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<document ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.64.100481" ID-Pensoft-Pub="2305-2562-2-149" ID-Pensoft-UUID="7D304C68B3385B7B8BF91C79A9C2ECBD" ID-ZooBank="6EC41DFDB88A4265BC1D269F6933D60F" ModsDocID="2305-2562-64-2-149" checkinTime="1683304273008" checkinUser="pensoft" docAuthor="Midgley, John, Bellingan, Terence &amp; Jordaens, Kurt" docDate="2023" docId="DE7EB38412995EB8839E2A297103693C" docLanguage="en" docName="AfrInvert 64(2): 149-163" docOrigin="African Invertebrates 64 (2)" docPubDate="2023-05-05" docSource="http://dx.doi.org/10.3897/afrinvertebr.64.100481" docTitle="Amphoterus braunsi van Doesburg 1956" docType="treatment" docVersion="1" id="7D304C68B3385B7B8BF91C79A9C2ECBD" lastPageNumber="149" masterDocId="7D304C68B3385B7B8BF91C79A9C2ECBD" masterDocTitle="A revision of the hover fly genus Amphoterus Bezzi, 1915 (Diptera, Syrphidae) with the description of one new species" masterLastPageNumber="163" masterPageNumber="149" pageNumber="149" updateTime="1683304273008" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the hover fly genus Amphoterus Bezzi, 1915 (Diptera, Syrphidae) with the description of one new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Midgley, John</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1203-3750</mods:nameIdentifier>
<mods:affiliation>KwaZulu-Natal Museum, Department Natural Sciences, 237 Jabu Ndlovu Street, Pietermaritzburg, KwaZulu-Natal, 3201 South Africa &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, 6139 South Africa</mods:affiliation>
<mods:nameIdentifier type="email">johnmidge@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bellingan, Terence</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3064-1744</mods:nameIdentifier>
<mods:affiliation>Department of Zoology and Entomology, Rhodes University, Makhanda, 6139 South Africa &amp; Albany Museum, Department of Entomology and Arachnology, 40 Somerset Street, Makhanda, Eastern Cape, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jordaens, Kurt</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4321-5944</mods:nameIdentifier>
<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section, Leuvense-steenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>64</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>149</mods:start>
<mods:end>163</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/afrinvertebr.64.100481</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/afrinvertebr.64.100481</mods:identifier>
<mods:identifier type="Pensoft-Pub">2305-2562-2-149</mods:identifier>
<mods:identifier type="ZooBank">6EC41DFDB88A4265BC1D269F6933D60F</mods:identifier>
<mods:identifier type="Pensoft-UUID">7D304C68B3385B7B8BF91C79A9C2ECBD</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C" httpUri="http://treatment.plazi.org/id/DE7EB38412995EB8839E2A297103693C" lastPageNumber="149" pageId="0" pageNumber="149">
<subSubSection pageId="0" pageNumber="149" type="nomenclature">
<paragraph pageId="0" pageNumber="149">
<taxonomicName ID-CoL="89YKH" LSID="DE7EB384-1299-5EB8-839E-2A297103693C" authority="van Doesburg, 1956" authorityName="van Doesburg" authorityYear="1956" class="Insecta" family="Syrphidae" genus="Amphoterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphoterus braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">Amphoterus braunsi van Doesburg, 1956</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="description">
<paragraph pageId="0" pageNumber="149">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Amphoterus braunsi, collected from sunny patch in indigenous forest at Rainbow Gorge, KwaZulu-Natal, South Africa. Specimen at AMGS." figureDoi="10.3897/afrinvertebr.64.100481.figure2" httpUri="https://binary.pensoft.net/fig/846531" pageId="0" pageNumber="149">Figs 2</figureCitation>
<figureCitation captionStart="Figures 3–6" captionStartId="F3" captionText="Figures 3 - 6. Amphoterus species, dorsal view 3 A. braunsi (♂) 4 A. braunsi (♀) 5 A. cribratus (♂) 6 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures3-6" httpUri="https://binary.pensoft.net/fig/846532" pageId="0" pageNumber="149">, 3</figureCitation>
<figureCitation captionStart="Figures 3–6" captionStartId="F3" captionText="Figures 3 - 6. Amphoterus species, dorsal view 3 A. braunsi (♂) 4 A. braunsi (♀) 5 A. cribratus (♂) 6 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures3-6" httpUri="https://binary.pensoft.net/fig/846532" pageId="0" pageNumber="149">, 4</figureCitation>
<figureCitation captionStart="Figures 7–10" captionStartId="F4" captionText="Figures 7 - 10. Amphoterus species, lateral view 7 A. braunsi (♂) 8 A. braunsi (♀) 9 A. cribratus (♂) 10 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures7-10" httpUri="https://binary.pensoft.net/fig/846533" pageId="0" pageNumber="149">, 7</figureCitation>
<figureCitation captionStart="Figures 7–10" captionStartId="F4" captionText="Figures 7 - 10. Amphoterus species, lateral view 7 A. braunsi (♂) 8 A. braunsi (♀) 9 A. cribratus (♂) 10 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures7-10" httpUri="https://binary.pensoft.net/fig/846533" pageId="0" pageNumber="149">, 8</figureCitation>
<figureCitation captionStart="Figures 11–14" captionStartId="F5" captionText="Figures 11 - 14. Amphoterus heads, frontal view 11 A. braunsi (♂) 12 A. braunsi (♀) 13 A. cribratus (♂) 14 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures11-14" httpUri="https://binary.pensoft.net/fig/846534" pageId="0" pageNumber="149">, 11</figureCitation>
<figureCitation captionStart="Figures 11–14" captionStartId="F5" captionText="Figures 11 - 14. Amphoterus heads, frontal view 11 A. braunsi (♂) 12 A. braunsi (♀) 13 A. cribratus (♂) 14 A. londti sp. nov. (♀)." figureDoi="10.3897/afrinvertebr.64.100481.figures11-14" httpUri="https://binary.pensoft.net/fig/846534" pageId="0" pageNumber="149">, 12</figureCitation>
<figureCitation captionStart="Figures 15–20" captionStartId="F6" captionText="Figures 15 - 20. Amphoterus male genitalia 15 - 17 A. braunsi 15 apical view 16 ventral view 17 lateral view 18 - 20 A. cribratus 18 apical view 19 ventral view 20 lateral view." figureDoi="10.3897/afrinvertebr.64.100481.figures15-20" httpUri="https://binary.pensoft.net/fig/846535" pageId="0" pageNumber="149">, 15-17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="reference_group">
<paragraph pageId="0" pageNumber="149">
<taxonomicName authorityName="van Doesburg" authorityYear="1956" class="Insecta" family="Syrphidae" genus="Amphoterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphoterus braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">Amphoterus braunsi</taxonomicName>
van Doesburg, 1956, 22: 517.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<taxonomicName authorityName="van Doesburg" authorityYear="1956" class="Insecta" family="Syrphidae" genus="Amphoterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphoterus braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">Amphoterus braunsi</taxonomicName>
van Doesburg -
<bibRefCitation author="Smith, KVG" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="149" pagination="488 - 510" refId="B12" refString="Smith, KVG, Vockeroth, JR, 1980. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth 1980</bibRefCitation>
: 497;
<bibRefCitation author="Dirickx, HG" journalOrPublisher="Museum d'histoire naturelle Geneve, Geneve" pageId="0" pageNumber="149" refId="B6" refString="Dirickx, HG, 1998. Catalogue synonymique et geographique des Syrphidae (Diptera) de la region Afrotropicale. Museum d'histoire naturelle Geneve, Geneve" title="Catalogue synonymique et geographique des Syrphidae (Diptera) de la region Afrotropicale." year="1998">Dirickx 1998</bibRefCitation>
: 4.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<taxonomicName baseAuthorityName="Doesburg" baseAuthorityYear="1956" class="Insecta" family="Syrphidae" genus="Eumerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumerus braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">Eumerus braunsi</taxonomicName>
(van Doesburg) -
<bibRefCitation author="Whittington, AE" journalOrPublisher="Studia Dipterologica" pageId="0" pageNumber="149" pagination="579 - 607" refId="B15" refString="Whittington, AE, 2003. The Afrotropical Syrphidae Fauna: An assessment. Studia Dipterologica 10: 579 - 607" title="The Afrotropical Syrphidae Fauna: An assessment." volume="10" year="2003">Whittington 2003</bibRefCitation>
, 10: 592. [citation only]
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="materials_examined">
<paragraph pageId="0" pageNumber="149">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="149">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Barberton, Dr Brauns" country="South Africa" location="Mpumalanga" specimenCount="1" specimenCount-male="1" stateProvince="Mpumalanga" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="South Africa">South Africa</collectingCountry>
•
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
,
<collectorName>Barberton</collectorName>
;
<collectorName>Dr Brauns</collectorName>
leg.; NMSA-Dip 051409, NMSA type number 731.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Other material</emphasis>
:
<materialsCitation collectingDate="1933-01-15" collectingDateMax="1933-01-30" collectingDateMin="1933-01-15" collectorName="R. E. Turner" country="South Africa" location="Katberg" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Cape">
<collectingCountry name="South Africa">South Africa</collectingCountry>
•
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="South Africa" name="Eastern Cape">Eastern Cape</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:7CD4D5F95A9F51B3518F34D659F7A0CD" country="South Africa" name="Katberg" stateProvince="Eastern Cape">Katberg</location>
;
<collectingDate value="1933-01-15" valueMax="1933-01-30" valueMin="1933-01-15">15-30 Jan. 1933</collectingDate>
;
<collectorName>R.E. Turner</collectorName>
leg. (NHMUK)
</materialsCitation>
. •
<materialsCitation collectingDate="1962-01-03" collectorName="B. &amp; P. Stuckenberg" country="South Africa" county="Karkloof" elevation="1500" location="Geekies Farm" municipality="Mount Alida" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingCounty>Karkloof</collectingCounty>
nr
<collectingMunicipality>Mount Alida</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:D25B8D1F722DA7048464A79683DEFA9A" country="South Africa" county="Karkloof" municipality="Mount Alida" name="Geekies Farm" stateProvince="KwaZulu-Natal">Geekies Farm</location>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="1962-01-03">3 Jan. 1962</collectingDate>
;
<collectorName>B. &amp; P. Stuckenberg</collectorName>
leg.; NMSA-Dip 052854 (NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="1971-01" collectorName="H. Townes" country="South Africa" location="Royal Natal National Park" specimenCount="3" specimenCount-female="3" stateProvince="KwaZulu-Natal">
<specimenCount type="female">3♀♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:8DA374259EA55E45181BF85FB622748F" country="South Africa" name="Royal Natal National Park" stateProvince="KwaZulu-Natal">Royal Natal National Park</location>
; 2828DB; late
<collectingDate value="1971-01">Jan. 1971</collectingDate>
;
<collectorName>H. Townes</collectorName>
leg.; NMSA-Dip 059707, 059708, 051873 (NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="1983-12-21" collectorName="B. R. Stuckenberg" country="South Africa" location="Geekies Farm" municipality="Karkloof" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Karkloof</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:A4011C6E04B4E582B39C0A84979E8024" country="South Africa" municipality="Karkloof" name="Geekies Farm" stateProvince="KwaZulu-Natal">Geekies Farm</location>
; 2930AB;
<collectingDate value="1983-12-21">21 Dec. 1983</collectingDate>
;
<collectorName>B.R. Stuckenberg</collectorName>
leg.; NMSA-Dip 056083 (NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="1987-12-10" collectorName="J. &amp; H. Londt, Mixed" country="South Africa" elevation="1260" latitude="-29.302776" location="Karkloof Nature Reserve" longLatPrecision="20" longitude="30.227777" specimenCount="2" specimenCount-female="2" stateProvince="KwaZulu-Natal">
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:B52328EB679B358C4D81AA9357802739" country="South Africa" latitude="-29.302776" longLatPrecision="20" longitude="30.227777" name="Karkloof Nature Reserve" stateProvince="KwaZulu-Natal">Karkloof Nature Reserve</location>
;
<geoCoordinate degrees="29" direction="south" minutes="18" orientation="latitude" precision="15" seconds="10" value="-29.302776">29°18'10&quot;S</geoCoordinate>
,
<geoCoordinate degrees="30" direction="east" minutes="13" orientation="longitude" precision="15" seconds="40" value="30.227777">30°13'40&quot;E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">1260 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="1987-12-10">10 Dec. 1987</collectingDate>
;
<collectorName>J. &amp; H. Londt</collectorName>
leg.;
<collectorName>Mixed</collectorName>
<taxonomicName class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" lsidName="Podocarpus" order="Pinales" pageId="0" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="149">Podocarpus</emphasis>
</taxonomicName>
forest edge; NMSA-Dip 059706, 064223 (NMSA)
</materialsCitation>
. •
<materialsCitation accessionNumber="OQ706113" collectingDate="2008-11-02" collectingDateMax="2008-11-07" collectingDateMin="2008-11-02" collectorName="G. Davies, GenBank" country="South Africa" latitude="-30.266666" location="Umzinto" longLatPrecision="1222" longitude="30.6" municipality="Vernon Crookes Nature Reserve" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Vernon Crookes Nature Reserve</collectingMunicipality>
near
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:2B8EDB93B9C324CBAA8805403B69AED3" country="South Africa" latitude="-30.266666" longLatPrecision="1222" longitude="30.6" municipality="Vernon Crookes Nature Reserve" name="Umzinto" stateProvince="KwaZulu-Natal">Umzinto</location>
;
<geoCoordinate degrees="30" direction="south" minutes="16" orientation="latitude" precision="925" value="-30.266666">30°16'S</geoCoordinate>
,
<geoCoordinate degrees="30" direction="east" minutes="36" orientation="longitude" precision="925" value="30.6">30°36'E</geoCoordinate>
;
<collectingDate value="2008-11-02" valueMax="2008-11-07" valueMin="2008-11-02">2-7 Nov. 2008</collectingDate>
;
<collectorName>G. Davies</collectorName>
leg.; RMCA DNA 165C02; NMSA-Dip 75216,
<collectorName>GenBank</collectorName>
:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OQ706113">OQ706113</accessionNumber>
(NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="2015-01-10" collectorName="M. Reemer" country="South Africa" elevation="1392" latitude="-29.260221" location="Benvie Farm" longLatPrecision="13" longitude="30.352222" municipality="Karkloof" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Karkloof</collectingMunicipality>
, near
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:B4159FA4258A4357704B27769D92356E" country="South Africa" latitude="-29.260221" longLatPrecision="13" longitude="30.352222" municipality="Karkloof" name="Benvie Farm" stateProvince="KwaZulu-Natal">Benvie Farm</location>
;
<geoCoordinate degrees="29" direction="south" minutes="15" orientation="latitude" precision="1" seconds="36.8" value="-29.260221">29°15'36.8&quot;S</geoCoordinate>
,
<geoCoordinate degrees="30" direction="east" minutes="21" orientation="longitude" precision="15" seconds="08" value="30.352222">30°21'08&quot;E</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.392" unit="m" value="1392.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.392" unit="m" value="1392.0">1392 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2015-01-10">10 Jan. 2015</collectingDate>
;
<collectorName>M. Reemer</collectorName>
leg.; MRSA078; (RMNH)
</materialsCitation>
. •
<materialsCitation accessionNumber="OQ706109" collectingDate="2017-12-03" collectorName="G. Stahls, E. Raettel, Stahls, GenBank" country="South Africa" latitude="-29.321667" location="Karkloof Canopy" longLatPrecision="20" longitude="30.261944" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
,
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:1D4497F663ADC74373A496D191226CD6" country="South Africa" latitude="-29.321667" longLatPrecision="20" longitude="30.261944" name="Karkloof Canopy" stateProvince="KwaZulu-Natal">Karkloof Canopy</location>
tour forest;
<geoCoordinate degrees="29" direction="south" minutes="19" orientation="latitude" precision="15" seconds="18" value="-29.321667">29°19'18&quot;S</geoCoordinate>
,
<geoCoordinate degrees="30" direction="east" minutes="15" orientation="longitude" precision="15" seconds="43" value="30.261944">30°15'43&quot;E</geoCoordinate>
;
<collectingDate value="2017-12-03">03 Dec. 2017</collectingDate>
;
<collectorName>
G.
<normalizedToken originalValue="Ståhls">Stahls</normalizedToken>
</collectorName>
&amp;
<collectorName>
E.
<normalizedToken originalValue="Rättel">Raettel</normalizedToken>
</collectorName>
leg.; http://id.luomus.fi/GJ.2928,
<collectorName>Stahls</collectorName>
_Y2423,
<collectorName>GenBank</collectorName>
:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OQ706109">OQ706109</accessionNumber>
(MZHC)
</materialsCitation>
. •
<materialsCitation collectingDate="2020-11-12" collectorName="J. Midgley" country="South Africa" latitude="-29.30356" location="Shawswood" longLatPrecision="1" longitude="30.30606" municipality="Karkloof" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal">
<specimenCount type="female">1♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Karkloof</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:FD54015286004188D04FD32E85E9EF1D" country="South Africa" latitude="-29.30356" longLatPrecision="1" longitude="30.30606" municipality="Karkloof" name="Shawswood" stateProvince="KwaZulu-Natal">Shawswood</location>
;
<geoCoordinate degrees="29.30356" direction="south" orientation="latitude" precision="1" value="-29.30356">-29.30356</geoCoordinate>
,
<geoCoordinate degrees="30.30606" direction="east" orientation="longitude" precision="1" value="30.30606">30.30606</geoCoordinate>
;
<collectingDate value="2020-11-12">12 Nov. 2020</collectingDate>
;
<collectorName>J. Midgley</collectorName>
leg.; NMSA-Dip 206396 (NMSA)
</materialsCitation>
. •
<materialsCitation accessionNumber="OQ706110, OQ706111, OQ706112" collectingDate="2020-11-24" collectorName="J. Midgley, GenBank" country="South Africa" latitude="-29.30356" location="Shawswood" longLatPrecision="4" longitude="7.137" municipality="Karkloof" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Karkloof</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:182A50CFF0DD793FCA1C4CD0E1E00F4C" country="South Africa" latitude="-29.30356" longLatPrecision="4" longitude="7.137" municipality="Karkloof" name="Shawswood" stateProvince="KwaZulu-Natal">Shawswood</location>
;
<geoCoordinate degrees="29.30356" direction="south" orientation="latitude" precision="1" value="-29.30356">-29.30356</geoCoordinate>
,
<geoCoordinate degrees="30.30606" direction="east" orientation="longitude" precision="1" value="30.30606">30.30606</geoCoordinate>
;
<collectingDate value="2020-11-24">24 Nov. 2020</collectingDate>
;
<collectorName>J. Midgley</collectorName>
leg.; RMCA DNA 1370E
<geoCoordinate degrees="06, 1370" direction="east" orientation="longitude" precision="5" value="6.137">06, 1370E</geoCoordinate>
<geoCoordinate degrees="07, 1370" direction="east" orientation="longitude" precision="5" value="7.137">07, 1370E</geoCoordinate>
08; NMSA-Dip 206389, 206390, 206391,
<collectorName>GenBank</collectorName>
:
<accessionNumber isEnumeration="true">OQ706110, OQ706111, OQ706112</accessionNumber>
(NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="2021-01-09" collectorName="L. Mva" country="South Africa" location="Shawswood" municipality="Karkloof" specimenCount="2" specimenCount-female="2" stateProvince="KwaZulu-Natal">
<specimenCount type="female">2♀♀</specimenCount>
;
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Karkloof</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:6B00A3495ED278C36EA47C5F691CE337" country="South Africa" municipality="Karkloof" name="Shawswood" stateProvince="KwaZulu-Natal">Shawswood</location>
;
<collectingDate value="2021-01-09">9 Jan. 2021</collectingDate>
;
<collectorName>L. Mva</collectorName>
leg.; NMSA-Dip 209856, 209857 (NMSA)
</materialsCitation>
. •
<materialsCitation collectingDate="2023-01-22" collectorName="T. Bellingan, K. Jordaens, J. Midgley" country="South Africa" latitude="-28.959932" location="Rainbow Gorge forest" longLatPrecision="1" longitude="29.226849" municipality="Didima" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Didima</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:1C8FF85B29073B1CF96AFF9614EBD303" country="South Africa" latitude="-28.959932" longLatPrecision="1" longitude="29.226849" municipality="Didima" name="Rainbow Gorge forest" stateProvince="KwaZulu-Natal">Rainbow Gorge forest</location>
,
<geoCoordinate degrees="28.959933" direction="south" orientation="latitude" precision="1" value="-28.959932">-28.959933</geoCoordinate>
,
<geoCoordinate degrees="29.226848" direction="east" orientation="longitude" precision="1" value="29.226849">29.226848</geoCoordinate>
;
<collectingDate value="2023-01-22">22 Jan. 2023</collectingDate>
;
<collectorName>T. Bellingan</collectorName>
,
<collectorName>K. Jordaens</collectorName>
&amp;
<collectorName>J. Midgley</collectorName>
leg. (RMCA)
</materialsCitation>
. •
<materialsCitation collectingDate="2023-01-23" collectorName="T. Bellingan, K. Jordaens, J. Midgley" country="South Africa" latitude="-28.959932" location="Rainbow Gorge forest" longLatPrecision="1" longitude="29.226849" municipality="Didima" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
<specimenCount type="male">1♂</specimenCount>
<specimenCount type="female">1♀</specimenCount>
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingMunicipality>Didima</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:DE7EB38412995EB8839E2A297103693C:749E547FFA1AFF712B7A93671E98A50D" country="South Africa" latitude="-28.959932" longLatPrecision="1" longitude="29.226849" municipality="Didima" name="Rainbow Gorge forest" stateProvince="KwaZulu-Natal">Rainbow Gorge forest</location>
,
<geoCoordinate degrees="28.959933" direction="south" orientation="latitude" precision="1" value="-28.959932">-28.959933</geoCoordinate>
,
<geoCoordinate degrees="29.226848" direction="east" orientation="longitude" precision="1" value="29.226849">29.226848</geoCoordinate>
;
<collectingDate value="2023-01-23">23 Jan. 2023</collectingDate>
;
<collectorName>T. Bellingan</collectorName>
,
<collectorName>K. Jordaens</collectorName>
&amp;
<collectorName>J. Midgley</collectorName>
leg. (AMGS)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/afrinvertebr.64.100481.figure2" httpUri="https://binary.pensoft.net/fig/846531" pageId="0" pageNumber="149" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">Figure 2.</emphasis>
Live
<taxonomicName authorityName="van Doesburg" authorityYear="1956" class="Insecta" family="Syrphidae" genus="Amphoterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphoterus braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis italics="true" pageId="0" pageNumber="149">Amphoterus braunsi</emphasis>
</taxonomicName>
, collected from sunny patch in indigenous forest at Rainbow Gorge,
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
,
<collectingCountry name="South Africa">South Africa</collectingCountry>
. Specimen at AMGS.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="description">
<paragraph pageId="0" pageNumber="149">Description.</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">Male</emphasis>
: length 9.5-9.7 mm, wing: 7.5-8 mm.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Head</emphasis>
: Eyes broadly dichoptic. Face dark brown; golden yellow pilose; evenly dense golden yellow pollinose. Gena dark brown; golden yellow pilose; sparse golden yellow pollinose. Oral opening occupies about
<normalizedToken originalValue="½">1/2</normalizedToken>
of width between eyes. Frons dark brown; golden yellow pilose; golden yellow pollinose, with a bare patch above the antennae and a bare band between the eyes about ⅔ of the way to form the antennal bases to the anterior ocellus, with a short suture from the angle of the eye extending about 1/6 of the width of the frons on the bare band. Ocellar triangle dark brown; bare anteriorly, with golden brown to dark brown pilosity posteriorly; lacking pollinosity. Ocellar triangle obtuse, distance between posterior ocelli 1.5 times the distance between anterior ocellus and posterior ocellus. Occiput dark brown; golden yellow to golden brown pilose, darker dorsally and paler laterally; golden yellow pollinose laterally, bare dorsally. Eye bare, facets of equal size across eye. Scape short, brown, with a few thick dark brown hairs at the dorsoapical and ventroapical margins, lateroapical margins bare. Antenna with pedicel elongate, about as long as postpedicel, 4.1 times as long as high, brown, covered in stout dark brown hairs, sparser basally, some white pollinosity near apex. Postpedicel elongate, 3.1 times as long as high, dorsal and ventral margin almost parallel basally, widest at
<normalizedToken originalValue="¾">3/4</normalizedToken>
of length, rounded apically; slightly darker brown than basal antennal segments and darker at apex; sparse white pollinose. Arista brown.
</paragraph>
<caption doi="10.3897/afrinvertebr.64.100481.figures3-6" httpUri="https://binary.pensoft.net/fig/846532" pageId="0" pageNumber="149" start="Figures 3–6" startId="F3">
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">Figures 3-6.</emphasis>
<taxonomicName authorityName="Bezzi" authorityYear="1915" class="Insecta" family="Syrphidae" genus="Amphoterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphoterus" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="149">Amphoterus</emphasis>
</taxonomicName>
species, dorsal view
<emphasis bold="true" pageId="0" pageNumber="149">3</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis italics="true" pageId="0" pageNumber="149">A. braunsi</emphasis>
</taxonomicName>
(♂)
<emphasis bold="true" pageId="0" pageNumber="149">4</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis italics="true" pageId="0" pageNumber="149">A. braunsi</emphasis>
</taxonomicName>
(♀)
<emphasis bold="true" pageId="0" pageNumber="149">5</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. cribratus" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="cribratus">
<emphasis italics="true" pageId="0" pageNumber="149">A. cribratus</emphasis>
</taxonomicName>
(♂)
<emphasis bold="true" pageId="0" pageNumber="149">6</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. londti" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="londti">
<emphasis italics="true" pageId="0" pageNumber="149">A. londti</emphasis>
</taxonomicName>
sp. nov. (♀).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Thorax</emphasis>
: Scutum dark brown, punctate; postpronotum, the lateral part of the transverse suture and the posterior part of the postalar callus paler, brownish yellow; golden yellow pilose, with denser patches at the posterior margin between the postalar calli, with some darker brown pilosity on the margin between the suture and the postalar callus and on the postalar callus; with a horizontal vitta of sparse golden yellow pollinosity from the postpronotum to in line with the posterior corner of the eye and a horizontal vitta of dense golden yellow pollinosity along the transverse suture and three vittae of sparse white pollinosity, one medial and two mediolateral, reaching from the anterior margin of the scutum to in line with the suture. Scutellum with flattened apical rim, dark brown, golden yellow pilose. Posterior anepisternum, anterior anepimeron and dorsal katepisternum with pale pilosity which is golden yellow dorsally gradually becoming white ventrally. Katatergite with thin, fine golden pilosity. Anterior anepisternum, katepisternum, katepimeron and meron with sparse white pollinosity, which extends slightly onto the posterior anepisternum and anepimeron. Metasternum with white pilosity.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Legs</emphasis>
: Reddish brown, except for the coxa, which is dark brown and the metafemur and metatibia, which have some darker markings; golden yellow pilose and a few darker pile on the tarsus. Metafemur somewhat thickened medially, with a shallow, darkened suture posteroventrally at middle of length; metatibia expanded apically, with a small groove ventrally at middle of length. Metabasitarsus large, as thick as metatibia and as long as other tarsal segments combined.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Wing</emphasis>
: Brown infuscated, darker anterior to the spurious vein, sometimes with a small hyaline patch at the base of cell
<emphasis italics="true" pageId="0" pageNumber="149">r1</emphasis>
. Cell
<emphasis italics="true" pageId="0" pageNumber="149">r1</emphasis>
open for about ⅓ of its length. Vein
<emphasis italics="true" pageId="0" pageNumber="149">R2+3</emphasis>
straight for most of its length, turning sharply upwards as it joins the wing margin. Vein
<emphasis italics="true" pageId="0" pageNumber="149">R4+5</emphasis>
straight. Crossvein
<emphasis italics="true" pageId="0" pageNumber="149">r-m</emphasis>
with a bend posteriorly, at about ⅔ of length of vein where the spurious vein would cross crossvein
<emphasis italics="true" pageId="0" pageNumber="149">r-m</emphasis>
. Vein
<emphasis italics="true" pageId="0" pageNumber="149">M1</emphasis>
recessive, with one or two appendices projecting towards the wing margin. Cell
<emphasis italics="true" pageId="0" pageNumber="149">dm</emphasis>
usually with a single appendix at posterior corner. Spurious vein weak, indistinct, only visible as a thickened fold. Entire wing microtrichose. Alula about 2.2 times as long as broad. Calypter yellowish, with long golden yellow pilosity. Haltere stem brown basally, becoming lighter distally, knob white.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Abdomen</emphasis>
: Punctate, dark brown, with yellow golden pilosity. Tergite (hereafter
<normalizedToken originalValue="“T”">&quot;T&quot;</normalizedToken>
) 2 somewhat trapezoidal, about half as long as wide, with longer pilosity at anterior corners, anterolateral and dorsomedial sections raised somewhat, with diagonal vittae of golden yellow pollinosity in the grooves between these sections. T3 parallel sided, about half as long as wide, with diagonal vittae of golden yellow pollinosity in shallower grooves than T2. T4 parallel sided anteriorly, rounded posteriorly, about as long as wide, evenly rounded dorsally, diagonal golden yellow pollinose vittae not in grooves.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Genitalia</emphasis>
: Hypandrium broad in lateral view, cerci trapezoid (Figs
<figureCitation captionStart="Figures 15–20" captionStartId="F6" captionText="Figures 15 - 20. Amphoterus male genitalia 15 - 17 A. braunsi 15 apical view 16 ventral view 17 lateral view 18 - 20 A. cribratus 18 apical view 19 ventral view 20 lateral view." figureDoi="10.3897/afrinvertebr.64.100481.figures15-20" httpUri="https://binary.pensoft.net/fig/846535" pageId="0" pageNumber="149">15-17</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">Female</emphasis>
(as for male, except as noted): length 8.3-10.4 mm, wing 6.7-8.5 mm.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Head</emphasis>
: Dense golden yellow pollinosity over most of the face, but with two vittae of less dense pollinosity running from base of antennae to outer oral margin. Ocellar triangle obtuse, distance between posterior ocelli 1.6 times the distance between anterior ocellus and posterior ocellus. Frons more extensively bare than in male, golden yellow pilosity limited to lateral margins and a thin vitta bellow ocellar triangle, without sutures. Pedicel similar colour brown to basal antennal segments.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Thorax</emphasis>
: Entire postalar calli pale.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Wing</emphasis>
: Vein
<emphasis italics="true" pageId="0" pageNumber="149">M1</emphasis>
often with a third appendix projecting into cell
<emphasis italics="true" pageId="0" pageNumber="149">r4+5</emphasis>
. Allula 2.1 times as long as broad.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Abdomen</emphasis>
: T2 parallel sided.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="comments">
<paragraph pageId="0" pageNumber="149">Comments.</paragraph>
<paragraph pageId="0" pageNumber="149">
Compared to other species in the genus, a large number of specimens of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. braunsi" order="Diptera" pageId="0" pageNumber="149" phylum="Arthropoda" rank="species" species="braunsi">
<emphasis italics="true" pageId="0" pageNumber="149">A. braunsi</emphasis>
</taxonomicName>
are available for study. This is likely due to higher collecting effort in South Africa than in other African countries and not necessarily an indication of relative abundance. Several collecting events resulted in multiple specimens, suggesting that adults of this species may be abundant at particular times or in particular habitats. In specimens where habitat information is available, individuals were collected from indigenous forests. From our own observations, they are collected in sunny patches with or without flowers, where they rest on leaves or flowers (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Amphoterus braunsi, collected from sunny patch in indigenous forest at Rainbow Gorge, KwaZulu-Natal, South Africa. Specimen at AMGS." figureDoi="10.3897/afrinvertebr.64.100481.figure2" httpUri="https://binary.pensoft.net/fig/846531" pageId="0" pageNumber="149">2</figureCitation>
). Detailed flower visiting information is not available at present.
</paragraph>
</subSubSection>
</treatment>
</document>
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