592 lines
89 KiB
XML
592 lines
89 KiB
XML
<document id="E396B7107B53C4DEDC26BF117F711FF1" ID-CLB-Dataset="9141" ID-DOI="10.11646/zootaxa.4845.2.6" ID-GBIF-Dataset="3e00a944-44df-4b5c-a5a2-fcbf2dc7e756" ID-ISSN="1175-5326" ID-Zenodo-Dep="4406586" ID-ZooBank="D6BDC2D9-83F3-4DCD-91D9-94B38E863C2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609423552866" checkinUser="plazi" docAuthor="Frolová, Pavlína & Ďuriš, Zdeněk" docDate="2020" docId="DA0C87D9FF8C9207FF03ACE0FEBAFC2C" docLanguage="en" docName="zootaxa.4845.2.6.pdf" docOrigin="Zootaxa 4845 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madangella koumacensis Frolová & Ďuriš 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="261" masterDocId="2635FFA1FF8E920FFF94AD1EFFC4FFAD" masterDocTitle="Madangella koumacensis, a new species of palaemonid shrimps (Decapoda Palaemonidae) from New Caledonia" masterLastPageNumber="263" masterPageNumber="253" pageNumber="255" updateTime="1698850482797" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2557C06138812D1166BAAA1385C0806E">Madangella koumacensis, a new species of palaemonid shrimps (Decapoda Palaemonidae) from New Caledonia</mods:title>
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<mods:namePart id="EB67845C474A212B8037A42730516C80">Frolová, Pavlína</mods:namePart>
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<mods:namePart id="34B01B57DAE3B6F482FD2054349954C9">Ďuriš, Zdeněk</mods:namePart>
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<mods:typeOfResource id="C5A767F2002EDED9FF413D8C1752EA28">text</mods:typeOfResource>
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<mods:title id="1B674D1CD58009E1FFBFDE4CEA972EC1">Zootaxa</mods:title>
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<mods:date id="4EB9AEF1B957DDE287A07B59284023FA">2020</mods:date>
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<treatment id="DA0C87D9FF8C9207FF03ACE0FEBAFC2C" ID-DOI="http://doi.org/10.5281/zenodo.4407325" ID-GBIF-Taxon="176349859" ID-Zenodo-Dep="4407325" LSID="urn:lsid:plazi:treatment:DA0C87D9FF8C9207FF03ACE0FEBAFC2C" httpUri="http://treatment.plazi.org/id/DA0C87D9FF8C9207FF03ACE0FEBAFC2C" lastPageId="8" lastPageNumber="261" pageId="2" pageNumber="255">
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<subSubSection id="1ABF6544FF8C920DFF03ACE0FDD2FDB4" box="[151,534,510,537]" pageId="2" pageNumber="255" type="nomenclature">
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<paragraph id="521A36CFFF8C920DFF03ACE0FDD2FDB4" blockId="2.[151,534,510,573]" box="[151,534,510,537]" pageId="2" pageNumber="255">
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<heading id="095281A3FF8C920DFF03ACE0FDD2FDB4" bold="true" box="[151,534,510,537]" fontSize="11" level="1" pageId="2" pageNumber="255" reason="1">
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<emphasis id="60D1EADDFF8C920DFF03ACE0FDD2FDB4" bold="true" box="[151,534,510,537]" pageId="2" pageNumber="255">
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<taxonomicName id="95A54D4CFF8C920DFF03ACE0FE08FDB4" ID-CoL="ab99a216-a064-4d11-831f-d2ccd2e4d5e4" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[151,460,510,537]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
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<emphasis id="60D1EADDFF8C920DFF03ACE0FE08FDB4" bold="true" box="[151,460,510,537]" italics="true" pageId="2" pageNumber="255">Madangella koumacensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7BE257A6FF8C920DFE47ACE1FDD2FDB4" box="[467,534,511,537]" pageId="2" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="1ABF6544FF8C920DFF03AF3DFECFFD90" box="[151,267,546,573]" pageId="2" pageNumber="255" type="description">
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<paragraph id="521A36CFFF8C920DFF03AF3DFECFFD90" blockId="2.[151,534,510,573]" box="[151,267,546,573]" pageId="2" pageNumber="255">
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(
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<figureCitation id="CA9E2A4AFF8C920DFF0BAF3DFEC6FD90" box="[159,258,546,573]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1815,1839]" captionStartId-1="3.[151,250,1899,1923]" captionStartId-2="4.[151,250,1672,1696]" captionStartId-3="5.[151,250,1846,1870]" captionStartId-4="6.[151,250,855,879]" captionTargetBox-0="[151,1430,1069,1781]" captionTargetBox-1="[193,1404,482,1852]" captionTargetBox-2="[162,1426,192,1639]" captionTargetBox-3="[160,1425,568,1809]" captionTargetBox-4="[189,1398,166,839]" captionTargetId-0="figure-336@2.[151,1436,1062,1788]" captionTargetId-1="figure-151@3.[175,1412,453,1876]" captionTargetId-2="figure-27@4.[151,1436,181,1649]" captionTargetId-3="figure-151@5.[151,1436,561,1823]" captionTargetId-4="figure-27@6.[189,1398,166,839]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." captionText-1="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." captionText-2="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." captionText-3="FIGURE 4. Madangella koumacensis sp. n., female holotype. A, left first pereiopod, lateral. B, right first pereiopod, proximal segments, lateral. C, same, chela and distal carpus, lateral. D, third pereiopod, lateral. E, same, dactylus and distal propodus, lateral. F, fourth pereiopod, lateral. G, same, dactylus and distal propodus, lateral. H, fifth pereiopod, lateral. I, same, dactylus and distal propodus, lateral. Scale bars, 1 mm." captionText-4="FIGURE 5. Madangella koumacensis sp. n., post mortem photo of the female holotype (photo by ZĎ)." figureDoi-0="http://doi.org/10.5281/zenodo.4406588" figureDoi-1="http://doi.org/10.5281/zenodo.4406590" figureDoi-2="http://doi.org/10.5281/zenodo.4406592" figureDoi-3="http://doi.org/10.5281/zenodo.4406594" figureDoi-4="http://doi.org/10.5281/zenodo.4406596" httpUri-0="https://zenodo.org/record/4406588/files/figure.png" httpUri-1="https://zenodo.org/record/4406590/files/figure.png" httpUri-2="https://zenodo.org/record/4406592/files/figure.png" httpUri-3="https://zenodo.org/record/4406594/files/figure.png" httpUri-4="https://zenodo.org/record/4406596/files/figure.png" pageId="2" pageNumber="255">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="1ABF6544FF8C920DFF03AF75FB62FD04" pageId="2" pageNumber="255" type="materials_examined">
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<paragraph id="521A36CFFF8C920DFF03AF75FB62FD04" blockId="2.[151,1437,618,1041]" pageId="2" pageNumber="255">
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<emphasis id="60D1EADDFF8C920DFF03AF75FE8EFD29" bold="true" box="[151,330,618,645]" pageId="2" pageNumber="255">Type material.</emphasis>
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<materialsCitation id="E2CD3C92FF8C920DFEC1AF75FB62FD04" ID-GBIF-Occurrence="3007910301" collectingDate="2019-10-28" collectorName="Stn KD" country="New Caledonia" elevation="8" latitude="-20.61" location="New Caledonia" longLatPrecision="126" longitude="164.265" pageId="2" pageNumber="255" specimenCount="1" typeStatus="holotype">
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<emphasis id="60D1EADDFF8C920DFEC1AF75FE06FD28" bold="true" box="[341,450,619,645]" pageId="2" pageNumber="255">
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<typeStatus id="8D1E886DFF8C920DFEC1AF75FE06FD28" box="[341,450,619,645]" pageId="2" pageNumber="255" type="holotype">Holotype</typeStatus>
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</emphasis>
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: ovigerous female MNHN-IU-2019-2713,
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<emphasis id="60D1EADDFF8C920DFC2DAF75FB7BFD28" box="[953,1215,619,645]" italics="true" pageId="2" pageNumber="255">Expedition Koumac2.3</emphasis>
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,
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<collectorName id="FF505319FF8C920DFB45AF75FAE8FD28" box="[1233,1324,618,645]" pageId="2" pageNumber="255">Stn KD</collectorName>
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503,
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<collectingCountry id="2AB2765FFF8C920DFAF3AF75FECFFD04" name="New Caledonia" pageId="2" pageNumber="255">New Caledonia</collectingCountry>
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,
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<geoCoordinate id="37915008FF8C920DFE83AF91FE51FD04" box="[279,405,655,681]" degrees="20" direction="south" minutes="36.6" orientation="latitude" pageId="2" pageNumber="255" precision="92" value="-20.61">20° 36.6’ S</geoCoordinate>
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,
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<geoCoordinate id="37915008FF8C920DFE36AF91FDF4FD04" box="[418,560,654,681]" degrees="164" direction="east" minutes="15.9" orientation="longitude" pageId="2" pageNumber="255" precision="92" value="164.265">164° 15.9’ E</geoCoordinate>
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, dredging, sandy-mud bottom, depth
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<quantity id="955D9B2AFF8C920DFC40AF91FBC5FD04" box="[980,1025,655,681]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="255" unit="m" value="8.0">
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<elevation id="D988D1FCFF8C920DFC40AF91FBC5FD04" box="[980,1025,655,681]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="255" unit="m" value="8.0">8 m</elevation>
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</quantity>
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,
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<date id="261B100FFF8C920DFB99AF91FB66FD04" box="[1037,1186,655,681]" pageId="2" pageNumber="255" value="2019-10-28">
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<collectingDate id="365FE9E7FF8C920DFB99AF91FB66FD04" box="[1037,1186,655,681]" pageId="2" pageNumber="255" value="2019-10-28">28 Oct. 2019</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1ABF6544FF8C9208FF53AFACFD7FFB9D" lastPageId="7" lastPageNumber="260" pageId="2" pageNumber="255" type="description">
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<paragraph id="521A36CFFF8C920DFF53AFACFDE9FBBC" blockId="2.[151,1437,618,1041]" pageId="2" pageNumber="255">
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<emphasis id="60D1EADDFF8C920DFF53AFACFE9DFD61" bold="true" box="[199,345,690,716]" pageId="2" pageNumber="255">Description.</emphasis>
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Small-sized palaemonid shrimp (
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<figureCitation id="CA9E2A4AFF8C920DFD48AFADFCDBFD60" box="[732,799,691,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255">Fig. 1</figureCitation>
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) with body slender, subcylindrical, glabrous; specimen incomplete, second pereiopods lost. Rostrum (
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<figureCitation id="CA9E2A4AFF8C920DFD3BAFC9FD3EFD5C" box="[687,762,727,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255">Figs 1</figureCitation>
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,
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<figureCitation id="CA9E2A4AFF8C920DFC9FAFC9FC96FD5C" box="[779,850,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="2" pageNumber="255">2A, B</figureCitation>
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) slightly upturned, short, with length about 0.6 of PoCL, reaching disto-dorsal margin of intermediate segment of antennular peduncle; rostral lamina moderately deep, strongly compressed, sides flat, without midrib; dorsal margin slightly convex, with 7 acute, anteriorly directed teeth, with interdental short setae, first rostral tooth more distant from epigastric tooth than from next tooth; ventral margin slightly sinuous, setose, with two distal teeth, first (proximal) tooth larger, second (subterminal) vestigial. Orbit feebly developed, postorbital ridge absent. Inferior orbital angle slightly produced in later view, obliquely truncate. Antennal, hepatic and epigastric teeth present, supraorbital tooth or tubercle lacking; antennal tooth marginal, reaching half of dorsal length of antennal basicerite, hepatic tooth as large as antennal one, situated posteriorly and slightly below latter.
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</paragraph>
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<caption id="06DA6647FF8C920DFF03AA09FC37F882" ID-DOI="http://doi.org/10.5281/zenodo.4406588" ID-Zenodo-Dep="4406588" box="[151,1011,1814,1839]" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255" startId="2.[151,250,1815,1839]" targetBox="[151,1430,1069,1781]" targetPageId="2">
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<paragraph id="521A36CFFF8C920DFF03AA09FC37F882" blockId="2.[151,1011,1814,1839]" box="[151,1011,1814,1839]" pageId="2" pageNumber="255">
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<emphasis id="60D1EADDFF8C920DFF03AA09FDA7F882" bold="true" box="[151,611,1814,1839]" pageId="2" pageNumber="255">
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FIGURE 1.
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<taxonomicName id="95A54D4CFF8C920DFE8DAA08FDE0F883" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[281,548,1814,1838]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
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<emphasis id="60D1EADDFF8C920DFE8DAA08FDE0F883" bold="true" box="[281,548,1814,1838]" italics="true" pageId="2" pageNumber="255">Madangella koumacensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7BE257A6FF8C920DFDBEAA09FDA7F882" box="[554,611,1815,1839]" pageId="2" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, female holotype, habitus, lateral view.
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</paragraph>
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</caption>
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<paragraph id="521A36CFFF8C920DFF53AA43FAABF836" blockId="2.[151,1437,1885,2020]" pageId="2" pageNumber="255">
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Fourth thoracic sternite (
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<figureCitation id="CA9E2A4AFF8C920DFE72AA43FDFBF8DA" box="[486,575,1885,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="2" pageNumber="255">Fig. 2C</figureCitation>
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) with short finger-like median process, fifth sternite with transverse ridge formed by pair of triangular sub-median teeth with rounded apices and small U-shaped incision between them.
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</paragraph>
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<paragraph id="521A36CFFF8C920CFF53AABBFAB4FF78" blockId="2.[151,1437,1885,2020]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="256" pageId="2" pageNumber="255">
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Pleonal segments (
|
||
<figureCitation id="CA9E2A4AFF8C920DFE0CAABBFE13F812" box="[408,471,1957,1983]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255">Fig. 1</figureCitation>
|
||
) smooth, with pleurae I-IV broadly rounded, and pleura V (
|
||
<figureCitation id="CA9E2A4AFF8C920DFBFAAABBFB70F812" box="[1134,1204,1957,1983]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="CA9E2A4AFF8C920DFB55AABBFB20F812" box="[1217,1252,1957,1983]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="2" pageNumber="255">2D</figureCitation>
|
||
) posteroventrally produced to acute angle. Sixth segment (
|
||
<figureCitation id="CA9E2A4AFF8C920DFDE3AAD7FD7AF84E" box="[631,702,1993,2019]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="2" pageNumber="255">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="CA9E2A4AFF8C920DFD5FAAD7FD2AF84E" box="[715,750,1993,2019]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="2" pageNumber="255">2D</figureCitation>
|
||
) dorsal length subequal to telson length and about half length of PoCL, depth 0.6 of dorsal length; posterolateral angles subtriangular, produced; posteroventral angle sinuate, distoventrally produced into pointed apex (
|
||
<figureCitation id="CA9E2A4AFF8D920CFDEDADA5FD10FF78" box="[633,724,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="3" pageNumber="256">Fig. 2D</figureCitation>
|
||
); posterior dorsal margin simple, not produced in midpoint.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF8D920CFF53ADC1FBE7FE00" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="256">
|
||
Telson (
|
||
<figureCitation id="CA9E2A4AFF8D920CFEB7ADC1FE5AFF54" box="[291,414,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="3" pageNumber="256">Figs 2E, F</figureCitation>
|
||
) subequal to sixth pleonal segment length and 3 times as long as maximal width; lateral margins feebly convex in anterior half, convergent posteriorly. Two pairs of submarginal dorsal spinules, small size, about 0.07 of telson length, situated at about 0.4 and 0.7 of telson length. Posterior margin about 0.5 of anterior width, subtriangular, with small rounded median point, with three pairs of marginal spiniform setae, lateral ones subequal to dorsal spines, intermediate pair longest, about 5 times longer than dorsal spines and about 0.3 of telson length, submedian pair setulose on sides, about 0.5 of intermediate spines length.
|
||
</paragraph>
|
||
<caption id="06DA6647FF8D920CFF03AA75FD84F842" ID-DOI="http://doi.org/10.5281/zenodo.4406590" ID-Zenodo-Dep="4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="3" pageNumber="256" startId="3.[151,250,1899,1923]" targetBox="[193,1404,482,1852]" targetPageId="3">
|
||
<paragraph id="521A36CFFF8D920CFF03AA75FD84F842" blockId="3.[151,1436,1899,2031]" pageId="3" pageNumber="256">
|
||
<emphasis id="60D1EADDFF8D920CFF03AA75FDF8F82E" bold="true" box="[151,572,1899,1923]" pageId="3" pageNumber="256">
|
||
FIGURE 2.
|
||
<taxonomicName id="95A54D4CFF8D920CFE8CAA75FE3AF82E" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[280,510,1899,1923]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="altirostris" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF8D920CFE8CAA75FE3AF82E" bold="true" box="[280,510,1899,1923]" italics="true" pageId="3" pageNumber="256">Madangella altirostris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7BE257A6FF8D920CFD90AA75FDF8F82E" box="[516,572,1899,1923]" pageId="3" pageNumber="256" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype.
|
||
<emphasis id="60D1EADDFF8D920CFD62AA75FCCCF82E" bold="true" box="[758,776,1899,1923]" pageId="3" pageNumber="256">A</emphasis>
|
||
, anterior carapace with rostrum, eyes and antennae, dorsal.
|
||
<emphasis id="60D1EADDFF8D920CFACBAA75FAB4F82E" bold="true" box="[1375,1392,1899,1923]" pageId="3" pageNumber="256">B</emphasis>
|
||
, anterior carapace with rostrum, lateral.
|
||
<emphasis id="60D1EADDFF8D920CFD99AA91FDDBF80A" bold="true" box="[525,543,1935,1959]" pageId="3" pageNumber="256">C</emphasis>
|
||
, thoracic sternites 4–6 and bases of pereiopods 1–3.
|
||
<emphasis id="60D1EADDFF8D920CFBA0AA91FB82F80A" bold="true" box="[1076,1094,1935,1959]" pageId="3" pageNumber="256">D</emphasis>
|
||
, fifth and sixth pleonal segments, lateral.
|
||
<emphasis id="60D1EADDFF8D920CFF77AAADFF30F866" bold="true" box="[227,244,1971,1995]" pageId="3" pageNumber="256">E</emphasis>
|
||
, telson, and left uropod, dorsal.
|
||
<emphasis id="60D1EADDFF8D920CFDAAAAADFD89F866" bold="true" box="[574,589,1971,1995]" pageId="3" pageNumber="256">F</emphasis>
|
||
, distal end of telson, dorsal.
|
||
<emphasis id="60D1EADDFF8D920CFCE0AAADFC4CF866" bold="true" box="[884,904,1971,1995]" pageId="3" pageNumber="256">G</emphasis>
|
||
, right antennula, dorsal.
|
||
<emphasis id="60D1EADDFF8D920CFB12AAADFB5EF866" bold="true" box="[1158,1178,1971,1995]" pageId="3" pageNumber="256">H</emphasis>
|
||
, right antenna, ventral.
|
||
<emphasis id="60D1EADDFF8D920CFA18AAADFA52F866" bold="true" box="[1420,1430,1971,1995]" pageId="3" pageNumber="256">I</emphasis>
|
||
, distal end of left uropodal exopod, dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="06DA6647FF8A920BFF03AB96FED3F945" ID-DOI="http://doi.org/10.5281/zenodo.4406592" ID-Zenodo-Dep="4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="4" pageNumber="257" startId="4.[151,250,1672,1696]" targetBox="[162,1426,192,1639]" targetPageId="4">
|
||
<paragraph id="521A36CFFF8A920BFF03AB96FED3F945" blockId="4.[151,1436,1672,1768]" pageId="4" pageNumber="257">
|
||
<emphasis id="60D1EADDFF8A920BFF03AB96FDA4F90C" bold="true" box="[151,608,1672,1697]" pageId="4" pageNumber="257">
|
||
FIGURE 3.
|
||
<taxonomicName id="95A54D4CFF8A920BFE8CAB96FDE6F90D" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[280,546,1672,1696]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF8A920BFE8CAB96FDE6F90D" bold="true" box="[280,546,1672,1696]" italics="true" pageId="4" pageNumber="257">Madangella koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7BE257A6FF8A920BFDBCAB97FDA4F90C" box="[552,608,1673,1697]" pageId="4" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, mouthparts (from left side).
|
||
<emphasis id="60D1EADDFF8A920BFBACAB96FB8EF90D" bold="true" box="[1080,1098,1672,1696]" pageId="4" pageNumber="257">A</emphasis>
|
||
, paragnaths,
|
||
<emphasis id="60D1EADDFF8A920BFB44AB97FB25F90C" bold="true" box="[1232,1249,1673,1697]" pageId="4" pageNumber="257">B</emphasis>
|
||
, mandible, medial aspect.
|
||
<emphasis id="60D1EADDFF8A920BFF77ABB2FF31F969" bold="true" box="[227,245,1708,1732]" pageId="4" pageNumber="257">C</emphasis>
|
||
, same, inner aspect.
|
||
<emphasis id="60D1EADDFF8A920BFE5CABB3FE1EF968" bold="true" box="[456,474,1709,1733]" pageId="4" pageNumber="257">D</emphasis>
|
||
, maxillula.
|
||
<emphasis id="60D1EADDFF8A920BFDC6ABB3FDA7F968" bold="true" box="[594,611,1709,1733]" pageId="4" pageNumber="257">E</emphasis>
|
||
, maxilla.
|
||
<emphasis id="60D1EADDFF8A920BFD5CABB3FD13F968" bold="true" box="[712,727,1709,1733]" pageId="4" pageNumber="257">F</emphasis>
|
||
, first maxilliped.
|
||
<emphasis id="60D1EADDFF8A920BFC1FABB2FC5BF969" bold="true" box="[907,927,1708,1732]" pageId="4" pageNumber="257">G</emphasis>
|
||
, second maxilliped.
|
||
<emphasis id="60D1EADDFF8A920BFBE4ABB3FB40F968" bold="true" box="[1136,1156,1709,1733]" pageId="4" pageNumber="257">H</emphasis>
|
||
, third maxilliped. Scale bar as indicated.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="521A36CFFF8A920AFF53AA09FE66FF78" blockId="4.[151,1436,1815,2021]" lastBlockId="5.[151,1437,151,537]" lastPageId="5" lastPageNumber="258" pageId="4" pageNumber="257">
|
||
Antennula (
|
||
<figureCitation id="CA9E2A4AFF8A920BFEDFAA09FE02F89C" box="[331,454,1815,1841]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="4" pageNumber="257">Fig. 2A, G</figureCitation>
|
||
) with peduncle normally developed, exceeding tip of rostrum by distal segment. Proximal segment about 2 times longer than wide (at midlength); stylocerite stout, acute, directed anterolaterally, failing to reach middle of medial margin length; distolateral margin of proximal segment slightly convex with minute acute distolateral tooth distinctly subterminal, distal margin oblique medially from latter tooth, medial lobe obsolete. Distal two peduncular segments elongate, with combined length 0.6 of proximal segment length; intermediate segment about 1.7 times longer than wide, distal segment subequal but slenderer than previous segment. Upper flagellum biramous, with 10–12 proximal segments fused, fused part long and slender; shorter free ramus 2-segmented; with 6 groups of aesthetascs.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF8B920AFF53ADC1FE9BFEC8" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="258">
|
||
Antenna (
|
||
<figureCitation id="CA9E2A4AFF8B920AFEA1ADC1FE48FF54" box="[309,396,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="5" pageNumber="258">Fig. 2H</figureCitation>
|
||
) with stout basicerite bearing short distolateral acute tooth. Carpocerite short, reaching to 0.3 of scaphocerite length. Scaphocerite elongate, extending well beyond antennular peduncle, about 3.5 times longer than wide and 0.8 of PoCL, lateral margin almost straight, with acute distolateral tooth slightly overreaching distal margin of lamina.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF8B920AFF53AC71FD43FE7C" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="258">
|
||
Eyes (
|
||
<figureCitation id="CA9E2A4AFF8B920AFE9AAC71FE92FE24" box="[270,342,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1815,1839]" captionTargetBox="[151,1430,1069,1781]" captionTargetId="figure-336@2.[151,1436,1062,1788]" captionTargetPageId="2" captionText="FIGURE 1. Madangella koumacensis sp. n., female holotype, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406588" httpUri="https://zenodo.org/record/4406588/files/figure.png" pageId="5" pageNumber="258">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="CA9E2A4AFF8B920AFEF1AC71FE4DFE24" box="[357,393,367,393]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="5" pageNumber="258">2A</figureCitation>
|
||
) well developed, large, not reaching end of proximal segment of antennular peduncle when extended anteriorly; cornea spherical, accessory pigment spot small, inconspicuous, situated dorsally on corneal margin. Eyestalk broad, shorter than cornea.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF8B920AFF53ACC5FB9FFDB4" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="258">
|
||
Mouthparts (
|
||
<figureCitation id="CA9E2A4AFF8B920AFECDACC5FE59FE58" box="[345,413,475,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="5" pageNumber="258">Fig. 3</figureCitation>
|
||
) from left side dissected; paragnaths (
|
||
<figureCitation id="CA9E2A4AFF8B920AFCC6ACC5FC6FFE58" box="[850,939,475,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="5" pageNumber="258">Fig. 3A</figureCitation>
|
||
) with alae broadly rounded, anteromedially oblique, feebly subdivided laterally, alae separated by deeper U-shaped median notch.
|
||
</paragraph>
|
||
<caption id="06DA6647FF8B920AFF03AA28FDA8F817" ID-DOI="http://doi.org/10.5281/zenodo.4406594" ID-Zenodo-Dep="4406594" httpUri="https://zenodo.org/record/4406594/files/figure.png" pageId="5" pageNumber="258" startId="5.[151,250,1846,1870]" targetBox="[160,1425,568,1809]" targetPageId="5">
|
||
<paragraph id="521A36CFFF8B920AFF03AA28FDA8F817" blockId="5.[151,1436,1846,1979]" pageId="5" pageNumber="258">
|
||
<emphasis id="60D1EADDFF8B920AFF03AA28FDA5F8E2" bold="true" box="[151,609,1846,1871]" pageId="5" pageNumber="258">
|
||
FIGURE 4.
|
||
<taxonomicName id="95A54D4CFF8B920AFE8CAA28FDE7F8E3" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[280,547,1846,1870]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF8B920AFE8CAA28FDE7F8E3" bold="true" box="[280,547,1846,1870]" italics="true" pageId="5" pageNumber="258">Madangella koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7BE257A6FF8B920AFDBDAA29FDA5F8E2" box="[553,609,1847,1871]" pageId="5" pageNumber="258" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype. A, left first pereiopod, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFBDAAA29FB9BF8E2" bold="true" box="[1102,1119,1847,1871]" pageId="5" pageNumber="258">B</emphasis>
|
||
, right first pereiopod, proximal segments, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFEC5AA44FEA7F8DF" bold="true" box="[337,355,1882,1906]" pageId="5" pageNumber="258">C</emphasis>
|
||
, same, chela and distal carpus, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFD6CAA45FCCEF8DE" bold="true" box="[760,778,1883,1907]" pageId="5" pageNumber="258">D</emphasis>
|
||
, third pereiopod, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFB84AA45FBE5F8DE" bold="true" box="[1040,1057,1883,1907]" pageId="5" pageNumber="258">E</emphasis>
|
||
, same, dactylus and distal propodus, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFF77AA61FF36F83A" bold="true" box="[227,242,1919,1943]" pageId="5" pageNumber="258">F</emphasis>
|
||
, fourth pereiopod, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFD97AA60FDD3F83B" bold="true" box="[515,535,1918,1942]" pageId="5" pageNumber="258">G</emphasis>
|
||
, same, dactylus and distal propodus, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFC4BAA61FC37F83A" bold="true" box="[991,1011,1919,1943]" pageId="5" pageNumber="258">H</emphasis>
|
||
, fifth pereiopod, lateral.
|
||
<emphasis id="60D1EADDFF8B920AFB64AA61FB3EF83A" bold="true" box="[1264,1274,1919,1943]" pageId="5" pageNumber="258">I</emphasis>
|
||
, same, dactylus and distal propodus, lateral. Scale bars, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="06DA6647FF889209FF03AE49FB7DFCDF" ID-DOI="http://doi.org/10.5281/zenodo.4406596" ID-Zenodo-Dep="4406596" box="[151,1209,855,882]" httpUri="https://zenodo.org/record/4406596/files/figure.png" pageId="6" pageNumber="259" startId="6.[151,250,855,879]" targetBox="[189,1398,166,839]" targetPageId="6">
|
||
<paragraph id="521A36CFFF889209FF03AE49FB7DFCDF" blockId="6.[151,1209,855,882]" box="[151,1209,855,882]" pageId="6" pageNumber="259">
|
||
<emphasis id="60D1EADDFF889209FF03AE49FDA7FCDD" bold="true" box="[151,611,855,880]" pageId="6" pageNumber="259">
|
||
FIGURE 5.
|
||
<taxonomicName id="95A54D4CFF889209FE8DAE49FDE0FCC2" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[281,548,855,879]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF889209FE8DAE49FDE0FCC2" bold="true" box="[281,548,855,879]" italics="true" pageId="6" pageNumber="259">Madangella koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7BE257A6FF889209FDBEAE46FDA7FCDD" box="[554,611,856,880]" pageId="6" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="60D1EADDFF889209FDE4AE46FD2AFCC2" box="[624,750,856,879]" italics="true" pageId="6" pageNumber="259">post mortem</emphasis>
|
||
photo of the female holotype (photo by ZĎ).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="521A36CFFF889209FF53AE80FBE3F8B5" blockId="6.[151,1437,926,2032]" pageId="6" pageNumber="259">
|
||
Mandible (
|
||
<figureCitation id="CA9E2A4AFF889209FED7AE80FE79FC15" box="[323,445,926,952]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3B, C</figureCitation>
|
||
) without palp; stout molar process distally truncate, with four large lobes, stout incisor process tapering distally, with three blunt teeth. Maxillula (
|
||
<figureCitation id="CA9E2A4AFF889209FCBCAEDCFCBBFC71" box="[808,895,962,988]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3D</figureCitation>
|
||
) with bilobed palp, outer lobe depressed, rounded distally; lower lobe short, strongly curved inward, with small apical seta; upper lacinia moderately broad, dorsal margin obtusely angulate, with about 9 stout serrulate terminal setae; lower lacinia tapering distally, dorsal margin sinuate and ventral margin convex, with about 8 stout serrulate terminal and few slender subterminal setae. Maxilla (
|
||
<figureCitation id="CA9E2A4AFF889209FF34A94CFF31FBC1" box="[160,245,1106,1132]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3E</figureCitation>
|
||
) with stout palp, tapering distally, without setae; basal endite slender, deeply bilobed, distal lobe slightly larger than proximal lobe, with about 13 and 10 slender distal setae, respectively; coxal margin feebly produced; scaphognathite strongly narrowed distally, moderately broad proximally, about 2.5 times longer than central width, with numerous plumose marginal setae. First maxilliped (
|
||
<figureCitation id="CA9E2A4AFF889209FCB7A9A0FCB3FB75" box="[803,887,1214,1240]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3F</figureCitation>
|
||
) with palp tapering distally, with single pappose preterminal seta; basal endite large, broad, anterolateral convex, with slender marginal and sub-marginal setae along distal and medial margins; coxal endite distinct, separated from basal endite by deep notch, with several slender setae; exopod well-developed, tapering distally, flagellum with 4 terminal setae, with small elongate caridean lobe proximolaterally; epipod large, feebly bilobed, subtriangular. Second maxilliped (
|
||
<figureCitation id="CA9E2A4AFF889209FBDEA850FB61FAC5" box="[1098,1189,1358,1384]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3G</figureCitation>
|
||
) as usual for family, endopod with dactylar segment about 3 times longer than wide, with numerous robust serrate setae along medial margin; propodal segment broadly rounded anteriorly, with numerous serrate setae; carpus short, with distodorsal pre-terminal seta, merus normal, ischium and basis fused, with suture feebly demarcated; exopod well-developed, with flagellum bearing 4 terminal setae; coxa strongly produced medially, subquadrate, with few setae; epipod simple, 1.5 times longer than greatest width. Third maxilliped (
|
||
<figureCitation id="CA9E2A4AFF889209FCF7AB1CFC7FF9B1" box="[867,955,1538,1564]" captionStart="FIGURE 3" captionStartId="4.[151,250,1672,1696]" captionTargetBox="[162,1426,192,1639]" captionTargetId="figure-27@4.[151,1436,181,1649]" captionTargetPageId="4" captionText="FIGURE 3. Madangella koumacensis sp. n., female holotype, mouthparts (from left side). A, paragnaths, B, mandible, medial aspect. C, same, inner aspect. D, maxillula. E, maxilla. F, first maxilliped. G, second maxilliped. H, third maxilliped. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.4406592" httpUri="https://zenodo.org/record/4406592/files/figure.png" pageId="6" pageNumber="259">Fig. 3H</figureCitation>
|
||
) slender and long, overreaching distal end of antennal carpocerite; ultimate segment tapering distally, about 6 times longer than broad basally, 0.7 times length of penultimate segment, with numerous serrate setae, penultimate segment about 0.7 of antepenultimate segment (ischiomerus), about 5 times longer than proximal width, with several long medial setae; ischiomerus fused with basis, with sutures feebly demarcated, about 6 times longer than proximal width, medial margin with setae, distal half of lateral margin with 8 spinules and few small setae; basis short, about 0.15 of ischiomeral length; exopod with flagellum reaching distal end of ischiomerus, with 4 long terminal setae; coxa with rounded lateral lobe, medially produced into subtriangular lobe; single reduced finger-like arthrobranch present.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF889209FF53AA3CFBDAF85D" blockId="6.[151,1437,926,2032]" pageId="6" pageNumber="259">
|
||
First pereiopod (
|
||
<figureCitation id="CA9E2A4AFF889209FE15AA3CFE00F891" box="[385,452,1826,1852]" captionStart="FIGURE 4" captionStartId="5.[151,250,1846,1870]" captionTargetBox="[160,1425,568,1809]" captionTargetId="figure-151@5.[151,1436,561,1823]" captionTargetPageId="5" captionText="FIGURE 4. Madangella koumacensis sp. n., female holotype. A, left first pereiopod, lateral. B, right first pereiopod, proximal segments, lateral. C, same, chela and distal carpus, lateral. D, third pereiopod, lateral. E, same, dactylus and distal propodus, lateral. F, fourth pereiopod, lateral. G, same, dactylus and distal propodus, lateral. H, fifth pereiopod, lateral. I, same, dactylus and distal propodus, lateral. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4406594" httpUri="https://zenodo.org/record/4406594/files/figure.png" pageId="6" pageNumber="259">Fig. 4</figureCitation>
|
||
A-C) slender, exceeding end of scaphocerite by chela and distal quarter of carpus; chela about 0.4 of PoCL; palm about 3.5 times longer than deep, with about 6 rows of short cleaning setae proximomedially; fingers as long as palm, with simple cutting edges and small acute hooked tips; carpus slender, 1.5 times chela length, about 9 times longer than distal width, tapering proximally; merus subequal to carpus length, about 12 times longer than central width, unarmed; ischium 0.6 of merus length, and slightly deeper; basis with series of submarginal setae along ventral margin; coxa with subquadrate distoventral setose lobe.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53AD89FD18FF1C" blockId="7.[151,1437,151,537]" box="[199,732,151,177]" pageId="7" pageNumber="260">
|
||
Both second pereiopods lacking from
|
||
<typeStatus id="8D1E886DFF899208FDE6AD89FD1CFF1C" box="[626,728,151,177]" pageId="7" pageNumber="260" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53ADA5FDC2FE24" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="260">
|
||
Ambulatory pereiopods (
|
||
<figureCitation id="CA9E2A4AFF899208FE4BADA5FDE6FF78" box="[479,546,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1846,1870]" captionTargetBox="[160,1425,568,1809]" captionTargetId="figure-151@5.[151,1436,561,1823]" captionTargetPageId="5" captionText="FIGURE 4. Madangella koumacensis sp. n., female holotype. A, left first pereiopod, lateral. B, right first pereiopod, proximal segments, lateral. C, same, chela and distal carpus, lateral. D, third pereiopod, lateral. E, same, dactylus and distal propodus, lateral. F, fourth pereiopod, lateral. G, same, dactylus and distal propodus, lateral. H, fifth pereiopod, lateral. I, same, dactylus and distal propodus, lateral. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4406594" httpUri="https://zenodo.org/record/4406594/files/figure.png" pageId="7" pageNumber="260">Fig. 4</figureCitation>
|
||
D-I) slender; third pereiopod exceeding scaphocerite by dactylus and propodus; dactylus long and slender, simple, feebly curved, about 0.6 of propodus length, 15 times longer than basal depth; propodus about 22 times longer than wide in midlength, sparsely setose dorsally; carpus about 0.6 times propodus length; merus 2.0 times longer than carpus, 20 times longer than wide, unarmed; ischium 0.5 of merus length and slightly deeper than latter; basis and coxa without special features. Fourth (
|
||
<figureCitation id="CA9E2A4AFF899208FC65AC55FBF1FEC8" box="[1009,1077,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1846,1870]" captionTargetBox="[160,1425,568,1809]" captionTargetId="figure-151@5.[151,1436,561,1823]" captionTargetPageId="5" captionText="FIGURE 4. Madangella koumacensis sp. n., female holotype. A, left first pereiopod, lateral. B, right first pereiopod, proximal segments, lateral. C, same, chela and distal carpus, lateral. D, third pereiopod, lateral. E, same, dactylus and distal propodus, lateral. F, fourth pereiopod, lateral. G, same, dactylus and distal propodus, lateral. H, fifth pereiopod, lateral. I, same, dactylus and distal propodus, lateral. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4406594" httpUri="https://zenodo.org/record/4406594/files/figure.png" pageId="7" pageNumber="260">Fig. 4</figureCitation>
|
||
F–G) and fifth pereiopods (
|
||
<figureCitation id="CA9E2A4AFF899208FAFBAC55FF61FE24" captionStart="FIGURE 4" captionStartId="5.[151,250,1846,1870]" captionTargetBox="[160,1425,568,1809]" captionTargetId="figure-151@5.[151,1436,561,1823]" captionTargetPageId="5" captionText="FIGURE 4. Madangella koumacensis sp. n., female holotype. A, left first pereiopod, lateral. B, right first pereiopod, proximal segments, lateral. C, same, chela and distal carpus, lateral. D, third pereiopod, lateral. E, same, dactylus and distal propodus, lateral. F, fourth pereiopod, lateral. G, same, dactylus and distal propodus, lateral. H, fifth pereiopod, lateral. I, same, dactylus and distal propodus, lateral. Scale bars, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4406594" httpUri="https://zenodo.org/record/4406594/files/figure.png" pageId="7" pageNumber="260">Fig. 4</figureCitation>
|
||
H–I) similar to third pereiopod.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53AC8DFE5AFDB4" blockId="7.[151,1437,151,537]" pageId="7" pageNumber="260">
|
||
Uropod (
|
||
<figureCitation id="CA9E2A4AFF899208FEB9AC8DFE58FE00" box="[301,412,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1899,1923]" captionTargetBox="[193,1404,482,1852]" captionTargetId="figure-151@3.[175,1412,453,1876]" captionTargetPageId="3" captionText="FIGURE 2. Madangella altirostris sp. n., female holotype. A, anterior carapace with rostrum, eyes and antennae, dorsal. B, anterior carapace with rostrum, lateral. C, thoracic sternites 4–6 and bases of pereiopods 1–3. D, fifth and sixth pleonal segments, lateral. E, telson, and left uropod, dorsal. F, distal end of telson, dorsal. G, right antennula, dorsal. H, right antenna, ventral. I, distal end of left uropodal exopod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4406590" httpUri="https://zenodo.org/record/4406590/files/figure.png" pageId="7" pageNumber="260">Fig. 2E, I</figureCitation>
|
||
) with protopod unarmed; exopod with lateral margin almost straight, terminating in small acute tooth reaching level of distal end of telson, with larger, curved mobile spine more medially, diaeresis deeply sinuate, distal end of exopod broadly rounded, semicircular; endopod broadest proximally, tapering distally, exceeding distal end of telson.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF03AF56FEB7FDCD" blockId="7.[151,371,584,608]" box="[151,371,584,608]" pageId="7" pageNumber="260">
|
||
<tableNote id="AF433741FF899208FF03AF56FEB7FDCD" box="[151,371,584,608]" pageId="7" pageNumber="260" targetBox="[163,1348,625,890]" targetPageId="7">Branchial formula:</tableNote>
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FE42AF6FFA86FCD7" pageId="7" pageNumber="260">
|
||
<table id="20A5C46FFF896DF0FF37AF6FFA80FCD7" box="[163,1348,625,890]" gridcols="9" gridrows="7" pageId="7" pageNumber="260">
|
||
<tr id="EC95348DFF896DF0FF37AF6FFA80FD24" box="[163,1348,625,649]" gridrow="0" pageId="7" pageNumber="260" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="AF445DF1FF896DF0FE22AF6FFD88FD24" box="[438,588,625,649]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="260">maxillipeds</th>
|
||
<th id="AF445DF1FF896DF0FBB7AF6FFB56FD24" box="[1059,1170,625,649]" gridcol="6" gridrow="0" pageId="7" pageNumber="260">pereiopods</th>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AF87FA80FD1C" box="[163,1348,665,689]" gridrow="1" pageId="7" pageNumber="260" rowspan-0="1">
|
||
<td id="AF445DF1FF896DF0FE22AF87FE07FD1C" box="[438,451,665,689]" gridcol="1" gridrow="1" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FD80AF87FD88FD1C" box="[532,588,665,689]" gridcol="2" gridrow="1" pageId="7" pageNumber="260">2</td>
|
||
<td id="AF445DF1FF896DF0FD1DAF87FD52FD1C" box="[649,662,665,689]" gridcol="3" gridrow="1" pageId="7" pageNumber="260">3</td>
|
||
<td id="AF445DF1FF896DF0FCFFAF87FCBCFD1C" box="[875,888,665,689]" gridcol="4" gridrow="1" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FC4DAF87FC22FD1C" box="[985,998,665,689]" gridcol="5" gridrow="1" pageId="7" pageNumber="260">2</td>
|
||
<td id="AF445DF1FF896DF0FBB7AF87FB56FD1C" box="[1059,1170,665,689]" gridcol="6" gridrow="1" pageId="7" pageNumber="260">3</td>
|
||
<td id="AF445DF1FF896DF0FB57AF87FB14FD1C" box="[1219,1232,665,689]" gridcol="7" gridrow="1" pageId="7" pageNumber="260">4</td>
|
||
<td id="AF445DF1FF896DF0FAA3AF87FA80FD1C" box="[1335,1348,665,689]" gridcol="8" gridrow="1" pageId="7" pageNumber="260">5</td>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AFDFFA80FD74" box="[163,1348,705,729]" gridrow="2" pageId="7" pageNumber="260">
|
||
<th id="AF445DF1FF896DF0FF37AFDFFEF7FD74" box="[163,307,705,729]" gridcol="0" gridrow="2" pageId="7" pageNumber="260">pleurobranchs</th>
|
||
<td id="AF445DF1FF896DF0FE22AFDFFE07FD74" box="[438,451,705,729]" gridcol="1" gridrow="2" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD80AFDFFD88FD74" box="[532,588,705,729]" gridcol="2" gridrow="2" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD1DAFDFFD52FD74" box="[649,662,705,729]" gridcol="3" gridrow="2" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FCFFAFDFFCBCFD74" box="[875,888,705,729]" gridcol="4" gridrow="2" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FC4DAFDFFC22FD74" box="[985,998,705,729]" gridcol="5" gridrow="2" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FBB7AFDFFB56FD74" box="[1059,1170,705,729]" gridcol="6" gridrow="2" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FB57AFDFFB14FD74" box="[1219,1232,705,729]" gridcol="7" gridrow="2" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FAA3AFDFFA80FD74" box="[1335,1348,705,729]" gridcol="8" gridrow="2" pageId="7" pageNumber="260">1</td>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AFF7FA80FCAF" box="[163,1348,745,770]" gridrow="3" pageId="7" pageNumber="260">
|
||
<th id="AF445DF1FF896DF0FF37AFF7FEF7FCAF" box="[163,307,745,770]" gridcol="0" gridrow="3" pageId="7" pageNumber="260">arthrobranchs</th>
|
||
<td id="AF445DF1FF896DF0FE22AFF7FE07FCAF" box="[438,451,745,770]" gridcol="1" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD80AFF7FD88FCAF" box="[532,588,745,770]" gridcol="2" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD1DAFF7FD52FCAF" box="[649,662,745,770]" gridcol="3" gridrow="3" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FCFFAFF7FCBCFCAF" box="[875,888,745,770]" gridcol="4" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FC4DAFF7FC22FCAF" box="[985,998,745,770]" gridcol="5" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FBB7AFF7FB56FCAF" box="[1059,1170,745,770]" gridcol="6" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FB57AFF7FB14FCAF" box="[1219,1232,745,770]" gridcol="7" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FAA3AFF7FA80FCAF" box="[1335,1348,745,770]" gridcol="8" gridrow="3" pageId="7" pageNumber="260">-</td>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AE0CFA80FC87" box="[163,1348,786,810]" gridrow="4" pageId="7" pageNumber="260">
|
||
<th id="AF445DF1FF896DF0FF37AE0CFEF7FC87" box="[163,307,786,810]" gridcol="0" gridrow="4" pageId="7" pageNumber="260">podobranchs</th>
|
||
<td id="AF445DF1FF896DF0FE22AE0CFE07FC87" box="[438,451,786,810]" gridcol="1" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD80AE0CFD88FC87" box="[532,588,786,810]" gridcol="2" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FD1DAE0CFD52FC87" box="[649,662,786,810]" gridcol="3" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FCFFAE0CFCBCFC87" box="[875,888,786,810]" gridcol="4" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FC4DAE0CFC22FC87" box="[985,998,786,810]" gridcol="5" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FBB7AE0CFB56FC87" box="[1059,1170,786,810]" gridcol="6" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FB57AE0CFB14FC87" box="[1219,1232,786,810]" gridcol="7" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FAA3AE0CFA80FC87" box="[1335,1348,786,810]" gridcol="8" gridrow="4" pageId="7" pageNumber="260">-</td>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AE24FA80FCFF" box="[163,1348,826,850]" gridrow="5" pageId="7" pageNumber="260">
|
||
<th id="AF445DF1FF896DF0FF37AE24FEF7FCFF" box="[163,307,826,850]" gridcol="0" gridrow="5" pageId="7" pageNumber="260">epipods</th>
|
||
<td id="AF445DF1FF896DF0FE22AE24FE07FCFF" box="[438,451,826,850]" gridcol="1" gridrow="5" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FD80AE24FD88FCFF" box="[532,588,826,850]" gridcol="2" gridrow="5" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FD1DAE24FD52FCFF" box="[649,662,826,850]" gridcol="3" gridrow="5" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FCFFAE24FCBCFCFF" box="[875,888,826,850]" gridcol="4" gridrow="5" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FC4DAE24FC22FCFF" box="[985,998,826,850]" gridcol="5" gridrow="5" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FBB7AE24FB56FCFF" box="[1059,1170,826,850]" gridcol="6" gridrow="5" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FB57AE24FB14FCFF" box="[1219,1232,826,850]" gridcol="7" gridrow="5" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FAA3AE24FA80FCFF" box="[1335,1348,826,850]" gridcol="8" gridrow="5" pageId="7" pageNumber="260">-</td>
|
||
</tr>
|
||
<tr id="EC95348DFF896DF0FF37AE7CFA80FCD7" box="[163,1348,866,890]" gridrow="6" pageId="7" pageNumber="260">
|
||
<th id="AF445DF1FF896DF0FF37AE7CFEF7FCD7" box="[163,307,866,890]" gridcol="0" gridrow="6" pageId="7" pageNumber="260">exopods</th>
|
||
<td id="AF445DF1FF896DF0FE22AE7CFE07FCD7" box="[438,451,866,890]" gridcol="1" gridrow="6" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FD80AE7CFD88FCD7" box="[532,588,866,890]" gridcol="2" gridrow="6" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FD1DAE7CFD52FCD7" box="[649,662,866,890]" gridcol="3" gridrow="6" pageId="7" pageNumber="260">1</td>
|
||
<td id="AF445DF1FF896DF0FCFFAE7CFCBCFCD7" box="[875,888,866,890]" gridcol="4" gridrow="6" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FC4DAE7CFC22FCD7" box="[985,998,866,890]" gridcol="5" gridrow="6" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FBB7AE7CFB56FCD7" box="[1059,1170,866,890]" gridcol="6" gridrow="6" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FB57AE7CFB14FCD7" box="[1219,1232,866,890]" gridcol="7" gridrow="6" pageId="7" pageNumber="260">-</td>
|
||
<td id="AF445DF1FF896DF0FAA3AE7CFA80FCD7" box="[1335,1348,866,890]" gridcol="8" gridrow="6" pageId="7" pageNumber="260">-</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53AEB4FD7FFB9D" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53AEB4FEBAFC69" bold="true" box="[199,382,938,964]" pageId="7" pageNumber="260">Measurements.</emphasis>
|
||
PoCL,
|
||
<quantity id="955D9B2AFF899208FE46AEB4FDEEFC69" box="[466,554,938,964]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="260" unit="mm" value="3.1">3.1 mm</quantity>
|
||
; rostrum length, 2.0 mm. Sixth pleomere dorsal length,
|
||
<quantity id="955D9B2AFF899208FB07AEB4FB2FFC69" box="[1171,1259,938,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="260" unit="mm" value="1.6">1.6 mm</quantity>
|
||
; depth, 1.0 mm. Total length of body (from tip of rostrum to posterior end of telson),
|
||
<quantity id="955D9B2AFF899208FC05AED0FC26FC45" box="[913,994,974,1000]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="260" unit="mm" value="14.0">14 mm</quantity>
|
||
. Telson length,
|
||
<quantity id="955D9B2AFF899208FB04AED0FB2CFC45" box="[1168,1256,974,1000]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="260" unit="mm" value="1.7">1.7 mm</quantity>
|
||
. First pereiopod segments length (fingers – palm – carpus – merus – ischium): 0.7 – 0.6 – 1.7 – 1.6 – 1.0 mm; third pereiopod (dactylus to ischium): 1.2 – 2.2 – 1.4 – 2.5 –
|
||
<quantity id="955D9B2AFF899208FDCAA908FD73FB9D" box="[606,695,1046,1072]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="260" unit="mm" value="1.4">1.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1ABF6544FF899208FF53A924FC37FB31" pageId="7" pageNumber="260" type="etymology">
|
||
<paragraph id="521A36CFFF899208FF53A924FA52FBF9" blockId="7.[151,1437,938,2009]" box="[199,1430,1082,1108]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A924FE82FBF9" bold="true" box="[199,326,1082,1108]" pageId="7" pageNumber="260">Etymology.</emphasis>
|
||
The species name is derived from Koumac, the regional centre in the
|
||
<collectingRegion id="9061F82DFF899208FBB3A924FB0DFBF9" box="[1063,1225,1082,1108]" country="New Caledonia" name="Province Nord" pageId="7" pageNumber="260">North Province</collectingRegion>
|
||
of
|
||
<collectingCountry id="2AB2765FFF899208FB78A924FA56FBF9" box="[1260,1426,1082,1108]" name="New Caledonia" pageId="7" pageNumber="260">New Caledonia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53A943FC37FB31" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A943FE9BFBDA" bold="true" box="[199,351,1117,1143]" pageId="7" pageNumber="260">Colouration.</emphasis>
|
||
Unknown; only faded traces of live colour composed of irregular reddish spots and yellowish dots over the carapace and pleon are evident in a
|
||
<emphasis id="60D1EADDFF899208FD2BA99CFC8FFB36" box="[703,843,1154,1179]" italics="true" pageId="7" pageNumber="260">post mortem</emphasis>
|
||
photo (
|
||
<figureCitation id="CA9E2A4AFF899208FC34A99CFC21FB31" box="[928,997,1153,1180]" captionStart="FIGURE 5" captionStartId="6.[151,250,855,879]" captionTargetBox="[189,1398,166,839]" captionTargetId="figure-27@6.[189,1398,166,839]" captionTargetPageId="6" captionText="FIGURE 5. Madangella koumacensis sp. n., post mortem photo of the female holotype (photo by ZĎ)." figureDoi="http://doi.org/10.5281/zenodo.4406596" httpUri="https://zenodo.org/record/4406596/files/figure.png" pageId="7" pageNumber="260">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1ABF6544FF899208FF53A9B8FC32FA34" pageId="7" pageNumber="260" type="biology_ecology">
|
||
<paragraph id="521A36CFFF899208FF53A9B8FB6CFB6D" blockId="7.[151,1437,938,2009]" box="[199,1192,1190,1216]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A9B8FEC2FB6D" bold="true" box="[199,262,1190,1216]" pageId="7" pageNumber="260">Host.</emphasis>
|
||
Unknown. Based on the slender simple ambulatory dactyli, potentially free-living.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53A9D7FD74FAA5" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A9D7FEEEFB4E" bold="true" box="[199,298,1225,1251]" pageId="7" pageNumber="260">Habitat.</emphasis>
|
||
The specimen was collected by dredging in shallow water between shore and a large reef platform inside a lagoon near Koumac, at a depth of
|
||
<specimenCount id="44A3FD46FF899208FDEAA9F0FD74FAA5" box="[638,688,1262,1288]" pageId="7" pageNumber="260" type="generic">
|
||
<quantity id="955D9B2AFF899208FDEAA9F0FD68FAA5" box="[638,684,1262,1288]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="260" unit="m" value="8.0">8 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53A80CFB6FFAD9" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A80CFE75FA86" bold="true" box="[199,433,1297,1324]" pageId="7" pageNumber="260">Molecular analysis.</emphasis>
|
||
The sequences of two mitochondrial genes were obtained for the species, namely 16S rRNA (561 bp) and COI (672 bp). The molecular comparison of both known species in the genus reveals their genetic divergences 17.6 % of 16S and 21.2 % of COI, supporting separate species recognition.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53A863FC32FA34" blockId="7.[151,1437,938,2009]" box="[199,1014,1405,1433]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A863FDE2FA35" bold="true" box="[199,550,1405,1432]" pageId="7" pageNumber="260">GenBank accession numbers.</emphasis>
|
||
<accessionNumber id="4DF6AB2CFF899208FDB9A863FD6FFA35" box="[557,683,1405,1432]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT816395" pageId="7" pageNumber="260">MT816395</accessionNumber>
|
||
(COI) and
|
||
<accessionNumber id="4DF6AB2CFF899208FCBFA860FC6DFA35" box="[811,937,1406,1432]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT819437" pageId="7" pageNumber="260">MT819437</accessionNumber>
|
||
(16S).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1ABF6544FF899207FF53A8BCFEBAFC2C" lastPageId="8" lastPageNumber="261" pageId="7" pageNumber="260" type="discussion">
|
||
<paragraph id="521A36CFFF899208FF53A8BCFF2DF939" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<emphasis id="60D1EADDFF899208FF53A8BCFEFFFA11" bold="true" box="[199,315,1442,1468]" pageId="7" pageNumber="260">Remarks.</emphasis>
|
||
The new species,
|
||
<taxonomicName id="95A54D4CFF899208FD9AA8BCFCE9FA11" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[526,813,1442,1468]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF899208FD9AA8BCFCE9FA11" box="[526,813,1442,1468]" italics="true" pageId="7" pageNumber="260">Madangella koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60D1EADDFF899208FCA1A8BCFCB2FA11" bold="true" box="[821,886,1442,1468]" pageId="7" pageNumber="260">
|
||
<taxonomicNameLabel id="7BE257A6FF899208FCA1A8BCFCB2FA11" box="[821,886,1442,1468]" pageId="7" pageNumber="260" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
clearly belongs to the previously monotypic genus
|
||
<taxonomicName id="95A54D4CFF899208FF51A8D8FE88FA4D" authorityName="Frolova & Duris" authorityYear="2018" box="[197,332,1478,1504]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF899208FF51A8D8FE88FA4D" box="[197,332,1478,1504]" italics="true" pageId="7" pageNumber="260">Madangella</emphasis>
|
||
</taxonomicName>
|
||
sharing with the
|
||
<typeStatus id="8D1E886DFF899208FD85A8D8FD85FA4D" box="[529,577,1478,1504]" pageId="7" pageNumber="260">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="95A54D4CFF899208FD33A8D8FCF9FA4D" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[679,829,1478,1504]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="altirostris">
|
||
<emphasis id="60D1EADDFF899208FD33A8D8FD7AFA72" box="[679,702,1478,1503]" italics="true" pageId="7" pageNumber="260">M</emphasis>
|
||
.
|
||
<emphasis id="60D1EADDFF899208FD59A8D8FCF9FA4D" box="[717,829,1478,1504]" italics="true" pageId="7" pageNumber="260">altirostris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60D1EADDFF899208FCA9A8D8FC85FA4D" box="[829,833,1478,1504]" italics="true" pageId="7" pageNumber="260">,</emphasis>
|
||
the following main diagnostic characters: (1) a short, slightly upturned rostrum, (2) epigastric tooth present, (3) well-developed antennal and hepatic teeth, (4) postorbital ridge and supraorbital tooth absent, (5) outer antennular flagellum with fused part slender, long, composed of over 10 elongate segments, and (6) ambulatory legs, and in particular the dactyli, long and slender. The mouthparts, including the mandible without the palp, are closely similar to those described and illustrated by
|
||
<bibRefCitation id="36344B3EFF899208FB75AB48FF21F939" author="Frolova, P. & Duris, Z." pageId="7" pageNumber="260" pagination="327 - 338" refId="ref5763" refString="Frolova, P. & Duris, Z. (2018) Madangella altirostris, a new genus and species of palaemonid shrimps (Crustacea: Decapoda: Palaemonidae) from Papua New Guinea. Zootaxa, 4438 (2), 327 - 338. https: // doi. org / 10.11646 / zootaxa. 4438.2.7" type="journal article" year="2018">Frolová & Ďuriš (2018)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF899208FF53AB80FB3CF874" blockId="7.[151,1437,938,2009]" pageId="7" pageNumber="260">
|
||
<taxonomicName id="95A54D4CFF899208FF53AB80FE20F915" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[199,484,1694,1720]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF899208FF53AB80FE20F915" box="[199,484,1694,1720]" italics="true" pageId="7" pageNumber="260">Madangella koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60D1EADDFF899208FE7EAB80FDE3F915" bold="true" box="[490,551,1694,1720]" pageId="7" pageNumber="260">
|
||
<taxonomicNameLabel id="7BE257A6FF899208FE7EAB80FDE3F915" box="[490,551,1694,1720]" pageId="7" pageNumber="260" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily distinguished from
|
||
<taxonomicName id="95A54D4CFF899208FC02AB80FBE2F915" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[918,1062,1694,1720]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="altirostris">
|
||
<emphasis id="60D1EADDFF899208FC02AB80FBE2F915" box="[918,1062,1694,1720]" italics="true" pageId="7" pageNumber="260">M. altirostris</emphasis>
|
||
</taxonomicName>
|
||
through the following characters: (1) rostrum reaching the distal margin of the intermediate segment of the antennal peduncle, tapering distally, and with 2 subterminal ventral teeth (
|
||
<emphasis id="60D1EADDFF899208FD9CABF8FDDBF952" box="[520,543,1766,1791]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
rostrum reaching distal margin of the basal antennular segment, broadly convex dorsally, with a single distoventral tooth in
|
||
<taxonomicName id="95A54D4CFF899208FDECAA14FCCEF889" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[632,778,1802,1828]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="altirostris">
|
||
<emphasis id="60D1EADDFF899208FDECAA14FCCEF889" box="[632,778,1802,1828]" italics="true" pageId="7" pageNumber="260">M. altirostris</emphasis>
|
||
</taxonomicName>
|
||
); (2) basal antennular segment with the distolateral margin oblique, and both distolateral tooth and lobe strongly reduced (
|
||
<emphasis id="60D1EADDFF899208FCC1AA30FCA8F8EA" box="[853,876,1838,1863]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
distolateral tooth and rounded lobe small but well developed, subequal); (3) first pereiopod carpus and merus being subequal (
|
||
<emphasis id="60D1EADDFF899208FC60AA4CFBCFF8C6" box="[1012,1035,1874,1899]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
carpus being distinctly longer); (4) ambulatory legs with distal stiff setae on the propodus, on both sides of the dactylar base (
|
||
<emphasis id="60D1EADDFF899208FB05AA68FB6CF822" box="[1169,1192,1910,1935]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
feebly setose distally on the propodus); (5) fifth pleomere being acutely produced posteroventrally (
|
||
<emphasis id="60D1EADDFF899208FB94AA84FBD3F81E" box="[1024,1047,1946,1971]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
broadly rounded); (7) telson being stouter, less than 3 times longer than wide anteriorly (
|
||
<emphasis id="60D1EADDFF899208FD65AAA0FCCCF87A" box="[753,776,1982,2007]" italics="true" pageId="7" pageNumber="260">vs</emphasis>
|
||
more elongate, 3.5 times longer than wide).
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF869207FF53AD89FAFEFD90" blockId="8.[151,1437,151,897]" pageId="8" pageNumber="261">
|
||
As mentioned in the Introduction, the genus
|
||
<taxonomicName id="95A54D4CFF869207FD22AD89FCF9FF1C" authorityName="Frolova & Duris" authorityYear="2018" box="[694,829,151,177]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FD22AD89FCF9FF1C" box="[694,829,151,177]" italics="true" pageId="8" pageNumber="261">Madangella</emphasis>
|
||
</taxonomicName>
|
||
belongs to a complex of palaemonid genera (the ‘CP’, or ‘Pon-1’, complex,
|
||
<emphasis id="60D1EADDFF869207FE18ADA5FE0EFF79" box="[396,458,187,212]" italics="true" pageId="8" pageNumber="261">sensu</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FE47ADA5FD66FF78" author="Horka, I. & De Grave, S. & Fransen, C. H. J. M. & Petrusek, A. & Duris, Z." box="[467,674,187,213]" pageId="8" pageNumber="261" pagination="26486" refId="ref6147" refString="Horka, I., De Grave, S., Fransen, C. H. J. M., Petrusek, A. & Duris, Z. (2016) Multiple host switching events shape the evolution of symbiotic palaemonid shrimps (Crustacea: Decapoda). Scientific Reports, 6, 26486. https: // doi. org / 10.1038 / srep 26486" type="journal article" year="2016">
|
||
Horká
|
||
<emphasis id="60D1EADDFF869207FDB5ADA5FD9FFF78" box="[545,603,187,213]" italics="true" pageId="8" pageNumber="261">et al.</emphasis>
|
||
, 2016
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="36344B3EFF869207FD48ADA5FC6EFF78" author="Chow, L. H. & De Grave, S. & Tsang, L. M." box="[732,938,187,213]" pageId="8" pageNumber="261" pagination="277 - 287" refId="ref5315" refString="Chow, L. H., De Grave, S. & Tsang, L. M. (2020) The family Anchistioididae Borradaile, 1915 Decapoda: Caridea) is a synonym of Palaemonidae Rafinesque, 1815 based on molecular and morphological evidence. Journal of Crustacean Biology, 40 (3), 277 - 287. https: // doi. org / 10.1093 / jcbiol / ruaa 012" type="journal article" year="2020">Chow et al., 2020</bibRefCitation>
|
||
, resp.) separated from the main diversity of symbiotic shrimps of the previous subfamily
|
||
<taxonomicName id="95A54D4CFF869207FD1AADC1FCDCFF54" authorityName="Kingsley" authorityYear="1879" box="[654,792,223,249]" class="Malacostraca" family="Pontoniinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="family">Pontoniinae</taxonomicName>
|
||
. Most species of its current genera (e.g.
|
||
<taxonomicName id="95A54D4CFF869207FB42ADC1FAFAFF54" authorityName="Clark" authorityYear="1919" box="[1238,1342,223,249]" class="Malacostraca" family="Palaemonidae" genus="Cuapetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FB42ADC1FAFAFF54" box="[1238,1342,223,249]" italics="true" pageId="8" pageNumber="261">Cuapetes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95A54D4CFF869207FADFADC1FE46FEB0" authority="Bruce, 1995" authorityName="Bruce" authorityYear="1995" class="Malacostraca" family="Palaemonidae" genus="Exoclimenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FADFADC1FF34FEB0" italics="true" pageId="8" pageNumber="261">Exoclimenella</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FF63AC1DFE46FEB0" author="Bruce, A. J." box="[247,386,258,285]" pageId="8" pageNumber="261" pagination="1 - 172" refId="ref5272" refString="Bruce, A. J. (1995) A synopsis of the Indo-West Pacific genera of the Pontoniinae (Crustacea: Decapoda: Pontoniinae). Theses Zoologicae, 25, 1 - 172. [imprint 1994, published 1995]" type="journal article" year="1995">Bruce, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95A54D4CFF869207FE19AC1DFE3CFEB0" authorityName="Dana" authorityYear="1852" box="[397,504,259,285]" class="Malacostraca" family="Palaemonidae" genus="Harpilius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FE19AC1DFE3CFEB0" box="[397,504,259,285]" italics="true" pageId="8" pageNumber="261">Harpilius</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="95A54D4CFF869207FDA1AC1DFCAEFEB0" authority="Bruce, 1995" authorityName="Bruce" authorityYear="1995" box="[565,874,258,285]" class="Malacostraca" family="Palaemonidae" genus="Periclimenella" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FDA1AC1DFD1EFEB0" box="[565,730,259,285]" italics="true" pageId="8" pageNumber="261">Periclimenella</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FD74AC1DFCAEFEB0" author="Bruce, A. J." box="[736,874,258,285]" pageId="8" pageNumber="261" pagination="1 - 172" refId="ref5272" refString="Bruce, A. J. (1995) A synopsis of the Indo-West Pacific genera of the Pontoniinae (Crustacea: Decapoda: Pontoniinae). Theses Zoologicae, 25, 1 - 172. [imprint 1994, published 1995]" type="journal article" year="1995">Bruce, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
) previously were placed in the polyphyletic genus
|
||
<taxonomicName id="95A54D4CFF869207FF03AC39FE73FEEC" authority="Costa, 1844" authorityName="Costa" authorityYear="1844" box="[151,439,295,321]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60D1EADDFF869207FF03AC39FEEDFEEC" box="[151,297,295,321]" italics="true" pageId="8" pageNumber="261">Periclimenes</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FEA5AC39FE73FEEC" author="Costa, O. G." box="[305,439,295,321]" pageId="8" pageNumber="261" pagination="285 - 292" refId="ref5506" refString="Costa, O. G. (1844) Su due nuovi generi di Crostacei decapodi macrouri. Annali delle Accademia degli Aspiranti Naturalisti, Napoli, 2, 285 - 292." type="journal article" year="1844">Costa, 1844</bibRefCitation>
|
||
</taxonomicName>
|
||
which still contains numerous species of yet unresolved taxonomic position. From those species, the present new species might be closely allied to,
|
||
<emphasis id="60D1EADDFF869207FCA7AC55FC5FFEC8" box="[819,923,331,357]" italics="true" pageId="8" pageNumber="261">inter alia</emphasis>
|
||
,
|
||
<taxonomicName id="95A54D4CFF869207FC3DAC55FAF1FEC8" authority="Kemp, 1922" authorityName="Kemp" authorityYear="1922" box="[937,1333,331,357]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="leptopus">
|
||
<emphasis id="60D1EADDFF869207FC3DAC55FB66FEC8" box="[937,1186,331,357]" italics="true" pageId="8" pageNumber="261">Periclimenes leptopus</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FB3EAC55FAF1FEC8" author="Kemp, S." box="[1194,1333,331,357]" pageId="8" pageNumber="261" pagination="113 - 288" refId="ref6287" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum, XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 288, pls. 1 - 9." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Andaman Islands, or
|
||
<taxonomicName id="95A54D4CFF869207FE1CAC71FD67FE24" authority="Bruce, 1969" authorityName="Bruce" authorityYear="1969" box="[392,675,367,393]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="exederens">
|
||
<emphasis id="60D1EADDFF869207FE1CAC71FDD7FE24" box="[392,531,367,393]" italics="true" pageId="8" pageNumber="261">P. exederens</emphasis>
|
||
<bibRefCitation id="36344B3EFF869207FD8EAC71FD67FE24" author="Bruce, A. J." box="[538,675,367,393]" pageId="8" pageNumber="261" pagination="253 - 278" refId="ref5229" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen, 43 (20), 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
from the South
|
||
<collectingCountry id="2AB2765FFF869207FCCFAC71FC5AFE24" box="[859,926,367,393]" name="China" pageId="8" pageNumber="261">China</collectingCountry>
|
||
Sea, due to their shared general morphology, especially the length and armature of the rostrum and the slender ambulatory pereiopods with elongate simple dactyli.
|
||
<taxonomicName id="95A54D4CFF869207FF59ACA9FE13FE7C" authorityName="Bruce" authorityYear="1969" box="[205,471,439,465]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="exederens">
|
||
<emphasis id="60D1EADDFF869207FF59ACA9FE13FE7C" box="[205,471,439,465]" italics="true" pageId="8" pageNumber="261">Periclimenes exederens</emphasis>
|
||
</taxonomicName>
|
||
is, however, easily distinguished from the new species by the hepatic tooth situated closer to the anterior margin of the carapace, the rounded pleurae of the fifth pleomere, and the longer fingers of the first pereiopods (see
|
||
<bibRefCitation id="36344B3EFF869207FEEBACE1FDCEFDB7" author="Bruce, A. J." box="[383,522,511,538]" pageId="8" pageNumber="261" pagination="253 - 278" refId="ref5229" refString="Bruce, A. J. (1969) Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen, 43 (20), 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
|
||
). Both species,
|
||
<taxonomicName id="95A54D4CFF869207FD23ACE1FC85FDB4" authorityName="Bruce" authorityYear="1969" box="[695,833,511,537]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="exederens">
|
||
<emphasis id="60D1EADDFF869207FD23ACE1FC85FDB4" box="[695,833,511,537]" italics="true" pageId="8" pageNumber="261">P. exederens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="95A54D4CFF869207FCE2ACE1FC2BFDB4" authorityName="Kemp" authorityYear="1922" box="[886,1007,511,537]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="leptopus">
|
||
<emphasis id="60D1EADDFF869207FCE2ACE1FC2BFDB4" box="[886,1007,511,537]" italics="true" pageId="8" pageNumber="261">P. leptopus</emphasis>
|
||
</taxonomicName>
|
||
, have the ambulatory dactyli failing to reach the median length of the propodus whilst in the present new species the dactyli are distinctly longer.
|
||
</paragraph>
|
||
<paragraph id="521A36CFFF869207FF53AF59FEBAFC2C" blockId="8.[151,1437,151,897]" pageId="8" pageNumber="261">
|
||
<taxonomicName id="95A54D4CFF869207FF53AF59FE06FDCC" authorityName="Kemp" authorityYear="1922" box="[199,450,583,609]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="leptopus">
|
||
<emphasis id="60D1EADDFF869207FF53AF59FE06FDCC" box="[199,450,583,609]" italics="true" pageId="8" pageNumber="261">Periclimenes leptopus</emphasis>
|
||
</taxonomicName>
|
||
seems to be morphologically closest to the present new species. However, its antennula has the basal segment with a produced lobe and a small lateral tooth distolaterally (
|
||
<bibRefCitation id="36344B3EFF869207FB18AF75FADAFD28" author="Kemp, S." box="[1164,1310,619,645]" pageId="8" pageNumber="261" pagination="113 - 288" refId="ref6287" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum, XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 288, pls. 1 - 9." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
|
||
), while the present new species has the basal segment obliquely truncated distolaterally and without a lobe, whilst possessing only a minute subdistal tooth. The new species also has a larger number of fused segments in the upper antennular flagellum. The affiliation of
|
||
<taxonomicName id="95A54D4CFF869207FE49AFC9FD9DFD5C" authorityName="Kemp" authorityYear="1922" box="[477,601,727,753]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="leptopus">
|
||
<emphasis id="60D1EADDFF869207FE49AFC9FD9DFD5C" box="[477,601,727,753]" italics="true" pageId="8" pageNumber="261">P. leptopus</emphasis>
|
||
</taxonomicName>
|
||
to the ‘Pon-I complex’ of genera cannot be confirmed due to the lack of the key character of the complex, the median tooth on the fourth thoracic sternite (mentioned by
|
||
<bibRefCitation id="36344B3EFF869207FB73AFE5FAB7FCB8" author="Kemp, S." box="[1255,1395,763,789]" pageId="8" pageNumber="261" pagination="113 - 288" refId="ref6287" refString="Kemp, S. (1922) Notes on Crustacea Decapoda in the Indian Museum, XV. Pontoniinae. Records of the Indian Museum, 24, 113 - 288, pls. 1 - 9." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
|
||
for some other taxa). A re-examination of
|
||
<typeStatus id="8D1E886DFF869207FDD7AE01FD6CFC94" box="[579,680,799,825]" pageId="8" pageNumber="261" type="syntype">syntypes</typeStatus>
|
||
, if extant, might help to clarify this question. Based on the observed morphological differences and lack of a sternal tooth in these related species, we suggest
|
||
<taxonomicName id="95A54D4CFF869207FB10AE5DFAFCFCF0" authority="Frolová & Ďuriš, 2020" authorityName="Frolová & Ďuriš" authorityYear="2020" box="[1156,1336,835,861]" class="Malacostraca" family="Palaemonidae" genus="Madangella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="koumacensis" status="sp. nov.">
|
||
<emphasis id="60D1EADDFF869207FB10AE5DFAFCFCF0" box="[1156,1336,835,861]" italics="true" pageId="8" pageNumber="261">M. koumacensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="60D1EADDFF869207FAD4AE5DFA44FCF0" bold="true" box="[1344,1408,835,861]" pageId="8" pageNumber="261">
|
||
<taxonomicNameLabel id="7BE257A6FF869207FAD4AE5DFA44FCF0" box="[1344,1408,835,861]" pageId="8" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to be a distinct species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |