190 lines
33 KiB
XML
190 lines
33 KiB
XML
<document id="9AD5FD6A122714D5B4E20C7E86670EF0" ID-CLB-Dataset="59767" ID-DOI="10.11646/zootaxa.5319.4.2" ID-GBIF-Dataset="b0dd118c-22d4-43bd-8dd8-bac4d810d1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203248" ID-ZooBank="20272586-E197-4DEA-811F-496572F121B7" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1690883685755" checkinUser="plazi" docAuthor="Kluge, Nikita, Sivaruban, T., Srinivasan, Pandiarajan, Barathy, S. & Isack, Rajasekaran" docDate="2023" docId="03E48787FFC7E657FF64FEF688F7FAF4" docLanguage="en" docName="zootaxa.5319.4.2.pdf" docOrigin="Zootaxa 5319 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ecdyonurus Eaton 1868, s. l." docType="treatment" docVersion="4" lastPageNumber="503" masterDocId="FFDDFFFFFFC5E655FFF3FF928B39FFF6" masterDocTitle="Redescription of the subgenus Rhithrogeniella Ulmer 1939 (Ephemeroptera, Heptageniidae, genus Ecdyonurus) based on reared specimens from India and Thailand" masterLastPageNumber="523" masterPageNumber="501" pageNumber="503" updateTime="1691065623882" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AC35406542EF5D3E5CA3A28243979BF0">Redescription of the subgenus Rhithrogeniella Ulmer 1939 (Ephemeroptera, Heptageniidae, genus Ecdyonurus) based on reared specimens from India and Thailand</mods:title>
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<mods:namePart id="C6A98F1712996FBD6143972E8AD72AA3">Kluge, Nikita</mods:namePart>
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<mods:affiliation id="E1F946314513BC6004D4E19005DA2F65">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia. Website: http: // insecta. bio. spbu. ru</mods:affiliation>
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<mods:namePart id="061509DC6988819FC3DB4B018273724F">Sivaruban, T.</mods:namePart>
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<mods:affiliation id="08417E262F82502FA4B81A3752B9D4B9">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:namePart id="F80186165D9AD69345E9FA29BA0E89E8">Srinivasan, Pandiarajan</mods:namePart>
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<mods:affiliation id="8DF1FF78EB1945237EEB17E0523FB21D">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:affiliation id="8B0C37E7DCBB73793AEBB3AF13020A48">Department of Zoology, Fatima College, Madurai- 625018, India.</mods:affiliation>
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<mods:namePart id="54AC44FF7205FBF8CFBBAFDC7ED20B59">Isack, Rajasekaran</mods:namePart>
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<mods:affiliation id="A02797C0BE8D242F7CFBC8E6382D5864">PG and Research department of Zoology, the American College, Madurai- 625002, India.</mods:affiliation>
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<mods:date id="FD8871E01210337BCD135BBABF8944F4">2023</mods:date>
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<mods:number id="F6A65A0230094F840DE12144B034BA1C">2023-07-27</mods:number>
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<treatment id="03E48787FFC7E657FF64FEF688F7FAF4" ID-DOI="http://doi.org/10.5281/zenodo.8211531" ID-Zenodo-Dep="8211531" LSID="urn:lsid:plazi:treatment:03E48787FFC7E657FF64FEF688F7FAF4" httpUri="http://treatment.plazi.org/id/03E48787FFC7E657FF64FEF688F7FAF4" lastPageNumber="503" pageId="2" pageNumber="503">
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<emphasis id="B939EA83FFC7E657FF64FEF68807FE88" bold="true" box="[151,830,355,382]" pageId="2" pageNumber="503">
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Genus
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<taxonomicName id="4C4D4D12FFC7E657FF1DFEF1892FFE88" ID-CoL="632Z8" ID-ENA="309576" authority="Eaton 1868" authorityName="Eaton" authorityYear="1868" box="[238,534,355,382]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus" sensu="lato">
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<emphasis id="B939EA83FFC7E657FF1DFEF18AB8FE88" bold="true" box="[238,385,355,382]" italics="true" pageId="2" pageNumber="503">Ecdyonurus</emphasis>
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<bibRefCitation id="EFDC4B60FFC7E657FE7BFEF6892FFE88" author="Eaton, A. E." box="[392,534,356,382]" pageId="2" pageNumber="503" pagination="141 - 144" refId="ref9395" refString="Eaton, A. E. (1868) Remarks upon the homologies of the ovipositor. Transactions of the Entomological Society of London, 1868, 141 - 144. https: // doi. org / 10.1111 / j. 1365 - 2311.1868. tb 00621. x" type="journal article" year="1868">Eaton 1868</bibRefCitation>
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<taxonomicNameLabel id="A20A57F8FFC7E657FDEEFEF6897FFE88" box="[541,582,356,382]" pageId="2" pageNumber="503" sensu="lato">s. l.</taxonomicNameLabel>
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<paragraph id="8BF23691FFC7E657FDB4FEF68807FE88" blockId="2.[151,830,355,418]" box="[583,830,356,382]" pageId="2" pageNumber="503">
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, or
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<taxonomicName id="4C4D4D12FFC7E657FD84FEF68836FE88" authorityName="Eaton" authorityYear="1868" box="[631,783,356,382]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">Ecdyonurus</taxonomicName>
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/fg1
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(
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<figureCitation id="13762A14FFC7E657FF6CFE1A8A24FE54" box="[159,285,392,418]" captionStart-0="FIGURES 1–10" captionStart-1="FIGURES 11–25" captionStart-10="FIGURES 91–96" captionStart-11="FIGURES 97–102" captionStart-12="FIGURES 103–110" captionStart-13="FIGURES 111–116" captionStart-2="FIGURES 26–33" captionStart-3="FIGURES 34–41" captionStart-4="FIGURES 42–52" captionStart-5="FIGURES 53–68" captionStart-6="FIGURES 69–72" captionStart-7="FIGURES 73–76" captionStart-8="FIGURES 77–84" captionStart-9="FIGURES 85–90" captionStartId-0="3.[151,264,1810,1834]" captionStartId-1="4.[151,264,1853,1877]" captionStartId-10="17.[151,264,1968,1992]" captionStartId-11="18.[151,264,1834,1858]" captionStartId-12="19.[151,264,1692,1716]" captionStartId-13="20.[151,264,1869,1893]" captionStartId-2="5.[151,264,1773,1797]" captionStartId-3="6.[151,264,1916,1940]" captionStartId-4="7.[151,264,1785,1809]" captionStartId-5="8.[151,264,1807,1831]" captionStartId-6="9.[151,264,1837,1861]" captionStartId-7="10.[151,264,1352,1376]" captionStartId-8="14.[151,264,1532,1556]" captionStartId-9="16.[151,264,1571,1595]" captionTargetBox-0="[151,1436,180,1772]" captionTargetBox-1="[151,1435,181,1829]" captionTargetBox-10="[177,1411,381,1945]" captionTargetBox-11="[151,1436,181,1810]" captionTargetBox-12="[154,1436,345,1669]" captionTargetBox-13="[151,1436,366,1846]" captionTargetBox-2="[151,1435,181,1749]" captionTargetBox-3="[151,1435,181,1893]" captionTargetBox-4="[151,1436,181,1762]" captionTargetBox-5="[164,1422,181,1784]" captionTargetBox-6="[151,1436,182,1813]" captionTargetBox-7="[151,1436,181,1328]" captionTargetBox-8="[151,1435,309,1509]" captionTargetBox-9="[164,1423,181,1546]" captionTargetId-0="figure-20@3.[151,1436,179,1784]" captionTargetId-1="figure-37@4.[151,1435,181,1829]" captionTargetId-10="figure-129@17.[177,1411,381,1945]" captionTargetId-11="figure-37@18.[151,1436,181,1810]" captionTargetId-12="figure-106@19.[151,1436,345,1669]" captionTargetId-13="figure-122@20.[151,1436,345,1846]" captionTargetId-2="figure-20@5.[151,1435,181,1749]" captionTargetId-3="figure-37@6.[151,1435,181,1893]" captionTargetId-4="figure-20@7.[151,1436,181,1762]" captionTargetId-5="figure-37@8.[164,1423,181,1784]" captionTargetId-6="figure-20@9.[151,1436,181,1813]" captionTargetId-7="figure-37@10.[151,1436,181,1328]" captionTargetId-8="figure-91@14.[151,1435,309,1509]" captionTargetId-9="figure-37@16.[164,1423,181,1546]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="17" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-13="20" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="14" captionTargetPageId-9="16" captionText-0="FIGURES 1–10. Ecdyonurus (Rhithrogeniella) ornatus from Kodaikanal (India). 