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<mods:title id="EF15334A10725DC3D51BD208BC56D9CC">On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic “ Trochosa ” species</mods:title>
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<mods:namePart id="3E04DD14BF226D3B1455E41D299E9318">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="2DBF660A880DD3A756A0421A6957912A">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan 685000, Russia. &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa &amp; Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland</mods:affiliation>
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<mods:namePart id="2876A2F045BBFA706A6EBE67B058E787">Nadolny, Anton A.</mods:namePart>
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<mods:affiliation id="9E7D8630916D5E8F3DD8CA035F6CE4D3">A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nakhimov Avenue 2, Sevastopol 299011. nadolnyanton @ mail. ru; https: // orcid. org / 0000 - 0001 - 5555 - 1721</mods:affiliation>
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<treatment id="3B770114A539FFCD2FACFF3DFA86FB2D" ID-DOI="http://doi.org/10.5281/zenodo.4412757" ID-GBIF-Taxon="177323835" ID-Zenodo-Dep="4412757" LSID="urn:lsid:plazi:treatment:3B770114A539FFCD2FACFF3DFA86FB2D" httpUri="http://treatment.plazi.org/id/3B770114A539FFCD2FACFF3DFA86FB2D" lastPageNumber="58" pageId="2" pageNumber="58">
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<taxonomicName id="74DECB81A539FFCD2FACFF3DFE0FFF58" ID-CoL="82S4" ID-ENA="74991" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[151,475,223,250]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2FACFF3DFE0FFF58" bold="true" box="[151,475,223,250]" pageId="2" pageNumber="58">
<emphasis id="81AA6C10A539FFCD2FACFF3DFEDCFF5B" bold="true" box="[151,264,223,250]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
C.L.
<bibRefCitation id="D74FCDF3A539FFCD2E75FF3DFE0FFF58" author="Koch, C. L." box="[334,475,223,249]" pageId="2" pageNumber="58" pagination="541 - 563" refId="ref12075" refString="Koch, C. L. (1847) s. n. In: Die Arachniden. Vierzehnter Band, Funfzehnter Band, Sechszehnter und letzter Band &amp; Sechszehnter und letzter Band. J. L. Lotzbeck, Nurnberg, pp. 89 - 210 + pls. 481 - 504 (f. 1343 - 1412), pp. 1 - 136 + pls. 505 - 540 (f. 1413 - 1504) &amp; pp. 1 - 80 + pls. 541 - 563 (f. 1505 - 1550) + pp. 1 - 64 (Index)." type="book chapter" year="1847">Koch, 1847</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="FBC4E389A539FFCD2FACFEC6FCDAFEFB" pageId="2" pageNumber="58" type="reference_group">
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<treatmentCitation id="327F9613A539FFCD2FACFEC6FE03FE9A" author="Koch, C. L." box="[151,471,291,316]" page="95" pageId="2" pageNumber="58" year="1847">
<taxonomicName id="74DECB81A539FFCD2FACFEC6FE03FE9A" ID-CoL="82S4" authority="C. L. Koch, 1847: 95" authorityName="C. L. Koch" authorityPageNumber="95" authorityYear="1847" box="[151,471,291,316]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2FACFEC6FF20FE9D" box="[151,244,292,316]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
C.L.
<bibRefCitation id="D74FCDF3A539FFCD2E08FEC6FE03FE9A" author="Koch, C. L." box="[307,471,291,316]" pageId="2" pageNumber="58" pagination="541 - 563" refId="ref12075" refString="Koch, C. L. (1847) s. n. In: Die Arachniden. Vierzehnter Band, Funfzehnter Band, Sechszehnter und letzter Band &amp; Sechszehnter und letzter Band. J. L. Lotzbeck, Nurnberg, pp. 89 - 210 + pls. 481 - 504 (f. 1343 - 1412), pp. 1 - 136 + pls. 505 - 540 (f. 1413 - 1504) &amp; pp. 1 - 80 + pls. 541 - 563 (f. 1505 - 1550) + pp. 1 - 64 (Index)." type="book chapter" year="1847">Koch, 1847: 95</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="327F9613A539FFCD2EDFFEC6FD1CFE9D" author="Engelhardt, W." box="[484,712,291,316]" page="224" pageId="2" pageNumber="58" year="1964">
<bibRefCitation id="D74FCDF3A539FFCD2EDFFEC6FD1CFE9D" author="Engelhardt, W." box="[484,712,291,316]" pageId="2" pageNumber="58" pagination="219 - 392" refId="ref11695" refString="Engelhardt, W. (1964) Die mitteleuropaischen Arten der Gattung Trochosa C. L. Koch, 1848 (Araneae, Lycosidae). Morphologie, Chemotaxonomie, Biologie, Autokologie. Zeitschrift fur Morphologie und Okologie der Tiere, 54 (3), 219 - 392. https: // doi. org / 10.1007 / BF 00390678" type="journal article" year="1964">Engelhardt 1964: 224</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="327F9613A539FFCD2DEEFEC6FBB4FE9D" author="Fuhn, I. E. &amp; Niculescu-Burlacu, F." box="[725,1120,292,316]" page="221" pageId="2" pageNumber="58" year="1971">
<bibRefCitation id="D74FCDF3A539FFCD2DEEFEC6FBB4FE9D" author="Fuhn, I. E. &amp; Niculescu-Burlacu, F." box="[725,1120,292,316]" pageId="2" pageNumber="58" pagination="1 - 253" refId="ref11811" refString="Fuhn, I. E. &amp; Niculescu-Burlacu, F. (1971) Fam. Lycosidae. Fauna Republicii Socialiste Romania, Arachnida, 5 (3), 1 - 253." type="journal article" year="1971">Fuhn &amp; Niculescu-Burlacu 1971: 221</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="327F9613A539FFCD2B56FEC6FA4CFE9D" author="Dondale, C. D. &amp; Redner, J. H." box="[1133,1432,292,316]" page="22" pageId="2" pageNumber="58" year="1990">
<bibRefCitation id="D74FCDF3A539FFCD2B56FEC6FA4CFE9D" author="Dondale, C. D. &amp; Redner, J. H." box="[1133,1432,292,316]" pageId="2" pageNumber="58" pagination="1 - 383" refId="ref11641" refString="Dondale, C. D. &amp; Redner, J. H. (1990) The wolf spiders, nursery web spiders, and lynx spiders of Canada and Alaska, Araneae: Lycosidae, Pisauridae, and Oxyopidae. The insects and arachnids of Canada, 17, 1 - 383." type="journal article" year="1990">Dondale &amp; Redner 1990: 22</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="327F9613A539FFCD2FFCFEA0FEAAFEFB" author="Zyuzin, A. A." box="[199,382,322,346]" page="422" pageId="2" pageNumber="58" year="1990">
<bibRefCitation id="D74FCDF3A539FFCD2FFCFEA0FEAAFEFB" author="Zyuzin, A. A." box="[199,382,322,346]" pageId="2" pageNumber="58" pagination="419 - 422" refId="ref14404" refString="Zyuzin, A. A. (1990) Studies on burrowing spiders of the family Lycosidae (Araneae). I. Preliminary data on structural and functional features. Acta Zoologica Fennica, 190, 419 - 422." type="journal article" year="1990">Zyuzin 1990: 422</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="327F9613A539FFCD2EBCFEA0FD81FEFB" author="Almquist, S." box="[391,597,322,346]" page="246" pageId="2" pageNumber="58" year="2005">
<bibRefCitation id="D74FCDF3A539FFCD2EBCFEA0FD81FEFB" author="Almquist, S." box="[391,597,322,346]" pageId="2" pageNumber="58" pagination="1 - 284" refId="ref11329" refString="Almquist, S. (2005) Swedish Araneae, part 1: families Atypidae to Hahniidae (Linyphiidae excluded). Insect Systematics &amp; Evolution, 62, 1 - 284." type="journal article" year="2005">Almquist 2005: 246</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="327F9613A539FFCD2D64FEA0FCDEFEFB" author="Tanaka, H." box="[607,778,322,346]" page="94" pageId="2" pageNumber="58" year="1988">
