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<document id="B0BE5307BD4C34B431B40DCEC085FD27" ID-DOI="10.5281/zenodo.179923" ID-GBIF-Dataset="d4476361-9dc4-449d-bcf8-133aa0dedc73" ID-ISSN="1175-5326" ID-Zenodo-Dep="179923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460013795156" checkinUser="plazi" docAuthor="Molineri, Carlos &amp; Malzacher, Peter" docDate="2007" docId="03E911354F20E612FF0283C1878AFEE5" docLanguage="en" docName="zt01660p031.pdf" docOrigin="Zootaxa 1660" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Caenis" docType="treatment" docVersion="9" lastPageNumber="3" masterDocId="FFD0694D4F22E617FF95816C8213FFE4" masterDocTitle="South American Caenis Stephens (Ephemeroptera, Caenidae), new species and stage descriptions" masterLastPageNumber="31" masterPageNumber="1" pageNumber="3" updateTime="1698233543632" updateUser="plazi">
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<mods:title id="9EF26B1689A9851893D20BC3E2DB821F">South American Caenis Stephens (Ephemeroptera, Caenidae), new species and stage descriptions</mods:title>
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<subSubSection id="C35AF3A84F20E615FF0283C18343FCE8" pageId="2" pageNumber="3" type="key">
<paragraph id="8BFFA0234F20E615FF0283C1807FFD23" blockId="2.[151,620,685,711]" box="[151,620,685,711]" pageId="2" pageNumber="3">
<heading id="D0B7174F4F20E615FF0283C1807FFD23" bold="true" box="[151,620,685,711]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B9347C314F20E615FF0283C1807FFD23" bold="true" box="[151,620,685,711]" pageId="2" pageNumber="3">
Key to
<taxonomicName id="4C40DBA04F20E615FF6583C18350FD23" ID-CoL="62JGW" ID-ENA="197147" box="[240,323,685,711]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9347C314F20E615FF6583C18350FD23" bold="true" box="[240,323,685,711]" italics="true" pageId="2" pageNumber="3">Caenis</emphasis>
</taxonomicName>
species from
<taxonomicName id="4C40DBA04F20E615FE7883C1807FFD23" ID-CoL="P9JS" box="[493,620,685,711]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="argentina">
<collectingCountry id="F357E0B34F20E615FE7883C1807FFD23" box="[493,620,685,711]" name="Argentina" pageId="2" pageNumber="3">Argentina</collectingCountry>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFFA0234F20E615FF02839E8343FCE8" blockId="2.[151,1437,754,1943]" box="[151,336,754,780]" pageId="2" pageNumber="3">Mature nymphs</paragraph>
</subSubSection>
<subSubSection id="C35AF3A84F20E615FF0282768789FC67" pageId="2" pageNumber="3" type="description">
<paragraph id="8BFFA0234F20E615FF0282768789FC67" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
1. Dark pigments on head present on mediolongitudinal line of occiput (
<figureCitation id="137BBCA64F20E615FB918276867FFCD0" box="[1028,1132,794,820]" captionStart="FIGURES 165 166" captionStartId="26.[151,269,1966,1990]" captionTargetBox="[302,1294,502,1958]" captionTargetId="figure@26.[300,1295,502,1964]" captionTargetPageId="26" captionText="FIGURES 165 166. Nymph, head color pattern: 165, C. argentina; 166, C. burmeisteri; 167, C. dominguezi; 168, C. tenella; 169, C. ludicra; 170, C. plaumanni; 171, C. uruzu." httpUri="https://zenodo.org/record/179935/files/figure.png" pageId="2" pageNumber="3">Fig. 166</figureCitation>
); coxal processes narrow, with smooth margins (
<figureCitation id="137BBCA64F20E615FE5A822E804CFCB8" box="[463,607,834,860]" captionStart="FIGURES 15 30" captionStartId="9.[151,269,1861,1885]" captionTargetBox="[173,1408,550,1838]" captionTargetId="figure@9.[164,1426,534,1864]" captionTargetPageId="9" captionText="FIGURES 15 30. Caenis burmeisteri. Male imago: 15, genitalia, v. v.; 16 18, detail of forceps, v. v. Nymph: 19, detail of anteromedian emargination of labrum; 20, labrum, d. v.; 21, hind coxa; 22, middle coxa; 23, foreleg; 24, middle leg; 25, hind leg; 26, microtrichia on submarginal ventral row of gill II; 27, gill II, d. v.; 28, foreclaw; 29, middle claw; 30, hind claw." httpUri="https://zenodo.org/record/179925/files/figure.png" pageId="2" pageNumber="3">Figs. 2122</figureCitation>
); tarsal claws III with 57 very large denticles (
<figureCitation id="137BBCA64F20E615FB08822E873DFCB8" box="[1181,1326,834,860]" captionStart="FIGURES 15 30" captionStartId="9.[151,269,1861,1885]" captionTargetBox="[173,1408,550,1838]" captionTargetId="figure@9.[164,1426,534,1864]" captionTargetPageId="9" captionText="FIGURES 15 30. Caenis burmeisteri. Male imago: 15, genitalia, v. v.; 16 18, detail of forceps, v. v. Nymph: 19, detail of anteromedian emargination of labrum; 20, labrum, d. v.; 21, hind coxa; 22, middle coxa; 23, foreleg; 24, middle leg; 25, hind leg; 26, microtrichia on submarginal ventral row of gill II; 27, gill II, d. v.; 28, foreclaw; 29, middle claw; 30, hind claw." httpUri="https://zenodo.org/record/179925/files/figure.png" pageId="2" pageNumber="3">Figs. 2829</figureCitation>
); operculate gills heavily maculated (
<figureCitation id="137BBCA64F20E615FD8D82068069FC60" box="[536,634,874,900]" captionStart="FIGURES 15 30" captionStartId="9.[151,269,1861,1885]" captionTargetBox="[173,1408,550,1838]" captionTargetId="figure@9.[164,1426,534,1864]" captionTargetPageId="9" captionText="FIGURES 15 30. Caenis burmeisteri. Male imago: 15, genitalia, v. v.; 16 18, detail of forceps, v. v. Nymph: 19, detail of anteromedian emargination of labrum; 20, labrum, d. v.; 21, hind coxa; 22, middle coxa; 23, foreleg; 24, middle leg; 25, hind leg; 26, microtrichia on submarginal ventral row of gill II; 27, gill II, d. v.; 28, foreclaw; 29, middle claw; 30, hind claw." httpUri="https://zenodo.org/record/179925/files/figure.png" pageId="2" pageNumber="3">Figs. 27</figureCitation>
,
<figureCitation id="137BBCA64F20E615FD13820680A7FC60" box="[646,692,874,900]" captionStart="FIGURES 158 161" captionStartId="24.[151,269,1805,1829]" captionTargetBox="[372,1214,197,1779]" captionTargetId="figure@24.[372,1216,197,1779]" captionTargetPageId="24" captionText="FIGURES 158 161. Nymph, general view: 158, C. argentina; 159, C. burmeisteri; 160, C. dominguezi; 161, C. ludicra." httpUri="https://zenodo.org/record/179933/files/figure.png" pageId="2" pageNumber="3">159</figureCitation>
,
<figureCitation id="137BBCA64F20E615FD54820680FDFC60" box="[705,750,874,900]" captionStart="FIGURES 172 178" captionStartId="27.[151,269,1913,1937]" captionTargetBox="[375,1213,194,1887]" captionTargetId="figure@27.[373,1213,194,1889]" captionTargetPageId="27" captionText="FIGURES 172 178. Nymph, abdominal terga: 172, C. argentina; 173, C. burmeisteri; 174, C. dominguezi; 175, C. ludicra; 176, C. tenella; 177, C. plaumanni; 178, C. uruzu." httpUri="https://zenodo.org/record/179936/files/figure.png" pageId="2" pageNumber="3">173</figureCitation>
) [PA, ARG (Corrientes,
<collectingCountry id="F357E0B34F20E615FB848206866EFC60" box="[1041,1149,874,900]" name="Taiwan" pageId="2" pageNumber="3">Formosa</collectingCountry>
)]
<emphasis id="B9347C314F20E615FB1982068789FC67" box="[1164,1434,874,899]" italics="true" pageId="2" pageNumber="3">
.............
<taxonomicName id="4C40DBA04F20E615FB6482068789FC67" box="[1265,1434,874,899]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="burmeisteri">C. burmeisteri</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<paragraph id="8BFFA0234F20E615FF0282FE8789FBC0" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
1´. Dark pigments on head variable but never present on medial line of occiput (
<figureCitation id="137BBCA64F20E615FBC982FE86DEFC48" box="[1116,1229,914,940]" captionStart="FIGURES 165 166" captionStartId="26.[151,269,1966,1990]" captionTargetBox="[302,1294,502,1958]" captionTargetId="figure@26.[300,1295,502,1964]" captionTargetPageId="26" captionText="FIGURES 165 166. Nymph, head color pattern: 165, C. argentina; 166, C. burmeisteri; 167, C. dominguezi; 168, C. tenella; 169, C. ludicra; 170, C. plaumanni; 171, C. uruzu." httpUri="https://zenodo.org/record/179935/files/figure.png" pageId="2" pageNumber="3">Figs. 165</figureCitation>
, 167171); coxal processes more or less elongated, with serrated margins (e.g.,
<figureCitation id="137BBCA64F20E615FC0B82D6861DFC30" box="[926,1038,954,980]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Figs. 12</figureCitation>
,
<figureCitation id="137BBCA64F20E615FB8F82D68675FC30" box="[1050,1126,954,980]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">3940</figureCitation>
,
<figureCitation id="137BBCA64F20E615FBE682D686CDFC30" box="[1139,1246,954,980]" captionStart="FIGURES 123 138" captionStartId="21.[151,269,1854,1878]" captionTargetBox="[164,1415,400,1819]" captionTargetId="figure@21.[151,1436,397,1828]" captionTargetPageId="21" captionText="FIGURES 123 138. Caenis tenella, nymph. 123, labrum, d. v.; 124, labium, v. v. (palpus and setae at the right, omitted); 125, pronotum, left half; 126, detail of posteromedian projection of abdominal tergum II; 127, middle coxa; 128, hind coxa; 129, setae on dorsal transverse row of fore femora; 130, foreleg; 131, middle leg; 132, hind leg; 133, microtrichia of submarginal ventral row of operculate gill; 134, operculate gill; 135, IX sternum, v. v. (a, female; b, male); 136, foreclaw; 137, middle claw; 138, hind claw." httpUri="https://zenodo.org/record/179931/files/figure.png" pageId="2" pageNumber="3">127128</figureCitation>
); tarsal claws I II with small denticles or without (e.g.,
<figureCitation id="137BBCA64F20E615FD0B828E8103FC18" box="[670,784,994,1020]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Figs. 34</figureCitation>
); operculate gills not maculated (
<figureCitation id="137BBCA64F20E615FB0E828E86EDFC18" box="[1179,1278,994,1020]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">Figs. 44</figureCitation>
,
<figureCitation id="137BBCA64F20E615FA9E828E8729FC18" box="[1291,1338,994,1020]" captionStart="FIGURES 123 138" captionStartId="21.[151,269,1854,1878]" captionTargetBox="[164,1415,400,1819]" captionTargetId="figure@21.[151,1436,397,1828]" captionTargetPageId="21" captionText="FIGURES 123 138. Caenis tenella, nymph. 123, labrum, d. v.; 124, labium, v. v. (palpus and setae at the right, omitted); 125, pronotum, left half; 126, detail of posteromedian projection of abdominal tergum II; 127, middle coxa; 128, hind coxa; 129, setae on dorsal transverse row of fore femora; 130, foreleg; 131, middle leg; 132, hind leg; 133, microtrichia of submarginal ventral row of operculate gill; 134, operculate gill; 135, IX sternum, v. v. (a, female; b, male); 136, foreclaw; 137, middle claw; 138, hind claw." httpUri="https://zenodo.org/record/179931/files/figure.png" pageId="2" pageNumber="3">134</figureCitation>
,
<figureCitation id="137BBCA64F20E615FAD2828E8760FC18" box="[1351,1395,994,1020]" captionStart="FIGURES 96 122" captionStartId="19.[151,269,1762,1786]" captionTargetBox="[233,1353,184,1724]" captionTargetId="figure@19.[230,1364,168,1753]" captionTargetPageId="19" captionText="FIGURES 96 122. Caenis uruzu. Male imago: 96, genitalia, v. v.; 97 98, details of forceps, v. v.; 99, lateral filaments of abdomen; 100, preesternal triangle; 101, detail of finger-like projection of abdominal tergum II, l. v. Nymph: 102, detail of anteromedian emargination of labrum; 103, labrum, d. v.; 104, detail of posteromedian projection of tergum II; 105, labium, v. v. (setae omitted on right side); 106 107, pronotal lateral margin; 108, middle coxa; 109, hind coxa; 110, operculate gill; 111, microtrichia on submarginal ventral row of operculate gill; 112, foreclaw; 113, middle claw; 114, hind claw; 115, setae on dorsum of middle and hind femora; 116, microtrichia on dorsum of operculate gill; 117, seta on posterior margin of hind tibia; 118, seta on hind tarsi; 119, setae on transverse dorsal row on fore femora; 120, fore leg; 121, middle leg; 122, hind leg." httpUri="https://zenodo.org/record/179930/files/figure.png" pageId="2" pageNumber="3">110</figureCitation>
) or slightly maculated (
<figureCitation id="137BBCA64F20E615FE3B8566800DFBC0" box="[430,542,1034,1060]" captionStart="FIGURES 158 161" captionStartId="24.[151,269,1805,1829]" captionTargetBox="[372,1214,197,1779]" captionTargetId="figure@24.[372,1216,197,1779]" captionTargetPageId="24" captionText="FIGURES 158 161. Nymph, general view: 158, C. argentina; 159, C. burmeisteri; 160, C. dominguezi; 161, C. ludicra." httpUri="https://zenodo.org/record/179933/files/figure.png" pageId="2" pageNumber="3">Figs. 158</figureCitation>
,
<figureCitation id="137BBCA64F20E615FDBF85668044FBC0" box="[554,599,1034,1060]" captionStart="FIGURES 172 178" captionStartId="27.[151,269,1913,1937]" captionTargetBox="[375,1213,194,1887]" captionTargetId="figure@27.[373,1213,194,1889]" captionTargetPageId="27" captionText="FIGURES 172 178. Nymph, abdominal terga: 172, C. argentina; 173, C. burmeisteri; 174, C. dominguezi; 175, C. ludicra; 176, C. tenella; 177, C. plaumanni; 178, C. uruzu." httpUri="https://zenodo.org/record/179936/files/figure.png" pageId="2" pageNumber="3">172</figureCitation>
) ............................................................................................................. 2
</paragraph>
<paragraph id="8BFFA0234F20E615FF02855E878AFB08" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
2. Small species (body length of male &lt;3.0 mm, female &lt;
<quantity id="4CB80DC64F20E615FCCD855E81ABFBA8" box="[856,952,1074,1100]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="3" unit="mm" value="3.8">3.8 mm</quantity>
); general coloration very pale, whitish or yellowish almost without grayish shading (
<figureCitation id="137BBCA64F20E615FD5585368135FB90" box="[704,806,1114,1140]" captionStart="FIGURES 162 164" captionStartId="25.[151,269,1409,1433]" captionTargetBox="[365,1220,182,1400]" captionTargetId="figure@25.[365,1221,182,1401]" captionTargetPageId="25" captionText="FIGURES 162 164. Nymph, general view: 162, C. tenella; 163, C. plaumanni; 164, C. uruzu." httpUri="https://zenodo.org/record/179934/files/figure.png" pageId="2" pageNumber="3">Fig. 162</figureCitation>
); foretibia with a medial row of long setae, as long or longer than width of tibia (
<figureCitation id="137BBCA64F20E615FD8585EE8069FB78" box="[528,634,1154,1180]" captionStart="FIGURES 123 138" captionStartId="21.[151,269,1854,1878]" captionTargetBox="[164,1415,400,1819]" captionTargetId="figure@21.[151,1436,397,1828]" captionTargetPageId="21" captionText="FIGURES 123 138. Caenis tenella, nymph. 123, labrum, d. v.; 124, labium, v. v. (palpus and setae at the right, omitted); 125, pronotum, left half; 126, detail of posteromedian projection of abdominal tergum II; 127, middle coxa; 128, hind coxa; 129, setae on dorsal transverse row of fore femora; 130, foreleg; 131, middle leg; 132, hind leg; 133, microtrichia of submarginal ventral row of operculate gill; 134, operculate gill; 135, IX sternum, v. v. (a, female; b, male); 136, foreclaw; 137, middle claw; 138, hind claw." httpUri="https://zenodo.org/record/179931/files/figure.png" pageId="2" pageNumber="3">Fig. 130</figureCitation>
); posteromedian projection of abdominal tergum II short and very broad (
<figureCitation id="137BBCA64F20E615FE8F85C68398FB20" box="[282,395,1194,1220]" captionStart="FIGURES 123 138" captionStartId="21.[151,269,1854,1878]" captionTargetBox="[164,1415,400,1819]" captionTargetId="figure@21.[151,1436,397,1828]" captionTargetPageId="21" captionText="FIGURES 123 138. Caenis tenella, nymph. 123, labrum, d. v.; 124, labium, v. v. (palpus and setae at the right, omitted); 125, pronotum, left half; 126, detail of posteromedian projection of abdominal tergum II; 127, middle coxa; 128, hind coxa; 129, setae on dorsal transverse row of fore femora; 130, foreleg; 131, middle leg; 132, hind leg; 133, microtrichia of submarginal ventral row of operculate gill; 134, operculate gill; 135, IX sternum, v. v. (a, female; b, male); 136, foreclaw; 137, middle claw; 138, hind claw." httpUri="https://zenodo.org/record/179931/files/figure.png" pageId="2" pageNumber="3">Figs. 126</figureCitation>
,
<figureCitation id="137BBCA64F20E615FE0285C683D4FB20" box="[407,455,1194,1220]" captionStart="FIGURES 162 164" captionStartId="25.[151,269,1409,1433]" captionTargetBox="[365,1220,182,1400]" captionTargetId="figure@25.[365,1221,182,1401]" captionTargetPageId="25" captionText="FIGURES 162 164. Nymph, general view: 162, C. tenella; 163, C. plaumanni; 164, C. uruzu." httpUri="https://zenodo.org/record/179934/files/figure.png" pageId="2" pageNumber="3">162</figureCitation>
); IX sternum with a deep V-shaped notch at hind margin and with very short lateral spines (
<figureCitation id="137BBCA64F20E615FEB585BE8391FB08" box="[288,386,1234,1260]" captionStart="FIGURES 123 138" captionStartId="21.[151,269,1854,1878]" captionTargetBox="[164,1415,400,1819]" captionTargetId="figure@21.[151,1436,397,1828]" captionTargetPageId="21" captionText="FIGURES 123 138. Caenis tenella, nymph. 123, labrum, d. v.; 124, labium, v. v. (palpus and setae at the right, omitted); 125, pronotum, left half; 126, detail of posteromedian projection of abdominal tergum II; 127, middle coxa; 128, hind coxa; 129, setae on dorsal transverse row of fore femora; 130, foreleg; 131, middle leg; 132, hind leg; 133, microtrichia of submarginal ventral row of operculate gill; 134, operculate gill; 135, IX sternum, v. v. (a, female; b, male); 136, foreclaw; 137, middle claw; 138, hind claw." httpUri="https://zenodo.org/record/179931/files/figure.png" pageId="2" pageNumber="3">Fig. 135</figureCitation>
ab) [URU, ARG (
<collectingCountry id="F357E0B34F20E615FDFC85BE80C7FB08" box="[617,724,1234,1260]" name="Taiwan" pageId="2" pageNumber="3">Formosa</collectingCountry>
, Misiones, Buenos Aires)] ...................................
<taxonomicName id="4C40DBA04F20E615FABD85BF878AFB08" box="[1320,1433,1235,1260]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tenella">
<emphasis id="B9347C314F20E615FABD85BF878AFB08" box="[1320,1433,1235,1260]" italics="true" pageId="2" pageNumber="3">C. tenella</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFFA0234F20E615FF0285968789FA50" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
2´. Medium to large species (male length&gt;
<quantity id="4CB80DC64F20E615FD0A859680EDFAF0" box="[671,766,1274,1300]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="3" unit="mm" value="3.5">3.5 mm</quantity>
, female&gt;
<quantity id="4CB80DC64F20E615FCE9859681CFFAF0" box="[892,988,1274,1300]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="3" unit="mm" value="4.1">4.1 mm</quantity>
); general coloration darker, yellowish brown to dark brown with grayish markings (
<figureCitation id="137BBCA64F20E615FD60844E81F1FAD8" box="[757,994,1314,1340]" captionStart="FIGURES 158 161" captionStartId="24.[151,269,1805,1829]" captionTargetBox="[372,1214,197,1779]" captionTargetId="figure@24.[372,1216,197,1779]" captionTargetPageId="24" captionText="FIGURES 158 161. Nymph, general view: 158, C. argentina; 159, C. burmeisteri; 160, C. dominguezi; 161, C. ludicra." httpUri="https://zenodo.org/record/179933/files/figure.png" pageId="2" pageNumber="3">Figs. 158, 160161</figureCitation>
,
<figureCitation id="137BBCA64F20E615FC67844E864EFAD8" box="[1010,1117,1314,1340]" captionStart="FIGURES 162 164" captionStartId="25.[151,269,1409,1433]" captionTargetBox="[365,1220,182,1400]" captionTargetId="figure@25.[365,1221,182,1401]" captionTargetPageId="25" captionText="FIGURES 162 164. Nymph, general view: 162, C. tenella; 163, C. plaumanni; 164, C. uruzu." httpUri="https://zenodo.org/record/179934/files/figure.png" pageId="2" pageNumber="3">163164</figureCitation>
); foretibia only with short setae (
<figureCitation id="137BBCA64F20E615FE848426837BFA80" box="[273,360,1354,1380]" captionStart="FIGURES 66 85" captionStartId="16.[151,269,1664,1688]" captionTargetBox="[184,1411,262,1604]" captionTargetId="figure@16.[151,1436,244,1640]" captionTargetPageId="16" captionText="FIGURES 66 85. Caenis ludicra, nymph. 66, middle coxa; 67, hind coxa; 68, hind claw; 69, middle claw; 70, foreclaw; 71, hind claw; 72, middle claw; 73, foreclaw; 74, microtrichia on dorsum of operculate gill; 75 76, microtrichia on submarginal ventral row of operculate gill; 77, seta on posterior margin of hind tibia; 78, seta on posterior margin of hind tarsi; 79, setae on dorsal transverse row of fore femora; 80, microtrichia on dorsum of hind femora; 81, seta on hind tarsi; 82, seta of posterior margin of hind femora; 83, foreleg; 84, middle leg; 85, hind leg." httpUri="https://zenodo.org/record/179928/files/figure.png" pageId="2" pageNumber="3">Fig. 83</figureCitation>
); posteromedian projection of abdominal tergum II longer and narrower (e.g.,
<figureCitation id="137BBCA64F20E615FB6984268740FA80" box="[1276,1363,1354,1380]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">Fig. 48</figureCitation>
); hind margin of IX sternum rounded, truncate or with an U-shaped indentation, lateral spines longer and stronger (e.g.,
<figureCitation id="137BBCA64F20E615FEA384F68399FA50" box="[310,394,1434,1460]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">Fig. 49</figureCitation>
) ......................................................................................................................................... 3
</paragraph>
<paragraph id="8BFFA0234F20E615FF0284AE8789FA38" blockId="2.[151,1437,754,1943]" box="[151,1434,1474,1500]" pageId="2" pageNumber="3">
3. Occiput without submedian marks (Fig. 167); operculate gills only with simple setae dorsally (
<figureCitation id="137BBCA64F20E615FA8884AE8761FA38" box="[1309,1394,1474,1500]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">Fig. 44</figureCitation>
). 4
</paragraph>
<paragraph id="8BFFA0234F20E615FF0284868789F9CB" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
3´. Occiput at least with a pair of submedian star-like dark markings (
<figureCitation id="137BBCA64F20E615FC7184868645F9E0" box="[996,1110,1514,1540]" captionStart="FIGURES 165 166" captionStartId="26.[151,269,1966,1990]" captionTargetBox="[302,1294,502,1958]" captionTargetId="figure@26.[300,1295,502,1964]" captionTargetPageId="26" captionText="FIGURES 165 166. Nymph, head color pattern: 165, C. argentina; 166, C. burmeisteri; 167, C. dominguezi; 168, C. tenella; 169, C. ludicra; 170, C. plaumanni; 171, C. uruzu." httpUri="https://zenodo.org/record/179935/files/figure.png" pageId="2" pageNumber="3">Figs. 165</figureCitation>
, 168171); operculate gills with simple setae and plumose or scale-shaped microtrichia1 dorsally (e.g.,
<figureCitation id="137BBCA64F20E615FBAC87798698F9CB" box="[1081,1163,1557,1583]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
,
<figureCitation id="137BBCA64F20E615FB02877986A7F9CB" box="[1175,1204,1557,1583]" captionStart="FIGURES 66 85" captionStartId="16.[151,269,1664,1688]" captionTargetBox="[184,1411,262,1604]" captionTargetId="figure@16.[151,1436,244,1640]" captionTargetPageId="16" captionText="FIGURES 66 85. Caenis ludicra, nymph. 66, middle coxa; 67, hind coxa; 68, hind claw; 69, middle claw; 70, foreclaw; 71, hind claw; 72, middle claw; 73, foreclaw; 74, microtrichia on dorsum of operculate gill; 75 76, microtrichia on submarginal ventral row of operculate gill; 77, seta on posterior margin of hind tibia; 78, seta on posterior margin of hind tarsi; 79, setae on dorsal transverse row of fore femora; 80, microtrichia on dorsum of hind femora; 81, seta on hind tarsi; 82, seta of posterior margin of hind femora; 83, foreleg; 84, middle leg; 85, hind leg." httpUri="https://zenodo.org/record/179928/files/figure.png" pageId="2" pageNumber="3">74</figureCitation>
)........................... 5
</paragraph>
<paragraph id="8BFFA0234F20E615FF0287518789F99B" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
4. Abdominal sternum IX with a median U-shaped indentation on posterior margin (
<figureCitation id="137BBCA64F20E615FB3987518715F9B3" box="[1196,1286,1597,1623]" captionStart="FIGURES 31 49" captionStartId="11.[151,269,1819,1843]" captionTargetBox="[156,1417,356,1818]" captionTargetId="figure@11.[151,1436,328,1852]" captionTargetPageId="11" captionText="FIGURES 31 49. Caenis dominguezi, nymph. 31, labrum and detail of anteromedian emargination, d. v.; 32, setae on dorsum of hind femora; 33, seta on hind tibiae; 34, setae of fore femoral dorso-transversal row; 35, pronotum, d. v.; 36, fore leg; 37, middle leg; 38, hind leg; 39, middle coxa; 40, hind coxa; 41, foreclaw; 42, middle claw; 43, hind claw; 44, operculate gill and detail of dorsal surface; 45 46, microtrichia of submarginal ventral row of operculate gill; 47, abdominal gill III; 48, abdominal segment II and operculate gill, l. v.; 49, sternum IX of a male nymph. Abrevviations: OG = operculate gill; T 2 = second abdominal tergum." httpUri="https://zenodo.org/record/179926/files/figure.png" pageId="2" pageNumber="3">Fig. 49</figureCitation>
) [BO, ARG (Córdoba, Catamarca, Santiago, Salta)] ................................................................................
<taxonomicName id="4C40DBA04F20E615FB65870A8789F99B" box="[1264,1434,1638,1663]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dominguezi">
<emphasis id="B9347C314F20E615FB65870A8789F99B" box="[1264,1434,1638,1663]" italics="true" pageId="2" pageNumber="3">C. dominguezi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFFA0234F20E615FF0287E18789F92B" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
4´Abdominal sternum IX with straight or slightly rounded posterior margin [CHI, ARG (Neuquén, Río Negro)]
<emphasis id="B9347C314F20E615FEB187DA8789F92B" box="[292,1434,1718,1743]" italics="true" pageId="2" pageNumber="3">
...........................................................................................................................................
<taxonomicName id="4C40DBA04F20E615FAB087DA8789F92B" box="[1317,1434,1718,1743]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gonseri">C. gonseri</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BFFA0234F20E615FF0287B18789F8A3" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
5. Dorsal surface of body and operculate gills with large, scale-shaped microtrichia (
<figureCitation id="137BBCA64F20E615FB1B87B186CDF913" box="[1166,1246,1757,1783]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Figs. 8</figureCitation>
i-j in
<bibRefCitation id="EFD1DDD24F20E615FAB587B18315F8FB" author="Malzacher" pageId="2" pageNumber="31" refString="Malzacher, P. (2001) South and Central American Caenis species with rounded forceps tips (Insecta: Ephemeroptera: Caenidae). Stuttgarter Beitrage zur Naturkunde Ser. A, 626, 1 - 20." type="journal article" year="2001">Malzacher 2001</bibRefCitation>
); lateral margin of segment IX, besides short and apically frayed spines, with small teeth (
<figureCitation id="137BBCA64F20E615FABA86698767F8FB" box="[1327,1396,1797,1823]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Fig. 8</figureCitation>
l in
<bibRefCitation id="EFD1DDD24F20E615FF5286418396F8A3" author="Malzacher" box="[199,389,1837,1863]" pageId="2" pageNumber="31" refString="Malzacher, P. (2001) South and Central American Caenis species with rounded forceps tips (Insecta: Ephemeroptera: Caenidae). Stuttgarter Beitrage zur Naturkunde Ser. A, 626, 1 - 20." type="journal article" year="2001">Malzacher 2001</bibRefCitation>
) [BRA, ARG (Misiones)]
<emphasis id="B9347C314F20E615FD2A86428789F8A3" box="[703,1434,1838,1863]" italics="true" pageId="2" pageNumber="3">
..............................................................................
<taxonomicName id="4C40DBA04F20E615FA9586428789F8A3" box="[1280,1434,1838,1863]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="plaumanni">C. plaumanni</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BFFA0234F20E615FF028639878AF873" blockId="2.[151,1437,754,1943]" pageId="2" pageNumber="3">
5. Dorsal surface of body and operculate gills without or with small ramified or bifurcate microtrichia (
<figureCitation id="137BBCA64F20E615FAF4863982CBF873" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
,
<figureCitation id="137BBCA64F20E615FF7186118317F873" box="[228,260,1917,1943]" captionStart="FIGURES 66 85" captionStartId="16.[151,269,1664,1688]" captionTargetBox="[184,1411,262,1604]" captionTargetId="figure@16.[151,1436,244,1640]" captionTargetPageId="16" captionText="FIGURES 66 85. Caenis ludicra, nymph. 66, middle coxa; 67, hind coxa; 68, hind claw; 69, middle claw; 70, foreclaw; 71, hind claw; 72, middle claw; 73, foreclaw; 74, microtrichia on dorsum of operculate gill; 75 76, microtrichia on submarginal ventral row of operculate gill; 77, seta on posterior margin of hind tibia; 78, seta on posterior margin of hind tarsi; 79, setae on dorsal transverse row of fore femora; 80, microtrichia on dorsum of hind femora; 81, seta on hind tarsi; 82, seta of posterior margin of hind femora; 83, foreleg; 84, middle leg; 85, hind leg." httpUri="https://zenodo.org/record/179928/files/figure.png" pageId="2" pageNumber="3">74</figureCitation>
,
<figureCitation id="137BBCA64F20E615FE858611832CF873" box="[272,319,1917,1943]" captionStart="FIGURES 96 122" captionStartId="19.[151,269,1762,1786]" captionTargetBox="[233,1353,184,1724]" captionTargetId="figure@19.[230,1364,168,1753]" captionTargetPageId="19" captionText="FIGURES 96 122. Caenis uruzu. Male imago: 96, genitalia, v. v.; 97 98, details of forceps, v. v.; 99, lateral filaments of abdomen; 100, preesternal triangle; 101, detail of finger-like projection of abdominal tergum II, l. v. Nymph: 102, detail of anteromedian emargination of labrum; 103, labrum, d. v.; 104, detail of posteromedian projection of tergum II; 105, labium, v. v. (setae omitted on right side); 106 107, pronotal lateral margin; 108, middle coxa; 109, hind coxa; 110, operculate gill; 111, microtrichia on submarginal ventral row of operculate gill; 112, foreclaw; 113, middle claw; 114, hind claw; 115, setae on dorsum of middle and hind femora; 116, microtrichia on dorsum of operculate gill; 117, seta on posterior margin of hind tibia; 118, seta on hind tarsi; 119, setae on transverse dorsal row on fore femora; 120, fore leg; 121, middle leg; 122, hind leg." httpUri="https://zenodo.org/record/179930/files/figure.png" pageId="2" pageNumber="3">116</figureCitation>
); lateral margin of segment IX without small teeth .................................................................... 6
</paragraph>
<paragraph id="8BFFA0234F20E612FF0286B0878AFEE5" blockId="2.[151,1003,2010,2036]" lastBlockId="5.[151,1437,152,634]" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<keyStep id="30B4B8864F20E615FF0286B081F8F817" box="[151,1003,2010,2036]" pageId="2" pageNumber="3">
<keyLead id="30B103164F20E615FF0286B081F8F817" box="[151,1003,2010,2036]" pageId="2" pageNumber="3">1. Microtrichia can only be seen under high magnification (e.g., x100x400)</keyLead>
</keyStep>
<subSection id="E2CFB9C84F27E612FF0281F4878AFEE5" pageId="5" pageNumber="6" type="multiple">
6.
<collectingCountry id="F357E0B34F27E612FF5381F48312FF56" box="[198,257,152,178]" name="India" pageId="5" pageNumber="6">Hind</collectingCountry>
margin of the head with relatively long, laterally directed setae; abdominal sterna with blackish sublateral spots and other distinct marks (
<figureCitation id="137BBCA64F27E612FD3481AC811EFF3E" box="[673,781,192,218]" captionStart="FIGURES 179 184" captionStartId="28.[151,269,1290,1314]" captionTargetBox="[151,1435,178,1265]" captionTargetId="figure@28.[151,1436,160,1266]" captionTargetPageId="28" captionText="FIGURES 179 184. Nymph, abdominal sterna: 179, C. argentina; 180, C. burmeisteri; 181, C. dominguezi; 182, C. tenella; 183, C. plaumanni; 184, C. uruzu." httpUri="https://zenodo.org/record/179937/files/figure.png" pageId="5" pageNumber="6">Fig. 179</figureCitation>
); coxal processes short, semicircular or semielliptical (
<figureCitation id="137BBCA64F27E612FF5A8184832DFEE6" box="[207,318,232,258]" captionStart="FIGURES 1 14" captionStartId="7.[151,269,1465,1489]" captionTargetBox="[203,1404,193,1426]" captionTargetId="figure@7.[174,1413,181,1438]" captionTargetPageId="7" captionText="FIGURES 1 14. Caenis argentina, nymph. 1, middle coxa, d. v.; 2, hind coxa, d. v; 3, foreclaw; 4, middle claw; 5, hind claw; 6, microtrichia on dorsum of gill II; 7 8, microtrichia on ventral submarginal row of gill II; 9, setae on longitudinal row of middle and hind tarsi; 10, setae on subapical dorsal row of fore femora; 11, setae on longitudinal row of foretarsi; 12, foreleg; 13, middle leg; 14, hind leg." httpUri="https://zenodo.org/record/179924/files/figure.png" pageId="5" pageNumber="6">Figs. 12</figureCitation>
) [ARG (Buenos Aires, Santiago, Catamarca, Tucumán)]
<emphasis id="B9347C314F27E612FC498184878AFEE5" box="[988,1433,232,257]" italics="true" pageId="5" pageNumber="6">
........................................
<taxonomicName id="4C40DBA04F27E612FA928184878AFEE5" box="[1287,1433,232,257]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="argentina">C. argentina</taxonomicName>
</emphasis>
</subSection>
</paragraph>
</treatment>
</document>