taxonomicName>
, using scanning electron microscopy. A)
<taxonomicName id="4C5D4D5BE47FFFC7FB5850C5FAB146A7" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1260,1401,1241,1264]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47FFFC7FB5850C5FAB146A7" box="[1260,1401,1241,1264]" italics="true" pageId="7" pageNumber="566">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47FFFC7FACA50C4FF644743" bold="true" pageId="7" pageNumber="566">
<taxonomicNameLabel id="A21A57B1E47FFFC7FACA50C4FF644743" pageId="7" pageNumber="566" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B)
<taxonomicName id="4C5D4D5BE47FFFC7FF6F50E1FE844743" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[219,332,1276,1300]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47FFFC7FF6F50E1FE844743" box="[219,332,1276,1300]" italics="true" pageId="7" pageNumber="566">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47FFFC7FEE050E0FE464743" bold="true" box="[340,398,1276,1300]" pageId="7" pageNumber="566">
<taxonomicNameLabel id="A21A57B1E47FFFC7FEE050E0FE464743" box="[340,398,1276,1300]" pageId="7" pageNumber="566" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; C)
<taxonomicName id="4C5D4D5BE47FFFC7FE0A50E1FDFD4743" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[446,565,1276,1300]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE47FFFC7FE0A50E1FDFD4743" box="[446,565,1276,1300]" italics="true" pageId="7" pageNumber="566">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47FFFC7FD8850E0FDBF4743" bold="true" box="[572,631,1276,1300]" pageId="7" pageNumber="566">
<taxonomicNameLabel id="A21A57B1E47FFFC7FD8850E0FDBF4743" box="[572,631,1276,1300]" pageId="7" pageNumber="566" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="4C5D4D5BE47FFFC7FD1350E0FC674743" baseAuthorityName="Karaman" baseAuthorityYear="1940" box="[679,943,1276,1300]" class="Insecta" family="Staphylinidae" genus="Pselaphogenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="treskanus">
<emphasis id="B929EACAE47FFFC7FD1350E0FC674743" box="[679,943,1276,1300]" italics="true" pageId="7" pageNumber="566">Pselaphogenius treskanus</emphasis>
</taxonomicName>
(Karaman, 1940). Scale bars correspond to 100 um.
</paragraph>
</caption>
<subSubSection id="C3476553E47FFFC8FF73517AFC834514" lastPageId="8" lastPageNumber="567" pageId="7" pageNumber="566" type="description">
<paragraph id="8BE236D8E47FFFC7FF73517AFE9C4447" blockId="7.[151,1437,1382,2021]" pageId="7" pageNumber="566">
<emphasis id="B929EACAE47FFFC7FF73517AFE9147D7" bold="true" box="[199,345,1382,1408]" pageId="7" pageNumber="566">Description.</emphasis>
Body length
<quantity id="4CA59B3DE47FFFC7FE5A517BFD8E47D6" box="[494,582,1383,1409]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="566" unit="mm" value="1.8">1.8 mm</quantity>
. Coloration largely reddish-brown, maxillary palpi and legs lighter, posterior margin of elytra much darker, almost black (
<figureCitation id="13662A5DE47FFFC7FD3F5196FD2C47F2" box="[651,740,1418,1445]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="7" pageNumber="566">Fig. 2C</figureCitation>
). Pilosity of body sparse; head, legs, and antennae covered by sparse, adjacent golden setae; elytra with few rows of setae, which are denser on elytral marginal line; with row of setae on distal margin of elytra; proximal margin of tergite II with very dense pilosity (
<figureCitation id="13662A5DE47FFFC7FBE851CEFB7B47BA" box="[1116,1203,1490,1517]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="7" pageNumber="566">Fig. 2C</figureCitation>
), remaining segment largely glabrous.
</paragraph>
<paragraph id="8BE236D8E47FFFC7FF735206FCCC455B" blockId="7.[151,1437,1382,2021]" pageId="7" pageNumber="566">
Head 1.5 times longer than wide, each eye composed of 9 ommatidia (
<figureCitation id="13662A5DE47FFFC7FBB95206FBBB4462" box="[1037,1139,1562,1589]" captionStart="FIGURE 3" captionStartId="4.[151,250,1764,1788]" captionTargetBox="[325,1263,345,1741]" captionTargetId="figure-99@4.[325,1263,345,1741]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron microscopy image of dorsal view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392301" httpUri="https://zenodo.org/record/8392301/files/figure.png" pageId="7" pageNumber="566">Figs. 3C</figureCitation>
,
<figureCitation id="13662A5DE47FFFC7FB305207FB604462" box="[1156,1192,1563,1589]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="7" pageNumber="566">4C</figureCitation>
); median gular ridge distinctly protruding distally, with subtriangular apex that is almost touching gular surface (
<figureCitation id="13662A5DE47FFFC7FB205222FB25440E" box="[1172,1261,1598,1625]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="7" pageNumber="566">Fig. 4C</figureCitation>
); antennae
<quantity id="4CA59B3DE47FFFC7FADF5223FF0B442A" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="7" pageNumber="566" unit="mm" value="0.97">0.97 mm</quantity>
long; scape cylindrical, 2.3 times longer than wide, and twice longer than pedicel; pedicel about as long as wide; pedicel and antennomeres 38 rounded, subequal in length; antennomeres 910 about 1.6 times longer than length of each antennomere 38; terminal antennomere (11) about 0.75 times as long as 910 combined, and 1.3 times longer than wide (
<figureCitation id="13662A5DE47FFFC7FEEB52D2FE7F44BE" box="[351,439,1742,1769]" captionStart="FIGURE 5" captionStartId="6.[151,250,1008,1032]" captionTargetBox="[361,1225,181,984]" captionTargetId="figure-26@6.[361,1225,181,984]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron microscopy image of antenna in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392305" httpUri="https://zenodo.org/record/8392305/files/figure.png" pageId="7" pageNumber="566">Fig. 5C</figureCitation>
). Maxillary palpomere 1 shortest, 23 times longer than 1; 3 extremely short and rounded; palpomere 4 1.6 times longer than 2, apex club-shaped.
</paragraph>
<paragraph id="8BE236D8E47FFFC7FF73530BFDD345CB" blockId="7.[151,1437,1382,2021]" pageId="7" pageNumber="566">
Pronotum
<quantity id="4CA59B3DE47FFFC7FE8F530BFE564566" box="[315,414,1815,1841]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="566" unit="mm" value="0.29">0.29 mm</quantity>
wide, smooth, about 1.2 times longer than wide; slightly constricted proximally and distally, broadest medially; about 0.8 of length of head; lacking median sulcus and carinae, median antebasal and lateral foveae absent; procoxal fovea and prothoracic-mesothoracic junction with dense tuft of setae. Metaventrite produced into acute, conical spine (
<figureCitation id="13662A5DE47FFFC7FE01539EFDC645CB" box="[437,526,1922,1948]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="7" pageNumber="566">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="8BE236D8E47FFFC7FF7353BAFBCA45B3" blockId="7.[151,1437,1382,2021]" pageId="7" pageNumber="566">Elytra about 1.7 times broader than long, and 1.7 times longer than pronotum; elytral surface smooth, lacking striae or distinct punctures; elytra with one basal fovea, and one sutural fovea.</paragraph>
<caption id="DF226650E470FFC8FF2350E8FECD472F" ID-DOI="http://doi.org/10.5281/zenodo.8392311" ID-Zenodo-Dep="8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="8" pageNumber="567" startId="8.[151,250,1268,1292]" targetBox="[208,1366,170,1243]" targetPageId="8" targetType="figure">
<paragraph id="8BE236D8E470FFC8FF2350E8FECD472F" blockId="8.[151,1437,1268,1400]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF2350E8FEDD475B" bold="true" box="[151,277,1268,1292]" pageId="8" pageNumber="567">FIGURE 7.</emphasis>
Photomicrographs showing the dorsal aedeagal surface of the examined
<taxonomicName id="4C5D4D5BE470FFC8FBBE50E8FBBF475B" authorityName="Jeannel" authorityYear="1950" box="[1034,1143,1268,1292]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="tribe" tribe="Pselaphini">Pselaphini</taxonomicName>
. A)
<taxonomicName id="4C5D4D5BE470FFC8FB1750E8FAFA475B" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1187,1330,1268,1292]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE470FFC8FB1750E8FAFA475B" box="[1187,1330,1268,1292]" italics="true" pageId="8" pageNumber="567">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE470FFC8FA8D50E8FABC475B" bold="true" box="[1337,1396,1268,1292]" pageId="8" pageNumber="567">
<taxonomicNameLabel id="A21A57B1E470FFC8FA8D50E8FABC475B" box="[1337,1396,1268,1292]" pageId="8" pageNumber="567" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B) same, schematic; C)
<taxonomicName id="4C5D4D5BE470FFC8FEC55104FE2C4767" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[369,484,1304,1328]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE470FFC8FEC55104FE2C4767" box="[369,484,1304,1328]" italics="true" pageId="8" pageNumber="567">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE470FFC8FE595104FDE04767" bold="true" box="[493,552,1304,1328]" pageId="8" pageNumber="567">
<taxonomicNameLabel id="A21A57B1E470FFC8FE595104FDE04767" box="[493,552,1304,1328]" pageId="8" pageNumber="567" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; D) same, schematic; E)
<taxonomicName id="4C5D4D5BE470FFC8FC805104FC654767" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[820,941,1304,1328]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE470FFC8FC805104FC654767" box="[820,941,1304,1328]" italics="true" pageId="8" pageNumber="567">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE470FFC8FC025104FC394767" bold="true" box="[950,1009,1304,1328]" pageId="8" pageNumber="567">
<taxonomicNameLabel id="A21A57B1E470FFC8FC025104FC394767" box="[950,1009,1304,1328]" pageId="8" pageNumber="567" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; F) same, schematic; G)
<taxonomicName id="4C5D4D5BE470FFC8FB485104FF3F4703" baseAuthorityName="Karaman" baseAuthorityYear="1940" class="Insecta" family="Staphylinidae" genus="Pselaphogenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="treskanus">
<emphasis id="B929EACAE470FFC8FB485104FF3F4703" italics="true" pageId="8" pageNumber="567">Pselaphogenius treskanus</emphasis>
</taxonomicName>
(Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um.
</paragraph>
</caption>
<paragraph id="8BE236D8E470FFC8FF7351B9FDF147B4" blockId="8.[151,1437,1445,2036]" pageId="8" pageNumber="567">Abdomen about as broad as elytra; tergite 1 (IV) (excluding paratergites) 1.4 times wider than long; sternite 2 (IV) about 2.3 times wider than long.</paragraph>
<paragraph id="8BE236D8E470FFC8FF7351F0FAFC447E" blockId="8.[151,1437,1445,2036]" pageId="8" pageNumber="567">
Aedeagus (
<figureCitation id="13662A5DE470FFC8FEF251F0FE724451" box="[326,442,1516,1542]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="8" pageNumber="567">Fig. 7E, F</figureCitation>
) with central spine of internal sac of aedeagus with membranous branch that overlays a second hook-like branch; second spine of aedeagus simple, located on top of denticle-bearing membrane.
</paragraph>
<paragraph id="8BE236D8E470FFC8FF73522EFCB54577" blockId="8.[151,1437,1445,2036]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF73522EFC79441B" bold="true" box="[199,945,1586,1612]" pageId="8" pageNumber="567">
Measurements of
<typeStatus id="54E6887AE470FFC8FE17522EFDC2441B" box="[419,522,1586,1612]" pageId="8" pageNumber="567">holotype</typeStatus>
(in mm)—W = width, L = length.
</emphasis>
Body, L—1.77; head, L—0.42, W—0.27; antenna, L—0.97; scapus, L—0.17, W—0.07; pedicel, L—0.09, W—0.06; antennomere III, L—0.07; antennomere IV, L—0.06; antennomere V, L—0.06; antennomere VI, L—0.06; antennomere VII, L—0.06; antennomere VIII, L—0.06; antennomere IX, L—0.1; antennomere X, L—0.1; antennomere XI, L—0.15, W—0.11; maxillary palpomere I, L—0.09; maxillary palpomere II, L—0.26; maxillary palpomere III, L—0.05; maxillary palpomere IV, L—0.44; pronotum, L—0.35, W—0.29; elytra, L—0.6, W—0.35; tergite IV, L—0.44, W—0.75; tergite IV (excluding paratergites) W—0.61; ventrite IV, L—0.37, W—0.87.
</paragraph>
<paragraph id="8BE236D8E470FFC8FF735335FC834514" blockId="8.[151,1437,1445,2036]" box="[199,843,1833,1859]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF735335FEED4514" bold="true" box="[199,293,1833,1859]" pageId="8" pageNumber="567">Female.</emphasis>
Similar to male, but with a smooth metaventrite.
</paragraph>
</subSubSection>
<subSubSection id="C3476553E470FFC8FF735350FE9F45FB" pageId="8" pageNumber="567" type="etymology">
<paragraph id="8BE236D8E470FFC8FF735350FAE14531" blockId="8.[151,1437,1445,2036]" box="[199,1321,1868,1894]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF735350FE864531" bold="true" box="[199,334,1868,1894]" pageId="8" pageNumber="567">Etymology.</emphasis>
The new species was named after the island of Euboea, where it appears to be endemic.
</paragraph>
<paragraph id="8BE236D8E470FFC8FF735373FE9F45FB" blockId="8.[151,1437,1445,2036]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF735373FEE245DE" bold="true" box="[199,298,1903,1929]" pageId="8" pageNumber="567">Habitat.</emphasis>
Specimens were collected in dense leaf-litter accumulations from a range of broadleaf deciduous forests (
<figureCitation id="13662A5DE470FFC8FF45538EFE8245FB" box="[241,330,1938,1964]" captionStart="FIGURE 8" captionStartId="9.[151,250,1185,1209]" captionTargetBox="[344,1243,181,1161]" captionTargetId="figure-17@9.[344,1243,181,1161]" captionTargetPageId="9" captionText="FIGURE 8. Habitat photos of examined species. A) Type locality of A. taygetensis sp. n. in Kastoreion; B) type locality of A. tymficus sp. n.; in Mt Tymfi; C) leaf litter where A. euboicus sp. n. was collected from in Mt Dirfis, the type locality; D) habitat in Mt Vitsi, where Pselaphogenius treskanus (Karaman, 1940) was collected from." figureDoi="http://doi.org/10.5281/zenodo.8392315" httpUri="https://zenodo.org/record/8392315/files/figure.png" pageId="8" pageNumber="567">Fig. 8C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E470FFC8FF7353A9FB104598" box="[199,1240,1973,2000]" pageId="8" pageNumber="567" type="distribution">
<paragraph id="8BE236D8E470FFC8FF7353A9FB104598" blockId="8.[151,1437,1445,2036]" box="[199,1240,1973,2000]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF7353A9FEA94598" bold="true" box="[199,353,1973,1999]" pageId="8" pageNumber="567">Distribution.</emphasis>
<collectingCountry id="F34A7648E470FFC8FED353AAFE7E4587" box="[359,438,1974,2000]" name="Greece" pageId="8" pageNumber="567">Greece</collectingCountry>
(so far endemic to the island of Euboea, on Mt Dirfis and its environs).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E470FFC8FF7353C6FA4245A4" box="[199,1418,2009,2036]" pageId="8" pageNumber="567" type="discussion">
<paragraph id="8BE236D8E470FFC8FF7353C6FA4245A4" blockId="8.[151,1437,1445,2036]" box="[199,1418,2009,2036]" pageId="8" pageNumber="567">
<emphasis id="B929EACAE470FFC8FF7353C6FEF345A3" bold="true" box="[199,315,2010,2036]" pageId="8" pageNumber="567">Remarks.</emphasis>
The sclerites of the internal sac of aedeagus of
<taxonomicName id="4C5D4D5BE470FFC8FCE453C6FC1B45A4" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[848,979,2009,2036]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE470FFC8FCE453C6FC1B45A4" box="[848,979,2009,2036]" italics="true" pageId="8" pageNumber="567">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE470FFC8FC6E53C6FBD145A3" bold="true" box="[986,1049,2010,2036]" pageId="8" pageNumber="567">
<taxonomicNameLabel id="A21A57B1E470FFC8FC6E53C6FBD145A3" box="[986,1049,2010,2036]" pageId="8" pageNumber="567" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are characteristic to this species.
</paragraph>
</subSubSection>
</treatment>
</document>