treatments-xml/data/E3/60/F9/E360F967FFBB7665FC2242B4C2AEE8CC.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5070618" ID-GBIF-Dataset="c66531b3-6276-44e8-b917-9492b5e0bff4" ID-GBIF-Taxon="183322870" ID-ISSN="1982-0224" ID-Zenodo-Dep="5070618" checkinTime="1614362859122" checkinUser="carolina" docAuthor="Armbruster, Jonathan W. &amp; Page, Lawrence M." docDate="2006" docId="E360F967FFBB7665FC2242B4C2AEE8CC" docLanguage="en" docName="NeotropIchthyol.4.4.401-409.pdf" docOrigin="Neotropical Ichthyology 4 (4)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252006000400003&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Pterygoplichthys punctatus Gunther 1864" docType="treatment" docVersion="6" lastPageId="5" lastPageNumber="406" masterDocId="1F59811FFFB97660FFBD4039C074EF7B" masterDocTitle="Redescription of Pterygoplichthys punctatus and description of a new species of Pterygoplichthys (Siluriformes: Loricariidae)" masterLastPageNumber="409" masterPageNumber="401" pageId="2" pageNumber="403" updateTime="1636468849756" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Redescription of Pterygoplichthys punctatus and description of a new species of Pterygoplichthys (Siluriformes: Loricariidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Armbruster, Jonathan W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Page, Lawrence M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2006</mods:date>
<mods:detail type="pubDate">
<mods:number>2006-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>401</mods:start>
<mods:end>409</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252006000400003&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252006000400003</mods:identifier>
<mods:identifier type="GBIF-Dataset">c66531b3-6276-44e8-b917-9492b5e0bff4</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4566982</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5070618" ID-GBIF-Taxon="183322870" ID-Zenodo-Dep="5070618" LSID="urn:lsid:plazi:treatment:E360F967FFBB7665FC2242B4C2AEE8CC" httpUri="http://treatment.plazi.org/id/E360F967FFBB7665FC2242B4C2AEE8CC" lastPageId="5" lastPageNumber="406" pageId="2" pageNumber="403">
<subSubSection pageId="2" pageNumber="403" type="nomenclature">
<paragraph blockId="2.[927,1393,653,708]" pageId="2" pageNumber="403">
<taxonomicName authority="Gunther 1864" authorityName="Gunther" authorityYear="1864" box="[927,1393,653,677]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="punctatus">
<emphasis bold="true" box="[927,1393,653,677]" pageId="2" pageNumber="403">
<emphasis bold="true" box="[927,1228,653,677]" italics="true" pageId="2" pageNumber="403">Pterygoplichthys punctatus</emphasis>
Günther 1864
</emphasis>
</taxonomicName>
<figureCitation box="[1131,1188,685,708]" captionStart="Fig" captionStartId="3.[151,190,1586,1609]" captionTargetBox="[151,1474,191,1563]" captionTargetId="figure-198@3.[151,1475,191,1563]" captionTargetPageId="3" captionText="Fig. 2. Dorsal, lateral, and ventral views of Pterygoplichthys punctatus, MCP 35755, 203.1 mm SL. Photos by J.W.Armbruster." figureDoi="http://doi.org/10.5281/zenodo.4566988" httpUri="https://zenodo.org/record/4566988/files/figure.png" pageId="2" pageNumber="403">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="403" type="materials_examined">
<paragraph blockId="2.[839,1482,746,1371]" pageId="2" pageNumber="403">
<emphasis bold="true" box="[839,1047,746,767]" pageId="2" pageNumber="403">Material Examined.</emphasis>
<collectingCountry box="[1054,1116,746,767]" name="Brazil" pageId="2" pageNumber="403">Brazil</collectingCountry>
,
<collectingRegion box="[1127,1256,746,767]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="403">Mato Grosso</collectingRegion>
.
<accessionNumber box="[1265,1391,746,767]" httpUri="http://www.ncbi.nlm.nih.gov/protein/AUM42928" pageId="2" pageNumber="403">
<collectionCode box="[1265,1321,746,767]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="2" pageNumber="403" type="University or college">AUM</collectionCode>
42928
</accessionNumber>
,
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.1669" pageId="2" pageNumber="403" unit="mm" value="1166.9">1, 166.9 mm</quantity>
SL and
<accessionNumber box="[967,1082,774,795]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF148742" pageId="2" pageNumber="403">
<collectionCode box="[967,998,774,795]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="2" pageNumber="403" type="Museum">UF</collectionCode>
148742
</accessionNumber>
, 2,
<quantity box="[1122,1275,774,795]" metricMagnitude="-1" metricUnit="m" metricValue="1.193" metricValueMax="1.6009999999999998" metricValueMin="0.7849999999999999" pageId="2" pageNumber="403" unit="mm" value="119.3" valueMax="160.1" valueMin="78.5">78.5-160.1 mm</quantity>
SL.
<accessionNumber box="[1327,1450,774,795]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP35755" pageId="2" pageNumber="403">
<collectionCode box="[1327,1378,774,795]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="403">MCP</collectionCode>
35755
</accessionNumber>
,
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.2030999999999998" pageId="2" pageNumber="403" unit="mm" value="1203.1">1, 203.1 mm</quantity>
SL,
<collectingCountry box="[984,1046,803,824]" name="Brazil" pageId="2" pageNumber="403">Brazil</collectingCountry>
,
<collectingRegion box="[1056,1185,803,824]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="403">Mato Grosso</collectingRegion>
, rio Bugre (rio Madeira drainage), about
<quantity box="[945,1003,832,853]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="403" unit="km" value="42.0">42 km</quantity>
N of rio Guaporé on highway BR-174,
<geoCoordinate box="[1365,1477,832,853]" degrees="14" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="403" precision="15" seconds="35" value="-14.859722">14°5135&quot;S</geoCoordinate>
,
<geoCoordinate box="[839,975,861,882]" degrees="059" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="403" precision="15" seconds="57" value="-59.299168">059°1757&quot;W</geoCoordinate>
,
<date box="[986,1117,861,882]" pageId="2" pageNumber="403" value="2004-07-12">12 July 2004</date>
.
<accessionNumber box="[1133,1254,861,882]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP35753" pageId="2" pageNumber="403">
<collectionCode box="[1133,1183,861,882]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="403">MCP</collectionCode>
35753
</accessionNumber>
, 3,
<quantity box="[1290,1440,861,882]" metricMagnitude="-2" metricUnit="m" metricValue="9.985" metricValueMax="11.78" metricValueMin="8.19" pageId="2" pageNumber="403" unit="mm" value="99.85" valueMax="117.8" valueMin="81.9">81.9-117.8 mm</quantity>
SL, creek, tributary of rio Novo which is tributary to rio Guaporé (rio Maderia drainage) on highway BR-174,
<geoCoordinate box="[1228,1345,918,939]" degrees="14" direction="south" minutes="13" orientation="latitude" pageId="2" pageNumber="403" precision="15" seconds="26" value="-14.223888">14°1326&quot;S</geoCoordinate>
,
<geoCoordinate box="[1354,1477,918,939]" degrees="59" direction="west" minutes="41" orientation="longitude" pageId="2" pageNumber="403" precision="15" seconds="27" value="-59.690834">59°4127&quot;W</geoCoordinate>
,
<date box="[839,976,947,968]" pageId="2" pageNumber="403" value="2004-07-12">12 July 2004</date>
.
<accessionNumber box="[989,1114,947,968]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP35754" pageId="2" pageNumber="403">
<collectionCode box="[989,1040,947,968]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="403">MCP</collectionCode>
35754
</accessionNumber>
, 2, 156.6-
<quantity box="[1219,1323,947,968]" metricMagnitude="-1" metricUnit="m" metricValue="1.9649999999999999" pageId="2" pageNumber="403" unit="mm" value="196.5">196.5 mm</quantity>
SL and
<collectionCode box="[1416,1481,947,968]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="403">MNRJ</collectionCode>
28496,
<quantity box="[911,1032,976,997]" metricMagnitude="0" metricUnit="m" metricValue="1.1671" pageId="2" pageNumber="403" unit="mm" value="1167.1">1, 167.1 mm</quantity>
SL, creek, tributary of rio Pindaituba, circa
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="403" unit="km" value="27.0">27 km</quantity>
N rio Guaporé on highway BR-174,
<geoCoordinate box="[1227,1344,1005,1026]" degrees="14" direction="south" minutes="59" orientation="latitude" pageId="2" pageNumber="403" precision="15" seconds="53" value="-14.998055">14°5953&quot;S</geoCoordinate>
,
<geoCoordinate box="[1353,1477,1005,1026]" degrees="59" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="403" precision="15" seconds="10" value="-59.28611">59°1710&quot;W</geoCoordinate>
,
<date box="[839,967,1034,1055]" pageId="2" pageNumber="403" value="2004-07-12">12 July 2004</date>
.
<collectionCode box="[975,1036,1034,1055]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="2" pageNumber="403">ANSP</collectionCode>
180225, 2, 160.0-
<quantity box="[1214,1312,1034,1055]" metricMagnitude="-1" metricUnit="m" metricValue="1.833" pageId="2" pageNumber="403" unit="mm" value="183.3">183.3 mm</quantity>
SL,
<accessionNumber box="[1358,1477,1034,1055]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP35756" pageId="2" pageNumber="403">
<collectionCode box="[1358,1408,1034,1055]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="403">MCP</collectionCode>
35756
</accessionNumber>
, 6, 131.3-
<quantity box="[925,1025,1062,1083]" metricMagnitude="-1" metricUnit="m" metricValue="1.9649999999999999" pageId="2" pageNumber="403" unit="mm" value="196.5">196.5 mm</quantity>
SL,
<collectionCode box="[1071,1136,1062,1083]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="403">MNRJ</collectionCode>
28497, 2, 165.8-
<quantity box="[1301,1400,1062,1083]" metricMagnitude="-1" metricUnit="m" metricValue="1.841" pageId="2" pageNumber="403" unit="mm" value="184.1">184.1 mm</quantity>
SL, and
<accessionNumber box="[839,950,1091,1112]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/UF149606" pageId="2" pageNumber="403">
<collectionCode box="[839,870,1091,1112]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="2" pageNumber="403" type="Museum">UF</collectionCode>
149606
</accessionNumber>
, 2, 154.3-
<quantity box="[1044,1143,1091,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.7719999999999998" pageId="2" pageNumber="403" unit="mm" value="177.2">177.2 mm</quantity>
SL, small river (tributary of the rio Guaporé) on highway BR-174, near Pontes e Lacerda and Comodoro,
<geoCoordinate box="[964,1086,1149,1170]" degrees="14" direction="south" minutes="55" orientation="latitude" pageId="2" pageNumber="403" precision="15" seconds="15" value="-14.920834">14°5515&quot;S</geoCoordinate>
,
<geoCoordinate box="[1100,1230,1149,1170]" degrees="59" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="403" precision="15" seconds="29" value="-59.29139">59°1729&quot;W</geoCoordinate>
,
<date box="[1244,1386,1149,1170]" pageId="2" pageNumber="403" value="2004-07-12">12 July 2004</date>
.
<collectionCode box="[1410,1482,1149,1170]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="403" type="Museum">USNM</collectionCode>
177716, 1,
<collectingCountry box="[953,1015,1178,1199]" name="Brazil" pageId="2" pageNumber="403">Brazil</collectingCountry>
, rio Urubu, 1958.
<accessionNumber box="[1210,1343,1178,1199]" httpUri="http://www.ncbi.nlm.nih.gov/protein/NMW76587" pageId="2" pageNumber="403">
<collectionCode box="[1210,1272,1178,1199]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="403">NMW</collectionCode>
76587
</accessionNumber>
, 1 (
<typeStatus box="[1383,1478,1178,1199]" pageId="2" pageNumber="403">Holotype</typeStatus>
, photographs provided by H. Wellendorf, not measured), rio Guaporé, Amazon system, San Vincente, rio Madeira drainage,
<collectingCountry name="Brazil" pageId="2" pageNumber="403">Brazil</collectingCountry>
,
<date box="[875,1025,1264,1285]" pageId="2" pageNumber="403" value="1827-01-08">8 January 1827</date>
, (= Arraial de São Vincente, a village that is no longer extant in northern
<collectingRegion box="[1089,1219,1293,1314]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="403">Mato Grosso</collectingRegion>
;
<bibRefCitation author="Papavero, N" box="[1229,1382,1293,1314]" pageId="2" pageNumber="403" refId="ref6504" refString="Papavero, N. 1971. Essays on the history of Neotropical dipterology, with special reference to collectors (1750 - 1905). Vol. I. Museu de Zoologia, Universidade de Sao Paulo, 216 pp." type="book" year="1971">
Papavero,
<quantity box="[1333,1382,1293,1314]" metricMagnitude="1" metricUnit="m" metricValue="5.00634" pageId="2" pageNumber="403" unit="in" value="1971.0">1971</quantity>
</bibRefCitation>
in
<bibRefCitation author="Weber, C" pageId="2" pageNumber="403" pagination="1 - 36" refId="ref6617" refString="Weber, C. 1992. Revision du genre Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae. Revue Francaise d'Aquariologie Herpetologie, 19: 1 - 36." type="journal article" year="1992">Weber, 1992</bibRefCitation>
).
<collectingCountry box="[910,985,1322,1343]" name="Bolivia" pageId="2" pageNumber="403">Bolivia</collectingCountry>
.
<collectionCode box="[997,1049,1322,1343]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="2" pageNumber="403">ZMA</collectionCode>
109.199,
<quantity box="[1152,1281,1322,1343]" metricMagnitude="0" metricUnit="m" metricValue="1.1248" pageId="2" pageNumber="403" unit="mm" value="1124.8">1, 124.8 mm</quantity>
SL, south of Todos Santos, in pool of standing water,
<date box="[1173,1320,1350,1371]" pageId="2" pageNumber="403" value="1966-08-08">8 August 1966</date>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403" type="diagnosis">
<paragraph blockId="2.[839,1482,1407,1943]" pageId="2" pageNumber="403">
<emphasis bold="true" box="[839,960,1408,1431]" pageId="2" pageNumber="403">Diagnosis.</emphasis>
<taxonomicName authorityName="Gunther" authorityYear="1864" box="[971,1279,1407,1432]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[971,1279,1407,1432]" italics="true" pageId="2" pageNumber="403">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
can be separated from all other
<taxonomicName authorityName="Gill" authorityYear="1858" box="[996,1179,1439,1463]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="genus">
<emphasis box="[996,1179,1439,1463]" italics="true" pageId="2" pageNumber="403">Pterygoplichthys</emphasis>
</taxonomicName>
except
<taxonomicName authorityName="Armbruster &amp; Page" authorityYear="2006" box="[1265,1365,1439,1464]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="weberi">
<emphasis box="[1265,1365,1439,1464]" italics="true" pageId="2" pageNumber="403">P. weberi</emphasis>
</taxonomicName>
by having the buccal papilla divided medially and lobulate (
<emphasis box="[1382,1412,1473,1496]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
entire and simple,
<figureCitation box="[977,1044,1504,1527]" captionStart="Fig" captionStartId="1.[831,870,1826,1849]" captionTargetBox="[831,1474,653,1797]" captionTargetId="figure-567@1.[831,1474,653,1797]" captionTargetPageId="1" captionText="Fig. 1. Mouth and buccal papilla (indicated by arrow) of a. Pterygoplichthys punctatus, MCP 35755, 203.1 mm SL, b. P. weberi, FMNH 96959, holotype, 183.9 mm SL, c. P. multiradiatus, AUM 18947, 153.3 mm SL. Photos by J.W. Armbruster and L.M. Page." figureDoi="http://doi.org/10.5281/zenodo.4566984" httpUri="https://zenodo.org/record/4566984/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
, or lobulate but not divided medially).
<taxonomicName authorityName="Gunther" authorityYear="1864" box="[839,1133,1535,1560]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[839,1133,1535,1560]" italics="true" pageId="2" pageNumber="403">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
can be separated from
<taxonomicName authorityName="Armbruster &amp; Page" authorityYear="2006" box="[1382,1481,1535,1560]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="weberi">
<emphasis box="[1382,1481,1535,1560]" italics="true" pageId="2" pageNumber="403">P. weberi</emphasis>
</taxonomicName>
by having the buccal papilla deeply divided at all ages (
<emphasis box="[1452,1481,1569,1592]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
shallowly divided in adult, more deeply divided in juvenile), by having many spots on the abdomen and the fin bases (
<emphasis box="[1452,1481,1633,1656]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
few spots), by having a narrower body (SL/cleithral width 3.6-4.0
<emphasis box="[921,950,1697,1720]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
3.3-3.4), and by having the lateral keel odontodes sloped posteriorly (
<emphasis box="[1055,1085,1729,1752]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
almost perpendicular to the plates) and shorter than the lateral-line plates (
<emphasis box="[1247,1276,1761,1784]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
longer than lateralline plates in adult).
</paragraph>
<paragraph blockId="2.[839,1482,1407,1943]" lastBlockId="3.[831,1474,1665,1721]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<taxonomicName authorityName="Gunther" authorityYear="1864" box="[876,1167,1823,1848]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[876,1167,1823,1848]" italics="true" pageId="2" pageNumber="403">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
can be further separated from
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1874" box="[839,968,1855,1880]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="scrophus">
<emphasis box="[839,855,1857,1880]" italics="true" pageId="2" pageNumber="403">P</emphasis>
.
<emphasis box="[869,968,1855,1879]" italics="true" pageId="2" pageNumber="403">scrophus</emphasis>
</taxonomicName>
<emphasis box="[968,1407,1855,1880]" italics="true" pageId="2" pageNumber="403">
,
<taxonomicName baseAuthorityName="Kner" baseAuthorityYear="1854" box="[980,1114,1855,1880]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="gibbiceps">P. gibbiceps</taxonomicName>
,
<taxonomicName baseAuthorityName="Weber" baseAuthorityYear="1991" box="[1126,1270,1857,1880]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="xinguensis">P. xinguensis</taxonomicName>
,
<taxonomicName baseAuthorityName="Kner" baseAuthorityYear="1854" box="[1282,1403,1855,1880]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="lituratus">P. lituratus</taxonomicName>
,
</emphasis>
and
<taxonomicName baseAuthorityName="Weber" baseAuthorityYear="1991" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="parnaibae">
<emphasis italics="true" pageId="2" pageNumber="403">P. parnaibae</emphasis>
</taxonomicName>
by lacking an elevated supraoccipital crest (
<emphasis box="[1452,1481,1889,1912]" italics="true" pageId="2" pageNumber="403">vs.</emphasis>
having an elevated supraoccipital forming an obvious crest higher than the plates in the nuchal region); from
<taxonomicName baseAuthorityName="Spix &amp; Agassiz" baseAuthorityYear="1829" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="etentaculatus">
<emphasis italics="true" pageId="3" pageNumber="404">P. etentaculatus</emphasis>
</taxonomicName>
by having fewer than five hypertrophied odontodes on the cheek plates (
<emphasis box="[494,524,1730,1753]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
more than five); from
<taxonomicName baseAuthorityName="Weber" baseAuthorityYear="1991" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="joselimaianus">
<emphasis italics="true" pageId="3" pageNumber="404">P. joselimaianus</emphasis>
</taxonomicName>
by having dark spots (
<emphasis box="[545,575,1762,1785]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
light spots); from
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1878" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="undecimalis">
<emphasis italics="true" pageId="3" pageNumber="404">P. undecimalis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Weber" authorityYear="1991" box="[343,481,1793,1817]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="zuliaensis">
<emphasis box="[343,481,1793,1817]" italics="true" pageId="3" pageNumber="404">P. zuliaensis</emphasis>
</taxonomicName>
by having a broadly round snout (
<emphasis box="[224,254,1826,1849]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
a narrow pointed snout,
<figureCitation box="[518,582,1826,1849]" captionStart="Fig" captionStartId="4.[158,197,1922,1945]" captionTargetBox="[156,1475,1202,1882]" captionTargetId="figure-639@4.[136,1497,1188,1895]" captionTargetPageId="4" captionText="Fig. 3. Dorsal profiles of snouts of a. Pterygoplichthys punctatus, UF 148742, 160.1 mm SL and b. P. zuliaensis,AUM 22586, 222.0mmSL." figureDoi="http://doi.org/10.5281/zenodo.4566992" httpUri="https://zenodo.org/record/4566992/files/figure.png" pageId="3" pageNumber="404">Fig. 3</figureCitation>
); from
<taxonomicName authorityName="Weber" authorityYear="1991" box="[661,794,1825,1849]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="zuliaensis">
<emphasis box="[661,794,1825,1849]" italics="true" pageId="3" pageNumber="404">P. zuliaensis</emphasis>
</taxonomicName>
by having a greater HL to interorbital width ratio (2.3-2.6
<emphasis box="[764,794,1858,1881]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
2.0-2.2); from
<taxonomicName baseAuthorityName="Hancock" baseAuthorityYear="1828" box="[309,482,1889,1913]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="multiradiatus">
<emphasis box="[309,482,1889,1913]" italics="true" pageId="3" pageNumber="404">P. multiradiatus</emphasis>
</taxonomicName>
by having usually 11 dorsalfin rays (only 3 of 27 with 12
<emphasis box="[456,485,1922,1945]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
12-13
<emphasis box="[557,597,1921,1945]" italics="true" pageId="3" pageNumber="404">fide</emphasis>
<bibRefCitation author="Weber, C" box="[602,738,1922,1945]" pageId="3" pageNumber="404" pagination="1 - 36" refId="ref6617" refString="Weber, C. 1992. Revision du genre Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae. Revue Francaise d'Aquariologie Herpetologie, 19: 1 - 36." type="journal article" year="1992">Weber, 1992</bibRefCitation>
); and from
<emphasis box="[215,712,1953,1977]" italics="true" pageId="3" pageNumber="404">
<taxonomicName authorityName="Eigenmann &amp; Kennedy" authorityYear="1903" box="[215,328,1954,1977]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="anisitsi">P. anisitsi</taxonomicName>
,
<taxonomicName baseAuthorityName="Weber" baseAuthorityYear="1991" box="[344,509,1953,1977]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="disjunctivus">P. disjunctivus</taxonomicName>
,
<taxonomicName baseAuthorityName="Hancock" baseAuthorityYear="1828" box="[525,708,1953,1977]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="multiradiatus">P. multiradiatus</taxonomicName>
,
</emphasis>
and
<taxonomicName baseAuthorityName="Castelnau" baseAuthorityYear="1855" box="[773,921,1665,1977]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="pardalis">
<emphasis box="[773,921,1665,1977]" italics="true" pageId="3" pageNumber="404">P. pardalis</emphasis>
</taxonomicName>
by having hypertrophied odontodes on the cheek plates of the adult (
<emphasis box="[1041,1071,1698,1721]" italics="true" pageId="3" pageNumber="404">vs.</emphasis>
no hypertrophied cheek odontodes).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566988" ID-Zenodo-Dep="4566988" box="[151,1475,1585,1609]" httpUri="https://zenodo.org/record/4566988/files/figure.png" pageId="3" pageNumber="404" startId="3.[151,190,1586,1609]" targetBox="[151,1474,191,1563]" targetPageId="3">
<paragraph blockId="3.[151,1475,1585,1609]" box="[151,1475,1585,1609]" pageId="3" pageNumber="404">
<emphasis bold="true" box="[151,219,1586,1609]" pageId="3" pageNumber="404">Fig. 2.</emphasis>
Dorsal, lateral, and ventral views of
<taxonomicName authority=", MCP" authorityName="MCP" box="[599,953,1585,1609]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[599,887,1585,1609]" italics="true" pageId="3" pageNumber="404">Pterygoplichthys punctatus</emphasis>
, MCP
</taxonomicName>
35755, 203.1 mm SL. Photos by J.W. Armbruster.
</paragraph>
</caption>
<subSubSection lastPageId="5" lastPageNumber="406" pageId="3" pageNumber="404" type="description">
<paragraph blockId="3.[831,1475,1762,1977]" lastBlockId="4.[158,801,834,1145]" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404">
<emphasis bold="true" box="[831,971,1762,1785]" pageId="3" pageNumber="404">Description.</emphasis>
Member of
<taxonomicName box="[1099,1259,1762,1785]" class="Actinopterygii" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="subFamily" subFamily="Hypostominae">Hypostominae</taxonomicName>
:
<taxonomicName authorityName="Armbruster" authorityYear="2004" box="[1267,1471,1762,1785]" class="Actinopterygii" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="401" phylum="Chordata" rank="tribe" tribe="Pterygoplichthyini">Pterygoplichthyini</taxonomicName>
:
<taxonomicName authorityName="Gill" authorityYear="1858" box="[831,1014,1793,1817]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
<emphasis box="[831,1014,1793,1817]" italics="true" pageId="3" pageNumber="404">Pterygoplichthys</emphasis>
</taxonomicName>
as diagnosed by
<bibRefCitation author="Armbruster, J" box="[1197,1403,1794,1817]" pageId="3" pageNumber="404" pagination="1 - 80" refId="ref6335" refString="Armbruster, J. W. 2004. Phylogenetic relationships of the suckermouth armored catfishes (Loricariidae) with emphasis on the Hypostominae and the Ancistrinae. Zoological Journal of the Linnean Society, 141: 1 - 80." type="journal article" year="2004">Armbruster (2004)</bibRefCitation>
. Morphometric data given in
<tableCitation box="[1085,1164,1826,1849]" captionStart="Table 1" captionStartId="4.[159,222,197,220]" captionTargetPageId="4" captionText="Table 1. Selected morphometrics of Pterygoplichthys punctatus (N=27 except D/K, N=24) and P. weberi (N=11 except D/K, N=7). The letters refer to codes for measurements in Boeseman (1968)." httpUri="http://table.plazi.org/id/3FB618F9FFBD7664FF2240FCC3E7EF87" pageId="3" pageNumber="404" tableUuid="3FB618F9FFBD7664FF2240FCC3E7EF87">Table 1</tableCitation>
. Largest specimen
<quantity box="[1367,1474,1826,1849]" metricMagnitude="-1" metricUnit="m" metricValue="2.0509999999999997" pageId="3" pageNumber="404" unit="mm" value="205.1">205.1 mm</quantity>
SL. Head forming arch from tip of snout to anterior margin of supraoccipital. Supraoccipital and nuchal region forming hump. Body depth decreases from origin of dorsal fin to dorsal procurrent caudal spines, and then increases to caudal fin. Ridge from anterodorsal corner of orbit runs ventral to nares, ending slightly anteroventral of anterior nare. Crest continuing posteriorly from supraorbital ridge with portion on sphenotic and pterotic bent strongly dorsally, posterior portion on pterotic angled dorsally, continuous with keel of mid-dorsal plate series. Dorsal rim of orbit slightly higher than interorbital space.
</paragraph>
<caption ID-Table-UUID="3FB618F9FFBD7664FF2240FCC3E7EF87" httpUri="http://table.plazi.org/id/3FB618F9FFBD7664FF2240FCC3E7EF87" pageId="4" pageNumber="405" startId="4.[159,222,197,220]" targetBox="[198,1433,278,790]" targetIsTable="true" targetPageId="4">
<paragraph blockId="4.[159,1482,196,252]" pageId="4" pageNumber="405">
<emphasis bold="true" box="[159,247,197,220]" pageId="4" pageNumber="405">Table 1.</emphasis>
Selected morphometrics of
<taxonomicName box="[550,842,196,220]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[550,842,196,220]" italics="true" pageId="4" pageNumber="405">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
(N=27 except D/K, N=24) and
<taxonomicName authorityName="Armbruster &amp; Page" authorityYear="2006" box="[1176,1274,196,220]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="weberi">
<emphasis box="[1176,1274,196,220]" italics="true" pageId="4" pageNumber="405">P. weberi</emphasis>
</taxonomicName>
(N=11 except D/K, N=7). The letters refer to codes for measurements in
<bibRefCitation author="Boeseman, M" box="[720,911,229,252]" pageId="4" pageNumber="405" pagination="1 - 89" refId="ref6423" refString="Boeseman, M. 1968. The genus Hypostomus Lacepede, 1803, and its Surinam representatives (Siluriformes, Loricariidae). Zoologische Verhandelingen (Leiden), 99: 1 - 89." type="journal article" year="1968">Boeseman (1968)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="405">
<table box="[198,1433,278,790]" gridcols="13" gridrows="21" pageId="4" pageNumber="405">
<tr box="[198,1433,278,298]" gridrow="0" pageId="4" pageNumber="405" rowspan-0="1">
<th box="[677,1035,278,298]" colspan="6" colspanRight="5" gridcol="1" gridrow="0" pageId="4" pageNumber="405">
<taxonomicName box="[732,979,278,298]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[732,979,278,298]" italics="true" pageId="4" pageNumber="405">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
</th>
<th box="[1075,1433,278,298]" colspan="6" colspanRight="5" gridcol="7" gridrow="0" pageId="4" pageNumber="405">
<taxonomicName authorityName="Armbruster &amp; Page" authorityYear="2006" box="[1143,1364,278,298]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="weberi">
<emphasis box="[1143,1364,278,298]" italics="true" pageId="4" pageNumber="405">Pterygoplichthys weberi</emphasis>
</taxonomicName>
</th>
</tr>
<tr box="[198,1433,303,322]" gridrow="1" pageId="4" pageNumber="405" rowspan-0="1">
<td box="[677,841,303,322]" colspan="3" colspanRight="2" gridcol="1" gridrow="1" pageId="4" pageNumber="405">Average ± SD</td>
<td box="[875,1035,303,322]" colspan="3" colspanRight="2" gridcol="4" gridrow="1" pageId="4" pageNumber="405">Range</td>
<td box="[1075,1239,303,322]" colspan="3" colspanRight="2" gridcol="7" gridrow="1" pageId="4" pageNumber="405">Average ± SD</td>
<td box="[1273,1433,303,322]" colspan="3" colspanRight="2" gridcol="10" gridrow="1" pageId="4" pageNumber="405">Range</td>
</tr>
<tr box="[198,1433,329,348]" gridrow="2" pageId="4" pageNumber="405">
<th box="[198,564,329,348]" gridcol="0" gridrow="2" pageId="4" pageNumber="405">SL (mm)</th>
<td box="[677,727,329,348]" gridcol="1" gridrow="2" pageId="4" pageNumber="405">145.7</td>
<td box="[758,771,329,348]" gridcol="2" gridrow="2" pageId="4" pageNumber="405">±</td>
<td box="[801,841,329,348]" gridcol="3" gridrow="2" pageId="4" pageNumber="405">43.3</td>
<td box="[875,915,329,348]" gridcol="4" gridrow="2" pageId="4" pageNumber="405">77.6</td>
<td box="[946,953,329,348]" gridcol="5" gridrow="2" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,329,348]" gridcol="6" gridrow="2" pageId="4" pageNumber="405">205.1</td>
<td box="[1075,1126,329,348]" gridcol="7" gridrow="2" pageId="4" pageNumber="405">149.6</td>
<td box="[1156,1169,329,348]" gridcol="8" gridrow="2" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,329,348]" gridcol="9" gridrow="2" pageId="4" pageNumber="405">42.7</td>
<td box="[1273,1313,329,348]" gridcol="10" gridrow="2" pageId="4" pageNumber="405">86.6</td>
<td box="[1344,1351,329,348]" gridcol="11" gridrow="2" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,329,348]" gridcol="12" gridrow="2" pageId="4" pageNumber="405">196.9</td>
</tr>
<tr box="[198,1433,354,373]" gridrow="3" pageId="4" pageNumber="405">
<th box="[198,564,354,373]" gridcol="0" gridrow="3" pageId="4" pageNumber="405">SL/Predorsal L. (SL/D)</th>
<td box="[677,727,354,373]" gridcol="1" gridrow="3" pageId="4" pageNumber="405">2.46</td>
<td box="[758,771,354,373]" gridcol="2" gridrow="3" pageId="4" pageNumber="405">±</td>
<td box="[801,841,354,373]" gridcol="3" gridrow="3" pageId="4" pageNumber="405">0.1</td>
<td box="[875,915,354,373]" gridcol="4" gridrow="3" pageId="4" pageNumber="405">2.29</td>
<td box="[946,953,354,373]" gridcol="5" gridrow="3" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,354,373]" gridcol="6" gridrow="3" pageId="4" pageNumber="405">2.65</td>
<td box="[1075,1126,354,373]" gridcol="7" gridrow="3" pageId="4" pageNumber="405">2.38</td>
<td box="[1156,1169,354,373]" gridcol="8" gridrow="3" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,354,373]" gridcol="9" gridrow="3" pageId="4" pageNumber="405">0.06</td>
<td box="[1273,1313,354,373]" gridcol="10" gridrow="3" pageId="4" pageNumber="405">2.30</td>
<td box="[1344,1351,354,373]" gridcol="11" gridrow="3" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,354,373]" gridcol="12" gridrow="3" pageId="4" pageNumber="405">2.52</td>
</tr>
<tr box="[198,1433,378,397]" gridrow="4" pageId="4" pageNumber="405">
<th box="[198,564,378,397]" gridcol="0" gridrow="4" pageId="4" pageNumber="405">SL/HL (SL/E)</th>
<td box="[677,727,378,397]" gridcol="1" gridrow="4" pageId="4" pageNumber="405">3.2</td>
<td box="[758,771,378,397]" gridcol="2" gridrow="4" pageId="4" pageNumber="405">±</td>
<td box="[801,841,378,397]" gridcol="3" gridrow="4" pageId="4" pageNumber="405">0.17</td>
<td box="[875,915,378,397]" gridcol="4" gridrow="4" pageId="4" pageNumber="405">2.89</td>
<td box="[946,953,378,397]" gridcol="5" gridrow="4" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,378,397]" gridcol="6" gridrow="4" pageId="4" pageNumber="405">3.42</td>
<td box="[1075,1126,378,397]" gridcol="7" gridrow="4" pageId="4" pageNumber="405">3.15</td>
<td box="[1156,1169,378,397]" gridcol="8" gridrow="4" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,378,397]" gridcol="9" gridrow="4" pageId="4" pageNumber="405">0.24</td>
<td box="[1273,1313,378,397]" gridcol="10" gridrow="4" pageId="4" pageNumber="405">2.74</td>
<td box="[1344,1351,378,397]" gridcol="11" gridrow="4" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,378,397]" gridcol="12" gridrow="4" pageId="4" pageNumber="405">3.65</td>
</tr>
<tr box="[198,1433,403,422]" gridrow="5" pageId="4" pageNumber="405">
<th box="[198,564,403,422]" gridcol="0" gridrow="5" pageId="4" pageNumber="405">SL/Cleithral W. (SL/F)</th>
<td box="[677,727,403,422]" gridcol="1" gridrow="5" pageId="4" pageNumber="405">3.72</td>
<td box="[758,771,403,422]" gridcol="2" gridrow="5" pageId="4" pageNumber="405">±</td>
<td box="[801,841,403,422]" gridcol="3" gridrow="5" pageId="4" pageNumber="405">0.16</td>
<td box="[875,915,403,422]" gridcol="4" gridrow="5" pageId="4" pageNumber="405">3.5</td>
<td box="[946,953,403,422]" gridcol="5" gridrow="5" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,403,422]" gridcol="6" gridrow="5" pageId="4" pageNumber="405">3.9</td>
<td box="[1075,1126,403,422]" gridcol="7" gridrow="5" pageId="4" pageNumber="405">3.32</td>
<td box="[1156,1169,403,422]" gridcol="8" gridrow="5" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,403,422]" gridcol="9" gridrow="5" pageId="4" pageNumber="405">0.1</td>
<td box="[1273,1313,403,422]" gridcol="10" gridrow="5" pageId="4" pageNumber="405">3.09</td>
<td box="[1344,1351,403,422]" gridcol="11" gridrow="5" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,403,422]" gridcol="12" gridrow="5" pageId="4" pageNumber="405">3.42</td>
</tr>
<tr box="[198,1433,428,447]" gridrow="6" pageId="4" pageNumber="405">
<th box="[198,564,428,447]" gridcol="0" gridrow="6" pageId="4" pageNumber="405">HL/Cleithral W. (E/F)</th>
<td box="[677,727,428,447]" gridcol="1" gridrow="6" pageId="4" pageNumber="405">1.16</td>
<td box="[758,771,428,447]" gridcol="2" gridrow="6" pageId="4" pageNumber="405">±</td>
<td box="[801,841,428,447]" gridcol="3" gridrow="6" pageId="4" pageNumber="405">0.03</td>
<td box="[875,915,428,447]" gridcol="4" gridrow="6" pageId="4" pageNumber="405">1.11</td>
<td box="[946,953,428,447]" gridcol="5" gridrow="6" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,428,447]" gridcol="6" gridrow="6" pageId="4" pageNumber="405">1.21</td>
<td box="[1075,1126,428,447]" gridcol="7" gridrow="6" pageId="4" pageNumber="405">1.06</td>
<td box="[1156,1169,428,447]" gridcol="8" gridrow="6" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,428,447]" gridcol="9" gridrow="6" pageId="4" pageNumber="405">0.06</td>
<td box="[1273,1313,428,447]" gridcol="10" gridrow="6" pageId="4" pageNumber="405">0.94</td>
<td box="[1344,1351,428,447]" gridcol="11" gridrow="6" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,428,447]" gridcol="12" gridrow="6" pageId="4" pageNumber="405">1.17</td>
</tr>
<tr box="[198,1433,452,471]" gridrow="7" pageId="4" pageNumber="405">
<th box="[198,564,452,471]" gridcol="0" gridrow="7" pageId="4" pageNumber="405">HL/Head D. (E/G)</th>
<td box="[677,727,452,471]" gridcol="1" gridrow="7" pageId="4" pageNumber="405">1.44</td>
<td box="[758,771,452,471]" gridcol="2" gridrow="7" pageId="4" pageNumber="405">±</td>
<td box="[801,841,452,471]" gridcol="3" gridrow="7" pageId="4" pageNumber="405">0.07</td>
<td box="[875,915,452,471]" gridcol="4" gridrow="7" pageId="4" pageNumber="405">1.35</td>
<td box="[946,953,452,471]" gridcol="5" gridrow="7" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,452,471]" gridcol="6" gridrow="7" pageId="4" pageNumber="405">1.56</td>
<td box="[1075,1126,452,471]" gridcol="7" gridrow="7" pageId="4" pageNumber="405">1.41</td>
<td box="[1156,1169,452,471]" gridcol="8" gridrow="7" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,452,471]" gridcol="9" gridrow="7" pageId="4" pageNumber="405">0.09</td>
<td box="[1273,1313,452,471]" gridcol="10" gridrow="7" pageId="4" pageNumber="405">1.27</td>
<td box="[1344,1351,452,471]" gridcol="11" gridrow="7" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,452,471]" gridcol="12" gridrow="7" pageId="4" pageNumber="405">1.55</td>
</tr>
<tr box="[198,1433,477,496]" gridrow="8" pageId="4" pageNumber="405">
<th box="[198,564,477,496]" gridcol="0" gridrow="8" pageId="4" pageNumber="405">HL/Snout L. (E/H)</th>
<td box="[677,727,477,496]" gridcol="1" gridrow="8" pageId="4" pageNumber="405">1.92</td>
<td box="[758,771,477,496]" gridcol="2" gridrow="8" pageId="4" pageNumber="405">±</td>
<td box="[801,841,477,496]" gridcol="3" gridrow="8" pageId="4" pageNumber="405">0.08</td>
<td box="[875,915,477,496]" gridcol="4" gridrow="8" pageId="4" pageNumber="405">1.82</td>
<td box="[946,953,477,496]" gridcol="5" gridrow="8" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,477,496]" gridcol="6" gridrow="8" pageId="4" pageNumber="405">2.02</td>
<td box="[1075,1126,477,496]" gridcol="7" gridrow="8" pageId="4" pageNumber="405">1.88</td>
<td box="[1156,1169,477,496]" gridcol="8" gridrow="8" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,477,496]" gridcol="9" gridrow="8" pageId="4" pageNumber="405">0.08</td>
<td box="[1273,1313,477,496]" gridcol="10" gridrow="8" pageId="4" pageNumber="405">1.80</td>
<td box="[1344,1351,477,496]" gridcol="11" gridrow="8" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,477,496]" gridcol="12" gridrow="8" pageId="4" pageNumber="405">2.04</td>
</tr>
<tr box="[198,1433,501,520]" gridrow="9" pageId="4" pageNumber="405">
<th box="[198,564,501,520]" gridcol="0" gridrow="9" pageId="4" pageNumber="405">HL/Orbit Dia. (E/I)</th>
<td box="[677,727,501,520]" gridcol="1" gridrow="9" pageId="4" pageNumber="405">7.61</td>
<td box="[758,771,501,520]" gridcol="2" gridrow="9" pageId="4" pageNumber="405">±</td>
<td box="[801,841,501,520]" gridcol="3" gridrow="9" pageId="4" pageNumber="405">0.59</td>
<td box="[875,915,501,520]" gridcol="4" gridrow="9" pageId="4" pageNumber="405">6.71</td>
<td box="[946,953,501,520]" gridcol="5" gridrow="9" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,501,520]" gridcol="6" gridrow="9" pageId="4" pageNumber="405">8.68</td>
<td box="[1075,1126,501,520]" gridcol="7" gridrow="9" pageId="4" pageNumber="405">6.98</td>
<td box="[1156,1169,501,520]" gridcol="8" gridrow="9" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,501,520]" gridcol="9" gridrow="9" pageId="4" pageNumber="405">0.59</td>
<td box="[1273,1313,501,520]" gridcol="10" gridrow="9" pageId="4" pageNumber="405">5.72</td>
<td box="[1344,1351,501,520]" gridcol="11" gridrow="9" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,501,520]" gridcol="12" gridrow="9" pageId="4" pageNumber="405">7.56</td>
</tr>
<tr box="[198,1433,526,545]" gridrow="10" pageId="4" pageNumber="405">
<th box="[198,564,526,545]" gridcol="0" gridrow="10" pageId="4" pageNumber="405">HL/Interorbital W. (E/J)</th>
<td box="[677,727,526,545]" gridcol="1" gridrow="10" pageId="4" pageNumber="405">2.36</td>
<td box="[758,771,526,545]" gridcol="2" gridrow="10" pageId="4" pageNumber="405">±</td>
<td box="[801,841,526,545]" gridcol="3" gridrow="10" pageId="4" pageNumber="405">0.08</td>
<td box="[875,915,526,545]" gridcol="4" gridrow="10" pageId="4" pageNumber="405">2.19</td>
<td box="[946,953,526,545]" gridcol="5" gridrow="10" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,526,545]" gridcol="6" gridrow="10" pageId="4" pageNumber="405">2.44</td>
<td box="[1075,1126,526,545]" gridcol="7" gridrow="10" pageId="4" pageNumber="405">2.48</td>
<td box="[1156,1169,526,545]" gridcol="8" gridrow="10" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,526,545]" gridcol="9" gridrow="10" pageId="4" pageNumber="405">0.11</td>
<td box="[1273,1313,526,545]" gridcol="10" gridrow="10" pageId="4" pageNumber="405">2.36</td>
<td box="[1344,1351,526,545]" gridcol="11" gridrow="10" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,526,545]" gridcol="12" gridrow="10" pageId="4" pageNumber="405">2.71</td>
</tr>
<tr box="[198,1433,550,569]" gridrow="11" pageId="4" pageNumber="405">
<th box="[198,564,550,569]" gridcol="0" gridrow="11" pageId="4" pageNumber="405">Predorsal L./Dorsal Spine L. (D/K)</th>
<td box="[677,727,550,569]" gridcol="1" gridrow="11" pageId="4" pageNumber="405">1.47</td>
<td box="[758,771,550,569]" gridcol="2" gridrow="11" pageId="4" pageNumber="405">±</td>
<td box="[801,841,550,569]" gridcol="3" gridrow="11" pageId="4" pageNumber="405">0.06</td>
<td box="[875,915,550,569]" gridcol="4" gridrow="11" pageId="4" pageNumber="405">1.41</td>
<td box="[946,953,550,569]" gridcol="5" gridrow="11" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,550,569]" gridcol="6" gridrow="11" pageId="4" pageNumber="405">1.55</td>
<td box="[1075,1126,550,569]" gridcol="7" gridrow="11" pageId="4" pageNumber="405">1.51</td>
<td box="[1156,1169,550,569]" gridcol="8" gridrow="11" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,550,569]" gridcol="9" gridrow="11" pageId="4" pageNumber="405">0.26</td>
<td box="[1273,1313,550,569]" gridcol="10" gridrow="11" pageId="4" pageNumber="405">1.25</td>
<td box="[1344,1351,550,569]" gridcol="11" gridrow="11" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,550,569]" gridcol="12" gridrow="11" pageId="4" pageNumber="405">1.93</td>
</tr>
<tr box="[198,1433,575,594]" gridrow="12" pageId="4" pageNumber="405">
<th box="[198,564,575,594]" gridcol="0" gridrow="12" pageId="4" pageNumber="405">SL/Dorsal Base L. (SL/L)</th>
<td box="[677,727,575,594]" gridcol="1" gridrow="12" pageId="4" pageNumber="405">2.91</td>
<td box="[758,771,575,594]" gridcol="2" gridrow="12" pageId="4" pageNumber="405">±</td>
<td box="[801,841,575,594]" gridcol="3" gridrow="12" pageId="4" pageNumber="405">0.1</td>
<td box="[875,915,575,594]" gridcol="4" gridrow="12" pageId="4" pageNumber="405">2.79</td>
<td box="[946,953,575,594]" gridcol="5" gridrow="12" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,575,594]" gridcol="6" gridrow="12" pageId="4" pageNumber="405">3.08</td>
<td box="[1075,1126,575,594]" gridcol="7" gridrow="12" pageId="4" pageNumber="405">2.79</td>
<td box="[1156,1169,575,594]" gridcol="8" gridrow="12" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,575,594]" gridcol="9" gridrow="12" pageId="4" pageNumber="405">0.10</td>
<td box="[1273,1313,575,594]" gridcol="10" gridrow="12" pageId="4" pageNumber="405">2.62</td>
<td box="[1344,1351,575,594]" gridcol="11" gridrow="12" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,575,594]" gridcol="12" gridrow="12" pageId="4" pageNumber="405">2.91</td>
</tr>
<tr box="[198,1433,599,618]" gridrow="13" pageId="4" pageNumber="405">
<th box="[198,564,599,618]" gridcol="0" gridrow="13" pageId="4" pageNumber="405">SL/Interdorsal L. (SL/M)</th>
<td box="[677,727,599,618]" gridcol="1" gridrow="13" pageId="4" pageNumber="405">6.57</td>
<td box="[758,771,599,618]" gridcol="2" gridrow="13" pageId="4" pageNumber="405">±</td>
<td box="[801,841,599,618]" gridcol="3" gridrow="13" pageId="4" pageNumber="405">0.63</td>
<td box="[875,915,599,618]" gridcol="4" gridrow="13" pageId="4" pageNumber="405">5.75</td>
<td box="[946,953,599,618]" gridcol="5" gridrow="13" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,599,618]" gridcol="6" gridrow="13" pageId="4" pageNumber="405">7.56</td>
<td box="[1075,1126,599,618]" gridcol="7" gridrow="13" pageId="4" pageNumber="405">7.17</td>
<td box="[1156,1169,599,618]" gridcol="8" gridrow="13" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,599,618]" gridcol="9" gridrow="13" pageId="4" pageNumber="405">1.09</td>
<td box="[1273,1313,599,618]" gridcol="10" gridrow="13" pageId="4" pageNumber="405">5.69</td>
<td box="[1344,1351,599,618]" gridcol="11" gridrow="13" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,599,618]" gridcol="12" gridrow="13" pageId="4" pageNumber="405">9.58</td>
</tr>
<tr box="[198,1433,624,643]" gridrow="14" pageId="4" pageNumber="405">
<th box="[198,564,624,643]" gridcol="0" gridrow="14" pageId="4" pageNumber="405">SL/Thorax L. (SL/N)</th>
<td box="[677,727,624,643]" gridcol="1" gridrow="14" pageId="4" pageNumber="405">4.06</td>
<td box="[758,771,624,643]" gridcol="2" gridrow="14" pageId="4" pageNumber="405">±</td>
<td box="[801,841,624,643]" gridcol="3" gridrow="14" pageId="4" pageNumber="405">0.09</td>
<td box="[875,915,624,643]" gridcol="4" gridrow="14" pageId="4" pageNumber="405">3.97</td>
<td box="[946,953,624,643]" gridcol="5" gridrow="14" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,624,643]" gridcol="6" gridrow="14" pageId="4" pageNumber="405">4.18</td>
<td box="[1075,1126,624,643]" gridcol="7" gridrow="14" pageId="4" pageNumber="405">3.94</td>
<td box="[1156,1169,624,643]" gridcol="8" gridrow="14" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,624,643]" gridcol="9" gridrow="14" pageId="4" pageNumber="405">0.30</td>
<td box="[1273,1313,624,643]" gridcol="10" gridrow="14" pageId="4" pageNumber="405">3.47</td>
<td box="[1344,1351,624,643]" gridcol="11" gridrow="14" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,624,643]" gridcol="12" gridrow="14" pageId="4" pageNumber="405">4.38</td>
</tr>
<tr box="[198,1433,648,667]" gridrow="15" pageId="4" pageNumber="405">
<th box="[198,564,648,667]" gridcol="0" gridrow="15" pageId="4" pageNumber="405">SL/Pectoral Spine L. (SL/O)</th>
<td box="[677,727,648,667]" gridcol="1" gridrow="15" pageId="4" pageNumber="405">3.23</td>
<td box="[758,771,648,667]" gridcol="2" gridrow="15" pageId="4" pageNumber="405">±</td>
<td box="[801,841,648,667]" gridcol="3" gridrow="15" pageId="4" pageNumber="405">0.15</td>
<td box="[875,915,648,667]" gridcol="4" gridrow="15" pageId="4" pageNumber="405">3.05</td>
<td box="[946,953,648,667]" gridcol="5" gridrow="15" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,648,667]" gridcol="6" gridrow="15" pageId="4" pageNumber="405">3.56</td>
<td box="[1075,1126,648,667]" gridcol="7" gridrow="15" pageId="4" pageNumber="405">3.09</td>
<td box="[1156,1169,648,667]" gridcol="8" gridrow="15" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,648,667]" gridcol="9" gridrow="15" pageId="4" pageNumber="405">0.16</td>
<td box="[1273,1313,648,667]" gridcol="10" gridrow="15" pageId="4" pageNumber="405">2.85</td>
<td box="[1344,1351,648,667]" gridcol="11" gridrow="15" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,648,667]" gridcol="12" gridrow="15" pageId="4" pageNumber="405">3.38</td>
</tr>
<tr box="[198,1433,673,692]" gridrow="16" pageId="4" pageNumber="405">
<th box="[198,564,673,692]" gridcol="0" gridrow="16" pageId="4" pageNumber="405">SL/Abdominal L. (SL/P)</th>
<td box="[677,727,673,692]" gridcol="1" gridrow="16" pageId="4" pageNumber="405">4.24</td>
<td box="[758,771,673,692]" gridcol="2" gridrow="16" pageId="4" pageNumber="405">±</td>
<td box="[801,841,673,692]" gridcol="3" gridrow="16" pageId="4" pageNumber="405">0.17</td>
<td box="[875,915,673,692]" gridcol="4" gridrow="16" pageId="4" pageNumber="405">3.97</td>
<td box="[946,953,673,692]" gridcol="5" gridrow="16" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,673,692]" gridcol="6" gridrow="16" pageId="4" pageNumber="405">4.48</td>
<td box="[1075,1126,673,692]" gridcol="7" gridrow="16" pageId="4" pageNumber="405">4.08</td>
<td box="[1156,1169,673,692]" gridcol="8" gridrow="16" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,673,692]" gridcol="9" gridrow="16" pageId="4" pageNumber="405">0.22</td>
<td box="[1273,1313,673,692]" gridcol="10" gridrow="16" pageId="4" pageNumber="405">3.78</td>
<td box="[1344,1351,673,692]" gridcol="11" gridrow="16" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,673,692]" gridcol="12" gridrow="16" pageId="4" pageNumber="405">4.51</td>
</tr>
<tr box="[198,1433,698,717]" gridrow="17" pageId="4" pageNumber="405">
<th box="[198,564,698,717]" gridcol="0" gridrow="17" pageId="4" pageNumber="405">SL/Pelvic Spine L. (SL/Q)</th>
<td box="[677,727,698,717]" gridcol="1" gridrow="17" pageId="4" pageNumber="405">3.72</td>
<td box="[758,771,698,717]" gridcol="2" gridrow="17" pageId="4" pageNumber="405">±</td>
<td box="[801,841,698,717]" gridcol="3" gridrow="17" pageId="4" pageNumber="405">0.19</td>
<td box="[875,915,698,717]" gridcol="4" gridrow="17" pageId="4" pageNumber="405">3.56</td>
<td box="[946,953,698,717]" gridcol="5" gridrow="17" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,698,717]" gridcol="6" gridrow="17" pageId="4" pageNumber="405">4.11</td>
<td box="[1075,1126,698,717]" gridcol="7" gridrow="17" pageId="4" pageNumber="405">3.7</td>
<td box="[1156,1169,698,717]" gridcol="8" gridrow="17" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,698,717]" gridcol="9" gridrow="17" pageId="4" pageNumber="405">0.30</td>
<td box="[1273,1313,698,717]" gridcol="10" gridrow="17" pageId="4" pageNumber="405">3.3</td>
<td box="[1344,1351,698,717]" gridcol="11" gridrow="17" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,698,717]" gridcol="12" gridrow="17" pageId="4" pageNumber="405">4.15</td>
</tr>
<tr box="[198,1433,722,741]" gridrow="18" pageId="4" pageNumber="405">
<th box="[198,564,722,741]" gridcol="0" gridrow="18" pageId="4" pageNumber="405">SL/Postanal L. (SL/R)</th>
<td box="[677,727,722,741]" gridcol="1" gridrow="18" pageId="4" pageNumber="405">2.97</td>
<td box="[758,771,722,741]" gridcol="2" gridrow="18" pageId="4" pageNumber="405">±</td>
<td box="[801,841,722,741]" gridcol="3" gridrow="18" pageId="4" pageNumber="405">0.13</td>
<td box="[875,915,722,741]" gridcol="4" gridrow="18" pageId="4" pageNumber="405">2.76</td>
<td box="[946,953,722,741]" gridcol="5" gridrow="18" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,722,741]" gridcol="6" gridrow="18" pageId="4" pageNumber="405">3.19</td>
<td box="[1075,1126,722,741]" gridcol="7" gridrow="18" pageId="4" pageNumber="405">3.16</td>
<td box="[1156,1169,722,741]" gridcol="8" gridrow="18" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,722,741]" gridcol="9" gridrow="18" pageId="4" pageNumber="405">0.21</td>
<td box="[1273,1313,722,741]" gridcol="10" gridrow="18" pageId="4" pageNumber="405">2.9</td>
<td box="[1344,1351,722,741]" gridcol="11" gridrow="18" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,722,741]" gridcol="12" gridrow="18" pageId="4" pageNumber="405">3.53</td>
</tr>
<tr box="[198,1433,747,766]" gridrow="19" pageId="4" pageNumber="405">
<th box="[198,564,747,766]" gridcol="0" gridrow="19" pageId="4" pageNumber="405">Interdorsal L./Caudal Peduncle D. (M/S)</th>
<td box="[677,727,747,766]" gridcol="1" gridrow="19" pageId="4" pageNumber="405">1.63</td>
<td box="[758,771,747,766]" gridcol="2" gridrow="19" pageId="4" pageNumber="405">±</td>
<td box="[801,841,747,766]" gridcol="3" gridrow="19" pageId="4" pageNumber="405">0.14</td>
<td box="[875,915,747,766]" gridcol="4" gridrow="19" pageId="4" pageNumber="405">1.42</td>
<td box="[946,953,747,766]" gridcol="5" gridrow="19" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,747,766]" gridcol="6" gridrow="19" pageId="4" pageNumber="405">1.86</td>
<td box="[1075,1126,747,766]" gridcol="7" gridrow="19" pageId="4" pageNumber="405">1.26</td>
<td box="[1156,1169,747,766]" gridcol="8" gridrow="19" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,747,766]" gridcol="9" gridrow="19" pageId="4" pageNumber="405">0.20</td>
<td box="[1273,1313,747,766]" gridcol="10" gridrow="19" pageId="4" pageNumber="405">0.93</td>
<td box="[1344,1351,747,766]" gridcol="11" gridrow="19" pageId="4" pageNumber="405">-</td>
<td box="[1383,1433,747,766]" gridcol="12" gridrow="19" pageId="4" pageNumber="405">1.49</td>
</tr>
<tr box="[198,1433,771,790]" gridrow="20" pageId="4" pageNumber="405">
<th box="[198,564,771,790]" gridcol="0" gridrow="20" pageId="4" pageNumber="405">Postanal L./Caudal Peduncle D. (R/S)</th>
<td box="[677,727,771,790]" gridcol="1" gridrow="20" pageId="4" pageNumber="405">3.61</td>
<td box="[758,771,771,790]" gridcol="2" gridrow="20" pageId="4" pageNumber="405">±</td>
<td box="[801,841,771,790]" gridcol="3" gridrow="20" pageId="4" pageNumber="405">0.3</td>
<td box="[875,915,771,790]" gridcol="4" gridrow="20" pageId="4" pageNumber="405">3.23</td>
<td box="[946,953,771,790]" gridcol="5" gridrow="20" pageId="4" pageNumber="405">-</td>
<td box="[984,1035,771,790]" gridcol="6" gridrow="20" pageId="4" pageNumber="405">4.07</td>
<td box="[1075,1126,771,790]" gridcol="7" gridrow="20" pageId="4" pageNumber="405">2.82</td>
<td box="[1156,1169,771,790]" gridcol="8" gridrow="20" pageId="4" pageNumber="405">±</td>
<td box="[1199,1239,771,790]" gridcol="9" gridrow="20" pageId="4" pageNumber="405">0.38</td>
<td box="[1273,1313,771,790]" gridcol="10" gridrow="20" pageId="4" pageNumber="405">2.32</td>
<td box="[1344,1433,771,790]" colspan="2" colspanRight="1" gridcol="11" gridrow="20" pageId="4" pageNumber="405">- 3.45</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[158,801,834,1145]" lastBlockId="4.[839,1482,834,1145]" pageId="4" pageNumber="405">Keels on lateral plates prominent, formed of ridges of bone and slightly hypertrophied odontodes. Keels present on all plate rows. One or two plates forming short, accessory row between dorsal and mid-dorsal plate series near head; keel on accessory plate row contiguous with but widely separated from keel on dorsal plate series beginning posteriorly to origin of dorsal fin. Keel on anterior plates of dorsal series beginning at midline just anteriorly to posterior margin of supraoccipital, angled to lateral edge of nuchal plate. Ridge on pterotic contiguous with keel on anterior three plates of middorsal plate series; keel on remaining plates of mid-dorsal plate series ventral to that of keel on anterior three plates.</paragraph>
<paragraph blockId="4.[839,1482,834,1145]" lastBlockId="5.[151,794,192,1463]" lastPageId="5" lastPageNumber="406" pageId="4" pageNumber="405">Nares separated by short flap of skin held erect in life. Dorsal, mid-dorsal, median and mid-ventral plate rows complete from head to caudal fin. Ventral plate row begins posteriorly to insertion of pelvic fin and continues to caudal fin. Base of caudal fin covered in elongate, roughly triangular plates. Ventral surface of body (except region dorsal to pectoral fin and ventral to mid-ventral plate row) covered in small plates. Plates on abdomen increase in number with standard length. Head covered in small plates. Frontal, nasal, sphenotic, infraorbitals, pterotic-supracleithrum, suprapreopercle, and supraoccipital supporting odontodes. Opercle with small patch of odontodes in juvenile, none in adult. Platelets covering anteroventral corner of opercle separated from opercle; platelets may be everted to approximately 90° from head. Evertible cheek plates with zero to three slightly hypertrophied odontodes (largest extending approximately to posterior end of opercle).</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566992" ID-Zenodo-Dep="4566992" httpUri="https://zenodo.org/record/4566992/files/figure.png" pageId="4" pageNumber="405" startId="4.[158,197,1922,1945]" targetBox="[156,1475,1202,1882]" targetPageId="4">
<paragraph blockId="4.[158,1481,1921,1977]" pageId="4" pageNumber="405">
<emphasis bold="true" box="[158,227,1922,1945]" pageId="4" pageNumber="405">Fig. 3.</emphasis>
Dorsal profiles of snouts of a.
<taxonomicName authority=", UF" authorityName="UF" box="[549,886,1921,1945]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[549,840,1921,1945]" italics="true" pageId="4" pageNumber="405">Pterygoplichthys punctatus</emphasis>
, UF
</taxonomicName>
148742, 160.1 mm SL and b.
<taxonomicName authority=", AUM" authorityName="AUM" box="[1199,1402,1921,1945]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="zuliaensis">
<emphasis box="[1199,1330,1921,1945]" italics="true" pageId="4" pageNumber="405">P. zuliaensis</emphasis>
, AUM
</taxonomicName>
22586, 222.0 mm SL.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566994" ID-Zenodo-Dep="4566994" httpUri="https://zenodo.org/record/4566994/files/figure.png" pageId="5" pageNumber="406" startId="5.[831,870,622,645]" targetBox="[831,1468,200,594]" targetPageId="5">
<paragraph blockId="5.[831,1475,621,837]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[831,904,622,645]" pageId="5" pageNumber="406">Fig. 4.</emphasis>
Localities for
<taxonomicName box="[1069,1366,621,645]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[1069,1366,621,645]" italics="true" pageId="5" pageNumber="406">Pterygoplichthys punctatus</emphasis>
</taxonomicName>
(squares) and
<taxonomicName authorityName="Armbruster &amp; Page" authorityYear="2006" box="[877,976,653,677]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="weberi">
<emphasis box="[877,976,653,677]" italics="true" pageId="5" pageNumber="406">P. weberi</emphasis>
</taxonomicName>
(dots). Open symbols are type localities. Localities for
<taxonomicName box="[956,1089,686,709]" class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="punctatus">
<emphasis box="[956,1089,686,709]" italics="true" pageId="5" pageNumber="406">P. punctatus</emphasis>
</taxonomicName>
from
<bibRefCitation author="Weber, C" box="[1157,1306,686,709]" pageId="5" pageNumber="406" pagination="1 - 36" refId="ref6617" refString="Weber, C. 1992. Revision du genre Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae. Revue Francaise d'Aquariologie Herpetologie, 19: 1 - 36." type="journal article" year="1992">Weber (1992)</bibRefCitation>
and specimens examined in this study. Because the rio Urubu locality of
<taxonomicName class="Actinopterygii" family="Loricariidae" genus="Pterygoplichthys" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="punctatus">
<emphasis italics="true" pageId="5" pageNumber="406">P. punctatus</emphasis>
</taxonomicName>
did not have precise locality information, the locality is indicated at random in the lower Urubu. Base map by M. J. Weitzman.
</paragraph>
</caption>
<paragraph blockId="5.[151,794,192,1463]" pageId="5" pageNumber="406">
Dorsal fin long and low in adult, higher in juvenile, consisting of small, V-shaped spinelet, fairly strong spine, usually 11 dorsal-fin rays (three of 27 with 12); adpressed dorsal fin does not reach adipose fin. Caudal fin forked, lower lobe longer than upper. Pectoral-fin spine strong, reaches posteriorly to pelvic-fin rays when depressed ventral to pelvic fin; cleithrum with exposed process dorsal to pectoral-fin rays and strongly angled dorsally; pectoral fin inserted on same plane as pelvic fin such that spine, when depressed parallel with body, lies on top of and in contact with pelvic fin. Pelvicfin spine thin, flexible, reaches barely to base of anal fin. Anal fin with relatively strong, unbranched first ray supporting odontodes. Adipose fin consisting of single median preadipose plate and strong, pointed spine; adipose-fin membrane not reaching procurrent caudal-fin spines. Dorsal fin II,11, pectoral fin I,6, pelvic fin I,5, anal fin I,4, caudal fin usually I,14,I (one specimen I,13,I). Jaws weakly angled, dentaries forming angle of approximately 90º. Teeth bicuspid, median cusp short, lateral cusp about half length of median cusp, stalk moderately long; 20-36 dentary teeth, 19-38 premaxillary teeth. Median plates 26-29 (mode 28), adipose-caudal plates four to six (mode five), anal-caudal plates 11-13 (mode 12), dorsal-adipose plates five to seven (mode six). Buccal papilla lobulate and divided medially (
<figureCitation box="[478,553,1440,1463]" captionStart="Fig" captionStartId="1.[831,870,1826,1849]" captionTargetBox="[831,1474,653,1797]" captionTargetId="figure-567@1.[831,1474,653,1797]" captionTargetPageId="1" captionText="Fig. 1. Mouth and buccal papilla (indicated by arrow) of a. Pterygoplichthys punctatus, MCP 35755, 203.1 mm SL, b. P. weberi, FMNH 96959, holotype, 183.9 mm SL, c. P. multiradiatus, AUM 18947, 153.3 mm SL. Photos by J.W. Armbruster and L.M. Page." figureDoi="http://doi.org/10.5281/zenodo.4566984" httpUri="https://zenodo.org/record/4566984/files/figure.png" pageId="5" pageNumber="406">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph blockId="5.[151,794,1504,1751]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[151,338,1504,1527]" pageId="5" pageNumber="406">Color in alcohol.</emphasis>
Body and fins light brown to gray and covered with black spots. Spots smallest on snout, progressively larger to caudal peduncle; spots on fins nearly as large as eye. Venter covered with discrete (uncoalesced) spots; spots smallest on throat and becoming progressively larger to interpelvic area. Spots proportionally larger and more distinct on juvenile. Caudal peduncle lighter ventrally than dorsally.
</paragraph>
<paragraph blockId="5.[151,794,1792,1975]" pageId="5" pageNumber="406">
<emphasis bold="true" box="[151,228,1792,1815]" pageId="5" pageNumber="406">Range.</emphasis>
Known from the rio Madeira drainage and the rio Urubu of
<collectingCountry box="[177,241,1824,1847]" name="Brazil" pageId="5" pageNumber="406">Brazil</collectingCountry>
(
<figureCitation box="[254,317,1824,1847]" captionStart="Fig" captionStartId="5.[831,870,622,645]" captionTargetBox="[831,1468,200,594]" captionTargetId="figure-781@5.[831,1474,192,594]" captionTargetPageId="5" captionText="Fig. 4. Localities for Pterygoplichthys punctatus (squares) and P. weberi (dots). Open symbols are type localities. Localities for P. punctatus from Weber (1992) and specimens examined in this study. Because the rio Urubu locality of P. punctatus did not have precise locality information, the locality is indicated at random in the lower Urubu. Base map by M. J. Weitzman." figureDoi="http://doi.org/10.5281/zenodo.4566994" httpUri="https://zenodo.org/record/4566994/files/figure.png" pageId="5" pageNumber="406">Fig. 4</figureCitation>
).
<bibRefCitation author="Weber, C" box="[338,481,1824,1847]" pageId="5" pageNumber="406" pagination="1 - 36" refId="ref6617" refString="Weber, C. 1992. Revision du genre Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae. Revue Francaise d'Aquariologie Herpetologie, 19: 1 - 36." type="journal article" year="1992">Weber (1992)</bibRefCitation>
additionally notes localities in the rio Purus and the rio
<collectingRegion box="[427,532,1856,1879]" country="Brazil" name="Tocantins" pageId="5" pageNumber="406">Tocantins</collectingRegion>
basin; however, we did not confirm these identifications. Also, not all specimens available to
<bibRefCitation author="Weber, C" box="[276,413,1920,1943]" pageId="5" pageNumber="406" pagination="637 - 643" refId="ref6586" refString="Weber, C. 1991. Nouveaux taxa dans Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae). Revue Suisse de Zoologie, 98: 637 - 643." type="journal article" year="1991">Weber (1991</bibRefCitation>
,
<bibRefCitation author="Weber, C" box="[423,477,1920,1943]" pageId="5" pageNumber="406" pagination="1 - 36" refId="ref6617" refString="Weber, C. 1992. Revision du genre Pterygoplichthys sensu lato (Pisces, Siluriformes, Loricariidae. Revue Francaise d'Aquariologie Herpetologie, 19: 1 - 36." type="journal article" year="1992">1992</bibRefCitation>
) from the upper Amazon were available to us, and their identity needs to be verified.
</paragraph>
</subSubSection>
</treatment>
</document>