149 lines
22 KiB
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149 lines
22 KiB
XML
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<mods:title id="DC4A681F05A6742CB3DB098896A0241A">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XII. Pandinurus hangarale sp. n. (Scorpionidae) from Somaliland and a review of type locality and true distribution of Pandinurus smithi (Pocock, 1897)</mods:title>
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<mods:namePart id="C0665896FCA4473586D375EC94BBE8C2">Kovařík, František</mods:namePart>
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<mods:namePart id="B51315810A4D186025A4A68A24379179">Lowe, Graeme</mods:namePart>
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<mods:namePart id="8456B6EB5E2FDDC2712768996447A5AB">Mazuch, Tomáš</mods:namePart>
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<mods:namePart id="0720F4A321EE4D414E37E75DA9302B7B">Awale, Ahmed Ibrahim</mods:namePart>
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<mods:title id="3BD07E33D4F117C69F43EF22B1BBE05F">Euscorpius</mods:title>
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<emphasis id="599AEAC9FF84FF8520E0FEF00E640CF2" bold="true" box="[390,534,373,400]" italics="true" pageId="3" pageNumber="2">Pandinurus</emphasis>
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<bibRefCitation id="0F7F4B2AFF84FF85237BFEF20EFE0CF3" author="FET" box="[541,652,375,401]" pageId="3" pageNumber="2" pagination="245 - 250" refId="ref5352" refString="FET, V. 1997. Notes on the taxonomy of some old world scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). The Journal of Arachnology, 25: 245 - 250." type="journal article" year="1997">Fet, 1997</bibRefCitation>
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(
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<figureCitation id="F3D52A5EFF84FF8520EAFE120E730CCC" box="[396,513,407,430]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–10" captionStart-10="Figures 34–35" captionStart-2="Figures 11–14" captionStart-3="Figures 15–21" captionStart-4="Figures 22–23" captionStart-5="Figure 24–25" captionStart-6="Figures 26–27" captionStart-7="Figures 28–29" captionStart-8="Figures 30–31" captionStart-9="Figures 32–33" captionStartId-0="4.[192,278,1596,1619]" captionStartId-1="5.[192,278,1374,1397]" captionStartId-10="14.[192,278,1566,1589]" captionStartId-2="6.[192,278,1680,1703]" captionStartId-3="7.[192,278,1722,1745]" captionStartId-4="8.[192,278,1792,1815]" captionStartId-5="9.[192,268,1782,1805]" captionStartId-6="10.[192,278,1825,1848]" captionStartId-7="11.[192,278,1851,1874]" captionStartId-8="12.[192,278,1768,1791]" captionStartId-9="13.[192,278,1830,1853]" captionTargetBox-0="[204,1420,204,1541]" captionTargetBox-1="[222,1400,191,1328]" captionTargetBox-10="[192,1437,911,1536]" captionTargetBox-2="[191,1440,167,1654]" captionTargetBox-3="[192,1439,209,1681]" captionTargetBox-4="[191,1440,191,1762]" captionTargetBox-5="[191,1440,213,1753]" captionTargetBox-6="[191,1440,191,1795]" captionTargetBox-7="[444,1169,208,1824]" captionTargetBox-8="[251,1378,191,1739]" captionTargetBox-9="[209,1417,195,1801]" captionTargetId-0="figure-11@4.[192,1439,192,1567]" captionTargetId-1="figure-6@5.[192,1437,184,1344]" captionTargetId-10="figure-348@14.[192,1437,911,1536]" captionTargetId-2="figure-11@6.[192,1439,168,1653]" captionTargetId-3="figure-6@7.[192,1439,192,1692]" captionTargetId-4="figure-11@8.[192,1439,192,1761]" captionTargetId-5="figure-6@9.[192,1439,213,1752]" captionTargetId-6="figure-11@10.[192,1439,192,1795]" captionTargetId-7="figure-6@11.[444,1169,208,1824]" captionTargetId-8="figure-11@12.[252,1377,192,1738]" captionTargetId-9="figure-6@13.[210,1416,196,1800]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="14" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="Figures 1–2: Pandinurus hangarale sp. n., male holotype in dorsal (1) and ventral (2) aspects. Scale bar: 10 mm." captionText-1="Figures 3–10: Pandinurus hangarale sp. n., male holotype, pedipalp segments. Chela dorsal (3), external (4) and ventrointernal (5). Patella dorsal (6), external (7) and ventral (8). Femur and trochanter dorsal (9) and ventral (10). Trichobothrial pattern is indicated." captionText-10="Figures 34–35: Pandinurus hangarale sp. n., male holotype, chromosomes (2n=120). Mitotic metaphase (34), meiotic metaphase I (35). Arrows indicate the largest chromosome pair. Scale bar: 10 µm." captionText-2="Figures 11–14: Pandinurus hangarale sp. n., male holotype. Carapace and tergites I–III (11), coxosternal area and sternites III–IV (12), left chelicera ventral (13) and dorsal (14)." captionText-3="Figures 15–21: Pandinurus hangarale sp. n., male holotype. Figures 15–17. Metasoma and telson lateral (15), dorsal (16), and ventral (17) views. Scale bar: 10 mm. Figures 18–21. Tarsomeres I and II of left legs I–IV, retrolateral aspects." captionText-4="Figures 22–23: Pandinurus hangarale sp. n., in vivo habitus. Male holotype shortly after ecdysis (22) and juvenile paratype (23)." captionText-5="Figure 24–25: Pandinurus hangarale sp. n., the type locality. Photo taken on 8. February 2017. In addition to P. hangarale sp. n. the first author recorded at this locality Gint calviceps (Pocock, 1900) and Parabuthus heterurus Pocock, 1897." captionText-6="Figures 26–27: Pandinurus smithi (Pocock, 1897). Figure 26. Female lectotype and original labels. Scale bar: 10 mm for the female lectotype. Figure 27. Male, at the locality, Ethiopia, 55 km S of Degehbur, 07°49'27.2"N 43°41'56.3"E, 1053 m a.s.l." captionText-7="Figures 28–29: Pandinurus hangarale sp. n., male holotype, left hemispermatophore. Dorsal (convex) (28) and ventral (concave) (29) views. Scale bar: 2 mm." captionText-8="Figures 30–31: Pandinurus hangarale sp. n., male holotype, left hemispermatophore, capsule region. Dorsal (convex) (30) and ventral (concave) (31) cross-stereoscopic views. Scale bar: 500 µm. Label abbreviations: bl, basal lobe; dt, dorsal trough; h, hook; ibr, internobasal reflection of sperm duct (= median transverse trough); il, inner lobe, ml, median lobe; tf, truncal flexure (= median transverse cleavage)." captionText-9="Figures 32–33: Pandinurus hangarale sp. n., male holotype, left hemispermatophore, capsule region. Internal (sperm duct margin) (32) and external (distal lamina margin) (33) cross-stereoscopic views. Scale bar: 500 µm. Label abbreviations: see legend for Figs. 30–31." pageId="3" pageNumber="2">Figs. 1–35</figureCitation>
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,
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<figureCitation id="F3D52A5EFF84FF85236DFE120E5A0CCC" box="[523,552,407,430]" captionStart="Figure 37" captionStartId="17.[192,268,1689,1712]" captionTargetBox="[192,1439,192,1659]" captionTargetId="figure-132@17.[192,1439,192,1659]" captionTargetPageId="17" captionText="Figure 37: A. Map with marked places where types of P. smithi were collected, originally published by Smith (1897). B. Actual map of Horn of Africa with marked discussed localities. 1–4. Localities cited by Pocock (1897) except locality "Ahdeh (Abdeh)" which we has not found (1. Turfa, type locality of P. smithi, 2. Silul, 3. Hargesa, 4. Boholgarshan.). 5–7. Localities where the first and the third authors collected Pandinurus specimens (5. Locality 11EV, Ethiopia, 55 km S of Degehbur, 07°49'27.2"N 43°41'56.3"E where was collected a male of P. smithi. 6. Locality 17SE, Somaliland, Toon village near Hargeisa, 09°23'30"N 44°07'10"E, the type locality of P. hangarale. 7. Woqooyi Galbeed, 09°11'56"N 44°09'50"E, between Hargeisa and Salahle, the locality of P. hangarale paratype)." pageId="3" pageNumber="2">37</figureCitation>
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,
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<tableCitation id="266C0360FF84FF852355FE120EF10CCC" box="[563,643,407,430]" captionStart="Table 1" captionStartId="14.[192,257,826,849]" captionText="Table 1: Comparative measurements of male holotype of Pandinurus hangarale sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Table 1</tableCitation>
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)
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</paragraph>
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<subSubSection id="23F46550FF84FF852196FE5D0EF20FA5" pageId="3" pageNumber="2" type="reference_group">
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<taxonomicName id="ACEE4D58FF84FF852196FE5D0EBA0C8D" authority="Vachon, 1974: 953" authorityName="Vachon" authorityPageNumber="953" authorityYear="1974" box="[240,712,472,495]" class="Arachnida" family="Scorpionidae" genus="Pandinus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Pandinurus">
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<emphasis id="599AEAC9FF84FF852196FE5D0D270C8D" box="[240,341,472,495]" italics="true" pageId="3" pageNumber="2">Pandinus</emphasis>
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(
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<emphasis id="599AEAC9FF84FF852001FE5D0D910C8D" box="[359,483,472,495]" italics="true" pageId="3" pageNumber="2">Pandinurus</emphasis>
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)
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<bibRefCitation id="0F7F4B2AFF84FF852090FE5D0EBA0C8D" author="VACHON" box="[502,712,472,495]" pageId="3" pageNumber="2" pagination="857 - 958" refId="ref6351" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e serie, 140 (Zoologie, 104): 857 - 958." type="journal article" year="1974">Vachon, 1974: 953</bibRefCitation>
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(
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<emphasis id="599AEAC9FF84FF8523BDFE5D0D7B0F6C" italics="true" pageId="3" pageNumber="2">nomen nudum</emphasis>
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,
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<typeStatus id="B4558879FF84FF852070FE720D360F6C" box="[278,324,503,526]" pageId="3" pageNumber="2">type</typeStatus>
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species not designated);
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<treatmentCitation id="EA4F10CAFF84FF852336FE720E9C0F6C" author="FET" box="[592,750,503,526]" page="248" pageId="3" pageNumber="2" year="1997">
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<bibRefCitation id="0F7F4B2AFF84FF852336FE720E9C0F6C" author="FET" box="[592,750,503,526]" pageId="3" pageNumber="2" pagination="245 - 250" refId="ref5352" refString="FET, V. 1997. Notes on the taxonomy of some old world scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). The Journal of Arachnology, 25: 245 - 250." type="journal article" year="1997">Fet, 1997: 248</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="EA4F10CAFF84FF85239FFE720D150F4F" author="FET" page="470" pageId="3" pageNumber="2" year="2000">
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<bibRefCitation id="0F7F4B2AFF84FF85239FFE720D150F4F" author="FET" pageId="3" pageNumber="2" refId="ref5390" refString="FET, V. 2000. Family Scorpionidae Latreille, 1802. Pp. 427 - 486 in Fet, V., W. D. Sissom, G. Lowe & M. E. Braunwalder. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 470–473</bibRefCitation>
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(in part) (complete reference list until 2000).
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</paragraph>
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<paragraph id="6B5136DBFF84FF852196FDD10EF20FA5" blockId="3.[192,802,472,711]" pageId="3" pageNumber="2">
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<taxonomicName id="ACEE4D58FF84FF852196FDD10ECA0F09" ID-CoL="6D6D" authority=": Kovarik et al., 2017: 42 - 100" authorityName="Kovarik" authorityPageNumber="42 - 100" authorityYear="2017" box="[240,696,596,619]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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||
<emphasis id="599AEAC9FF84FF852196FDD10D1F0F09" box="[240,365,596,619]" italics="true" pageId="3" pageNumber="2">Pandinurus</emphasis>
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||
: Kovařík et al., 2017: 42–100
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</taxonomicName>
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||
(
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||
<figureCitation id="F3D52A5EFF84FF8523AFFDD10CAF0FEB" captionStart-0="Figures 3–10" captionStart-1="Figures 11–14" captionStart-2="Figures 15–21" captionStartId-0="5.[192,278,1374,1397]" captionStartId-1="6.[192,278,1680,1703]" captionStartId-2="7.[192,278,1722,1745]" captionTargetBox-0="[222,1400,191,1328]" captionTargetBox-1="[191,1440,167,1654]" captionTargetBox-2="[192,1439,209,1681]" captionTargetId-0="figure-6@5.[192,1437,184,1344]" captionTargetId-1="figure-11@6.[192,1439,168,1653]" captionTargetId-2="figure-6@7.[192,1439,192,1692]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figures 3–10: Pandinurus hangarale sp. n., male holotype, pedipalp segments. Chela dorsal (3), external (4) and ventrointernal (5). Patella dorsal (6), external (7) and ventral (8). Femur and trochanter dorsal (9) and ventral (10). Trichobothrial pattern is indicated." captionText-1="Figures 11–14: Pandinurus hangarale sp. n., male holotype. Carapace and tergites I–III (11), coxosternal area and sternites III–IV (12), left chelicera ventral (13) and dorsal (14)." captionText-2="Figures 15–21: Pandinurus hangarale sp. n., male holotype. Figures 15–17. Metasoma and telson lateral (15), dorsal (16), and ventral (17) views. Scale bar: 10 mm. Figures 18–21. Tarsomeres I and II of left legs I–IV, retrolateral aspects." pageId="3" pageNumber="2">Figs. 5– 18</figureCitation>
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, 144–167, 175–181, 183, 185, 187, 194–394, 396, Table 2) (historical and taxonomic comments, complete reference list), Kovařík et al., 2017: 1–14.
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||
</paragraph>
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||
</subSubSection>
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<materialsCitation id="DB863C86FF84FF852196FD740EA50E6A" box="[240,727,753,776]" collectionCode="SPECIES" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
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.
|
||
<taxonomicName id="ACEE4D58FF84FF8520E8FD740EA50E6A" authority="Pocock, 1888" authorityName="Pocock" authorityYear="1888" box="[398,727,753,776]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="exitialis">
|
||
<emphasis id="599AEAC9FF84FF8520E8FD740E4E0E6A" box="[398,572,753,776]" italics="true" pageId="3" pageNumber="2">Scorpio exitialis</emphasis>
|
||
Pocock, 1888
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23F46550FF84FF852196FCB40D7E0B4A" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="6B5136DBFF84FF852196FCB40D7E0B4A" blockId="3.[192,802,817,1576]" pageId="3" pageNumber="2">
|
||
DIAGNOSIS. Total length
|
||
<quantity id="AC169B3EFF84FF85236EFCB40EFD0E2A" box="[520,655,817,840]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.35" metricValueMin="0.7" pageId="3" pageNumber="2" unit="mm" value="102.5" valueMax="135.0" valueMin="70.0">70–135 mm</quantity>
|
||
. External trichobothria on patella number 16–21 (5–6
|
||
<emphasis id="599AEAC9FF84FF8523ECFCD50ED10E05" box="[650,675,848,871]" italics="true" pageId="3" pageNumber="2">eb</emphasis>
|
||
, 3–6
|
||
<emphasis id="599AEAC9FF84FF852384FCD50F770E05" box="[738,773,848,871]" italics="true" pageId="3" pageNumber="2">esb</emphasis>
|
||
, 2
|
||
<emphasis id="599AEAC9FF84FF8521A6FCEA0CAD0EE4" box="[192,223,879,902]" italics="true" pageId="3" pageNumber="2">em</emphasis>
|
||
, 3–4
|
||
<emphasis id="599AEAC9FF84FF852046FCEA0D4C0EE4" box="[288,318,879,902]" italics="true" pageId="3" pageNumber="2">est</emphasis>
|
||
, 3
|
||
<emphasis id="599AEAC9FF84FF852005FCEA0D040EE4" box="[355,374,879,902]" italics="true" pageId="3" pageNumber="2">et</emphasis>
|
||
); ventral trichobothria on patella number 29–59; accessory external trichobothrium
|
||
<emphasis id="599AEAC9FF84FF8523BBFC080E840EC6" box="[733,758,909,932]" italics="true" pageId="3" pageNumber="2">ea</emphasis>
|
||
on chela absent or present, internal trichobothria on chela number 1–4; ventral trichobothria on chela number 10– 19. Pedipalp chela manus lobiform. Dorsoexternal carina on pedipalp chela absent. Male has pronounced median lobe on movable finger of pedipalp and larger telson than female. Pectines with fulcra. Pectinal teeth number 12–22. Sternum subpentagonal, longer than wide. Carapace without distinct carinae. Dentate margins of movable and fixed fingers of pedipalp chela with distinct granules in two parallel rows present in distal half of fingers. Proximal half of fingers almost without granules in males and with distinct granules in a row in juveniles and females. These granules do not cover whole median lobe in males, but are usually represented by only several granules on top of lobe. Tergites I–VI of mesosoma bear one carina. Stridulation organ located on pedipalp coxae and first pair of legs, but can be reduced. Metasomal segments I–IV with paired, parallel ventral submedian carinae present. Telson without subaculear tubercle. Legs with one pedal spur, retrolateral spur absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |