167 lines
14 KiB
XML
167 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.363.6482" ID-GBIF-Dataset="36e5551d-3881-45a2-810a-386e517c86fc" ID-PMC="PMC3867181" ID-Pensoft-Pub="1313-2970-363-1" ID-PubMed="24363601" ID-ZBK="780695A9CB2E4FECA0560BA6F456185E" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-363-1" ModsDocOrigin="ZooKeys 363" ModsDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" checkinTime="1451246560986" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2013" docId="F94D03C1E6A214EF60370E70CCDC866C" docLanguage="en" docName="ZooKeys 363: 1-161" docOrigin="ZooKeys 363" docSource="http://dx.doi.org/10.3897/zookeys.363.6482" docTitle="Hydrochasma williamsae Mathis & Zatwarnicki, 2013, sp. n." docType="treatment" docUuid="01EE2E2E-0913-4C01-8A71-525CEC06E796" docUuidSource="ZooBank" docVersion="4" lastPageNumber="33" masterDocId="FFF2FFCA0333765855689824FFCDB839" masterDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="31" updateTime="1668157177207" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>363</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.363.6482</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.363.6482</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-363-1</mods:identifier>
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<mods:identifier type="ZBK">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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<mods:identifier type="ZooBank">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050137" LSID="urn:lsid:zoobank.org:act:01EE2E2E-0913-4C01-8A71-525CEC06E796" httpUri="http://treatment.plazi.org/id/F94D03C1E6A214EF60370E70CCDC866C" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
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<subSubSection pageId="30" pageNumber="31" type="nomenclature">
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<paragraph pageId="30" pageNumber="31">
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<pageBreakToken pageId="30" pageNumber="31" start="start">10</pageBreakToken>
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.
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<taxonomicName LSID="http://zoobank.org/01EE2E2E-0913-4C01-8A71-525CEC06E796" class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma williamsae" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="williamsae">Hydrochasma williamsae</taxonomicName>
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<taxonomicNameLabel pageId="30" pageNumber="31">sp. n.</taxonomicNameLabel>
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Figs 1, 11, 57-61
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="diagnosis">
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<paragraph pageId="30" pageNumber="31">Diagnosis.</paragraph>
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<paragraph lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 1.50-2.30 mm.
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<pageBreakToken pageId="31" pageNumber="32" start="start">Head</pageBreakToken>
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: Subglobose, very broad ventrally, oral opening comparatively large. Antennal coloration variable, entirely yellow to nearly evenly divided between yellowish and dark gray, dorsal and anterior surfaces of pedicel and basal flagellomere extensively dark gray. Parafacial silvery white, concolorous with facial coloration; parafacial silvery white, concolorous with face; mesonotum yellowish to golden brown; gena-to-eye ratio 0.25-0.27. Thorax: Pleural area gray (Fig. 1). Wing with costal vein ratio 0.63-0.65; M vein ratio 0.52-0.53. Forefemur bearing a distinctive, comb-like row of stout setulae along anteroventral and posteroventral surfaces; tibiae mostly gray; hindtibia bearing a large, spur-like seta ventroapically (Fig. 11). Abdomen: Tergites 1-4 with broad, medial brown stripe, uneven lateral margin, lacking wedge-shaped, gray to silvery gray areas laterally (Fig. 1); tergite 5 of male truncate apically, mostly gray with faint brown, medial stripe. Male terminalia (Figs 57-60): Combined structures generally moderately elongate and appearing angulate, in posterior view height about 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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width, generally setulose, especially medially at midlength but also along ventral margins; epandrium with dorsal arch above cerci attenuated, not connected, in posterior view (Fig. 57) with cercal cavity forming a broad U, with arms of U robustly developed, ventral portion of arms with each lateral half robustly developed, tapered to dorsal apex, apex pointed medially, dorsal portion of epandrium squarish, especially prominent dorsolateral, rounded angles; ventral portion of epandrium ventrad of midlength, shallow, V-shaped notch, as 2 robust, ventral projections that are tapered to a narrow point ventroapically, forming a deeply and narrowly incised medial incision or narrowly U-shaped pocket, in lateral view (Fig. 58) with very irregular anterior and posterior margins, generally robust, apex narrowly pointed; cerci moderately long, height more than twice width, widely semi-hemispherical (Fig. 57), pointed dorsally, in lateral view (Fig. 58) evenly lunate, not attached lateroventrally or ventrally with epandrium; aedeagus in lateral view (Fig. 60) elongate, length slightly more than 3.5
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<normalizedToken originalValue="×">x</normalizedToken>
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width, tubular, shallowly curved, especially apically, apex narrowly pointed, in ventral view (Fig. 59) elongate, widest sub-basally, thereafter apically tapered to narrowly truncate apex; phallapodeme in lateral view (Fig. 60) irregularly triangular, short, with extended, medial keel short and narrow, not skewed, apex of keel rounded, in ventral view (Fig. 59) a slightly elongate, moderately robust T with keel portion as a symmetrical, lateral bulge; gonite in lateral view (Fig. 60) moderately narrow, moderately elongate, bar-like, nearly straight, end toward aedeagus tapered to point, in ventral view (Fig. 59) small, width only slightly longer than length, tapered medially; hypandrium in lateral view (Fig. 60) like a deep, globularly rounded pocket, wide, tapered to point posteriorly, anterior margin broadly rounded, in ventral view (Fig. 59) very broadly and robustly developed, width only slightly longer than length, posterior margin very shallowly emarginated, anterior margin broadly rounded.
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</paragraph>
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<caption pageId="31" pageNumber="32">
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<paragraph pageId="31" pageNumber="32">
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Figures 57-60.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma williamsae" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="williamsae">Hydrochasma williamsae</taxonomicName>
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sp. n. (USA. New Mexico. Grant: Mimbres River) 57 epandrium and cerci, posterior view 58 same, lateral view 59 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 60 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="type material">
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<paragraph pageId="31" pageNumber="32">Type material.</paragraph>
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<paragraph pageId="31" pageNumber="32">
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The holotype male of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma williamsae" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="williamsae">Hydrochasma williamsae</taxonomicName>
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is labeled "BELIZE. Stann Cr[ee]k. Dist[rict]. MullinsRiver[,] (17km N Dangriga)[,] 29 March 1988, Wayne N. Mathis/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma williamsae" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="williamsae">Hydrochasma williamsae</taxonomicName>
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Mathis & Zatwarnicki, USNM [red]/USNM ENT 00094378 [plastic bar code label]." The holotype is double mounted (minuten in a block of plastic), is in excellent condition, and is deposited in the USNM. Sixteen paratypes (5♂, 11♀; USNM) bear the same label data as the holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="type locality">
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<paragraph pageId="31" pageNumber="32">Type locality.</paragraph>
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<paragraph pageId="31" pageNumber="32">
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Belize. Stann Creek: Mullins River (17 km N Dangriga;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="17.103333">17°06.2'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-88.29667">88°17.8'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="other specimens examined">
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<paragraph pageId="31" pageNumber="32">Other specimens examined.</paragraph>
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<paragraph pageId="31" pageNumber="32">
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Nearctic: UNITED STATES. NEW MEXICO. Grant: Mimbres River (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="32.73">32°43.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-107.86667">107°52'W</geoCoordinate>
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; 1665 m), 13-22 Aug 2007, 2009, D. and W. N. Mathis, T. Zatwarnicki (2♂; USNM).
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</paragraph>
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<paragraph pageId="31" pageNumber="32">
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TEXAS. Llano: Enchanted Rock (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="30.506666">30°30.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-98.81834">98°49.1'W</geoCoordinate>
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), 15 Jun 1953, W. W. Wirth (1♂; USNM).
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</paragraph>
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<paragraph pageId="31" pageNumber="32">
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Neotropical. COSTA RICA. Cartago: La Suiza (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="9.858334">09°51.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.625">83°37.5'W</geoCoordinate>
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), 28 Jun 2001, W. N. Mathis (2♂, 4♀; USNM).
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: Talamanca (
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<normalizedToken originalValue="Estación">Estacion</normalizedToken>
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Gandoca;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="9.623333">9°37.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.695">82°41.7'W</geoCoordinate>
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), 18 May 2004, W. Porras (1♂; INBIO).
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</paragraph>
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<paragraph pageId="31" pageNumber="32">
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HONDURAS.
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<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
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: Omoa (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="16.796667">16°47.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-87.973335">87°58.4'W</geoCoordinate>
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), 26 Sep 1995, D. and W. N. Mathis (1♂, 1♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="32" pageNumber="33" type="distribution">
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<paragraph pageId="32" pageNumber="33">
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<pageBreakToken pageId="32" pageNumber="33" start="start">Distribution</pageBreakToken>
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</paragraph>
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<paragraph pageId="32" pageNumber="33">
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(Fig. 61). Nearctic: United States (New Mexico, Texas). Neotropical: Belize (Stann Creek), Costa Rica (Cartago,
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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), Honduras (
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<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
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).
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</paragraph>
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<caption pageId="32" pageNumber="33">
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<paragraph pageId="32" pageNumber="33">
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Figure 61. Distribution of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma williamsae" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="williamsae">Hydrochasma williamsae</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="32" pageNumber="33" type="etymology">
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<paragraph pageId="32" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="32" pageNumber="33">The specific epithet, williamsae, is a genitive patronym to honor our colleague and cherished friend, the late Ms Hollis Barton Williams, who provided technical support to us for nearly 35 years. Holly passed away August 23, 2009.</paragraph>
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</subSubSection>
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<subSubSection pageId="32" pageNumber="33" type="remarks">
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<paragraph pageId="32" pageNumber="33">Remarks.</paragraph>
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<paragraph pageId="32" pageNumber="33">
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This species, as noted previously in the
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<normalizedToken originalValue="“Remarks”">"Remarks"</normalizedToken>
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section of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma faciale" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="faciale">Hydrochasma faciale</taxonomicName>
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, is distinguished by structures of the male terminalia, especially the generally short and robust epandrium in posterior and lateral views. The more angulate dorsolateral shoulders (best seen in posterior view) and the tapered and pointed ventral epandrial process are unique to this species and distinguish it from congeners.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |