172 lines
18 KiB
XML
172 lines
18 KiB
XML
<document id="93DD54D33AACA532B6D705A815C35C2A" ID-CLB-Dataset="207550" ID-DOI="10.11646/zootaxa.5219.5.4" ID-GBIF-Dataset="7391a03c-6a6e-4f02-9e7e-546c391f1a67" ID-ISSN="1175-5326" ID-Zenodo-Dep="7438997" ID-ZooBank="ECC4C9BF-2E15-4C6A-8C69-C3B8E30A400B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670916363146" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Perkovsky, Evgeny E., Anisyutkin, Leonid N. & Dubovikoff, Dmitry A." docDate="2022" docId="03F487CED60FFFB71DE0FE70FBF9FC26" docLanguage="en" docName="zootaxa.5219.5.4.pdf" docOrigin="Zootaxa 5219 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Raphidioptera Linnaeus 1758" docType="treatment" docVersion="7" lastPageNumber="461" masterDocId="FFCDFFB6D60BFFB21D77FFAAFF81FFCB" masterDocTitle="First larvae of Raphidioptera from Eocene Sakhalinian and Rovno ambers" masterLastPageNumber="466" masterPageNumber="456" pageNumber="460" updateTime="1699413421191" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BD2D8EE3203CC3B2CD514435CA3C3E8A">
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<mods:title id="1F6A19C789E842CFFA4BEC7492261C69">First larvae of Raphidioptera from Eocene Sakhalinian and Rovno ambers</mods:title>
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<mods:name id="44816183620E73A96A76E3A9D0B9D0D2" type="personal">
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<mods:namePart id="68F40104AE8C7CD22203068BD075AAAD">Makarkin, Vladimir N.</mods:namePart>
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<mods:nameIdentifier id="856793DC473365456070F530FE365809" type="ORCID">0000-0002-1304-0461</mods:nameIdentifier>
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<mods:affiliation id="4F5BC5BFA5BD940FED8A5142FF58140B">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. vnmakarkin @ mail. ru; https: // orcid. org / 0000 - 0002 - 1304 - 046</mods:affiliation>
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<mods:nameIdentifier id="6B3501318CE90CA8707DE100B49A70B0" type="email">vnmakarkin@mail.ru</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="E7D4127ABCA69D5DDDF579AE69A6D373">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="47B1F58EA71E86095738D84C6FD6E2F2">Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, ul. Bogdana Khmel’nitskogo 15, Kiev, 01030 Ukraine; Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, Moscow, 117647, Russia</mods:affiliation>
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<mods:name id="ED14C4AF7471694B1FE8DFD8A7F1BA35" type="personal">
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<mods:roleTerm id="11A6A3301170BDE646E51A17F2096061">Author</mods:roleTerm>
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<mods:namePart id="C4F3E153DDC301E8198462BA1E052F32">Anisyutkin, Leonid N.</mods:namePart>
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<mods:nameIdentifier id="5C63F9C7360688756AFF98520BC7FFF0" type="ORCID">0000-0002-7587-9078</mods:nameIdentifier>
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<mods:affiliation id="6FB22FB7B5812712CA5C0A0C2DC7ED98">Zoological Institute of Russian Academy of Sciences, Universitetskaya nab., 1, Saint-Petersburg, 199034, Russia. https: // orcid. org / 0000 - 0002 - 7587 - 9078</mods:affiliation>
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</mods:role>
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<mods:namePart id="E89579B3CFF130C8119A58CBDD28701F">Dubovikoff, Dmitry A.</mods:namePart>
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<mods:affiliation id="6ED97E65D22DA7F6150C621F61E6B3FD">Saint-Petersburg State University, 16 - liniya, 29, Saint-Petersburg, 199178, Russia.</mods:affiliation>
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<mods:title id="618D67EFD92D0432F05AE255E18E0821">Zootaxa</mods:title>
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<mods:part id="32054C11319F997F4802C67A55789F70">
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<mods:date id="2F16B76CAFFC090CB72485F7CD1AE4BF">2022</mods:date>
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<mods:number id="E81DF0AE0CFAB0801792E4692CCF9572">2022-12-13</mods:number>
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<mods:number id="065CEDE83F7230800BFAC4CA152A0BE0">5219</mods:number>
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<mods:identifier id="107D2AAD0951B63F19E482AF6DA994DE" type="DOI">10.11646/zootaxa.5219.5.4</mods:identifier>
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<mods:identifier id="1CC7854F2A1F8F716283C6B38D7EF72D" type="GBIF-Dataset">7391a03c-6a6e-4f02-9e7e-546c391f1a67</mods:identifier>
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<treatment id="03F487CED60FFFB71DE0FE70FBF9FC26" ID-DOI="http://doi.org/10.5281/zenodo.7439003" ID-GBIF-Taxon="206230605" ID-Zenodo-Dep="7439003" LSID="urn:lsid:plazi:treatment:03F487CED60FFFB71DE0FE70FBF9FC26" httpUri="http://treatment.plazi.org/id/03F487CED60FFFB71DE0FE70FBF9FC26" lastPageId="5" lastPageNumber="461" pageId="4" pageNumber="460">
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<subSubSection id="C3476553D60FFFB61DE0FE70FDDDFE3E" box="[151,604,474,501]" pageId="4" pageNumber="460" type="nomenclature">
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<paragraph id="8BE236D8D60FFFB61DE0FE70FDDDFE3E" blockId="4.[151,604,474,537]" box="[151,604,474,501]" pageId="4" pageNumber="460">
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<heading id="D0AA81B4D60FFFB61DE0FE70FDDDFE3E" bold="true" box="[151,604,474,501]" fontSize="11" level="1" pageId="4" pageNumber="460" reason="1">
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<emphasis id="B929EACAD60FFFB61DE0FE70FDDDFE3E" bold="true" box="[151,604,474,501]" pageId="4" pageNumber="460">
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<taxonomicName id="4C5D4D5BD60FFFB61DE0FE70FECFFE3F" ID-CoL="6236X" ID-ENA="50482" authorityName="Linnaeus" authorityYear="1758" box="[151,334,474,500]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="460" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
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fam. gen. sp. indet. B
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476553D60FFFB61DE0FE55FF58FDD2" box="[151,217,511,537]" pageId="4" pageNumber="460" type="description">
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<paragraph id="8BE236D8D60FFFB61DE0FE55FF58FDD2" blockId="4.[151,604,474,537]" box="[151,217,511,537]" pageId="4" pageNumber="460">
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<figureCitation id="13662A5DD60FFFB61DE0FE55FF58FDD2" box="[151,217,511,537]" captionStart="FIGURE 3" captionStartId="4.[152,255,1897,1921]" captionTargetBox="[151,1436,561,1874]" captionTargetId="figure-143@4.[151,1436,561,1874]" captionTargetPageId="4" captionText="FIGURE 3. Raphidioptera fam. gen. sp. indet. B, specimen SIZK K-8213. A, dorsal view; B, ventral view; C, head, dorsal view; D, anterior part of the head (ventral view); E, right foreleg. a1, 1st antennomere; a2, 2nd antennomere; cl, claw; cs, coronal suture; cx, coxa; fe, femur; fs, frontal suture; he, head; lp, labial palpus; msn, mesonotum; mp, maxillary palpus; mtn, metanotum; pn, pronotum; ta, tarsus; ti, tibia; tr, trochanter. Scale bars = 1 mm (A–D), 0.5 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.7439007" httpUri="https://zenodo.org/record/7439007/files/figure.png" pageId="4" pageNumber="460">Fig. 3</figureCitation>
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</paragraph>
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</subSubSection>
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<caption id="DF226650D60FFFB61DEFF8C3FB02F825" ID-DOI="http://doi.org/10.5281/zenodo.7439007" ID-Zenodo-Dep="7439007" httpUri="https://zenodo.org/record/7439007/files/figure.png" pageId="4" pageNumber="460" startId="4.[152,255,1897,1921]" targetBox="[151,1436,561,1874]" targetPageId="4">
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<paragraph id="8BE236D8D60FFFB61DEFF8C3FB02F825" blockId="4.[151,1436,1897,2030]" pageId="4" pageNumber="460">
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<emphasis id="B929EACAD60FFFB61DEFF8C3FE9BF849" bold="true" box="[152,282,1897,1922]" pageId="4" pageNumber="460">FIGURE 3.</emphasis>
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<taxonomicName id="4C5D4D5BD60FFFB61C55F8C0FE33F849" authorityName="Linnaeus" authorityYear="1758" box="[290,434,1898,1922]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="460" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
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fam. gen. sp. indet. B, specimen SIZK K-8213. A, dorsal view; B, ventral view; C, head, dorsal view; D, anterior part of the head (ventral view); E, right foreleg. a1, 1st antennomere; a2, 2nd antennomere; cl, claw; cs, coronal suture; cx, coxa; fe, femur; fs, frontal suture; he, head; lp, labial palpus; msn, mesonotum; mp, maxillary palpus; mtn, metanotum; pn, pronotum; ta, tarsus; ti, tibia; tr, trochanter. Scale bars = 1 mm (A–D), 0.5 mm (E).
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</paragraph>
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</caption>
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<subSubSection id="C3476553D60EFFB71DE0FF3CFAA1FED6" pageId="5" pageNumber="461" type="materials_examined">
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<paragraph id="8BE236D8D60EFFB71DE0FF3CFDC6FF32" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">
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<emphasis id="B929EACAD60EFFB71DE0FF3CFE00FF7B" bold="true" box="[151,385,150,176]" pageId="5" pageNumber="461">Material examined.</emphasis>
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<materialsCitation id="3B353C85D60EFFB71CFFFF3DFDC2FF32" ID-GBIF-Occurrence="3989533303" collectionCode="SIZK" country="Ukraine" county="Syninclusions" location="Schmalhausen Institute of Zoology" municipality="An" pageId="5" pageNumber="461" specimenCount="1" stateProvince="Kyiv">
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Specimen
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<collectionCode id="ED4CAE1DD60EFFB71C89FF3DFDBDFF7A" box="[510,572,151,177]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="5" pageNumber="461">SIZK</collectionCode>
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K-8213, deposited in the
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<location id="8E826003D60EFFB71E16FF3DFB6DFF7A" LSID="urn:lsid:plazi:treatment:03F487CED60FFFB71DE0FE70FBF9FC26:8E826003D60EFFB71E16FF3DFB6DFF7A" box="[865,1260,151,177]" country="Ukraine" county="Syninclusions" municipality="An" name="Schmalhausen Institute of Zoology" pageId="5" pageNumber="461" stateProvince="Kyiv">Schmalhausen Institute of Zoology</location>
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,
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<collectingRegion id="4999F83AD60EFFB7198FFF3DFAAEFF7A" box="[1272,1327,151,177]" country="Ukraine" name="Kyiv" pageId="5" pageNumber="461">Kiev</collectingRegion>
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,
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<collectingCountry id="F34A7648D60EFFB7184CFF3DFA18FF7A" box="[1339,1433,151,177]" name="Ukraine" pageId="5" pageNumber="461">Ukraine</collectingCountry>
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.
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<collectingMunicipality id="6B86ACA2D60EFFB71DE0FF11FF38FF1E" box="[151,185,187,213]" pageId="5" pageNumber="461">An</collectingMunicipality>
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incomplete mature larva lacking the posterior part; details of the head are unclear in part because of a milky covering.
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<collectingCounty id="62834E54D60EFFB71C70FF75FE28FF32" box="[263,425,223,249]" pageId="5" pageNumber="461">Syninclusions</collectingCounty>
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: stellate hairs
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE236D8D60EFFB71DB0FEA8FAA1FED6" blockId="5.[151,1437,150,1005]" box="[199,1312,258,285]" pageId="5" pageNumber="461">
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<emphasis id="B929EACAD60EFFB71DB0FEA8FE46FED7" bold="true" box="[199,455,258,285]" pageId="5" pageNumber="461">Locality and horizon.</emphasis>
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<materialsCitation id="3B353C85D60EFFB71CB9FEA9FC66FED6" ID-GBIF-Occurrence="3989533301" box="[462,999,259,285]" country="Ukraine" location="Sarny District" pageId="5" pageNumber="461" specimenCount="1" stateProvince="Rivne">
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<collectingCountry id="F34A7648D60EFFB71CB9FEA9FDACFED6" box="[462,557,259,285]" name="Ukraine" pageId="5" pageNumber="461">Ukraine</collectingCountry>
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:
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<collectingRegion id="4999F83AD60EFFB71F40FEA9FD5FFED6" box="[567,734,259,285]" country="Ukraine" name="Rivne" pageId="5" pageNumber="461">Rovno Region</collectingRegion>
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:
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<location id="8E826003D60EFFB71F9FFEA9FC08FED6" LSID="urn:lsid:plazi:treatment:03F487CED60FFFB71DE0FE70FBF9FC26:8E826003D60EFFB71F9FFEA9FC08FED6" box="[744,905,259,285]" country="Ukraine" name="Sarny District" pageId="5" pageNumber="461" stateProvince="Rivne">Sarny District</location>
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: Klesov
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</materialsCitation>
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;
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<collectingRegion id="4999F83AD60EFFB71E85FEA9FBBCFED6" box="[1010,1085,259,285]" country="Ukraine" name="Rivne" pageId="5" pageNumber="461">Rovno</collectingRegion>
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amber, late Eocene.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476553D60EFFB71DB0FE8CFF64FC96" pageId="5" pageNumber="461" type="description">
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<paragraph id="8BE236D8D60EFFB71DB0FE8CFB36FE8A" blockId="5.[151,1437,150,1005]" box="[199,1207,294,321]" pageId="5" pageNumber="461">
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<emphasis id="B929EACAD60EFFB71DB0FE8CFED8FE8B" bold="true" box="[199,345,294,320]" pageId="5" pageNumber="461">Description.</emphasis>
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Larva. Preserved length
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<emphasis id="B929EACAD60EFFB71F06FE8DFD0AFE8B" box="[625,651,295,320]" italics="true" pageId="5" pageNumber="461">ca</emphasis>
|
||
.
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<quantity id="4CA59B3DD60EFFB71FEDFE8CFC81FE8A" box="[666,768,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="461" unit="mm" value="12.5">12.5 mm</quantity>
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, estimated complete length
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<emphasis id="B929EACAD60EFFB7194FFE8DFBD3FE8B" box="[1080,1106,295,320]" italics="true" pageId="5" pageNumber="461">ca</emphasis>
|
||
.
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<quantity id="4CA59B3DD60EFFB71916FE8CFB32FE8A" box="[1121,1203,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="461" unit="mm" value="15.0">15 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE236D8D60EFFB71DB0FEE1FEC8FDF6" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">
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Head relatively broad, not narrowed posteriorly,
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<emphasis id="B929EACAD60EFFB71FA8FEE1FD78FEAF" box="[735,761,331,356]" italics="true" pageId="5" pageNumber="461">ca</emphasis>
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. 2.0 mm long (including mandibles),
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<quantity id="4CA59B3DD60EFFB719EFFEE0FB7BFEAE" box="[1176,1274,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="461" unit="mm" value="1.25">1.25 mm</quantity>
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wide (
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<emphasis id="B929EACAD60EFFB71833FEE1FADFFEAF" box="[1348,1374,331,356]" italics="true" pageId="5" pageNumber="461">ca</emphasis>
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. 1.60 times as long as wide). Ecdysial cleavage lines poorly discernible, consisting of frontal and coronal sutures; frontal sutures diverge at a relatively sharp angle. Mandible protruding, poorly discernible. Maxillary palpi relatively long; four distal elongate palpomeres; basal palpomere not discernible. Labial palpi short, three-segmented. Antennal tubercles probably short, but not clearly discernible. Antenna: first antennomere rather short and stout; second antennomere very short, narrow; other antennomeres broken. Stemmata poorly discernible; at least three stemmata visible dorsally.
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</paragraph>
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<paragraph id="8BE236D8D60EFFB71DB0FDEDFA19FD62" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">
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Thorax covered with long scarce setae. Pronotum relatively short, widest anteriorly;
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<quantity id="4CA59B3DD60EFFB719FEFDECFB6FFDAA" box="[1161,1262,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="461" unit="mm" value="1.5">1.50 mm</quantity>
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long,
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<quantity id="4CA59B3DD60EFFB71840FDECFA1CFDAA" box="[1335,1437,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="461" unit="mm" value="1.15">1.15 mm</quantity>
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wide (1.30 times as long as wide); shorter than head (0.79 times as long as head length). Mesonotum nearly quadrate (0.92 times as long as wide), widest caudally. Metanotum transverse (0.70 times as long as wide), widest caudally.
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</paragraph>
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<paragraph id="8BE236D8D60EFFB71DB0FD19FDC0FCDE" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">All legs similar in structure, short, covered with long scarce setae. Coxa, trochanter very short, stout. Femur elongate, stout. Tibia nearly as long as femur, rather stout. Tarsus elongate, narrowed toward apex (conical); claws small, slightly curved, basally dilated.</paragraph>
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<paragraph id="8BE236D8D60EFFB71DB0FCB5FF64FC96" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">Abdomen wider than thorax; seven preserved segments transverse (their length less than width). Setae long, scarce.</paragraph>
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</subSubSection>
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<subSubSection id="C3476553D60EFFB71DB0FCCDFBF9FC26" pageId="5" pageNumber="461" type="discussion">
|
||
<paragraph id="8BE236D8D60EFFB71DB0FCCDFBF9FC26" blockId="5.[151,1437,150,1005]" pageId="5" pageNumber="461">
|
||
<emphasis id="B929EACAD60EFFB71DB0FCCDFEBAFC4A" bold="true" box="[199,315,871,897]" pageId="5" pageNumber="461">Remarks.</emphasis>
|
||
The natural color of the larva is altered, with maculation presumed absent. All setae are covered by numerous minute droplets of water. The larva was found in a relatively small piece of amber, so it might be preserved in resin from various parts of the amber tree (
|
||
<emphasis id="B929EACAD60EFFB71E7AFC05FCAEFC03" box="[781,815,943,968]" italics="true" pageId="5" pageNumber="461">e.g</emphasis>
|
||
., any part of the trunk or stout twigs) (
|
||
<bibRefCitation id="EFCC4B29D60EFFB71993FC05FF55FC26" author="Perkovsky, E. E. & Rasnitsyn, A. P. & Vlaskin, A. P. & Rasnitsyn, S. P." pageId="5" pageNumber="461" pagination="293 - 301" refId="ref7531" refString="Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Rasnitsyn, S. P. (2012) Contribution to the study of the structure of amber forest communities based on analysis of syninclusions in the Rovno Amber (Late Eocene of Ukraine). Paleontological Journal, 46, 293 - 301. https: // doi. org / 10.1134 / S 0031030112030136" type="journal article" year="2012">
|
||
Perkovsky
|
||
<emphasis id="B929EACAD60EFFB71813FC05FA1CFC02" box="[1380,1437,943,969]" italics="true" pageId="5" pageNumber="461">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). The presence of stellate hairs indicates that this resin was exposed to the open air.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |