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<mods:title id="2FFB01786379609C212F3015CE3D6625">Systematics of the family Carditidae (Bivalvia: Archiheterodonta) in the Cenozoic of Argentina</mods:title>
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<paragraph id="8BE2369DB802B469FF6875B4FD4DF831" blockId="5.[151,573,1941,1967]" box="[151,573,1941,1967]" pageId="5" pageNumber="56">
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<emphasis id="B929EA8FB802B469FF6875B4FD4DF831" bold="true" box="[151,573,1941,1967]" pageId="5" pageNumber="56">
Family
<taxonomicName id="4C5D4D1EB802B469FF0975B4FD4DF831" authority="Ferussac, 1822" authorityName="Ferussac" authorityYear="1822" box="[246,573,1941,1967]" class="Bivalvia" family="Carditidae" kingdom="Animalia" order="Carditoida" pageId="5" pageNumber="56" phylum="Mollusca" rank="family">
Carditidae
<bibRefCitation id="EFCC4B6CB802B469FE7A75B4FD4DF831" author="Ferussac" box="[389,573,1941,1967]" pageId="5" pageNumber="56" refString="Ferussac, A. E. de (1822) Tableaux systematiques des animaux mollusques. A. Bertrand, Paris, 111 pp." type="book" year="1822">Férussac, 1822</bibRefCitation>
</taxonomicName>
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<paragraph id="8BE2369DB802B46AFF6875FDFC60FEFA" blockId="5.[151,1436,2012,2037]" lastBlockId="6.[151,1436,151,536]" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
<emphasis id="B929EA8FB802B469FF6875FDFE7BF86B" bold="true" box="[151,267,2012,2037]" pageId="5" pageNumber="56">Remarks.</emphasis>
Some authors only recognize a few genera such as
<taxonomicName id="4C5D4D1EB802B469FCB475FDFBFDF86B" authority="Lamarck, 1801" authorityName="Lamarck" authorityYear="1801" box="[843,1165,2012,2037]" class="Bivalvia" family="Carditidae" genus="Venericardia" higherTaxonomySource="CoL" kingdom="Animalia" order="Carditida" pageId="5" pageNumber="56" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB802B469FCB475FDFCA8F86B" box="[843,984,2012,2037]" italics="true" pageId="5" pageNumber="56">Venericardia</emphasis>
<bibRefCitation id="EFCC4B6CB802B469FC1D75FDFBFDF86B" author="Lamarck" box="[994,1165,2012,2037]" pageId="5" pageNumber="56" refString="Lamarck, J. B. P. A. de M., C. de (1801) Systeme des animaux sans vertebres ou tableau general des classes, des orders et des genres de ces animaux. L'Auteur, Paris, 432 pp." type="book" year="1801">Lamarck, 1801</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5D4D1EB802B46AFB3775FDFFA2FF2E" authority="Bruguiere, 1792" authorityName="Bruguiere" authorityYear="1792" class="Bivalvia" family="Carditidae" genus="Cardita" kingdom="Animalia" lastPageId="6" lastPageNumber="57" order="Carditoida" pageId="5" pageNumber="56" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB802B469FB3775FDFA6EF86B" box="[1224,1310,2012,2037]" italics="true" pageId="5" pageNumber="56">Cardita</emphasis>
<bibRefCitation id="EFCC4B6CB802B46AFADA75FDFFA2FF2E" author="Bruguiere" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56" refString="Bruguiere, J. G. (1792) Encyclopedie methodique. Tableau encyclopedique et methodique des trois regnes de la nature. Vers, coquilles, mollusques et polypiers. Chez Panckoucke, Imprimeur-Libraire, hotel de Thou, rue des Poitevins, Paris, 180 pp." type="book" year="1792">Bruguière, 1792</bibRefCitation>
</taxonomicName>
, and a large number of subgenera (e.g.
<taxonomicName id="4C5D4D1EB801B46AFD6072B6FB03FF2E" authority="Stenzel &amp; Krause, 1957" authorityName="Stenzel &amp; Krause" authorityYear="1957" box="[671,1139,151,176]" class="Bivalvia" family="Carditidae" genus="Claibornicardia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB801B46AFD6072B6FC24FF2E" box="[671,852,151,176]" italics="true" pageId="6" pageNumber="57">Claibornicardia</emphasis>
<bibRefCitation id="EFCC4B6CB801B46AFCA272B6FB03FF2E" author="Stenzel" box="[861,1139,151,176]" pageId="6" pageNumber="57" refString="Stenzel, H. B. &amp; Krause, E. K. (1957) In: Stenzel, H. B., Krause, E. K. &amp; Twining, S. T. Pelecypoda from the type locality of the Stone City beds (Middle Eocene) of Texas. Texas University Publication, 5704, 1 - 237." type="journal article" year="1957">Stenzel &amp; Krause, 1957</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D4D1EB801B46AFB7E72B8FAE8FF2E" authority="Stewart, 1930" authorityName="Stewart" authorityYear="1930" box="[1153,1432,152,176]" class="Bivalvia" family="Carditidae" genus="Venericor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB801B46AFB7E72B8FB9BFF2E" box="[1153,1259,153,176]" italics="true" pageId="6" pageNumber="57">Venericor</emphasis>
<bibRefCitation id="EFCC4B6CB801B46AFB0872B9FAE8FF2E" author="Stewart" box="[1271,1432,152,176]" pageId="6" pageNumber="57" refString="Stewart, R. B. (1930) Gabb's California Cretaceous and Tertiary type Lamellibranchs. Special Publication of the Academy of Natural Sciences of Philadelphia, 3, 1 - 314." type="journal article" year="1930">Stewart, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5D4D1EB801B46AFF68729DFECAFF4A" authority="Sacco, 1899" authorityName="Sacco" authorityYear="1899" box="[151,442,188,213]" class="Bivalvia" family="Carditidae" genus="Megacardita" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB801B46AFF68729DFE57FF4B" box="[151,295,188,213]" italics="true" pageId="6" pageNumber="57">Megacardita</emphasis>
<bibRefCitation id="EFCC4B6CB801B46AFED2729DFECAFF4A" author="Sacco" box="[301,442,188,212]" pageId="6" pageNumber="57" refString="Sacco, F. (1899) I Molluschi dei terreni terziarii del Piemonti e della Liguria. Parte XXVII (Unionidae, Carditidae, Astartidae, Crassatellidae, Lasaeidae, Galeommidae, Cardiidae, Limnocardiidae e Chamidae). Carlo Clausen, Torino, 102 pp." type="book" year="1899">Sacco, 1899</bibRefCitation>
</taxonomicName>
); however, some of those taxa are ranked as genera by other authors (for comparisons of different approaches see
<bibRefCitation id="EFCC4B6CB801B46AFE5072C1FD45FF66" author="Sacco" box="[431,565,224,248]" pageId="6" pageNumber="57" refString="Sacco, F. (1899) I Molluschi dei terreni terziarii del Piemonti e della Liguria. Parte XXVII (Unionidae, Carditidae, Astartidae, Crassatellidae, Lasaeidae, Galeommidae, Cardiidae, Limnocardiidae e Chamidae). Carlo Clausen, Torino, 102 pp." type="book" year="1899">Sacco 1899</bibRefCitation>
;
<bibRefCitation id="EFCC4B6CB801B46AFDBF72FEFDC3FF66" author="Dall" box="[576,691,223,248]" pageId="6" pageNumber="57" refString="Dall, W. H. (1903) Contributions to the Tertiary fauna of Florida with special reference to the Silex Beds of the Caloosahatchie River. Part IV. Transactions of the Wagner Free Institute of Science of Philadelphia, 3 (4), 1219 - 1654." type="journal article" year="1903">Dall 1903</bibRefCitation>
;
<bibRefCitation id="EFCC4B6CB801B46AFD4072FEFC27FF66" author="Chavan" box="[703,855,223,248]" pageId="6" pageNumber="57" refString="Chavan, A. (1969) Superfamily Carditacea Fleming. 1920. In: Moore, R. C. (Ed.), Treatise on Invertebrate Paleontology. Part N. Mollusca 6. Bivalvia 2. Geological Society of America, University of Kansas, Lawrence, pp. 491 - 518. [pp. N 491 - N 518]" type="journal article" year="1969">Chavan 1969</bibRefCitation>
;
<bibRefCitation id="EFCC4B6CB801B46AFC9B72C1FC81FF66" author="Moore" box="[868,1009,224,248]" pageId="6" pageNumber="57" refString="Moore, E. J. (1992) Tertiary marine pelecypods of California and Baja California: Erycinidae through Carditidae. United States Geological Survey, Professional Paper, 1228 - E, 1 - 37." type="book chapter" year="1992">Moore 1992</bibRefCitation>
,
<bibRefCitation id="EFCC4B6CB801B46AFC0372FEFA30FF66" author="McClure" box="[1020,1344,223,248]" pageId="6" pageNumber="57" refString="McClure, K. &amp; Lockwood, R. (2015) Relationships among Venericardia (Bivalvia: Carditidae) on the U. S. Coastal Plain during the Paleogene. Journal of Paleontology, 89 (3), 522 - 531. https: // doi. org / 10.1017 / jpa. 2015.23" type="journal article" year="2015">McClure &amp; Lockwood 2015</bibRefCitation>
). As the use of subgenera implies closer phylogenetic relationship within each group than between these groups (
<bibRefCitation id="EFCC4B6CB801B46AFAB67325FFA3FEDE" author="Dubois" pageId="6" pageNumber="57" refString="Dubois, A. (1982) Les notiens de genre, sous-genre et groupe d'especes en zoologie a la lumiere de la systematique evolutive. Italian Journal of Zoology, 16, 9 - 65." type="journal article" year="1982">Dubois 1982</bibRefCitation>
), we will regard all supraspecific groups as genera, discarding
<emphasis id="B929EA8FB801B46AFC567308FB71FEDE" box="[937,1025,297,320]" italics="true" pageId="6" pageNumber="57">a priori</emphasis>
any phylogenetic inferences until a thorough phylogenetic study of the family is carried out.
</paragraph>
<paragraph id="8BE2369DB801B46AFF38734EFB55FD86" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="57">
<bibRefCitation id="EFCC4B6CB801B46AFF38734EFE04FE16" author="Chavan" box="[199,372,367,392]" pageId="6" pageNumber="57" refString="Chavan, A. (1969) Superfamily Carditacea Fleming. 1920. In: Moore, R. C. (Ed.), Treatise on Invertebrate Paleontology. Part N. Mollusca 6. Bivalvia 2. Geological Society of America, University of Kansas, Lawrence, pp. 491 - 518. [pp. N 491 - N 518]" type="journal article" year="1969">Chavan (1969)</bibRefCitation>
outlined a classification with seven subfamilies mainly based on shell outline and hinge morphology. As a result, members of the venericardids of
<bibRefCitation id="EFCC4B6CB801B46AFCC973B5FCA2FE32" author="Harris" box="[822,978,404,429]" pageId="6" pageNumber="57" refString="Harris, G. D. (1919) Pelecypoda of the Saint Maurice and Claiborne Stages. Bulletin of the American Paleontology, 6 (31), 1 - 260." type="journal article" year="1919">Harris (1919)</bibRefCitation>
,
<bibRefCitation id="EFCC4B6CB801B46AFC1F73B5FA7BFE32" author="Gardner" box="[992,1291,404,429]" pageId="6" pageNumber="57" refString="Gardner, J. &amp; Bowles, E. (1939) The Venericardia planicosta Group in the Gulf Province. United States Geological Survey, Professional Paper, 189 - F, 143 - 215." type="book chapter" year="1939">Gardner &amp; Bowles (1939)</bibRefCitation>
and
<bibRefCitation id="EFCC4B6CB801B46AFAB973B5FF91FE4E" author="Heaslip" pageId="6" pageNumber="57" refString="Heaslip, W. G. (1968) Cenozoic evolution of the alticostate venericards in Gulf and East Coastal North America. Palaeontographica Americana, 6 (39), 55 - 135." type="journal article" year="1968">Heaslip (1968)</bibRefCitation>
were indistinctly included in the subfamilies
<taxonomicName id="4C5D4D1EB801B46AFD1F7396FCE0FE4E" authorityName="Chavan" authorityYear="1969" box="[736,912,439,464]" class="Bivalvia" family="Carditidae" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="subFamily" subFamily="Carditamerinae">Carditamerinae</taxonomicName>
,
<taxonomicName id="4C5D4D1EB801B46AFC647396FB3BFE4E" authorityName="Chavan" authorityYear="1969" box="[923,1099,439,464]" class="Bivalvia" family="Carditidae" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="subFamily" subFamily="Venericardiinae">Venericardiinae</taxonomicName>
or
<taxonomicName id="4C5D4D1EB801B46AFB8F7396FA70FE4E" authorityName="Chavan" authorityYear="1969" box="[1136,1280,439,464]" class="Bivalvia" family="Carditidae" kingdom="Animalia" order="Carditoida" pageId="6" pageNumber="57" phylum="Mollusca" rank="subFamily" subFamily="Carditesinae">Carditesinae</taxonomicName>
. In this paper, we reject Chavans (1969) approach because it ignores the phylogenetic proposals by
<bibRefCitation id="EFCC4B6CB801B46AFB8D73FDFA7CFE6A" author="Harris" box="[1138,1292,476,501]" pageId="6" pageNumber="57" refString="Harris, G. D. (1919) Pelecypoda of the Saint Maurice and Claiborne Stages. Bulletin of the American Paleontology, 6 (31), 1 - 260." type="journal article" year="1919">Harris (1919)</bibRefCitation>
and
<bibRefCitation id="EFCC4B6CB801B46AFAB873FDFF95FD86" author="Heaslip" pageId="6" pageNumber="57" refString="Heaslip, W. G. (1968) Cenozoic evolution of the alticostate venericards in Gulf and East Coastal North America. Palaeontographica Americana, 6 (39), 55 - 135." type="journal article" year="1968">Heaslip (1968)</bibRefCitation>
. A comparison among genera discussed in the text is presented in Table 2.
</paragraph>
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