treatments-xml/data/10/D5/9B/10D59B32B7EC564FAEB367B515C5B0D0.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1130.87730" ID-GBIF-Dataset="0591791c-72cb-46dc-bc36-363e2ebc81f5" ID-PMC="PMC9836693" ID-Pensoft-Pub="1313-2970-1130-79" ID-Pensoft-UUID="C525797A12CE5FF6AF35DD3628B5F2B2" ID-PubMed="36761014" ID-ZooBank="57F636A8D488446F9210CBFA33A78998" ModsDocID="1313-2970-1130-79" checkinTime="1668727489642" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Ruiz, Gustavo R. S., Ng, Paul Y. C., Vishnudas, Ettukandathil Haridas &amp; Sudhikumar, Ambalaparambil V." docDate="2022" docId="10D59B32B7EC564FAEB367B515C5B0D0" docLanguage="en" docName="ZooKeys 1130: 79-102" docOrigin="ZooKeys 1130" docPubDate="2022-11-17" docSource="http://dx.doi.org/10.3897/zookeys.1130.87730" docTitle="Kelawakaju Maddison &amp; Ruiz 2022, gen. nov." docType="treatment" docUuid="1A91FAF6-5C6F-4AAB-A770-2395DE6CCAF3" docUuidSource="ZooBank" docVersion="4" id="C525797A12CE5FF6AF35DD3628B5F2B2" lastPageNumber="79" masterDocId="C525797A12CE5FF6AF35DD3628B5F2B2" masterDocTitle="Kelawakaju gen. nov., a new Asian lineage of marpissine jumping spiders (Araneae, Salticidae, Marpissina)" masterLastPageNumber="102" masterPageNumber="79" pageNumber="79" updateTime="1732486332407" updateUser="ExternalLinkService">
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<mods:title id="AFEE9C7A3F4B61861EB4C88A13DEC6AE">Kelawakaju gen. nov., a new Asian lineage of marpissine jumping spiders (Araneae, Salticidae, Marpissina)</mods:title>
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<mods:namePart id="663562281022F5453823042B8B742250">Maddison, Wayne P.</mods:namePart>
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<mods:affiliation id="690484546CD1D05EBA5BC6EC54FC4605">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
<mods:nameIdentifier id="35961D5BCE0511419F57CED560D5A0DD" type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
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<mods:namePart id="35385012EE87EDB5E11E98FC1E484B3A">Ruiz, Gustavo R. S.</mods:namePart>
<mods:affiliation id="A2922E5BE8AD61BF7794F68D30A963B8">Instituto de Cie ̂ ncias Biolo ́ gicas, Universidade Federal do Para ́, Rua Augusto Corre ̂ a, 01, CEP 66075 - 110, Bele ́ m, PA, Brazil</mods:affiliation>
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<mods:namePart id="1A16290849D54BEAEECBD0EB2FD73BAA">Ng, Paul Y. C.</mods:namePart>
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<mods:namePart id="BCDF5A3887AEFA79D1F711E88AB8EB41">Vishnudas, Ettukandathil Haridas</mods:namePart>
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<mods:affiliation id="8872C73C7241A2AC4640A5EDAAFEE144">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, 680 125, India</mods:affiliation>
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<mods:namePart id="E71F7B3C829C8982B7D4262395829A67">Sudhikumar, Ambalaparambil V.</mods:namePart>
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<mods:affiliation id="EB1C5374329AD0F54399048DD72386B2">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, 680 125, India</mods:affiliation>
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<mods:date id="A5AFBDBBBFC98642CBB414995CCD5CA3">2022</mods:date>
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<treatment id="10D59B32B7EC564FAEB367B515C5B0D0" ID-GBIF-Taxon="205097765" LSID="urn:lsid:zoobank.org:act:1A91FAF6-5C6F-4AAB-A770-2395DE6CCAF3" httpUri="http://treatment.plazi.org/id/10D59B32B7EC564FAEB367B515C5B0D0" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Salticidae" scope_order="Araneae">
<subSubSection id="0A4BA17103640B1200CDFA7B88135724" pageId="0" pageNumber="79" type="nomenclature">
<paragraph id="692621D69F667612227A0978D954B871" pageId="0" pageNumber="79">
<taxonomicName id="6094F00E61706CAF805AE991227029D7" ID-CoL="C7L93" ID-ENA="3042499" LSID="https://zoobank.org/1A91FAF6-5C6F-4AAB-A770-2395DE6CCAF3" authority="Maddison &amp; Ruiz" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus" status="gen. nov.">Kelawakaju Maddison &amp; Ruiz</taxonomicName>
<taxonomicNameLabel id="22BC4DC4B13103FFB5CDC3D61DACB350" pageId="0" pageNumber="79">gen. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="9072820C26ABFF24CCA2003FC1E391B8" pageId="0" pageNumber="79">Type species.</paragraph>
<paragraph id="6E028E461BBBAAE52DC6ADD649C76484" pageId="0" pageNumber="79">
<taxonomicName id="B6E1A53E2EA251F37AC37180F9AA828F" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="B05064F82DF8F83365F1E02BB3303DC3" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
Maddison &amp; Ruiz, sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="3DE7C6D3D4FA8DB894F56B8FD462037E" pageId="0" pageNumber="79" type="species included">
<paragraph id="602DF9B10F89E6FCC0272E73D656B5BF" pageId="0" pageNumber="79">Species included.</paragraph>
<paragraph id="DFA8BCDB52DAEF70088CC9731F08CAFF" pageId="0" pageNumber="79">
<taxonomicName id="A8F3B7A8D00BC77EC642F46568024943" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="95531B8C43FF9844DB2086F94D0C4C96" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
species group:
</paragraph>
<paragraph id="63777074893C869E7BE33AFF0727B438" pageId="0" pageNumber="79">
<taxonomicName id="A4941046060171D88CE8834577F88352" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="A579EF9C95D2DF0EAC67E5628CDD54B8" italics="true" pageId="0" pageNumber="79">Kelawakaju mulu</emphasis>
</taxonomicName>
Maddison &amp; Ruiz, sp. nov.
</paragraph>
<paragraph id="C03B42877102F43AE3DE898C1AB02532" pageId="0" pageNumber="79">
<taxonomicName id="DC67F965E9C357CC71837E488FFCACF5" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju intexta" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="intexta">
<emphasis id="411CC2141A291C9BA654EDDEBB02EBCA" italics="true" pageId="0" pageNumber="79">Kelawakaju intexta</emphasis>
</taxonomicName>
Maddison &amp; Ruiz, sp. nov.
</paragraph>
<paragraph id="DB4156C2FBD958C8EFF30235A6B59171" pageId="0" pageNumber="79">
<taxonomicName id="0074D63F7C9326CBFE53FA12D9C27CC5" class="Arachnida" kingdom="Animalia" lsidName="K. singapura" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="5C064BC8C946F68802F04452D67BCE6D" italics="true" pageId="0" pageNumber="79">K. singapura</emphasis>
</taxonomicName>
species group:
</paragraph>
<paragraph id="6BEC12C70A6AB0FFC2ACC5480A2834FC" pageId="0" pageNumber="79">
<taxonomicName id="522444A9FE5E67A882728C64DB1114E3" authorityName="Maddison &amp; Ng" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju singapura" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="69FFCAC3E5802FAC211226A9FD2B639A" italics="true" pageId="0" pageNumber="79">Kelawakaju singapura</emphasis>
</taxonomicName>
Maddison &amp; Ng, sp. nov.
</paragraph>
<paragraph id="51AF0AE5330932C37D912B1218E74720" pageId="0" pageNumber="79">
<taxonomicName id="3A05B1F299EBE321DFCE9B153F645DF9" class="Arachnida" kingdom="Animalia" lsidName="K. frenata" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="F812DFE53FF0ECB7C426F6422266718E" italics="true" pageId="0" pageNumber="79">K. frenata</emphasis>
</taxonomicName>
species group:
</paragraph>
<paragraph id="E8F122CA13FB583CF218718B7B0BDC25" pageId="0" pageNumber="79">
<taxonomicName id="A2A23FC7EFFDA74CF38B2E5F5264EC31" authorityName="Maddison &amp; Ruiz &amp; Ng &amp; Vishnudas &amp; Sudhikumar" authorityYear="2022" baseAuthorityName="Simon" baseAuthorityYear="1901" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju frenata" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="BD36886E36C33D20BF083F55865EF9F8" italics="true" pageId="0" pageNumber="79">Kelawakaju frenata</emphasis>
</taxonomicName>
(Simon, 1901)
</paragraph>
<paragraph id="1DB43453500996C00285AEBD20324658" pageId="0" pageNumber="79">
<taxonomicName id="53E04DEEB089B227E85FECBBDB6A2D05" authorityName="Maddison &amp; Ng" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju leucomelas" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="leucomelas">
<emphasis id="1B70D2DFF01F157A33411B646E7BF136" italics="true" pageId="0" pageNumber="79">Kelawakaju leucomelas</emphasis>
</taxonomicName>
Maddison &amp; Ng, sp. nov.
</paragraph>
<paragraph id="6E2E210819453BBAA51679FFB7E4C71C" pageId="0" pageNumber="79">
<taxonomicName id="4D6F9FCA068251D379976D07B3BC2D1B" authorityName="Vishnudas, Maddison, &amp; Sudhikumar" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju sahyadri" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="sahyadri">
<emphasis id="A5E688F181CCAC729D12CA155FF94C64" italics="true" pageId="0" pageNumber="79">Kelawakaju sahyadri</emphasis>
</taxonomicName>
Vishnudas, Maddison, &amp; Sudhikumar, sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="4AA5ECF6FB47CDB42AE7DC61816248E5" pageId="0" pageNumber="79" type="etymology">
<paragraph id="A36FCAA4EB0F580B3C1F020467599287" pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph id="AA47DEDB37E5BF4E2CDC39AFA23CBD90" pageId="0" pageNumber="79">
The name means tree spider in the Berawan language from the area of Long Terawan, Sarawak (
<emphasis id="033741AD8E4D5AFE760768C8D1331FC2" italics="true" pageId="0" pageNumber="79">kelawak</emphasis>
= spider; 
<emphasis id="5B710EAFDECE75126A964896659F71BE" italics="true" pageId="0" pageNumber="79">kaju</emphasis>
or 
<emphasis id="434163DA6B3E9C2A635EDEE0AE73AB4E" italics="true" pageId="0" pageNumber="79">kajuh</emphasis>
= tree; Syria Lejau Malang, pers. comm.), where the first specimens of 
<taxonomicName id="EA3C2DC080F054A9CFE5B611C0AF205D" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="B08E8088037B9CCF07B5C97EC878B52D" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
were found. To be treated grammatically as feminine.
</paragraph>
</subSubSection>
<subSubSection id="3FAAD3E78198DCE4741C7B3E05FAE2AE" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="96F69DDDC68B1D1844F79D28CC5D7532" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="9A1D996DDEC9040E16A9955EEFE3ABC9" pageId="0" pageNumber="79">
Elongate and flat-bodied salticids, unusual among marpissines for the elongated or enlarged male chelicerae. Retrolateral tibial apophysis of palp long, blade-like, more or less straight and parallel to axis of palp. Embolus relatively short among marpissines, arising more or less terminally on the bulb (9-12 
<normalizedToken id="AF0CD830DD5FEA47E04EBCD40BBC345D" originalValue="o’clock">o'clock</normalizedToken>
in ventral view of left palp). Markings cryptic on tree trunks, either mottled or with low-contrast longitudinal bands.
</paragraph>
</subSubSection>
<subSubSection id="B0C95FC683872B715F7686AF88F75846" pageId="0" pageNumber="79" type="description">
<paragraph id="67D0BE6B2B993BE7C9597A86ABF1711B" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="8F9C9B93262CD1DFC4B2CA926C72CCDC" pageId="0" pageNumber="79">
<emphasis id="2BFF6521B1587A9F045F48F3430F0092" bold="true" italics="true" pageId="0" pageNumber="79">Carapace</emphasis>
flat, narrower (
<taxonomicName id="A6B52D7389DBCCA2DE839622C0EA9E9C" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="7D65ACD5EEB187DDA6DB4E05C84B37F6" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
group, Figs 
<figureCitation id="CC6D3357E65DA266344917C0CDCBAA4E" captionStart="Figures 10–15" captionStartId="F3" captionText="Figures 10 - 15. Carapaces of Kelawakaju males 10 K. mulu holotype 11 K. intexta holotype 12 K. singapura holotype 13 K. frenata from Guangxi, Dongxing 14 K. leucomelas holotype 15 K. sahyadri holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures10-15" httpUri="https://binary.pensoft.net/fig/771008" pageId="0" pageNumber="79">10</figureCitation>
, 
<figureCitation id="F1CC196FCE5431423912497B62EB29AD" captionStart="Figures 10–15" captionStartId="F3" captionText="Figures 10 - 15. Carapaces of Kelawakaju males 10 K. mulu holotype 11 K. intexta holotype 12 K. singapura holotype 13 K. frenata from Guangxi, Dongxing 14 K. leucomelas holotype 15 K. sahyadri holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures10-15" httpUri="https://binary.pensoft.net/fig/771008" pageId="0" pageNumber="79">11</figureCitation>
) to broader (
<taxonomicName id="39B2552297CB23B1A9737BEE4BC570BE" class="Arachnida" kingdom="Animalia" lsidName="K. leucomelas" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="leucomelas">
<emphasis id="839CE94E0F32CEECCA4BB5066D3561E8" italics="true" pageId="0" pageNumber="79">K. leucomelas</emphasis>
</taxonomicName>
, Fig. 
<figureCitation id="72C5EDDA8428DBFD08CAED4E88206E11" captionStart="Figures 10–15" captionStartId="F3" captionText="Figures 10 - 15. Carapaces of Kelawakaju males 10 K. mulu holotype 11 K. intexta holotype 12 K. singapura holotype 13 K. frenata from Guangxi, Dongxing 14 K. leucomelas holotype 15 K. sahyadri holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures10-15" httpUri="https://binary.pensoft.net/fig/771008" pageId="0" pageNumber="79">14</figureCitation>
). Lower part of the thorax in some species with 1-3 distinct narrow vertical lines of pale scales (
<taxonomicName id="BC6D6E6D0AD5DE6141B91F355B2218CF" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="F21B1ABC9639BC552677F39B2BC5FDF9" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
: Fig. 
<figureCitation id="F55EB6D0762EC811D7774BBE57CA50E2" captionStart="Figures 23–28" captionStartId="F5" captionText="Figures 23 - 28. Kelawakaju mulu 23 - 25 holotype male SWK 12 - 2610 26 - 28 paratype female SWK 12 - 2639. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures23-28" httpUri="https://binary.pensoft.net/fig/771010" pageId="0" pageNumber="79">27</figureCitation>
; 
<taxonomicName id="37409A38CE9FE0FF6190226427222040" class="Arachnida" kingdom="Animalia" lsidName="K. intexta" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="intexta">
<emphasis id="F035CFCBAA6E849E3063E9FECF9E9A70" italics="true" pageId="0" pageNumber="79">K. intexta</emphasis>
</taxonomicName>
: Fig. 