<bibRefCitation id="D74FCDF3A539FFCD2D64FEA0FCDEFEFB" author="Tanaka, H." box="[607,778,322,346]" pageId="2" pageNumber="58" pagination="93 - 113" refId="ref14073" refString="Tanaka, H. (1988) Lycosid spiders of Japan II. The genus Trochosa C. L. Koch. Acta Arachnologica, 36, 93 - 113. https: // doi. org / 10.2476 / asjaa. 36.93" type="journal article" year="1988">Tanaka 1988: 94</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="FBC4E389A539FFCD2FACFE62FD78FE39" box="[151,684,383,408]" pageId="2" pageNumber="58" type="type_taxon">
<paragraph id="B361B002A539FFCD2FACFE62FD78FE39" blockId="2.[151,684,383,408]" box="[151,684,383,408]" pageId="2" pageNumber="58">
<emphasis id="81AA6C10A539FFCD2FACFE62FECAFE36" bold="true" box="[151,286,383,408]" pageId="2" pageNumber="58">
<typeStatus id="6C650EA0A539FFCD2FACFE62FF18FE39" box="[151,204,384,408]" pageId="2" pageNumber="58">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="74DECB81A539FFCD2E11FE62FD7DFE39" authority="De Geer, 1778" authorityName="De Geer" authorityYear="1778" box="[298,681,384,408]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="58" phylum="Mollusca" rank="subSpecies" species="lupus" subSpecies="ruricola">
<emphasis id="81AA6C10A539FFCD2E11FE62FDD8FE39" box="[298,524,384,408]" italics="true" pageId="2" pageNumber="58">Aranea lupus ruricola</emphasis>
<bibRefCitation id="D74FCDF3A539FFCD2D29FE62FD7DFE39" author="De Geer, C." box="[530,681,384,408]" pageId="2" pageNumber="58" pagination="176 - 324" refId="ref11568" refString="De Geer, C. (1778) Des araignees. In: Memoires pour servir a l'histoire des insectes. Tome septieme. Pierre Hesselberg, Stockholm, pp. 176 - 324, pls. 11 - 19 + 38 - 39." type="book chapter" year="1778">De Geer, 1778</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="FBC4E389A539FFCD2FACFE27FCA0FC30" pageId="2" pageNumber="58" type="diagnosis">
<paragraph id="B361B002A539FFCD2FACFE27FCA0FC30" blockId="2.[151,1437,453,1164]" pageId="2" pageNumber="58">
<emphasis id="81AA6C10A539FFCD2FACFE27FEDEFE7E" bold="true" box="[151,266,453,479]" pageId="2" pageNumber="58">Diagnosis</emphasis>
.
<taxonomicName id="74DECB81A539FFCD2E21FE24FE55FE41" authorityName="C. L. Koch" authorityYear="1847" box="[282,385,454,480]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2E21FE24FE55FE41" box="[282,385,454,480]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
resembles
<taxonomicName id="74DECB81A539FFCD2D39FE25FCDEFE41" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[514,778,454,480]" class="Arachnida" family="Lycosidae" genus="Pirata" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2D39FE25FD9EFE41" box="[514,586,455,480]" italics="true" pageId="2" pageNumber="58">Pirata</emphasis>
Sundevall, 1833
</taxonomicName>
and the related genera
<taxonomicName id="74DECB81A539FFCD2B2DFE24FACDFE41" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[1046,1305,454,480]" class="Arachnida" family="Lycosidae" genus="Piratula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2B2DFE24FBA7FE41" box="[1046,1139,454,480]" italics="true" pageId="2" pageNumber="58">Piratula</emphasis>
Roewer, 1960
</taxonomicName>
and
<taxonomicName id="74DECB81A539FFCD2A68FE24FE3BFDA5" authority="Dondale &amp; Redner, 1981" authorityName="Dondale &amp; Redner" authorityYear="1981" class="Arachnida" family="Lycosidae" genus="Trebacosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2A68FE24FF1EFDA5" italics="true" pageId="2" pageNumber="58">Trebacosa</emphasis>
Dondale &amp; Redner, 1981
</taxonomicName>
by the carapace pattern: a pair of longitudinal stripes in the anterior part of a light median band. However, the genus can be easily separated from the others due to its large size, and the parallel longitudinal stripes in
<taxonomicName id="74DECB81A539FFCD2EAAFDD0FE2CFDED" authorityName="C. L. Koch" authorityYear="1847" box="[401,504,562,588]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2EAAFDD0FE2CFDED" box="[401,504,562,588]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
(
<figureCitation id="2BE5AC87A539FFCD2D3CFDD0FD93FDED" box="[519,583,562,588]" captionStart="FIGURE 1" captionStartId="3.[151,250,1319,1343]" captionTargetBox="[157,1436,181,1295]" captionTargetId="figure-23@3.[151,1436,181,1295]" captionTargetPageId="3" captionText="FIGURE 1. Males (A–B) and females (C–D) of Trochosa hispanica from the Crimea.A, C–D living and B preserved in ethanol. Scale bar 2 mm" figureDoi="http://doi.org/10.5281/zenodo.4412759" httpUri="https://zenodo.org/record/4412759/files/figure.png" pageId="2" pageNumber="58">Fig. 1</figureCitation>
) which are converging posteriorly in
<taxonomicName id="74DECB81A539FFCD2CD6FDD1FBE1FDED" authorityName="Sundevall" authorityYear="1833" box="[1005,1077,563,588]" class="Arachnida" family="Lycosidae" genus="Pirata" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2CD6FDD1FBE1FDED" box="[1005,1077,563,588]" italics="true" pageId="2" pageNumber="58">Pirata</emphasis>
</taxonomicName>
and related genera (see
<bibRefCitation id="D74FCDF3A539FFCD2A79FDD0FEC7FDD1" author="Omelko, M. M. &amp; Marusik, Y. M. &amp; Koponen, S." pageId="2" pageNumber="58" pagination="195 - 232" refId="ref13295" refString="Omelko, M. M., Marusik, Y. M. &amp; Koponen, S. (2011) A survey of the east Palearctic Lycosidae (Aranei). 8. The genera Pirata Sundevall, 1833 and Piratula Roewer, 1960 in the Russian Far East. Arthropoda Selecta, 20, 195 - 232. https: // doi. org / 10.15298 / arthsel. 20.3.05" type="journal article" year="2011">
Omelko
<emphasis id="81AA6C10A539FFCD2FACFDB5FF1DFDD1" box="[151,201,598,624]" italics="true" pageId="2" pageNumber="58">et al</emphasis>
. 2011
</bibRefCitation>
).
<taxonomicName id="74DECB81A539FFCD2E1DFDB4FE59FDD1" authorityName="C. L. Koch" authorityYear="1847" box="[294,397,598,624]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2E1DFDB4FE59FDD1" box="[294,397,598,624]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
can be differentiated from the genera mentioned before by the copulatory organs, which are similar to those of
<taxonomicName id="74DECB81A539FFCD2E48FD99FD89FD34" authority="Simon, 1885" authorityName="Simon" authorityYear="1885" box="[371,605,634,661]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2E48FD99FE6BFD35" box="[371,447,635,660]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
Simon, 1885
</taxonomicName>
. Members of
<taxonomicName id="74DECB81A539FFCD2DC4FD98FCB2FD35" authorityName="C. L. Koch" authorityYear="1847" box="[767,870,634,660]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2DC4FD98FCB2FD35" box="[767,870,634,660]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
and
<taxonomicName id="74DECB81A539FFCD2C9AFD99FC39FD35" authorityName="Simon" authorityYear="1885" box="[929,1005,635,660]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2C9AFD99FC39FD35" box="[929,1005,635,660]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
</taxonomicName>
differ from other genera by having a triangular tegular apophysis (ventral view), an elliptically curved embolus with a basal transparent membrane, a pointed synembolus, and the epigyne has an anchor-shaped septum and a pair of deep anterior hoods.
<taxonomicName id="74DECB81A539FFCD2A3CFD20FABAFD7D" authorityName="C. L. Koch" authorityYear="1847" box="[1287,1390,706,732]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2A3CFD20FABAFD7D" box="[1287,1390,706,732]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="74DECB81A539FFCD2EA8FD05FE0BFCA1" authorityName="Simon" authorityYear="1885" box="[403,479,743,768]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2EA8FD05FE0BFCA1" box="[403,479,743,768]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
</taxonomicName>
by: 1) the lack of strong macrosetae on the cymbial apex (
<emphasis id="81AA6C10A539FFCD2B54FD05FB52FCA1" box="[1135,1158,743,768]" italics="true" pageId="2" pageNumber="58">vs</emphasis>
. present); 2) the epigyne having a small fovea, and the septal stem covering more than half of the fovea (in
<taxonomicName id="74DECB81A539FFCD2B0BFCE9FBA8FC85" authorityName="Simon" authorityYear="1885" box="[1072,1148,779,804]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2B0BFCE9FBA8FC85" box="[1072,1148,779,804]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
</taxonomicName>
, the fovea is larger, equal to three times the width of the septal stem); 3) the width of the septal base is equal to the length of the septum (in the West Palaearctic,
<taxonomicName id="74DECB81A539FFCD2EBDFCB1FE06FCCD" authorityName="Simon" authorityYear="1885" box="[390,466,851,876]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2EBDFCB1FE06FCCD" box="[390,466,851,876]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
</taxonomicName>
’s septum length is 1.2–1.4 times longer than the width of the septal base); 4) the septal stem widens anteriorly (in
<taxonomicName id="74DECB81A539FFCD2EF8FC95FDDBFC31" authorityName="Simon" authorityYear="1885" box="[451,527,887,912]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2EF8FC95FDDBFC31" box="[451,527,887,912]" italics="true" pageId="2" pageNumber="58">Hogna</emphasis>
</taxonomicName>
, the septum edges are parallel).
</paragraph>
</subSubSection>
<subSubSection id="FBC4E389A539FFCD2FFCFC7BFA86FB2D" pageId="2" pageNumber="58" type="discussion">
<paragraph id="B361B002A539FFCD2FFCFC7BFA86FB2D" blockId="2.[151,1437,453,1164]" pageId="2" pageNumber="58">
<emphasis id="81AA6C10A539FFCD2FFCFC7BFE9CFC12" bold="true" box="[199,328,921,947]" pageId="2" pageNumber="58">Comments</emphasis>
. A search for published figures and
<typeStatus id="6C650EA0A539FFCD2DCDFC78FCE5FC15" box="[758,817,922,948]" pageId="2" pageNumber="58">types</typeStatus>
of species described by
<bibRefCitation id="D74FCDF3A539FFCD2B6EFC78FADAFC14" author="Roewer, C. F." box="[1109,1294,922,949]" pageId="2" pageNumber="58" pagination="751 - 782" refId="ref13685" refString="Roewer, C. F. (1955 a) Die Araneen der Osterreichischen Iran-Expedition 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Series I, 164, 751 - 782." type="journal article" year="1955">Roewer (1955a)</bibRefCitation>
reveals that some arachnologists are not aware of the key characters of the genus. For example,
<bibRefCitation id="D74FCDF3A539FFCD2B6EFC5CFADFFC78" author="Roewer, C. F." box="[1109,1291,958,985]" pageId="2" pageNumber="58" pagination="751 - 782" refId="ref13685" refString="Roewer, C. F. (1955 a) Die Araneen der Osterreichischen Iran-Expedition 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Series I, 164, 751 - 782." type="journal article" year="1955">Roewer (1955a)</bibRefCitation>
described
<taxonomicName id="74DECB81A539FFCD2AB3FC5DFED0FC5D" authorityName="Simon" authorityYear="1870" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="81AA6C10A539FFCD2AB3FC5DFED0FC5D" italics="true" pageId="2" pageNumber="58">T. hispanica</emphasis>
</taxonomicName>
under three different names in three different genera but none of them in
<taxonomicName id="74DECB81A539FFCD2B74FC00FB62FC5D" authorityName="C. L. Koch" authorityYear="1847" box="[1103,1206,994,1020]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2B74FC00FB62FC5D" box="[1103,1206,994,1020]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
. Simultaneously, he placed a species,
<taxonomicName id="74DECB81A539FFCD2E63FBE5FD0CFB80" authority="(Roewer, 1955 a)" baseAuthorityName="Roewer" baseAuthorityYear="1955" box="[344,728,1030,1057]" class="Arachnida" family="Lycosidae" genus="Hippasa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="81AA6C10A539FFCD2E63FBE5FDC2FB81" box="[344,534,1030,1056]" italics="true" pageId="2" pageNumber="58">Hippasa loeffleri</emphasis>
(
<bibRefCitation id="D74FCDF3A539FFCD2D18FBE4FD04FB80" author="Roewer, C. F." box="[547,720,1030,1057]" pageId="2" pageNumber="58" pagination="751 - 782" refId="ref13685" refString="Roewer, C. F. (1955 a) Die Araneen der Osterreichischen Iran-Expedition 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Series I, 164, 751 - 782." type="journal article" year="1955">Roewer, 1955a</bibRefCitation>
)
</taxonomicName>
, belonging to a different subfamily, as well as
<taxonomicName id="74DECB81A539FFCD2BE7FBE4FEA7FBE5" authority="Mcheidze, 1946" authorityName="Mcheidze" authorityYear="1946" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="tbilisiensis">
<emphasis id="81AA6C10A539FFCD2BE7FBE4FF61FBE5" italics="true" pageId="2" pageNumber="58">Arctosa tbilisiensis</emphasis>
<bibRefCitation id="D74FCDF3A539FFCD2F86FBC8FEA7FBE5" author="Mcheidze, T." box="[189,371,1066,1092]" pageId="2" pageNumber="58" pagination="285 - 302" refId="ref12896" refString="Mcheidze, T. (1946) Novyye vidy paukov v Gruzii. Bulletin du Museum de Georgie, 13 (A), 285 - 302." type="journal article" year="1946">Mcheidze, 1946</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="74DECB81A539FFCD2EA6FBC8FC94FBE4" authority="Roewer, 1955 a" authorityName="Roewer" authorityYear="1955" box="[413,832,1066,1093]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="impercussa">
<emphasis id="81AA6C10A539FFCD2EA6FBC8FD58FBE5" box="[413,652,1066,1092]" italics="true" pageId="2" pageNumber="58">Trochosa impercussa</emphasis>
<bibRefCitation id="D74FCDF3A539FFCD2DAFFBC8FC94FBE4" author="Roewer, C. F." box="[660,832,1066,1093]" pageId="2" pageNumber="58" pagination="751 - 782" refId="ref13685" refString="Roewer, C. F. (1955 a) Die Araneen der Osterreichischen Iran-Expedition 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Series I, 164, 751 - 782." type="journal article" year="1955">Roewer, 1955a</bibRefCitation>
</taxonomicName>
) in
<taxonomicName id="74DECB81A539FFCD2C55FBC8FC01FBE5" authorityName="C. L. Koch" authorityYear="1847" box="[878,981,1066,1092]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2C55FBC8FC01FBE5" box="[878,981,1066,1092]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
. In the revision of Australian
<taxonomicName id="74DECB81A539FFCD2A15FBC8FEE8FBC9" authority=", McKay (1979)" authorityName=", McKay" authorityYear="1979" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="81AA6C10A539FFCD2A15FBC8FA41FBE5" box="[1326,1429,1066,1092]" italics="true" pageId="2" pageNumber="58">Trochosa</emphasis>
,
<bibRefCitation id="D74FCDF3A539FFCD2FACFBACFEE8FBC9" author="McKay, R. J." box="[151,316,1102,1128]" pageId="2" pageNumber="58" pagination="277 - 298" refId="ref12858" refString="McKay, R. J. (1979) The wolf spiders of Australia (Araneae: Lycosidae): 13. The genus Trochosa. Memoirs of the Queensland Museum, 19, 277 - 298." type="journal article" year="1979">McKay (1979)</bibRefCitation>
</taxonomicName>
treated 12 species, although 11 of them are now transferred to other genera. The remaining
<taxonomicName id="74DECB81A539FFCD2A0EFBACFE55FB2D" authority="McKay, 1979" authorityName="McKay" authorityYear="1979" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="glarea">
<emphasis id="81AA6C10A539FFCD2A0EFBACFF0BFB2D" italics="true" pageId="2" pageNumber="58">Trochosa glarea</emphasis>
<bibRefCitation id="D74FCDF3A539FFCD2FDDFB90FE55FB2D" author="McKay, R. J." box="[230,385,1138,1164]" pageId="2" pageNumber="58" pagination="277 - 298" refId="ref12858" refString="McKay, R. J. (1979) The wolf spiders of Australia (Araneae: Lycosidae): 13. The genus Trochosa. Memoirs of the Queensland Museum, 19, 277 - 298." type="journal article" year="1979">McKay, 1979</bibRefCitation>
</taxonomicName>
, known only from the female, has an epigyne that differs from those of the generotype.
</paragraph>
</subSubSection>
</treatment>
</document>
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