408 lines
56 KiB
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408 lines
56 KiB
XML
<document id="16039352A98E32BDCEC0A4C33B5C9B1B" ID-CLB-Dataset="3948" ID-DOI="10.5852/ejt.2021.775.1551" ID-GBIF-Dataset="cc10c087-bec3-4947-81d7-b3e5580a5c94" ID-ISSN="2118-9773" ID-Zenodo-Dep="5589689" ID-ZooBank="976D7020-5904-4951-97CE-B4FE58DA12A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634822039012" checkinUser="felipe" docAuthor="González-Alvarado, Arturo & Vaz-De, Fernando Z." docDate="2021" docId="03F48795DC5FFFB4A6C9A50FFBACFE80" docLanguage="en" docName="ejt-775_86-106.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum gilli Gonzalez-Alvarado & Vaz-de-Mello 2021" docType="treatment" docVersion="8" lastPageNumber="93" masterDocId="FFCDFFEDDC5DFFB3A45AA753FFCDFFC7" masterDocTitle="Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)" masterLastPageNumber="106" masterPageNumber="86" pageNumber="88" updateTime="1699464201688" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="2438AA000D5C528691D7AAA7CF6C2210">
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<mods:title id="2B52F0BF3EA214CCADFB13F80B46ADEE">Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)</mods:title>
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<mods:name id="F093ADFD6A3E315EA3CC07C0A8BB3548" type="personal">
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<mods:roleTerm id="4D5A88BA568BEE4E3EBB61998E9CF33C">Author</mods:roleTerm>
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<mods:namePart id="E82B7BE807DDAD7A511443234D74C5A3">González-Alvarado, Arturo</mods:namePart>
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<mods:affiliation id="58AE12B0BE4A24E77C257038EEA12179">Programa de Pós-graduação em Ecologia e Conservação da Biodiversidade, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="7FEC6F406D212D34EE3CD054B6060306" type="email">fagakorn@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="577BA1196739D76ED62BB44F34DCC98B">Author</mods:roleTerm>
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<mods:namePart id="6F0D3EF4BD1F84B258DA86ECC9B1CF96">Vaz-De, Fernando Z.</mods:namePart>
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<mods:affiliation id="B7BE6E8330B9D10C317E2E0E571951AA">Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="1EE21314026C94D32CF1CA9DF311E73E" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="F0E505BB07F224694FD3654161747827">European Journal of Taxonomy</mods:title>
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<mods:part id="63273995E3281CAC322BC63FB0701228">
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<mods:date id="EE0313625281289C9CC2219FA5CD7242">2021</mods:date>
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<mods:number id="46F2B45D6C6AC4125507DA4AADDFAF86">2021-10-20</mods:number>
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<mods:number id="41AB901BC04A19FD1375A4EF41F58446">775</mods:number>
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<treatment id="03F48795DC5FFFB4A6C9A50FFBACFE80" ID-DOI="http://doi.org/10.5281/zenodo.5590367" ID-GBIF-Taxon="189151487" ID-Zenodo-Dep="5590367" LSID="urn:lsid:plazi:treatment:03F48795DC5FFFB4A6C9A50FFBACFE80" httpUri="http://treatment.plazi.org/id/03F48795DC5FFFB4A6C9A50FFBACFE80" lastPageId="7" lastPageNumber="93" pageId="2" pageNumber="88">
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<subSubSection id="C3476508DC5FFFB1A6C9A50FFC6DFDB0" box="[659,928,604,631]" pageId="2" pageNumber="88" type="nomenclature">
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<paragraph id="8BE23683DC5FFFB1A6C9A50FFC6DFDB0" blockId="2.[659,928,604,631]" box="[659,928,604,631]" pageId="2" pageNumber="88">
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<heading id="D0AA81EFDC5FFFB1A6C9A50FFC6DFDB0" box="[659,928,604,631]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="88" reason="2">
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The
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<taxonomicName id="4C5D4D00DC5FFFB1A692A50FFD35FDB0" ID-CoL="7VN7Q" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[712,760,604,631]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="88" phylum="Arthropoda" rank="species" species="gilli">
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<emphasis id="B929EA91DC5FFFB1A692A50FFD35FDB0" bold="true" box="[712,760,604,631]" italics="true" pageId="2" pageNumber="88">gilli</emphasis>
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</taxonomicName>
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species group
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476508DC5FFFB0A4E7A52CFC4EFD9C" lastPageId="3" lastPageNumber="89" pageId="2" pageNumber="88" type="description">
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<paragraph id="8BE23683DC5FFFB1A4E7A52CFE83FD5E" blockId="2.[189,334,639,665]" box="[189,334,639,665]" pageId="2" pageNumber="88">
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<heading id="D0AA81EFDC5FFFB1A4E7A52CFE83FD5E" bold="true" box="[189,334,639,665]" fontSize="11" level="3" pageId="2" pageNumber="88" reason="3">
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<emphasis id="B929EA91DC5FFFB1A4E7A52CFE83FD5E" bold="true" box="[189,334,639,665]" pageId="2" pageNumber="88">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE23683DC5FFFB1A4E7A5FCFB50FD0E" blockId="2.[189,1181,686,713]" box="[189,1181,686,713]" pageId="2" pageNumber="88">
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BODY (
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<figureCitation id="13662A06DC5FFFB1A555A5FDFE9EFD0F" box="[271,339,686,713]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="2" pageNumber="88">Fig. 1</figureCitation>
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). Medium-sized species, length
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<quantity id="4CA59B66DC5FFFB1A68BA5FDFC81FD0E" box="[721,844,686,713]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="8.4" metricValueMin="8.0" pageId="2" pageNumber="88" unit="mm" value="8.2" valueMax="8.4" valueMin="8.0">8–8.4 mm</quantity>
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, humeral width
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<quantity id="4CA59B66DC5FFFB1A053A5FDFB54FD0E" box="[1033,1177,686,713]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="2" pageNumber="88" unit="mm" value="5.15" valueMax="5.2" valueMin="5.1">5.1–5.2 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE23683DC5FFFB1A4E7A5A3FE4FFCB4" blockId="2.[189,1400,752,883]" pageId="2" pageNumber="88">
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HEAD (
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<figureCitation id="13662A06DC5FFFB1A54AA5A3FE95FCCD" box="[272,344,752,778]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="2" pageNumber="88">Fig. 2</figureCitation>
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). Eyes large, inter-ocular distance seven to nine times width of one eye. Clypeal median emargination broadly u-shaped. Clypeal teeth separated approximately by 1.5 times the basal width of a tooth. Anterior margin of the clypeus, between clypeal teeth, concave and expanded posteriorly into triangular shape.
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</paragraph>
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<paragraph id="8BE23683DC5FFFB1A4E7A4C8FDC4FB85" blockId="2.[189,1400,923,1090]" pageId="2" pageNumber="88">
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PRONOTUM (
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<figureCitation id="13662A06DC5FFFB1A513A4C8FE5DFC72" box="[329,400,923,949]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="2" pageNumber="88">Fig. 3</figureCitation>
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). Margin between anterior and medial angle subconcave. Medial angle of pronotum rounded to projected. Punctures almost with the same size, basal punctures only slightly larger than discal punctures. Pronotal disc with shiny points well-defined (
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<figureCitation id="13662A06DC5FFFB1A7C0A4B2FC3DFC3C" box="[922,1008,993,1019]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="2" pageNumber="88">Fig. 3F</figureCitation>
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, arrows), separated between each other and separated or contiguous to punctures; or irregular (
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<figureCitation id="13662A06DC5FFFB1A7CEA357FC3EFBD9" box="[916,1011,1028,1054]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="2" pageNumber="88">Fig. 3G</figureCitation>
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, arrows), contiguous between to each other and to punctures.
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</paragraph>
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<paragraph id="8BE23683DC5FFFB1A4E7A33AFE51FA5A" blockId="2.[187,1400,1129,1437]" pageId="2" pageNumber="88">
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ELYTRA (
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<figureCitation id="13662A06DC5FFFB1A57FA33AFE40FB43" box="[293,397,1129,1156]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1892,1918]" captionStartId-1="7.[189,232,1892,1918]" captionTargetBox-0="[219,1369,314,1798]" captionTargetBox-1="[201,1386,717,1836]" captionTargetId-0="figure-77@6.[219,1369,314,1798]" captionTargetId-1="figure-170@7.[201,1386,717,1836]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." captionText-1="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi-0="http://doi.org/10.5281/zenodo.5589701" figureDoi-1="http://doi.org/10.5281/zenodo.5589705" httpUri-0="https://zenodo.org/record/5589701/files/figure.png" httpUri-1="https://zenodo.org/record/5589705/files/figure.png" pageId="2" pageNumber="88">Fig. 4–5</figureCitation>
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). Interstriae with shiny points mixed with the punctures. Interstriae VI and VII with basal carina almost identical in size. Carina of ninth interstria reaching middle of elytral length or slightly surpassing it. Striae I–VIII inconspicuous including apically, only in some parts striae slightly visible and very narrow or striae I–VII conspicuous and narrow, width of third stria
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<emphasis id="B929EA91DC5FFFB1A064A381FBADFB2B" bold="true" box="[1086,1120,1234,1260]" pageId="2" pageNumber="88">1/30</emphasis>
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<superScript id="7C289BCBDC5FFFB1A03BA381FBA3FB27" attach="left" box="[1121,1134,1234,1248]" fontSize="6" pageId="2" pageNumber="88">th</superScript>
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or
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<emphasis id="B929EA91DC5FFFB1A0CEA381FB7AFB2B" bold="true" box="[1172,1207,1234,1260]" pageId="2" pageNumber="88">1/40</emphasis>
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<superScript id="7C289BCBDC5FFFB1A0EDA381FB09FB27" attach="left" box="[1207,1220,1234,1248]" fontSize="6" pageId="2" pageNumber="88">th</superScript>
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of the distance between stria II and III. If conspicuous, first stria widest and either striae III–VII subequal in width or striae III–VII ill-defined, successively narrower and more ill-defined, with VII almost inconspicuous, and stria VIII conspicuous apical and laterally or only laterally; in both discontinuous in some parts and reaching the apex of carina of the ninth interstria. Apex of elytra with tubercles on interstriae III–VII or III, V–VII (
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<figureCitation id="13662A06DC5FFFB1A51FA2D1FE40FA5A" box="[325,397,1410,1437]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="2" pageNumber="88">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE23683DC5FFFB1A4E7A297FE86F949" blockId="2.[188,1399,1476,1678]" pageId="2" pageNumber="88">
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METAVENTRITE (
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<figureCitation id="13662A06DC5FFFB1A528A297FE75FA18" box="[370,440,1476,1503]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="2" pageNumber="88">Fig. 6</figureCitation>
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). With weak posterior excavation, occupying approximately the metaventral basal fourth. Disc with conspicuous punctures at 8 × magnification. Disc punctures at least twice smaller than punctures on anterior-lateral area of metaventral process and dispersed, separated at least by three diameters of a puncture. Punctures on anterior-lateral area of metaventral process separated by less than one diameter. Anterior-medial area of metaventral process with few and smaller punctures than anteriorlateral ones.
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</paragraph>
|
||
<paragraph id="8BE23683DC5FFFB1A4E7A1E5FDC6F935" blockId="2.[188,1398,1717,1779]" pageId="2" pageNumber="88">LEGS. Ventral surface of protibia with a weak carina. Posterior edge of metafemur with two margins, one dorsal and other one ventral.</paragraph>
|
||
<paragraph id="8BE23683DC5FFFB0A4E7A048FE66FE77" blockId="2.[188,1400,1818,2021]" lastBlockId="3.[188,1399,265,432]" lastPageId="3" lastPageNumber="89" pageId="2" pageNumber="88">
|
||
MALE. Protibial spur broad and foliaceus. Mesofemur modified, with slight sinuosity on apical third.Apex of mesotibia wider than female and on ventral-internal margin with a small or large spatulate expansion. Metatrochanter modified or not, if modified with an expansion on distal third. Metafemur with steep tapering on basal third, with expansion on posterior-ventral margin, before that steep tapering. Internal margin of metatibia with small or large tubercles. Ventrite I expanded posteriorly, expansion reaching from the middle of ventrite IV to almost the distal margin of ventrite V; width of expansion on ventrite III variable, narrower to wider than distance between clypeal teeth. Genitalia (
|
||
<figureCitation id="13662A06DC5EFFB0A025A65AFB22FEE4" box="[1151,1263,265,291]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,232,1822,1848]" captionStartId-1="10.[189,232,1857,1883]" captionTargetBox-0="[206,1383,744,1716]" captionTargetBox-1="[233,1362,650,1777]" captionTargetId-0="figure-193@9.[203,1383,744,1716]" captionTargetId-1="figure-181@10.[218,1370,644,1787]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." captionText-1="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi-0="http://doi.org/10.5281/zenodo.5589711" figureDoi-1="http://doi.org/10.5281/zenodo.5589713" httpUri-0="https://zenodo.org/record/5589711/files/figure.png" httpUri-1="https://zenodo.org/record/5589713/files/figure.png" pageId="3" pageNumber="89">Figs 7–8</figureCitation>
|
||
). Paramera subtriangular, with dorsal and ventral edges straight in lateral view. Apex of paramera rounded in dorsal view. Paramera with short and thin apical-dorsal notch (
|
||
<figureCitation id="13662A06DC5EFFB0A73CA61CFC7CFEAD" box="[870,945,335,362]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="3" pageNumber="89">Fig. 7</figureCitation>
|
||
, arrows). Medial area of endophallus with one endophallite (
|
||
<figureCitation id="13662A06DC5EFFB0A590A621FDDDFE4A" box="[458,528,370,397]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="3" pageNumber="89">Fig. 8</figureCitation>
|
||
). Basal circular shape endophallite with ring very thin and handle strongly broadened medially.
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC5EFFB0A4E7A68BFC4EFD9C" blockId="3.[189,1399,471,603]" pageId="3" pageNumber="89">
|
||
FEMALE (
|
||
<figureCitation id="13662A06DC5EFFB0A573A684FEBEFE36" box="[297,371,471,498]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[268,1315,559,1863]" captionTargetId="figure-129@12.[268,1320,553,1873]" captionTargetPageId="12" captionText="Fig. 9. Habitus A, C and ventral view B, D of the only two females known of the gilli species group. A–B. Paratype of Deltochilum gilli González-Alvarado & Vaz-de-Mello, 2021. C–D. Paratype D. jocelynae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589715" httpUri="https://zenodo.org/record/5589715/files/figure.png" pageId="3" pageNumber="89">Fig. 9</figureCitation>
|
||
). Protibial spur thinner than that of male and spiniform. Apex of mesotibia narrower than that of male. Metatrochanter not modified. Meso- and metafemur not modified. Internal margin of metatibia without tubercles. Metasternal disc with posterior excavation smaller than male. Ventrite V as wide as ventrite VI medially. Ventrite VI narrowed medially.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF22660BDC5EFFB0A4E7A037FAB9F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589691" ID-Zenodo-Dep="5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="3" pageNumber="89" startId="3.[189,232,1892,1918]" targetBox="[222,1362,786,1773]" targetPageId="3">
|
||
<paragraph id="8BE23683DC5EFFB0A4E7A037FAB9F82E" blockId="3.[189,1399,1892,2025]" pageId="3" pageNumber="89">
|
||
<emphasis id="B929EA91DC5EFFB0A4E7A037FEC2F8B8" bold="true" box="[189,271,1892,1919]" pageId="3" pageNumber="89">Fig. 1.</emphasis>
|
||
Habitus of members of the
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A635A036FD53F8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[623,670,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC5EFFB0A635A036FD53F8B8" box="[623,670,1893,1919]" italics="true" pageId="3" pageNumber="89">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B929EA91DC5EFFB0A73BA037FCBBF8B9" bold="true" box="[865,886,1892,1918]" pageId="3" pageNumber="89">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A07FA036FB3EF8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1061,1267,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC5EFFB0A07FA036FB3EF8B8" box="[1061,1267,1893,1919]" italics="true" pageId="3" pageNumber="89">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888.
|
||
<emphasis id="B929EA91DC5EFFB0A75FA0DBFCDAF865" bold="true" box="[773,791,1928,1954]" pageId="3" pageNumber="89">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A7E5A0DAFB98F865" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[959,1109,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5EFFB0A7E5A0DAFB98F865" box="[959,1109,1928,1954]" italics="true" pageId="3" pageNumber="89">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5EFFB0A007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="3" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CNCI 379889.
|
||
<emphasis id="B929EA91DC5EFFB0A4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="3" pageNumber="89">C</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A524A0FFFDF8F802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5EFFB0A524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="3" pageNumber="89">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5EFFB0A5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="3" pageNumber="89">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5EFFB0A61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="3" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041746.
|
||
<emphasis id="B929EA91DC5EFFB0A7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="3" pageNumber="89">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A0F6A0FFFABBF802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5EFFB0A0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="3" pageNumber="89">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5EFFB0A089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="3" pageNumber="89">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5EFFB0A4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="3" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041744.
|
||
<emphasis id="B929EA91DC5EFFB0A606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="3" pageNumber="89">E</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5EFFB0A748A09CFC19F82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[786,980,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="89" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5EFFB0A748A09CFCEAF82F" box="[786,807,1999,2024]" italics="true" pageId="3" pageNumber="89">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5EFFB0A76CA09CFC19F82F" box="[822,980,1999,2024]" italics="true" pageId="3" pageNumber="89">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5EFFB0A786A09CFBF4F82E" box="[988,1081,1999,2025]" pageId="3" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041745.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3476508DC59FFB7A4E7A65BFC61FEB2" pageId="4" pageNumber="90" type="reference_group">
|
||
<paragraph id="8BE23683DC59FFB7A4E7A65BFE90FEE5" blockId="4.[189,349,264,290]" box="[189,349,264,290]" pageId="4" pageNumber="90">
|
||
<heading id="D0AA81EFDC59FFB7A4E7A65BFE90FEE5" bold="true" box="[189,349,264,290]" fontSize="11" level="3" pageId="4" pageNumber="90" reason="3">
|
||
<emphasis id="B929EA91DC59FFB7A4E7A65BFE90FEE5" bold="true" box="[189,349,264,290]" pageId="4" pageNumber="90">Composition</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC59FFB7A4E7A664FC61FEB2" blockId="4.[189,1398,311,373]" pageId="4" pageNumber="90">
|
||
<taxonomicName id="4C5D4D00DC59FFB7A4E7A664FC54FE96" ID-CoL="7VN7Q" authority="Gonzalez-Alvarado & Vaz-de-Mello, 2021" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[189,921,311,337]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC59FFB7A4E7A664FE46FE96" box="[189,395,311,337]" italics="true" pageId="4" pageNumber="90">Deltochilum gilli</emphasis>
|
||
<bibRefCitation id="EFCC4B72DC59FFB7A5CCA664FC54FE96" author="Gonzalez-Alvarado A. & Vaz-de-Mello F. Z." box="[406,921,311,337]" pageId="4" pageNumber="90" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. & Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado & Vaz-de-Mello, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5D4D00DC59FFB7A7F0A66BFB89FE96" ID-CoL="8SSPY" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[938,1092,311,338]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A7F0A66BFC72FE96" box="[938,959,312,337]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A788A664FB89FE96" box="[978,1092,311,337]" italics="true" pageId="4" pageNumber="90">jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A015A66BFB7DFE95" box="[1103,1200,312,338]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5D4D00DC59FFB7A0E5A66BFABBFE96" ID-CoL="8SSP2" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1215,1398,312,338]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A0E5A66BFB19FE96" box="[1215,1236,312,337]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A0BDA66BFABBFE96" box="[1255,1398,312,337]" italics="true" pageId="4" pageNumber="90">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A4E7A608FED7FEB2" box="[189,282,347,373]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5D4D00DC59FFB7A57CA608FE20FEB3" ID-CoL="8SSNC" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[294,493,347,373]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A57CA608FEF6FEB3" box="[294,315,347,372]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A510A608FE20FEB3" box="[330,493,347,372]" italics="true" pageId="4" pageNumber="90">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A5AEA608FD82FEB2" box="[500,591,347,373]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5D4D00DC59FFB7A6D2A608FC87FEB3" ID-CoL="8SSQ6" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[648,842,347,373]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A6D2A608FD50FEB3" box="[648,669,347,372]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A6F6A608FC87FEB3" box="[684,842,347,372]" italics="true" pageId="4" pageNumber="90">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A70BA608FC61FEB2" box="[849,940,347,373]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF22660BDC59FFB7A4E7A037FAB8F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589695" ID-Zenodo-Dep="5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="4" pageNumber="90" startId="4.[189,232,1892,1918]" targetBox="[237,1351,525,1826]" targetPageId="4">
|
||
<paragraph id="8BE23683DC59FFB7A4E7A037FAB8F82E" blockId="4.[189,1399,1892,2025]" pageId="4" pageNumber="90">
|
||
<emphasis id="B929EA91DC59FFB7A4E7A037FEDCF8B8" bold="true" box="[189,273,1892,1919]" pageId="4" pageNumber="90">Fig. 2.</emphasis>
|
||
Head of members of the
|
||
<taxonomicName id="4C5D4D00DC59FFB7A605A036FD42F8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[607,655,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC59FFB7A605A036FD42F8B8" box="[607,655,1893,1919]" italics="true" pageId="4" pageNumber="90">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B929EA91DC59FFB7A70DA037FCA1F8B9" bold="true" box="[855,876,1892,1918]" pageId="4" pageNumber="90">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC59FFB7A07BA036FB3CF8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1057,1265,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC59FFB7A07BA036FB3CF8B8" box="[1057,1265,1893,1919]" italics="true" pageId="4" pageNumber="90">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888.
|
||
<emphasis id="B929EA91DC59FFB7A75FA0DBFCD5F865" bold="true" box="[773,792,1928,1954]" pageId="4" pageNumber="90">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC59FFB7A7E5A0DAFB98F865" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[959,1109,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A7E5A0DAFB98F865" box="[959,1109,1928,1954]" italics="true" pageId="4" pageNumber="90">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CNCI 379889.
|
||
<emphasis id="B929EA91DC59FFB7A4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="4" pageNumber="90">C</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC59FFB7A524A0FFFDF8F802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="4" pageNumber="90">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041746.
|
||
<emphasis id="B929EA91DC59FFB7A7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="4" pageNumber="90">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC59FFB7A0F6A0FFFABBF802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="4" pageNumber="90">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041744.
|
||
<emphasis id="B929EA91DC59FFB7A606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="4" pageNumber="90">E</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC59FFB7A748A09CFC18F82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[786,981,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="90" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC59FFB7A748A09CFCEAF82F" box="[786,807,1999,2024]" italics="true" pageId="4" pageNumber="90">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC59FFB7A76DA09CFC18F82F" box="[823,981,1999,2024]" italics="true" pageId="4" pageNumber="90">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC59FFB7A787A09CFBF7F82E" box="[989,1082,1999,2025]" pageId="4" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041745.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3476508DC58FFB4A4E7A65AFBACFE80" lastPageId="7" lastPageNumber="93" pageId="5" pageNumber="91" type="discussion">
|
||
<paragraph id="8BE23683DC58FFB6A4E7A65AFEE2FEE4" blockId="5.[189,303,265,291]" box="[189,303,265,291]" pageId="5" pageNumber="91">
|
||
<heading id="D0AA81EFDC58FFB6A4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="5" pageNumber="91" reason="3">
|
||
<emphasis id="B929EA91DC58FFB6A4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" pageId="5" pageNumber="91">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE23683DC58FFB6A4E7A664FAB5FEB3" blockId="5.[189,1400,311,373]" pageId="5" pageNumber="91">
|
||
This species group is only known from few (eight) specimens and their biology is poorly known. The specimens were collected above
|
||
<quantity id="4CA59B66DC58FFB6A664A609FD44FEB2" box="[574,649,346,373]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="91" unit="m" value="680.0">680 m</quantity>
|
||
a.s.l. and the highest record is
|
||
<quantity id="4CA59B66DC58FFB6A7AEA609FB86FEB2" box="[1012,1099,346,373]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="5" pageNumber="91" unit="m" value="1411.0">1411 m</quantity>
|
||
a.s.l. (
|
||
<specimenCount id="9D5BFD0ADC58FFB6A0C3A609FAF6FEB3" box="[1177,1339,346,372]" pageId="5" pageNumber="91" type="generic">one specimen</specimenCount>
|
||
does
|
||
</paragraph>
|
||
<caption id="DF22660BDC58FFB6A4E7A04DFBD7F82F" ID-DOI="http://doi.org/10.5281/zenodo.5589697" ID-Zenodo-Dep="5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="5" pageNumber="91" startId="5.[189,232,1822,1848]" targetBox="[217,1370,471,1774]" targetPageId="5">
|
||
<paragraph id="8BE23683DC58FFB6A4E7A04DFBD7F82F" blockId="5.[188,1400,1822,2025]" pageId="5" pageNumber="91">
|
||
<emphasis id="B929EA91DC58FFB6A4E7A04DFEC3F8FF" bold="true" box="[189,270,1822,1848]" pageId="5" pageNumber="91">Fig. 3.</emphasis>
|
||
Pronotum of members of the
|
||
<taxonomicName id="4C5D4D00DC58FFB6A624A04DFD63F8FF" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[638,686,1822,1848]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC58FFB6A624A04DFD63F8FF" box="[638,686,1822,1848]" italics="true" pageId="5" pageNumber="91">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B929EA91DC58FFB6A731A04DFCB3F8FF" bold="true" box="[875,894,1822,1848]" pageId="5" pageNumber="91">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A070A04DFB39F8FF" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1066,1268,1822,1848]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC58FFB6A070A04DFB39F8FF" box="[1066,1268,1822,1848]" italics="true" pageId="5" pageNumber="91">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888.
|
||
<emphasis id="B929EA91DC58FFB6A75FA011FCD5F89B" bold="true" box="[773,792,1858,1884]" pageId="5" pageNumber="91">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A7E5A011FB9BF89C" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[959,1110,1857,1883]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A7E5A011FB9BF89C" box="[959,1110,1857,1883]" italics="true" pageId="5" pageNumber="91">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A007A011FB71F89B" box="[1117,1212,1858,1884]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CNCI 379889.
|
||
<emphasis id="B929EA91DC58FFB6A4E4A037FF1FF8B9" bold="true" box="[190,210,1892,1918]" pageId="5" pageNumber="91">C</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A524A035FDF8F8B8" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[382,565,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A524A035FE5EF8B8" box="[382,403,1894,1919]" italics="true" pageId="5" pageNumber="91">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC58FFB6A5FCA035FDF8F8B8" box="[422,565,1894,1919]" italics="true" pageId="5" pageNumber="91">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A61AA036FD6DF8B8" box="[576,672,1893,1919]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041746.
|
||
<emphasis id="B929EA91DC58FFB6A7B1A036FBCDF8B8" bold="true" box="[1003,1024,1893,1919]" pageId="5" pageNumber="91">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A0F6A035FABBF8B8" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1196,1398,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A0F6A035FB0CF8B8" box="[1196,1217,1894,1919]" italics="true" pageId="5" pageNumber="91">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC58FFB6A089A035FABBF8B8" box="[1235,1398,1894,1919]" italics="true" pageId="5" pageNumber="91">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A4E7A0DBFED6F865" box="[189,283,1928,1954]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041744
|
||
<emphasis id="B929EA91DC58FFB6A603A0DBFDA1F865" bold="true" box="[601,620,1928,1954]" pageId="5" pageNumber="91">E</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A748A0DAFC19F865" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[786,980,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A748A0DAFCEAF865" box="[786,807,1929,1954]" italics="true" pageId="5" pageNumber="91">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC58FFB6A76CA0DAFC19F865" box="[822,980,1929,1954]" italics="true" pageId="5" pageNumber="91">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A787A0DBFBF6F865" box="[989,1083,1928,1954]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041745.
|
||
<emphasis id="B929EA91DC58FFB6A4E4A0F8FF02F802" bold="true" box="[190,207,1963,1989]" pageId="5" pageNumber="91">F</emphasis>
|
||
. Miscrosculpture of pronotal disc, Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A6AEA0FFFC44F802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[756,905,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A6AEA0FFFC44F802" box="[756,905,1963,1989]" italics="true" pageId="5" pageNumber="91">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A7CAA0FFFC27F801" box="[912,1002,1964,1990]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B929EA91DC58FFB6A7B4A0F8FBC8F802" bold="true" box="[1006,1029,1963,1989]" pageId="5" pageNumber="91">G</emphasis>
|
||
. Miscrosculpture pronotal disc, Holotype of
|
||
<taxonomicName id="4C5D4D00DC58FFB6A50BA09CFDC9F82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[337,516,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC58FFB6A50BA09CFEABF82F" box="[337,358,1999,2024]" italics="true" pageId="5" pageNumber="91">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC58FFB6A52FA09CFDC9F82F" box="[373,516,1999,2024]" italics="true" pageId="5" pageNumber="91">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC58FFB6A656A09CFDAAF82E" box="[524,615,1999,2025]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
White arrows show the shiny points.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF22660BDC5BFFB5A4E7A037FAB8F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589701" ID-Zenodo-Dep="5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="6" pageNumber="92" startId="6.[189,232,1892,1918]" targetBox="[219,1369,314,1798]" targetPageId="6">
|
||
<paragraph id="8BE23683DC5BFFB5A4E7A037FAB8F82E" blockId="6.[189,1399,1892,2025]" pageId="6" pageNumber="92">
|
||
<emphasis id="B929EA91DC5BFFB5A4E7A037FEDCF8B8" bold="true" box="[189,273,1892,1919]" pageId="6" pageNumber="92">Fig. 4.</emphasis>
|
||
Elytra of members of the
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A63EA036FD59F8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[612,660,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC5BFFB5A63EA036FD59F8B8" box="[612,660,1893,1919]" italics="true" pageId="6" pageNumber="92">gilli</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B929EA91DC5BFFB5A700A037FCA2F8B9" bold="true" box="[858,879,1892,1918]" pageId="6" pageNumber="92">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A078A036FB3CF8B8" authorityName="Gonzalez-Alvarado & Vaz-de-Mello" authorityYear="2021" box="[1058,1265,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="gilli">
|
||
<emphasis id="B929EA91DC5BFFB5A078A036FB3CF8B8" box="[1058,1265,1893,1919]" italics="true" pageId="6" pageNumber="92">Deltochilum gilli</emphasis>
|
||
</taxonomicName>
|
||
González- Alvarado & Vaz-de-Mello, 2021, CNCI 379888.
|
||
<emphasis id="B929EA91DC5BFFB5A75FA0DBFCDAF865" bold="true" box="[773,791,1928,1954]" pageId="6" pageNumber="92">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A7E5A0DAFB9BF865" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[959,1110,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5BFFB5A7E5A0DAFB9BF865" box="[959,1110,1928,1954]" italics="true" pageId="6" pageNumber="92">D. jocelynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5BFFB5A007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="6" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CNCI 379889.
|
||
<emphasis id="B929EA91DC5BFFB5A4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="6" pageNumber="92">C</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A524A0FFFDF8F802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5BFFB5A524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="6" pageNumber="92">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5BFFB5A5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="6" pageNumber="92">nonstriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5BFFB5A61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="6" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041746.
|
||
<emphasis id="B929EA91DC5BFFB5A7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="6" pageNumber="92">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A0F6A0FFFABBF802" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5BFFB5A0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="6" pageNumber="92">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5BFFB5A089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="6" pageNumber="92">quasistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5BFFB5A4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="6" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041744.
|
||
<emphasis id="B929EA91DC5BFFB5A606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="6" pageNumber="92">E</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C5D4D00DC5BFFB5A748A09CFC18F82F" authority="González-Alvarado & Vaz-De, 2021" authorityName="González-Alvarado & Vaz-De" authorityYear="2021" box="[786,981,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
|
||
<emphasis id="B929EA91DC5BFFB5A748A09CFCEAF82F" box="[786,807,1999,2024]" italics="true" pageId="6" pageNumber="92">D</emphasis>
|
||
.
|
||
<emphasis id="B929EA91DC5BFFB5A76DA09CFC18F82F" box="[823,981,1999,2024]" italics="true" pageId="6" pageNumber="92">tenuistriatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A57EADC5BFFB5A787A09CFBF7F82E" box="[989,1082,1999,2025]" pageId="6" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, CMNEN WSD00041745.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE23683DC5AFFB4A4E7A65AFBACFE80" blockId="7.[189,1398,265,327]" pageId="7" pageNumber="93">
|
||
not have altitude information). They were collected using pitfall traps baited with human dung (
|
||
<specimenCount id="9D5BFD0ADC5AFFB4A112A65AFEF1FE80" pageId="7" pageNumber="93" type="generic">five specimens</specimenCount>
|
||
), carrion (
|
||
<specimenCount id="9D5BFD0ADC5AFFB4A5E8A67FFD9BFE80" box="[434,598,300,327]" pageId="7" pageNumber="93" type="generic">one specimen</specimenCount>
|
||
) as well as via window trap (
|
||
<specimenCount id="9D5BFD0ADC5AFFB4A7F7A67FFB99FE80" box="[941,1108,300,327]" pageId="7" pageNumber="93" type="generic">one specimen</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |