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<document id="3F2EA665D65FEEE2B42B5C72EB5BB828" ID-CLB-Dataset="7088" ID-DOI="10.11646/zootaxa.4963.3.11" ID-GBIF-Dataset="37bbbcdd-7455-4fd2-854c-49f010b3b27c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4704278" ID-ZooBank="05C2E47F-426D-42EF-B07E-D586C63B6527" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618904250788" checkinUser="plazi" docAuthor="Liu, Hsing-Che &amp; Fikáček, Martin" docDate="2021" docId="03C29A73FFCF410BFF4BA47D069BFDB9" docLanguage="en" docName="zootaxa.4963.3.11.pdf" docOrigin="Zootaxa 4963 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius (Microlaccobius) roseiceps Regimbart 1903" docType="treatment" docVersion="6" lastPageNumber="596" masterDocId="FFFBE20BFFC74102FFDCA234052DFFA0" masterDocTitle="Review of the Taiwanese Laccobius Erichson, 1837 (Coleoptera: Hydrophilidae) with description of a new species" masterLastPageNumber="599" masterPageNumber="587" pageNumber="595" updateTime="1698937256683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E71D4B7C7BC3EF2880B21EA540137CA5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EBD953537A97C1D2CABDB853C7F1EA23">
<mods:title id="C66FE5450FB82A4C6349049BACCA9DF7">Review of the Taiwanese Laccobius Erichson, 1837 (Coleoptera: Hydrophilidae) with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="DA9C0C6D5189FB88DB00D45950226222" type="personal">
<mods:role id="6AEFB71A6B5D41B3A0399FAD9EBB33F0">
<mods:roleTerm id="09C469A4EE127F81533A65852273A0FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12B60A86D5644FA8CAE8791491AC023D">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="DF4263A27AE27B659F3F576088B1A112">Taiwanese Society of Insect Natural History, No. 8, Wufeng Rd., Hsinchu County 310, Taiwan.</mods:affiliation>
</mods:name>
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<mods:namePart id="D5D48DDB39A0CE094510DE67D10FB9D7">Fikáček, Martin</mods:namePart>
<mods:nameIdentifier id="0C9BD70ED0E4FD97FDD75700CCAE86E9" type="ORCID">0000-0002-2078-6798</mods:nameIdentifier>
<mods:affiliation id="30ACEDEB5156D50D266EFD5CC1DD7945">Department of Biological Sciences, National Sun Yat-sen University, No. 70 Lien-hai Rd., Kaohsiung City 80424, Taiwan. &amp; Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 - Horní Počernice, Czech Republic. &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic. mfikacek @ gmail. com; https: // orcid. org / 0000 - 0002 - 2078 - 6798</mods:affiliation>
<mods:nameIdentifier id="9BDB2F52C304DD1ACC76A2C2AB099BDB" type="email">mfikacek@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="0474207A7E4FE032A470E2DCD6A4867E">Zootaxa</mods:title>
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<mods:part id="8DDF2F10CADDC3990AD16EF85DEBE5F7">
<mods:date id="36ECC0061E3BE5558C0DA976BE04CA89">2021</mods:date>
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<mods:number id="EB206ED61B63C1BDB5D1AD2A1AE8A73B">2021-04-20</mods:number>
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<mods:detail id="082D67DCB7D12580C0D44573D336F047" type="volume">
<mods:number id="9F4C62B6EC991F38D6E02FFB992C3B1E">4963</mods:number>
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<mods:classification id="1D48EA8D24B7275436609D505F3A7644">journal article</mods:classification>
<mods:identifier id="E222826B9F1B01D861EB9EB2FD866A9A" type="CLB-Dataset">7088</mods:identifier>
<mods:identifier id="A5DEE8CCA8E13F314BB6045AE7D8EF8A" type="DOI">10.11646/zootaxa.4963.3.11</mods:identifier>
<mods:identifier id="852F866ED24E3FE31DC4B840619E3A2F" type="GBIF-Dataset">37bbbcdd-7455-4fd2-854c-49f010b3b27c</mods:identifier>
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<treatment id="03C29A73FFCF410BFF4BA47D069BFDB9" ID-DOI="http://doi.org/10.5281/zenodo.4704303" ID-GBIF-Taxon="181874752" ID-Zenodo-Dep="4704303" LSID="urn:lsid:plazi:treatment:03C29A73FFCF410BFF4BA47D069BFDB9" httpUri="http://treatment.plazi.org/id/03C29A73FFCF410BFF4BA47D069BFDB9" lastPageId="9" lastPageNumber="596" pageId="8" pageNumber="595">
<subSubSection id="C37178EEFFCF410AFF4BA47D06C3F9C2" box="[151,1006,1609,1636]" pageId="8" pageNumber="595" type="nomenclature">
<paragraph id="8BD42B65FFCF410AFF4BA47D06C3F9C2" blockId="8.[151,1006,1609,1673]" box="[151,1006,1609,1636]" pageId="8" pageNumber="595">
<heading id="D09C9C09FFCF410AFF4BA47D06C3F9C2" bold="true" box="[151,1006,1609,1636]" fontSize="11" level="1" pageId="8" pageNumber="595" reason="1">
<emphasis id="B91FF777FFCF410AFF4BA47D06C3F9C2" bold="true" box="[151,1006,1609,1636]" pageId="8" pageNumber="595">
<taxonomicName id="4C6B50E6FFCF410AFF4BA47D0690F9C2" ID-CoL="3RLRH" authority="Regimbart, 1903 R Aflo" authorityName="Regimbart" authorityYear="1903" box="[151,957,1609,1636]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="595" phylum="Arthropoda" rank="species" species="roseiceps" subGenus="Microlaccobius">
<emphasis id="B91FF777FFCF410AFF4BA47D043EF9C4" bold="true" box="[151,275,1609,1636]" italics="true" pageId="8" pageNumber="595">Laccobius</emphasis>
(
<emphasis id="B91FF777FFCF410AFEF8A47D04F2F9C4" bold="true" box="[292,479,1609,1636]" italics="true" pageId="8" pageNumber="595">Microlaccobius</emphasis>
)
<emphasis id="B91FF777FFCF410AFE2DA47E074DF9C4" bold="true" box="[497,608,1610,1636]" italics="true" pageId="8" pageNumber="595">roseiceps</emphasis>
Régimbart, 1903 RẦflǿ
</taxonomicName>
牙ḃ
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFCF410AFF4BA45A0456F929" box="[151,379,1646,1673]" pageId="8" pageNumber="595" type="description">
<paragraph id="8BD42B65FFCF410AFF4BA45A0456F929" blockId="8.[151,1006,1609,1673]" box="[151,379,1646,1673]" pageId="8" pageNumber="595">
(
<figureCitation id="135037E0FFCF410AFF43A45A040DF928" box="[159,288,1646,1672]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1423,1447]" captionTargetBox="[163,1412,192,1363]" captionTargetId="figure-24@4.[151,1436,181,1400]" captionTargetPageId="4" captionText="FIGURES 10–18. Dosal view of the habitus and male genitalia of Taiwanese Laccobius species (10–13: habitus, 14–18: male genitalia). 10, 14: L. politus Gentili, 1979. 11, 15: L. formosus Gentili, 1979. 12, 16: L. hammondi Gentili, 1984. 13, 17: L. roseiceps Régimbart, 1903. 18: L. fragilis Nakane, 1966, after Nakane (1966)." figureDoi="http://doi.org/10.5281/zenodo.4704291" httpUri="https://zenodo.org/record/4704291/files/figure.png" pageId="8" pageNumber="595">Figs 13, 17</figureCitation>
,
<figureCitation id="135037E0FFCF410AFEF0A45A0466F928" box="[300,331,1646,1672]" captionStart="FIGURES 19–22" captionStartId="6.[151,264,984,1008]" captionTargetBox="[165,1406,237,940]" captionTargetId="figure-24@6.[151,1436,181,960]" captionTargetPageId="6" captionText="FIGURES 19–22. Pronotum of Taiwanese Laccobius (Microlaccobius) species. 19: L. formosus Gentili, 1979. 20: L. hammondi Gentili, 1984. 21: L. hoi sp. nov. 22: L. roseiceps Régimbart, 1903." figureDoi="http://doi.org/10.5281/zenodo.4704295" httpUri="https://zenodo.org/record/4704295/files/figure.png" pageId="8" pageNumber="595">22</figureCitation>
,
<figureCitation id="135037E0FFCF410AFE8AA45A045EF929" box="[342,371,1646,1673]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1431,1455]" captionTargetBox="[184,1404,189,1402]" captionTargetId="figure-24@8.[177,1411,181,1408]" captionTargetPageId="8" captionText="FIGURES 23–26. Habitat of Taiwanese Laccobius species. 23: Nantou County, Puli: habitat of L. formosus. 24: Taichung City, Wufeng: habitat of L. hammondi and L. hoi sp. nov. 25: Pingtung County, Ligang: habitat of L. formosus, L. hoi sp. nov. and L. roseiceps. 26: New Taipei City, Pingxi: habitat of L. politus." figureDoi="http://doi.org/10.5281/zenodo.4704299" httpUri="https://zenodo.org/record/4704299/files/figure.png" pageId="8" pageNumber="595">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFCF410AFF4BA48607D5F96A" box="[151,760,1714,1738]" pageId="8" pageNumber="595" type="reference_group">
<paragraph id="8BD42B65FFCF410AFF4BA48607D5F96A" blockId="8.[151,760,1714,1738]" box="[151,760,1714,1738]" pageId="8" pageNumber="595">
<taxonomicName id="4C6B50E6FFCF410AFF4BA48607D9F96A" ID-CoL="3RLRH" authority="Regimbart, 1903: 59" authorityName="Regimbart" authorityPageNumber="59" authorityYear="1903" box="[151,756,1714,1738]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="595" phylum="Arthropoda" rank="species" species="roseiceps" subGenus="Microlaccobius">
<emphasis id="B91FF777FFCF410AFF4BA48605D2F96A" box="[151,255,1714,1738]" italics="true" pageId="8" pageNumber="595">Laccobius</emphasis>
(
<emphasis id="B91FF777FFCF410AFED2A4860481F96A" box="[270,428,1714,1738]" italics="true" pageId="8" pageNumber="595">Microlaccobius</emphasis>
)
<emphasis id="B91FF777FFCF410AFE67A4870737F96A" box="[443,538,1715,1738]" italics="true" pageId="8" pageNumber="595">roseiceps</emphasis>
Régimbart, 1903: 59
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFCF410AFF4BA4CC00AAF8DE" pageId="8" pageNumber="595" type="materials_examined">
<paragraph id="8BD42B65FFCF410AFF4BA4CC00AAF8DE" blockId="8.[151,1437,1784,2027]" pageId="8" pageNumber="595">
<emphasis id="B91FF777FFCF410AFF4BA4CC04A8F8B2" bold="true" box="[151,389,1784,1810]" pageId="8" pageNumber="595">Material examined.</emphasis>
<materialsCitation id="3B032138FFCF410AFE53A4CC060CF896" ID-GBIF-Occurrence="3107229309" collectingDate="2019-04-26" collectionCode="HCLC" country="Taiwan" elevation="32" latitude="22.7982" location="Erchong River" longLatPrecision="7" longitude="120.5011" municipality="Ligang Township" pageId="8" pageNumber="595" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Pingtung">
<emphasis id="B91FF777FFCF410AFE53A4CC07A0F8B3" bold="true" box="[399,653,1784,1811]" pageId="8" pageNumber="595">
<collectingCountry id="F37C6BF5FFCF410AFE53A4CC0728F8B2" box="[399,517,1784,1810]" name="Taiwan" pageId="8" pageNumber="595">TAIWAN</collectingCountry>
:
<collectingRegion id="49AFE587FFCF410AFDC9A4CD07A9F8B1" box="[533,644,1785,1811]" country="Taiwan" name="Pingtung" pageId="8" pageNumber="595">PINGTUNG</collectingRegion>
:
</emphasis>
<specimenCount id="9D6DE0ECFFCF410AFD44A4CD07DBF8B2" box="[664,758,1784,1811]" pageId="8" pageNumber="595" type="male">3 males</specimenCount>
,
<specimenCount id="9D6DE0ECFFCF410AFCD9A4CC0658F8B2" box="[773,885,1784,1810]" pageId="8" pageNumber="595" type="female">6 females</specimenCount>
(
<collectionCode id="ED7AB3A0FFCF410AFC55A4CD06F6F8B3" box="[905,987,1785,1811]" pageId="8" pageNumber="595">HCLC</collectionCode>
):
<collectingMunicipality id="6BB0B11FFFCF410AFC2DA4CC0194F8B2" box="[1009,1209,1784,1810]" pageId="8" pageNumber="595">Ligang Township</collectingMunicipality>
[
<emphasis id="B91FF777FFCF410AFB12A4CD000FF8B0" bold="true" box="[1230,1314,1785,1808]" pageId="8" pageNumber="595">¶ffiė</emphasis>
],
<location id="8EB47DBEFFCF410AFAE1A4CC05F8F896" LSID="urn:lsid:plazi:treatment:03C29A73FFCF410BFF4BA47D069BFDB9:8EB47DBEFFCF410AFAE1A4CC05F8F896" country="Taiwan" latitude="22.7982" longLatPrecision="7" longitude="120.5011" municipality="Ligang Township" name="Erchong River" pageId="8" pageNumber="595" stateProvince="Pingtung">Erchong River</location>
[
<emphasis id="B91FF777FFCF410AFF3AA5290417F894" bold="true" box="[230,314,1821,1844]" pageId="8" pageNumber="595">二AEë</emphasis>
],
<geoCoordinate id="EE5F4DA2FFCF410AFE8EA52904E0F897" box="[338,461,1821,1847]" degrees="22.7982" direction="north" orientation="latitude" pageId="8" pageNumber="595" precision="5" value="22.7982">22.7982°N</geoCoordinate>
<geoCoordinate id="EE5F4DA2FFCF410AFE09A5280771F897" box="[469,604,1820,1847]" degrees="120.5011" direction="east" orientation="longitude" pageId="8" pageNumber="595" precision="5" value="120.5011">120.5011°E</geoCoordinate>
,
<quantity id="4C938680FFCF410AFDB4A52907B4F897" box="[616,665,1821,1847]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="595" unit="m" value="32.0">
<elevation id="0046CC56FFCF410AFDB4A52907B4F897" box="[616,665,1821,1847]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="595" unit="m" value="32.0">32m</elevation>
</quantity>
<date id="FFD50DA5FFCF410AFD7DA528060CF896" box="[673,801,1820,1846]" pageId="8" pageNumber="595" value="2019-04-26">
<collectingDate id="EF91F44DFFCF410AFD7DA528060CF896" box="[673,801,1820,1846]" pageId="8" pageNumber="595" value="2019-04-26">26.IV.2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B032138FFCF410AFCF0A52806FAF896" ID-GBIF-Occurrence="3107229322" box="[812,983,1820,1846]" country="Taiwan" location="Hsing-Che Liu" pageId="8" pageNumber="595" specimenCount="1" stateProvince="Pingtung">
<location id="8EB47DBEFFCF410AFCF0A52806FAF896" LSID="urn:lsid:plazi:treatment:03C29A73FFCF410BFF4BA47D069BFDB9:8EB47DBEFFCF410AFCF0A52806FAF896" box="[812,983,1820,1846]" country="Taiwan" name="Hsing-Che Liu" pageId="8" pageNumber="595" stateProvince="Pingtung">Hsing-Che Liu</location>
</materialsCitation>
.
<materialsCitation id="3B032138FFCF410AFC3FA5290473F8DE" ID-GBIF-Occurrence="3107229301" collectingDate="2020-06-24" collectionCode="HCLC" collectorName="Hsing-Che Liu &amp; Uitsiann Ong" country="Taiwan" elevation="229" latitude="22.996017" location="Laonong River" longLatPrecision="1" longitude="120.638176" municipality="Liouguei District" pageId="8" pageNumber="595" specimenCount="1" stateProvince="Kaohsiung">
<emphasis id="B91FF777FFCF410AFC3FA5290141F897" bold="true" box="[995,1132,1821,1847]" pageId="8" pageNumber="595">
<collectingRegion id="49AFE587FFCF410AFC3FA529014EF895" box="[995,1123,1821,1847]" country="Taiwan" name="Kaohsiung" pageId="8" pageNumber="595">KAOHSIUNG</collectingRegion>
:
</emphasis>
<specimenCount id="9D6DE0ECFFCF410AFBA8A52901EEF897" box="[1140,1219,1821,1847]" pageId="8" pageNumber="595" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED7AB3A0FFCF410AFB0EA5290009F897" box="[1234,1316,1821,1847]" pageId="8" pageNumber="595">HCLC</collectionCode>
):
<collectingMunicipality id="6BB0B11FFFCF410AFAEBA52805C6F8FA" pageId="8" pageNumber="595">Liouguei District</collectingMunicipality>
[
<emphasis id="B91FF777FFCF410AFF23A575047EF8F8" bold="true" box="[255,339,1857,1880]" pageId="8" pageNumber="595">六AE區</emphasis>
],
<location id="8EB47DBEFFCF410AFEB2A5750734F8FA" LSID="urn:lsid:plazi:treatment:03C29A73FFCF410BFF4BA47D069BFDB9:8EB47DBEFFCF410AFEB2A5750734F8FA" box="[366,537,1856,1883]" country="Taiwan" latitude="22.996017" longLatPrecision="1" longitude="120.638176" municipality="Liouguei District" name="Laonong River" pageId="8" pageNumber="595" stateProvince="Kaohsiung">Laonong River</location>
[
<emphasis id="B91FF777FFCF410AFDF1A57507ACF8F8" bold="true" box="[557,641,1857,1880]" pageId="8" pageNumber="595">Ž濃ë</emphasis>
],
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<geoCoordinate id="EE5F4DA2FFCF410AFCE1A57406CCF8FA" box="[829,993,1856,1882]" degrees="120.638178" direction="east" orientation="longitude" pageId="8" pageNumber="595" precision="1" value="120.638176">120.638178°E</geoCoordinate>
,
<quantity id="4C938680FFCF410AFC33A575011FF8FB" box="[1007,1074,1857,1883]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="8" pageNumber="595" unit="m" value="229.0">
<elevation id="0046CC56FFCF410AFC33A575011FF8FB" box="[1007,1074,1857,1883]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="8" pageNumber="595" unit="m" value="229.0">229m</elevation>
</quantity>
,
<date id="FFD50DA5FFCF410AFB9DA57501E9F8FB" box="[1089,1220,1857,1883]" pageId="8" pageNumber="595" value="2020-06-24">
<collectingDate id="EF91F44DFFCF410AFB9DA57501E9F8FB" box="[1089,1220,1857,1883]" pageId="8" pageNumber="595" value="2020-06-24">24.VI.2020</collectingDate>
</date>
.
<collectorName id="269E4EB3FFCF410AFB0EA5740051F8FA" box="[1234,1404,1856,1882]" pageId="8" pageNumber="595">Hsing-Che Liu</collectorName>
&amp;
<collectorName id="269E4EB3FFCF410AFF4BA5500400F8DF" box="[151,301,1892,1919]" pageId="8" pageNumber="595">Uitsiann Ong</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B032138FFCF410AFEB9A55100AAF8DE" ID-GBIF-Occurrence="3107229326" box="[357,1415,1892,1919]" collectingDate="2020-07-05" collectingDateMax="2020-07-10" collectingDateMin="2020-07-05" collectionCode="HCLC, NCHU" collectorName="Hsing-Che Liu" country="Taiwan" location="Main Island" pageId="8" pageNumber="595" specimenCount="22" specimenCount-male="2" stateProvince="Kinmen">
<emphasis id="B91FF777FFCF410AFEB9A55104EAF8DF" bold="true" box="[357,455,1893,1919]" pageId="8" pageNumber="595">
<collectingRegion id="49AFE587FFCF410AFEB9A5510493F8DC" box="[357,446,1893,1919]" country="Taiwan" name="Kinmen" pageId="8" pageNumber="595">KINMEN</collectingRegion>
:
</emphasis>
<specimenCount id="9D6DE0ECFFCF410AFE12A551070AF8DE" box="[462,551,1892,1919]" pageId="8" pageNumber="595" type="male">2 males</specimenCount>
,
<specimenCount id="9D6DE0ECFFCF410AFDEEA55107A2F8DF" box="[562,655,1893,1919]" pageId="8" pageNumber="595" type="generic">20 spec.</specimenCount>
(
<collectionCode id="ED7AB3A0FFCF410AFD41A55107C1F8DF" box="[669,748,1893,1919]" pageId="8" pageNumber="595">HCLC</collectionCode>
,
<collectionCode id="ED7AB3A0FFCF410AFD2BA5510661F8DF" box="[759,844,1893,1919]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="8" pageNumber="595">NCHU</collectionCode>
):
<location id="8EB47DBEFFCF410AFC83A55006C5F8DE" LSID="urn:lsid:plazi:treatment:03C29A73FFCF410BFF4BA47D069BFDB9:8EB47DBEFFCF410AFC83A55006C5F8DE" box="[863,1000,1892,1918]" country="Taiwan" name="Main Island" pageId="8" pageNumber="595" stateProvince="Kinmen">Main Island</location>
,
<date id="FFD50DA5FFCF410AFC2FA5500189F8DF" box="[1011,1188,1892,1919]" pageId="8" pageNumber="595" value="2020-07-05" valueMax="2020-07-10" valueMin="2020-07-05">
<collectingDate id="EF91F44DFFCF410AFC2FA5500189F8DF" box="[1011,1188,1892,1919]" pageId="8" pageNumber="595" value="2020-07-05" valueMax="2020-07-10" valueMin="2020-07-05">5-10. VII. 2020</collectingDate>
</date>
,
<collectorName id="269E4EB3FFCF410AFB73A550007BF8DE" box="[1199,1366,1892,1918]" pageId="8" pageNumber="595">Hsing-Che Liu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFCF410BFF1BA5BC0660FE29" lastPageId="9" lastPageNumber="596" pageId="8" pageNumber="595" type="diagnosis">
<paragraph id="8BD42B65FFCF410BFF1BA5BC0640FF59" blockId="8.[151,1437,1784,2027]" lastBlockId="9.[151,1437,150,537]" lastPageId="9" lastPageNumber="596" pageId="8" pageNumber="595">
<emphasis id="B91FF777FFCF410AFF1BA5BC046FF802" bold="true" box="[199,322,1928,1954]" pageId="8" pageNumber="595">Diagnosis.</emphasis>
Small species, body length
<quantity id="4C938680FFCF410AFD55A5BD0638F803" box="[649,789,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="8" pageNumber="595" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7–2.2 mm</quantity>
, body width 1.0–
<quantity id="4C938680FFCF410AFC3CA5BD0117F803" box="[992,1082,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="595" unit="mm" value="1.3">1.3 mm</quantity>
. Body elongate oval (
<figureCitation id="135037E0FFCF410AFAE5A5BC00A3F803" box="[1337,1422,1928,1955]" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1423,1447]" captionTargetBox="[163,1412,192,1363]" captionTargetId="figure-24@4.[151,1436,181,1400]" captionTargetPageId="4" captionText="FIGURES 10–18. Dosal view of the habitus and male genitalia of Taiwanese Laccobius species (10–13: habitus, 14–18: male genitalia). 10, 14: L. politus Gentili, 1979. 11, 15: L. formosus Gentili, 1979. 12, 16: L. hammondi Gentili, 1984. 13, 17: L. roseiceps Régimbart, 1903. 18: L. fragilis Nakane, 1966, after Nakane (1966)." figureDoi="http://doi.org/10.5281/zenodo.4704291" httpUri="https://zenodo.org/record/4704291/files/figure.png" pageId="8" pageNumber="595">Fig. 13</figureCitation>
), weakly convex. Head dark, with distinct preocular spots. Pronotum pale, with small dark oval spot in median, spot leaving large lateral portions of the pronotum yellow. (
<figureCitation id="135037E0FFCF410AFCD1A5E40642F84B" box="[781,879,2000,2027]" captionStart="FIGURES 19–22" captionStartId="6.[151,264,984,1008]" captionTargetBox="[165,1406,237,940]" captionTargetId="figure-24@6.[151,1436,181,960]" captionTargetPageId="6" captionText="FIGURES 19–22. Pronotum of Taiwanese Laccobius (Microlaccobius) species. 19: L. formosus Gentili, 1979. 20: L. hammondi Gentili, 1984. 21: L. hoi sp. nov. 22: L. roseiceps Régimbart, 1903." figureDoi="http://doi.org/10.5281/zenodo.4704295" httpUri="https://zenodo.org/record/4704295/files/figure.png" pageId="8" pageNumber="595">Figs. 22</figureCitation>
). Scutellar shield dark. Elytra pale, with densely arranged small dark spots, larger spots present medially and laterally. Leg pale or yellowish.
<emphasis id="B91FF777FFCE410BFB6AA2A20073FF11" bold="true" box="[1206,1374,150,177]" italics="true" pageId="9" pageNumber="596">Male genitalia</emphasis>
(
<figureCitation id="135037E0FFCE410BFAB3A2A3059BFF75" captionStart="FIGURES 10–18" captionStartId="4.[151,264,1423,1447]" captionTargetBox="[163,1412,192,1363]" captionTargetId="figure-24@4.[151,1436,181,1400]" captionTargetPageId="4" captionText="FIGURES 10–18. Dosal view of the habitus and male genitalia of Taiwanese Laccobius species (10–13: habitus, 14–18: male genitalia). 10, 14: L. politus Gentili, 1979. 11, 15: L. formosus Gentili, 1979. 12, 16: L. hammondi Gentili, 1984. 13, 17: L. roseiceps Régimbart, 1903. 18: L. fragilis Nakane, 1966, after Nakane (1966)." figureDoi="http://doi.org/10.5281/zenodo.4704291" httpUri="https://zenodo.org/record/4704291/files/figure.png" pageId="9" pageNumber="596">Fig. 17</figureCitation>
): Aedeagus
<quantity id="4C938680FFCE410BFEE3A28F04B8FF75" box="[319,405,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="596" unit="mm" value="0.4">0.4 mm</quantity>
long. Median lobe wide, its apex rounded, with a pair of diverging lines running from apex; parameres slender, larger than median lobe, slightly bent inward.
</paragraph>
<paragraph id="8BD42B65FFCE410BFF1BA3360660FE29" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="596">
<emphasis id="B91FF777FFCE410BFF1BA33604E2FEBC" bold="true" box="[199,463,258,284]" pageId="9" pageNumber="596">Differential diagnosis.</emphasis>
<taxonomicName id="4C6B50E6FFCE410BFE04A3370791FEBC" authorityName="Regimbart" authorityYear="1903" box="[472,700,259,285]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="roseiceps">
<emphasis id="B91FF777FFCE410BFE04A3370791FEBC" box="[472,700,259,285]" italics="true" pageId="9" pageNumber="596">Laccobius roseiceps</emphasis>
</taxonomicName>
may be easily recognized by the pronotum with small oval dark spot. In this character, it may resemble
<taxonomicName id="4C6B50E6FFCE410BFD94A31307A4FEE1" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[584,649,295,321]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFCE410BFD94A31307A4FEE1" box="[584,649,295,321]" italics="true" pageId="9" pageNumber="596">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFCE410BFD53A31307C8FEE1" box="[655,741,295,321]" pageId="9" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
from which it mainly differs in smaller body size (
<quantity id="4C938680FFCE410BFAC9A31300B0FEE1" box="[1301,1437,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="9" pageNumber="596" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7–2.2 mm</quantity>
in
<taxonomicName id="4C6B50E6FFCE410BFF69A37F0411FEC4" authorityName="Regimbart" authorityYear="1903" box="[181,316,331,356]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="roseiceps">
<emphasis id="B91FF777FFCE410BFF69A37F0411FEC4" box="[181,316,331,356]" italics="true" pageId="9" pageNumber="596">L. roseiceps</emphasis>
</taxonomicName>
,
<quantity id="4C938680FFCE410BFE97A37E04FEFEC5" box="[331,467,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="9" pageNumber="596" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
in
<taxonomicName id="4C6B50E6FFCE410BFE24A37F0717FEC5" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[504,570,331,357]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFCE410BFE24A37F0717FEC5" box="[504,570,331,357]" italics="true" pageId="9" pageNumber="596">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFCE410BFD9EA37F07B3FEC5" box="[578,670,331,357]" pageId="9" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
). The structure of the median lobe (with the diverging lines) is very typical and hard-to-confuse with any other
<taxonomicName id="4C6B50E6FFCE410BFDA7A35B0660FE29" box="[635,845,367,393]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91FF777FFCE410BFDA7A35B07C3FE29" box="[635,750,367,393]" italics="true" pageId="9" pageNumber="596">Laccobius</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFCE410BFF1BA3A6069BFDB9" pageId="9" pageNumber="596" type="distribution">
<paragraph id="8BD42B65FFCE410BFF1BA3A60149FE71" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="596">
<emphasis id="B91FF777FFCE410BFF1BA3A6044CFE0C" bold="true" box="[199,353,402,428]" pageId="9" pageNumber="596">Distribution.</emphasis>
<collectingCountry id="F37C6BF5FFCE410BFEBAA3A70484FE0D" box="[358,425,403,429]" name="China" pageId="9" pageNumber="596">China</collectingCountry>
(
<collectingCountry id="F37C6BF5FFCE410BFE69A3A70711FE0D" box="[437,572,403,429]" name="Hong Kong" pageId="9" pageNumber="596">Hong Kong</collectingCountry>
),
<collectingCountry id="F37C6BF5FFCE410BFD92A3A707A3FE0D" box="[590,654,403,429]" name="Japan" pageId="9" pageNumber="596">Japan</collectingCountry>
(Honshu, Kyushu, Tanegashima),
<collectingCountry id="F37C6BF5FFCE410BFBD6A3A70171FE0D" box="[1034,1116,403,429]" name="Taiwan" pageId="9" pageNumber="596">Taiwan</collectingCountry>
(
<collectingRegion id="49AFE587FFCE410BFBB5A3A701E8FE0D" box="[1129,1221,403,429]" country="Taiwan" name="Kinmen" pageId="9" pageNumber="596">Kinmen</collectingRegion>
islands) (
<bibRefCitation id="EFFA5694FFCE410BFAF1A3A7043FFE71" author="Nakajima, J. &amp; Hayashi, M. &amp; Ishida, K. &amp; Kitano, T. &amp; Yoshitomi, H." pageId="9" pageNumber="596" refId="ref8671" refString="Nakajima, J., Hayashi, M., Ishida, K., Kitano, T. &amp; Yoshitomi, H. (2020) Aquatic Coleoptera and Hemiptera of Japan. Bun-ichi Sogo Shuppan, Tokyo, 351 pp." type="book" year="2020">
Nakajima
<emphasis id="B91FF777FFCE410BFF4BA38305E5FE71" box="[151,200,439,465]" italics="true" pageId="9" pageNumber="596">et al</emphasis>
. 2020
</bibRefCitation>
;
<bibRefCitation id="EFFA5694FFCE410BFEC1A38304CCFE71" author="Przewozny, M." box="[285,481,439,465]" pageId="9" pageNumber="596" refId="ref8776" refString="Przewozny, M. (2020) Catalogue of Palearctic Hydrophiloidea (Coleoptera). Internet version 2020 - 01 - 01. Available from: http: // www. waterbeetles. eu (accessed 9 April 2020)" type="url" year="2020">Przewoźny, 2020</bibRefCitation>
,
<bibRefCitation id="EFFA5694FFCE410BFE30A38307B9FE71" author="Liu, H. - C. &amp; Ma, C. - H. &amp; Fikacek, M." box="[492,660,439,465]" pageId="9" pageNumber="596" pagination="39 - 62" refId="ref8526" refString="Liu, H. - C., Ma, C. - H. &amp; Fikacek, M. (2021) Water scavenger beetles of Kinmen County, Taiwan (Coleoptera: Hydrophilidae). Taiwan Journal of Biodiversity, 32 (1), 39 - 62." type="journal article" year="2021">
Liu
<emphasis id="B91FF777FFCE410BFDC5A3830766FE71" box="[537,587,439,465]" italics="true" pageId="9" pageNumber="596">et al</emphasis>
. 2021
</bibRefCitation>
). First record for
<collectingCountry id="F37C6BF5FFCE410BFC84A3830687FE71" box="[856,938,439,465]" name="Taiwan" pageId="9" pageNumber="596">Taiwan</collectingCountry>
island (
<figureCitation id="135037E0FFCE410BFBDFA383017AFE71" box="[1027,1111,439,465]" captionStart="FIGURES 27" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[151,1431,309,1911]" captionTargetId="figure-86@10.[151,1435,309,1921]" captionTargetPageId="10" captionText="FIGURES 27. Map of Taiwan showing the distribution of the Laccobius species." figureDoi="http://doi.org/10.5281/zenodo.4704305" httpUri="https://zenodo.org/record/4704305/files/figure.png" pageId="9" pageNumber="596">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BD42B65FFCE410BFF1BA3EE069BFDB9" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="596">
<emphasis id="B91FF777FFCE410BFF1BA3EE0466FE54" bold="true" box="[199,331,474,500]" pageId="9" pageNumber="596">Bionomics.</emphasis>
All specimens were collected from very slowly running streams. In southern
<collectingCountry id="F37C6BF5FFCE410BFB1CA3EF0038FE55" box="[1216,1301,475,501]" name="Taiwan" pageId="9" pageNumber="596">Taiwan</collectingCountry>
, the species co-occurred syntopically with
<taxonomicName id="4C6B50E6FFCE410BFE31A3CB0703FDB9" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[493,558,511,537]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFCE410BFE31A3CB0703FDB9" box="[493,558,511,537]" italics="true" pageId="9" pageNumber="596">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFCE410BFDE9A3CB07A1FDB9" box="[565,652,511,537]" pageId="9" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6B50E6FFCE410BFD1DA3CB066BFDB9" authorityName="Gentili" authorityYear="1979" box="[705,838,511,537]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B91FF777FFCE410BFD1DA3CB066BFDB9" box="[705,838,511,537]" italics="true" pageId="9" pageNumber="596">L. formosus</emphasis>
</taxonomicName>
(
<figureCitation id="135037E0FFCE410BFC89A3CB0684FDB9" box="[853,937,510,537]" captionStart="FIGURES 23–26" captionStartId="8.[151,264,1431,1455]" captionTargetBox="[184,1404,189,1402]" captionTargetId="figure-24@8.[177,1411,181,1408]" captionTargetPageId="8" captionText="FIGURES 23–26. Habitat of Taiwanese Laccobius species. 23: Nantou County, Puli: habitat of L. formosus. 24: Taichung City, Wufeng: habitat of L. hammondi and L. hoi sp. nov. 25: Pingtung County, Ligang: habitat of L. formosus, L. hoi sp. nov. and L. roseiceps. 26: New Taipei City, Pingxi: habitat of L. politus." figureDoi="http://doi.org/10.5281/zenodo.4704299" httpUri="https://zenodo.org/record/4704299/files/figure.png" pageId="9" pageNumber="596">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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