<figureCitation id="54E3F218BC2BADE5475CC4A069C2C7A0" captionStart="Figures 29–34" captionStartId="F6" captionText="Figures 29 - 34. Kelawakaju intexta 29 - 31 holotype male SWK 12 - 3752 32 - 34 paratype male SWK 12 - 0523. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures29-34" httpUri="https://binary.pensoft.net/fig/771011" pageId="0" pageNumber="79">32</figureCitation>
; 
<taxonomicName id="D0B30CD96E0AEBA5EBCB13713B4014A7" class="Arachnida" kingdom="Animalia" lsidName="K. singapura" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="singapura">
<emphasis id="76D3A911AA18C9BD39B7D533A8ED4866" italics="true" pageId="0" pageNumber="79">K. singapura</emphasis>
</taxonomicName>
: Figs 
<figureCitation id="EA410FD7F30560570DB264E4AC77ABBA" captionStart="Figures 35–45" captionStartId="F7" captionText="Figures 35 - 45. Kelawakaju singapura 35 holotype male JK. 21.08.02.001 embolus, ventral 36 paratype male JK. 90.10.21.0002 palp, ventral 37 same, retrolateral 38 paratype female AS 19.0023 epigyne, ventral 39 same, vulva, dorsal 40 - 42 holotype male 43 paratype female JK. 21.05.14.0001 44, 45 paratype female AS 19.0023. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures35-45" httpUri="https://binary.pensoft.net/fig/771012" pageId="0" pageNumber="79">43</figureCitation>
, 
<figureCitation id="3C02F75F9CE4D8ECA0DAC82A923EFE81" captionStart="Figures 35–45" captionStartId="F7" captionText="Figures 35 - 45. Kelawakaju singapura 35 holotype male JK. 21.08.02.001 embolus, ventral 36 paratype male JK. 90.10.21.0002 palp, ventral 37 same, retrolateral 38 paratype female AS 19.0023 epigyne, ventral 39 same, vulva, dorsal 40 - 42 holotype male 43 paratype female JK. 21.05.14.0001 44, 45 paratype female AS 19.0023. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures35-45" httpUri="https://binary.pensoft.net/fig/771012" pageId="0" pageNumber="79">45</figureCitation>
), resembling similar stripes in the baviine 
<taxonomicName id="0F2D284BE8E92D2EA8A2E9A82B6966F8" authorityName="Thorell" authorityYear="1895" class="Arachnida" family="Salticidae" genus="Piranthus" kingdom="Animalia" lsidName="Piranthus" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="EF9F3BEE5B8293DD72C708D4302CE2EA" italics="true" pageId="0" pageNumber="79">Piranthus</emphasis>
</taxonomicName>
Thorell, 1895 (
<bibRefCitation id="3D99289F875FFD5A13778EA570DC4E0A" DOI="https://doi.org/10.3897/zookeys.1004.57526" author="Maddison, WP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="79" pagination="27 - 97" refId="B15" refString="Maddison, WP, Beattie, I, Marathe, K, Ng, PYC, Kanesharatnam, N, Benjamin, SP, Kunte, K, 2020. A phylogenetic and taxonomic review of baviine jumping spiders (Araneae: Salticidae: Baviini). ZooKeys 1004: 27 - 97, DOI: https://doi.org/10.3897/zookeys.1004.57526" title="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae: Salticidae: Baviini)." url="https://doi.org/10.3897/zookeys.1004.57526" volume="1004" year="2020">Maddison et al. 2020</bibRefCitation>
: fig. 263) and the gophoine 
<taxonomicName id="430F37D10236B0A58AC79422005B33C5" authorityName="Simon" authorityYear="1900" class="Arachnida" family="Salticidae" genus="Cotinusa" kingdom="Animalia" lsidName="Cotinusa" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="EA9C81D930861A4C4CE3621BBC6A147D" italics="true" pageId="0" pageNumber="79">Cotinusa</emphasis>
</taxonomicName>
Simon, 1900 (
<bibRefCitation id="99974144CFE44D8E153C819ACB18BA35" DOI="https://doi.org/10.1080/21658005.2016.1199502" author="Rubio, GD" journalOrPublisher="Zoology and Ecology" pageId="0" pageNumber="79" pagination="265 - 274" refId="B17" refString="Rubio, GD, Baigorria, JE, 2016. New species and the first known male of the jumping spider Ceriomura Simon (Araneae: Salticidae: Gophoini) with the faunistic revision of gophoines from Misiones, Argentina. Zoology and Ecology 26 (4): 265 - 274, DOI: https://doi.org/10.1080/21658005.2016.1199502" title="New species and the first known male of the jumping spider Ceriomura Simon (Araneae: Salticidae: Gophoini) with the faunistic revision of gophoines from Misiones, Argentina." url="https://doi.org/10.1080/21658005.2016.1199502" volume="26" year="2016">Rubio and Baigorria 2016</bibRefCitation>
). 
<emphasis id="EEA5253789FCF3B789963C682B2948CB" bold="true" italics="true" pageId="0" pageNumber="79">Chelicerae</emphasis>
with seta-bearing tubercles on paturon of males and some females (Figs 
<figureCitation id="913394F28E5E605AEF37AC2E4004D3CD" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">2-4</figureCitation>
). Males of all but two species have narrow stripes of white scales on the front face of the chelicerae, forming an inverted V (Figs 
<figureCitation id="5F9A7FECA21576AA8D2A14A8F684FFF9" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">3</figureCitation>
, 
<figureCitation id="E2DC12B7F6245891777AB851086D2CA9" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">4</figureCitation>
, 
<figureCitation id="3BC89F507D9258B2B47F20556151AD69" captionStart="Figures 73–78" captionStartId="F9" captionText="Figures 73 - 78. Kelawakaju frenata 73, 74 male from Tai Tam County Park, Hong Kong (© 2020 Artur Tomaszek) 75 female from Guangdong, Gaotan Town 76 male from Guangxi, Dongxing, dorsal 77 female from Dongxing, dorsal 78 same, ventral. Specimen in 73, 74 not examined microscopically; inferred as K. frenata by appearance and locality." figureDoi="10.3897/zookeys.1130.87730.figures73-78" httpUri="https://binary.pensoft.net/fig/771014" pageId="0" pageNumber="79">74</figureCitation>
, 
<figureCitation id="E5739B8060D121517A9A862F5C6E0932" captionStart="Figures 79–81" captionStartId="F10" captionText="Figures 79 - 81. Kelawakaju leucomelas 79 - 80 male from the type locality 81 female from same locality. Photographs © Chris Ang 2021. Specimens not examined microscopically; inferred as K. leucomelas by appearance and locality." figureDoi="10.3897/zookeys.1130.87730.figures79-81" httpUri="https://binary.pensoft.net/fig/771015" pageId="0" pageNumber="79">79</figureCitation>
, 
<figureCitation id="71AD6EC52CCA69C74337BA1989E172E2" captionStart="Figures 82–89" captionStartId="F11" captionText="Figures 82 - 89. Kelawakaju sahyadri 82 - 85 male holotype AS 19.4895 86 - 88 female paratype AS 19.4934 89 female paratype from Kerala." figureDoi="10.3897/zookeys.1130.87730.figures82-89" httpUri="https://binary.pensoft.net/fig/771016" pageId="0" pageNumber="79">83</figureCitation>
). Two promarginal teeth and one retromarginal tooth (sometimes with a second cusp, Fig. 
<figureCitation id="B7105BEFE0B70C488E0242EDC999664F" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">6</figureCitation>
). 
<emphasis id="0BB05354230F5896381E30530DFC1FC0" bold="true" italics="true" pageId="0" pageNumber="79">
<normalizedToken id="B85C791843FE25B44CE623E903CE452C" originalValue="Palp’s">Palp's</normalizedToken>
</emphasis>
RTA a long blade. Embolus appears freely movable, separate from functional tegulum. Cymbium modified at ventral-retrolateral-proximal corner (e.g., Figs 
<figureCitation id="C8300A35A109886538D591A311B63EA6" captionStart="Figures 16–22" captionStartId="F4" captionText="Figures 16 - 22. Kelawakaju mulu species group, genitalia 16 - 20 K. mulu 16 holotype male palp, ventral 17 same, retrolateral 18 paratype female SWK 12 - 2639 epigyne, ventral 19 same, vulva, dorsal 20 same, ventral 21, 22 K. intexta holotype male palp 21 retrolateral 22 ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures16-22" httpUri="https://binary.pensoft.net/fig/771009" pageId="0" pageNumber="79">17</figureCitation>
, 
<figureCitation id="7AE9D039B724E5F68D4D9ADBBB46B728" captionStart="Figures 16–22" captionStartId="F4" captionText="Figures 16 - 22. Kelawakaju mulu species group, genitalia 16 - 20 K. mulu 16 holotype male palp, ventral 17 same, retrolateral 18 paratype female SWK 12 - 2639 epigyne, ventral 19 same, vulva, dorsal 20 same, ventral 21, 22 K. intexta holotype male palp 21 retrolateral 22 ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures16-22" httpUri="https://binary.pensoft.net/fig/771009" pageId="0" pageNumber="79">21</figureCitation>
, 
<figureCitation id="E4FF6F9EBCF7DC4B5D4FF3DF7BBDB83F" captionStart="Figures 46–72" captionStartId="F8" captionText="Figures 46 - 72. Kelawakaju frenata species group, genitalia 46 - 54 K. frenata 46 male from Dongxing, palp, ventral 47 same, retrolateral 48 same, dorsal 49 male from Hong Kong, embolus, ventral 50 Simon's (1901 a) figure, reversed 51 female d 224 from Dongxing, epigyne, ventral 52 second female from Dongxing, epigyne, ventral 53 same, vulva, ventral 54 same, dorsal 55 - 63 K. leucomelas 55 holotype male palp, ventral 56 same, retrolateral 57 same, embolus, oblique 58 paratype male JK 13.12.10.0001, embolus, oblique 59 male JK. 19.08.18.0010, embolus, oblique 60 same, palp, retrolateral 61 same, ventral 62 female paratype JK. 20.11.13.0001, epigyne, ventral 63 same, vulva, dorsal 64 - 72 K. sahyadri 64 holotype male, palp, ventral 65 same, retrolateral 66 same, embolus, oblique 67 paratype male from Kerala, embolus, oblique 68 same, ventral 69 same, palp, ventral 70 same, retrolateral 71 paratype female AS 19.4934 epigyne, ventral 72 same vulva, dorsal. Oblique views of embolus are between ventral and prolateral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures46-72" httpUri="https://binary.pensoft.net/fig/771013" pageId="0" pageNumber="79">47</figureCitation>
). 
<emphasis id="3E0C2AE8734905E1DBEC2677D35243F4" bold="true" italics="true" pageId="0" pageNumber="79">Abdomen</emphasis>
long and narrow.
</paragraph>
<paragraph id="5214AE96B1C99A154AE87E3DE41C6EAA" pageId="0" pageNumber="79">We recognize three species groups in the genus.</paragraph>
</subSubSection>
<subSubSection id="955CB805FF9BB4762D32B52C6B5937FB" pageId="0" pageNumber="79" type="kelawakaju mulu species group">
<paragraph id="D406C28085992319F16A7CC59714872B" pageId="0" pageNumber="79">
<taxonomicName id="C565F095C22E452EAB9E81BE5CAFAE1F" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="5630AA4BB20371DDF639107E0C7328CC" italics="true" pageId="0" pageNumber="79">Kelawakaju mulu</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph id="A754DC7AF87F8FC3B32FFD2864BDD384" pageId="0" pageNumber="79">
The 
<taxonomicName id="CA475778B4543C222E2A308F1B23A2D5" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="F109AE2D263D37904A3D90969C57FD80" italics="true" pageId="0" pageNumber="79">Kelawakaju mulu</emphasis>
</taxonomicName>
species group includes 
<taxonomicName id="1BC4248EB974752C5D34D987A41089AF" class="Arachnida" kingdom="Animalia" lsidName="K. mulu" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="mulu">
<emphasis id="0FAF1AA5208181FB4B6595853B0CB141" italics="true" pageId="0" pageNumber="79">K. mulu</emphasis>
</taxonomicName>
, 
<taxonomicName id="680631165D57D89F05F2358A9B8DDC61" class="Arachnida" kingdom="Animalia" lsidName="K. intexta" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="intexta">
<emphasis id="E2AD9A765557C357DE4ECCDDA83DA125" italics="true" pageId="0" pageNumber="79">K. intexta</emphasis>
</taxonomicName>
, and a third as-yet-undescribed species from Singapore. They are smaller-bodied than other 
<taxonomicName id="4BD1A510EE91680C96756FD31B307CFE" authorityName="Maddison &amp; Ruiz" authorityYear="2022" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="772E72274D32B766EA933F8056ABB011" italics="true" pageId="0" pageNumber="79">Kelawakaju</emphasis>
</taxonomicName>
, with mottled markings, and narrow chelicerae that project forward in the male. The embolus is narrow and forms a smooth curve bending toward the retrolateral. The lower part of the thorax has three vertical stripes of pale scales on each side. Epigynal openings are delicate and the edges difficult to discern (Fig. 
<figureCitation id="303620906D78A6563B067946FA9F6338" captionStart="Figures 16–22" captionStartId="F4" captionText="Figures 16 - 22. Kelawakaju mulu species group, genitalia 16 - 20 K. mulu 16 holotype male palp, ventral 17 same, retrolateral 18 paratype female SWK 12 - 2639 epigyne, ventral 19 same, vulva, dorsal 20 same, ventral 21, 22 K. intexta holotype male palp 21 retrolateral 22 ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1130.87730.figures16-22" httpUri="https://binary.pensoft.net/fig/771009" pageId="0" pageNumber="79">18</figureCitation>
). Retromarginal tooth of chelicera with small second cusp basally (Figs 
<figureCitation id="C85C8E369BCD06746AFEE6E0DD46A4DD" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">5</figureCitation>
, 
<figureCitation id="77FB25501D9A26A3F4B81699FA3C3369" captionStart="Figures 2–9" captionStartId="F2" captionText="Figures 2 - 9. Chelicerae of Kelawakaju species 2 - 4 oblique view with carapace 5 - 9 ventral view 2 K. mulu male holotype 3 K. intexta male holotype 4 K. frenata male from Guangxi 5 K. mulu male holotype 6 K. intexta male holotype 7 K. singapura male holotype 8 K. frenata male from Guangxi 9 K. frenata female from Guangxi. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1130.87730.figures2-9" httpUri="https://binary.pensoft.net/fig/771007" pageId="0" pageNumber="79">6</figureCitation>
). Members of this group may prefer more shaded habitats than those of the 
<taxonomicName id="53934D013F1FEAEAADB0BFC2E5BB8C8B" authorityName="Maddison &amp; Ruiz &amp; Ng &amp; Vishnudas &amp; Sudhikumar" authorityYear="2022" baseAuthorityName="Simon" baseAuthorityYear="1901" class="Arachnida" family="Salticidae" genus="Kelawakaju" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelawakaju frenata" order="Araneae" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="33D8101A62143237314E5D9F651E6ADF" italics="true" pageId="0" pageNumber="79">Kelawakaju frenata</emphasis>
</taxonomicName>
group, having been found only inside forests.
</paragraph>
</subSubSection>
</treatment>
</document>