1–3, larvae; 4–10, tergalii I–VII." captionText-1="FIGURES 11–25. Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae. 11–14, specimen from Agumbe (India); 15–18, 19–21 and 22–25, specimens from Pai (Thailand) (11, 15, 19, 22, head; 12, 16, 20, 23, half of pronotum and mesonotum; 13, 17, 24, hind leg; 14, 18, 21, 25, abdomen)." captionText-10="FIGURES 91–96. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from Pai (Thailand). 91–92, reared imago, specimen [XIII](4)2015; 93–94, reared imago, specimen [XIX](1)2015; 95–96, reared subimago, specimen [XVIII](4)2015." captionText-11="FIGURES 97–102. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs of reared imagines from Kwai-Yai (Thailand). 97–98, specimen [III](11)2015; 99–100, specimen [III](37)A2015; 101–102, specimen [IV](2)2015." captionText-12="FIGURES 103–110. 103–109, Ecdyonurus (Rhithrogeniella) sp. 2: 103, male imago, 104, abdomen of male imago; 105, male subimago; 106, female subimago; 107, thorax of male subimago, ventral view; 108, male subimaginal exuviae; 109, enlarged part of its ventral arc of prealar bridge; 110, the same of E. (Rh.) ornatus." captionText-13="FIGURES 111–116. Ecdyonurus (Rhithrogeniella) sp. 2: 111, genitalia of male imago; 112, penis of male subimago; 113–114, penes of male imagines; 115–116, eggs." captionText-2="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." captionText-3="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." captionText-4="FIGURES 42–52. Larvae. 42–50, Ecdyonurus (Rhithrogeniella) ornatus, larval exuviae: 42–44, specimen from Agumbe (India), posterior margins of abdominal terga I, VIII and X; 45, specimen from Pai (Thailand), posterior margins of abdominal tergum VI; 46–48, cercus of specimen from Agumbe: 46, proximal portion; 47, middle portion in median-ventral view, to show bases of primary swimming setae on inner (median) side; 48, the same portion with focus on lateral-dorsal side, to show bases of secondary swimming setae on outer (lateral) side; 49–50, cercus of specimen from Kodaikanal (India). 51–52, Ecdyonurus (Rhithrogeniella) joernensis (Russia, Orenburg province, Tashla, 16.VIII.1986, coll. N. Kluge): two portions of cercus, dorsal view." captionText-5="FIGURES 53–68. Ecdyonurus (Rhithrogeniella) ornatus. 53–56, male abdomina to show hypodermal coloration: 53, subimago extracted from mature larva; 54–55, imagines; 56, subimago; 57, male imago; 58–60, fore and hind wings; 61, hind wing; 62– 63, fore and middle legs of male imago; 64, male imago with subimaginal exuviae; 65, 1st tarsal segment of male subimaginal exuviae; 66–67, exuviae of half of subimaginal mesonotum; 68, cercus of male imago (53, 54, 57–60, 66, from Pai; 56, 61, 67, from Kwai-Yai; 56, 61, 67, from Agumbe; 64–65, from Kodaikanal). Abbreviations: cz.a, anterior chromozone; cz.lp, lateroparapsidal chromozone; cz.ps, chromozone of parascutellum." captionText-6="FIGURES 69–72. Ecdyonurus (Rhithrogeniella) ornatus, genitalia of male imagines, ventral view. 69–70, specimen from Kodaikanal (India) in tense condition; 71, specimen from Agumbe (India); 72, specimen from Pai (Thailand)." captionText-7="FIGURES 73–76. Ecdyonurus (Rhithrogeniella) ornatus, development of penis. 73–74, imaginal penis and its subimaginal exuviae with minute points of titillators; 75, subimaginal penis extracted from mature larva, with imaginal titillators developing under subimaginal cuticle; 76, subimaginal penis extracted from mature larva, without tissues under subimaginal titillators (73–74, 76, from Pai; 75, from Kodaikanal. Abbreviations: tit.i, imaginal titillator; tit.s, subimaginal titillator)." captionText-8="FIGURES 77–84. Ecdyonurus (Rhithrogeniella) ornatus, female imagines. 77–78, specimen from Agumbe (India); 79–81, specimens from Kwai-Yai (Thailand); 82–84, specimens from Pai (Thailand)." captionText-9="FIGURES 85–90. Ecdyonurus (Rhithrogeniella) ornatus, fully developed eggs from India. 85–86, reared imago from Agumbe, specimen [XIII](2)2013; 87–88, reared imago from Agumbe, specimen [IX](6)2013; 89–90, mature larva from Kodaikanal." figureDoi-0="http://doi.org/10.5281/zenodo.8203250" figureDoi-1="http://doi.org/10.5281/zenodo.8203254" figureDoi-10="http://doi.org/10.5281/zenodo.8203272" figureDoi-11="http://doi.org/10.5281/zenodo.8203274" figureDoi-12="http://doi.org/10.5281/zenodo.8203276" figureDoi-13="http://doi.org/10.5281/zenodo.8203280" figureDoi-2="http://doi.org/10.5281/zenodo.8203256" figureDoi-3="http://doi.org/10.5281/zenodo.8203258" figureDoi-4="http://doi.org/10.5281/zenodo.8203260" figureDoi-5="http://doi.org/10.5281/zenodo.8203262" figureDoi-6="http://doi.org/10.5281/zenodo.8203264" figureDoi-7="http://doi.org/10.5281/zenodo.8203266" figureDoi-8="http://doi.org/10.5281/zenodo.8203268" figureDoi-9="http://doi.org/10.5281/zenodo.8203270" httpUri-0="https://zenodo.org/record/8203250/files/figure.png" httpUri-1="https://zenodo.org/record/8203254/files/figure.png" httpUri-10="https://zenodo.org/record/8203272/files/figure.png" httpUri-11="https://zenodo.org/record/8203274/files/figure.png" httpUri-12="https://zenodo.org/record/8203276/files/figure.png" httpUri-13="https://zenodo.org/record/8203280/files/figure.png" httpUri-2="https://zenodo.org/record/8203256/files/figure.png" httpUri-3="https://zenodo.org/record/8203258/files/figure.png" httpUri-4="https://zenodo.org/record/8203260/files/figure.png" httpUri-5="https://zenodo.org/record/8203262/files/figure.png" httpUri-6="https://zenodo.org/record/8203264/files/figure.png" httpUri-7="https://zenodo.org/record/8203266/files/figure.png" httpUri-8="https://zenodo.org/record/8203268/files/figure.png" httpUri-9="https://zenodo.org/record/8203270/files/figure.png" pageId="2" pageNumber="503">Figs 1–116</figureCitation>
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)
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</paragraph>
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</emphasis>
|
||
<subSubSection id="C357651AFFC7E657FF64FE438E08FE1D" box="[151,1329,464,491]" pageId="2" pageNumber="503" type="materials_examined">
|
||
<paragraph id="8BF23691FFC7E657FF64FE438E08FE1D" blockId="2.[151,1329,464,491]" box="[151,1329,464,491]" pageId="2" pageNumber="503">
|
||
<materialsCitation id="3B253CCCFFC7E657FF64FE438E14FE1D" box="[151,1325,464,491]" collectionCode="ICZN" pageId="2" pageNumber="503" specimenCode="ICZN 2015" specimenCount="1" typeStatus="neotype">
|
||
Type species:
|
||
<taxonomicName id="4C4D4D12FFC7E657FEC6FE428992FE1D" authority="Fabricius 1775" authorityName="Fabricius" authorityYear="1775" box="[309,683,464,491]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="venosa">
|
||
<emphasis id="B939EA83FFC7E657FEC6FE428AC4FE1C" box="[309,509,464,490]" italics="true" pageId="2" pageNumber="503">Ephemera venosa</emphasis>
|
||
Fabricius 1775
|
||
</taxonomicName>
|
||
(with
|
||
<typeStatus id="54F68833FFC7E657FD06FE438877FE1D" box="[757,846,465,491]" pageId="2" pageNumber="503">neotype</typeStatus>
|
||
designated by
|
||
<specimenCode id="DBEB9EEAFFC7E657FC05FE428F43FE1D" box="[1014,1146,464,491]" collectionCode="ICZN" pageId="2" pageNumber="503">ICZN 2015</specimenCode>
|
||
: Opinion 2356)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357651AFFC7E657FF64FD8A88F7FAF4" pageId="2" pageNumber="503" type="diagnosis">
|
||
<paragraph id="8BF23691FFC7E657FF64FD8A8903FDC4" blockId="2.[151,1437,536,1283]" box="[151,570,536,563]" pageId="2" pageNumber="503">
|
||
<emphasis id="B939EA83FFC7E657FF64FD8A8A33FDC4" bold="true" box="[151,266,536,562]" pageId="2" pageNumber="503">Diagnosis</emphasis>
|
||
(according to
|
||
<bibRefCitation id="EFDC4B60FFC7E657FE5EFD8A890BFDC4" author="Kluge, N. J." box="[429,562,536,562]" pageId="2" pageNumber="503" refId="ref9854" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FDAE8A37FD34" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">
|
||
Metathoracic nerve ganglion greatly transferred anteriorly, nearer to mesothoracic ganglion, being connected with it by short unpaired connective; due to this, imaginal and subimaginal median impression of furcasternum parallel-sided or widened toward its anterior part (not narrowed in anterior part, in contrast to all other
|
||
<taxonomicName id="4C4D4D12FFC7E657FB02FD168EACFD68" authorityName="Needham" authorityYear="1901" box="[1265,1429,644,670]" class="Insecta" family="Heptageniidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="family">Heptageniidae</taxonomicName>
|
||
) (
|
||
<figureCitation id="13762A14FFC7E657FF6CFD3A8A3BFD34" box="[159,258,680,706]" captionStart="FIGURES 103–110" captionStartId="19.[151,264,1692,1716]" captionTargetBox="[154,1436,345,1669]" captionTargetId="figure-106@19.[151,1436,345,1669]" captionTargetPageId="19" captionText="FIGURES 103–110. 103–109, Ecdyonurus (Rhithrogeniella) sp. 2: 103, male imago, 104, abdomen of male imago; 105, male subimago; 106, female subimago; 107, thorax of male subimago, ventral view; 108, male subimaginal exuviae; 109, enlarged part of its ventral arc of prealar bridge; 110, the same of E. (Rh.) ornatus." figureDoi="http://doi.org/10.5281/zenodo.8203276" httpUri="https://zenodo.org/record/8203276/files/figure.png" pageId="2" pageNumber="503">Fig. 107</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FD5E89AEFCA4" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">
|
||
Left and right mandibles have incisors dissimilar: incisor of left mandible (with mola projected distally) pointed, with a row of denticles on inner margin; incisor of right mandible (with mola projected proximally) with large denticle at some distance from apex and row of smaller denticles proximad of it; denticles on outer margins of both incisors at a distance from apex (
|
||
<figureCitation id="13762A14FFC7E657FDFBFCAA89B5FCA4" box="[520,652,824,850]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Figs 29–30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FCCE8EA3FC80" blockId="2.[151,1437,536,1283]" box="[199,1434,860,886]" pageId="2" pageNumber="503">
|
||
Superlinguae of peculiar shape uniform for all representatives, with rounded apico-lateral projection (
|
||
<figureCitation id="13762A14FFC7E657FACAFCCE8EB7FC80" box="[1337,1422,860,886]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FC1289F2FC48" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">
|
||
On maxilla, ventral row of setae transformed to field of irregularly situated setae (unique apomorphy) (
|
||
<figureCitation id="13762A14FFC7E657FAB2FC128B8FFC48" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Figs 28, 33</figureCitation>
|
||
). Proximal dentiseta always bifurcate (
|
||
<figureCitation id="13762A14FFC7E657FD99FC368987FC48" box="[618,702,932,958]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Fig. 33</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FC5A8FE9FC14" blockId="2.[151,1437,536,1283]" box="[199,1232,968,995]" pageId="2" pageNumber="503">
|
||
Vestige of segment 3 of maxillary palp triangular, larger than in
|
||
<taxonomicName id="4C4D4D12FFC7E657FC65FC5A8F20FC15" box="[918,1049,968,995]" class="Insecta" family="Heptageniidae" genus="Heptagenia" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">Heptagenia</taxonomicName>
|
||
/f6=g5 (
|
||
<figureCitation id="13762A14FFC7E657FB9CFC5A8FFDFC14" box="[1135,1220,968,994]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FC7E880BFBF0" blockId="2.[151,1437,536,1283]" box="[199,818,1004,1031]" pageId="2" pageNumber="503">
|
||
Glossae rhomboid, with inner margin convex (
|
||
<figureCitation id="13762A14FFC7E657FD22FC7E881DFBF0" box="[721,804,1004,1030]" captionStart="FIGURES 26–33" captionStartId="5.[151,264,1773,1797]" captionTargetBox="[151,1435,181,1749]" captionTargetId="figure-20@5.[151,1435,181,1749]" captionTargetPageId="5" captionText="FIGURES 26–33. Ecdyonurus (Rhithrogeniella) ornatus, larval mouthparts. 26, labrum; 27, anterior margin of labrum bent ventrally; 28, maxilla; 29–30, apices of left and right mandibles; 31, glossa and paraglossa of labium; 32, hypopharynx and superlingua; 33, apex of maxilla, dorsal view. Abbreviations: c1, c2, c3, three maxillary canines; ds1, distal (first) dentiseta; ds2, proximal (second) dentiseta; i-d, inner-dorsal row of setae; i-v, first (largest) seta of inner-ventral setal row; in, incisor; kd, kinetodontium; prs, prostheca." figureDoi="http://doi.org/10.5281/zenodo.8203256" httpUri="https://zenodo.org/record/8203256/files/figure.png" pageId="2" pageNumber="503">Fig. 31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FB828A24FB84" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">Tergalii are differentiated as following: tergalius I narrowed, with concave anal margin (i.e. more or less banana-like); tergalii II–VII widened, with anal margin most convex (i.e. more or less roundish-triangular). Unique apomorphy.</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FBEE89D2FB4D" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">
|
||
Outer (posterior) margin of larval femur with regular row of long stout setae situated less dorsally, than similar setae in
|
||
<taxonomicName id="4C4D4D12FFC7E657FF00FB328ABBFB4C" authorityName="Eaton" authorityYear="1881" box="[243,386,1184,1210]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">Rhithrogena</taxonomicName>
|
||
/fg1 and
|
||
<taxonomicName id="4C4D4D12FFC7E657FE12FB32897FFB4C" authorityName="Eaton" authorityYear="1885" box="[481,582,1184,1210]" class="Insecta" family="Heptageniidae" genus="Cinygma" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EA83FFC7E657FE12FB32897FFB4C" box="[481,582,1184,1210]" italics="true" pageId="2" pageNumber="503">Cinygma</emphasis>
|
||
</taxonomicName>
|
||
. (
|
||
<figureCitation id="13762A14FFC7E657FDAFFB3289E6FB4D" box="[604,735,1184,1211]" captionStart="FIGURES 34–41" captionStartId="6.[151,264,1916,1940]" captionTargetBox="[151,1435,181,1893]" captionTargetId="figure-37@6.[151,1435,181,1893]" captionTargetPageId="6" captionText="FIGURES 34–41. Ecdyonurus (Rhithrogeniella) ornatus, exuviae of larval legs (reared specimen from Kodaikanal). 34, fore femur; 35, outer margin of fore femur; 36–38, fore, middle and hind legs; 39, hind tibia; 40, hind tarsus; 41, claw." figureDoi="http://doi.org/10.5281/zenodo.8203258" httpUri="https://zenodo.org/record/8203258/files/figure.png" pageId="2" pageNumber="503">Figs 34–35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23691FFC7E657FF34FB5688F7FAF4" blockId="2.[151,1437,536,1283]" pageId="2" pageNumber="503">
|
||
<emphasis id="B939EA83FFC7E657FF34FB568A5EFB28" bold="true" box="[199,359,1220,1246]" pageId="2" pageNumber="503">Composition.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C4D4D12FFC7E657FE03FB56894FFB28" authorityName="Eaton" authorityYear="1868" box="[496,630,1220,1246]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B939EA83FFC7E657FE03FB56894FFB28" box="[496,630,1220,1246]" italics="true" pageId="2" pageNumber="503">Ecdyonurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A57F8FFC7E657FD8CFB578990FB28" box="[639,681,1220,1247]" pageId="2" pageNumber="503" sensu="lato">s. l.</taxonomicNameLabel>
|
||
is divided into several subordinated taxa which can be treated as subgenera (
|
||
<bibRefCitation id="EFDC4B60FFC7E657FEEAFB7A8A99FAF4" author="Kluge, N. J." box="[281,416,1256,1282]" pageId="2" pageNumber="503" refId="ref9854" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
|
||
), with the subgenus
|
||
<taxonomicName id="4C4D4D12FFC7E657FD76FB7A880DFAF4" authorityName="Ulmer" authorityYear="1939" box="[645,820,1256,1282]" pageId="2" pageNumber="503" rank="subGenus" subGenus="Rhithrogeniella">
|
||
<emphasis id="B939EA83FFC7E657FD76FB7A880DFAF4" box="[645,820,1256,1282]" italics="true" pageId="2" pageNumber="503">Rhithrogeniella</emphasis>
|
||
</taxonomicName>
|
||
among them.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |