813 lines
98 KiB
XML
813 lines
98 KiB
XML
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008103BFFBEFDF2DAFB9AD56C0A" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus Erichson 1847" docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="131" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>49</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>154</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331143" ID-GBIF-Taxon="167368579" ID-Zenodo-Dep="4331143" LSID="urn:lsid:plazi:treatment:F91EE008103BFFBEFDF2DAFB9AD56C0A" httpUri="http://treatment.plazi.org/id/F91EE008103BFFBEFDF2DAFB9AD56C0A" lastPageId="12" lastPageNumber="141" pageId="2" pageNumber="131">
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<subSubSection box="[542,841,168,192]" pageId="2" pageNumber="131" type="nomenclature">
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<paragraph blockId="2.[542,841,168,192]" box="[542,841,168,192]" pageId="2" pageNumber="131">
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<heading box="[542,841,168,192]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="131" reason="2">
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<taxonomicName ID-CoL="42PQ" ID-ENA="518467" authority="Erichson 1847" authorityName="Erichson" authorityYear="1847" box="[542,841,168,192]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[542,778,168,192]" italics="true" pageId="2" pageNumber="131">
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<emphasis bold="true" box="[542,673,168,192]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
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<bibRefCitation author="Erichson WF" box="[682,778,168,192]" pageId="2" pageNumber="131" refId="ref9841" refString="Erichson WF. 1847. Naturgeschichte des Insecten Deutschland. I. Coleoptera. Berlin: Nicolaischen Buchhandlung." type="book" year="1847">Erichson</bibRefCitation>
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</emphasis>
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1847
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</taxonomicName>
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</heading>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection pageId="2" pageNumber="131" type="type_taxon">
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<paragraph blockId="2.[212,1172,232,607]" box="[212,658,232,257]" pageId="2" pageNumber="131">
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||
<heading box="[212,658,232,257]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
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<typeStatus box="[212,269,233,257]" pageId="2" pageNumber="131">Type</typeStatus>
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species: D.
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<taxonomicName authority="Boheman 1857" authorityName="Boheman" authorityYear="1857" box="[408,651,232,256]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
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<emphasis box="[408,472,232,256]" italics="true" pageId="2" pageNumber="131">pullus</emphasis>
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<bibRefCitation author="Boheman CH" box="[481,651,232,256]" pageId="2" pageNumber="131" refId="ref9783" refString="Boheman CH. 1857. Insecta Caffrariae annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilio sussultus descripsit. Vol. 2. Coleoptera. Holmiae: Ex officina Norstedtiana." type="book" year="1857">Boheman 1857</bibRefCitation>
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</taxonomicName>
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||
.
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</heading>
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||
</paragraph>
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<paragraph blockId="2.[212,1172,232,607]" pageId="2" pageNumber="131">
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,264,288]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[212,343,264,288]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
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</taxonomicName>
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was established by
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<bibRefCitation author="Erichson WF" box="[570,749,264,288]" pageId="2" pageNumber="131" refId="ref9841" refString="Erichson WF. 1847. Naturgeschichte des Insecten Deutschland. I. Coleoptera. Berlin: Nicolaischen Buchhandlung." type="book" year="1847">Erichson (1847)</bibRefCitation>
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for ‘one small south African species, which differs from
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<taxonomicName authorityName="Hope" authorityYear="1833" box="[424,549,296,320]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[424,549,296,320]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
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</taxonomicName>
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in narrower hind legs’. Ten years later
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<bibRefCitation author="Boheman CH" box="[987,1172,296,320]" pageId="2" pageNumber="131" refId="ref9783" refString="Boheman CH. 1857. Insecta Caffrariae annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilio sussultus descripsit. Vol. 2. Coleoptera. Holmiae: Ex officina Norstedtiana." type="book" year="1857">Boheman (1857)</bibRefCitation>
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gave the name to this species,
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<taxonomicName authorityName="Boheman" authorityYear="1857" box="[565,666,328,352]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
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<emphasis box="[565,666,328,352]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
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</taxonomicName>
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, and so fixed the
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<typeStatus box="[879,927,328,352]" pageId="2" pageNumber="131">type</typeStatus>
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species of the genus.
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<bibRefCitation author="Sharp D" box="[212,358,359,383]" pageId="2" pageNumber="131" pagination="33 - 54" refId="ref10290" refString="Sharp D. 1876. Descriptions of some new genera and species of Scarabaeidae from tropical Asia and Malasia. Col Hefte. 13: 33 - 54." type="journal article" year="1876">Sharp (1876)</bibRefCitation>
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described
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<taxonomicName authorityName="Sharp" authorityYear="1876" box="[481,676,360,384]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
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<emphasis box="[481,676,360,384]" italics="true" pageId="2" pageNumber="131">Coptorhina parva</emphasis>
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</taxonomicName>
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from northern
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<collectingCountry box="[857,920,359,383]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
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.
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<bibRefCitation author="Arrow GJ" box="[933,1087,359,383]" pageId="2" pageNumber="131" refId="ref9736" refString="Arrow GJ. 1931. The Fauna of British India III, Coprinae. London: Taylor & Francis." type="book" year="1931">Arrow (1931)</bibRefCitation>
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moved
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<taxonomicName authorityName="Sharp" authorityYear="1876" box="[212,300,392,416]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
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<emphasis box="[212,300,392,416]" italics="true" pageId="2" pageNumber="131">C.parva</emphasis>
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</taxonomicName>
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to
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[352,483,392,416]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[352,483,392,416]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
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</taxonomicName>
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and described the second species from
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<collectingCountry box="[975,1037,391,415]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
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,
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<taxonomicName box="[1056,1164,392,415]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="striatus">
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<emphasis box="[1056,1164,392,415]" italics="true" pageId="2" pageNumber="131">D.striatus</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Paulian R" box="[212,380,423,447]" pageId="2" pageNumber="131" pagination="162 - 164" refId="ref10236" refString="Paulian R. 1934. Quelques Panelini asiatiques nouveaux ou peu connus. Bull Soc Ent Fr. 39: 162 - 164." type="journal article" year="1934">Paulian (1934)</bibRefCitation>
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described the third species from
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<collectingCountry box="[775,837,423,447]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
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,
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<taxonomicName authorityName="Paulian" authorityYear="1934" box="[854,975,424,448]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="cardoni">
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<emphasis box="[854,975,424,448]" italics="true" pageId="2" pageNumber="131">D. cardoni</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Janssens A" box="[994,1172,423,447]" pageId="2" pageNumber="131" pagination="1 - 104" refId="ref10112" refString="Janssens A. 1939. Coprini. Exploration du Parc National Albert, Mission GF de Witte. Institut des parcs nationaux du Congo Belge. 29: 1 - 104." type="journal article" year="1939">Janssens (1939)</bibRefCitation>
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described the second species from Africa,
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<taxonomicName box="[691,790,455,479]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="gilleti">
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<emphasis box="[691,790,455,479]" italics="true" pageId="2" pageNumber="131">D. gilleti</emphasis>
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</taxonomicName>
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, and gave a diagnostic key to all species known by then.
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<bibRefCitation author="Frey G" box="[482,614,487,512]" pageId="2" pageNumber="131" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey (1963)</bibRefCitation>
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described
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<taxonomicName authorityName="Frey" authorityYear="1963" box="[736,868,488,511]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="janssensi">
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<emphasis box="[736,868,488,511]" italics="true" pageId="2" pageNumber="131">D. janssensi</emphasis>
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</taxonomicName>
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from
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<collectingCountry box="[940,1041,487,511]" name="Ethiopia" pageId="2" pageNumber="131">Ethiopia</collectingCountry>
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. Except for this primary taxonomic literature, members of the genus were listed in the catalogues and monographs (
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<bibRefCitation author="Peringuey L" box="[421,597,551,575]" pageId="2" pageNumber="131" pagination="187 - 265" refId="ref10260" refString="Peringuey L. 1901. Tribe: COPRINI (Part 2). Trans South Afr Phil Soc. 12: 187 - 265. doi: 10.1080 / 21560382.1901.9525981" type="journal article" year="1901">Péringuey 1901</bibRefCitation>
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;
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<bibRefCitation author="Gillet JE" box="[615,742,551,575]" pageId="2" pageNumber="131" pagination="1 - 100" refId="ref10076" refString="Gillet JE. 1911. Scarabaeidae: Coprinae, I. In: Junk W, editor. Coleopterorum Catalogus auspicis et auxilio. Berlin: A. Schenkling; 38: 1 - 100." type="journal article" year="1911">Gillet 1911</bibRefCitation>
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;
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<bibRefCitation author="Balthasar V" box="[759,930,551,575]" pageId="2" pageNumber="131" refId="ref9756" refString="Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region; Coleoptera: Lamellicornia. Prague: Tschechoslowakische Akademie der Wissenschaften." type="book" year="1963">Balthasar 1963</bibRefCitation>
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;
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<bibRefCitation author="Ferreira M" box="[947,1102,551,575]" pageId="2" pageNumber="131" pagination="5 - 1088" refId="ref9860" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Ent Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
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;
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<bibRefCitation author="Kral D & Lobl I" pageId="2" pageNumber="131" refId="ref10144" refString="Kral D, Lobl I. 2006. Ateuchini. In: Lobl I, Smetana A, editor, Catalogue of Palaearctic Coleoptera, V. 3. Stensrup: Apollo Books." type="journal volume" year="2006">Král and Löbl 2006</bibRefCitation>
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||
;
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<bibRefCitation author="Davis ALV & Frolov AV & Scholtz CH" box="[394,587,583,607]" pageId="2" pageNumber="131" refId="ref9818" refString="Davis ALV, Frolov AV, Scholtz CH. 2008. The African dung beetle genera. Pretoria: Protea Book House." type="book" year="2008">Davis et al. 2008</bibRefCitation>
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) but few additional data were given.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="131" type="diagnosis">
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<paragraph blockId="2.[212,1171,682,872]" box="[212,320,682,705]" pageId="2" pageNumber="131">
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||
<heading box="[212,320,682,705]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
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||
<emphasis box="[212,320,682,705]" italics="true" pageId="2" pageNumber="131">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph blockId="2.[212,1171,682,872]" pageId="2" pageNumber="131">
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,721,745]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[212,343,721,745]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
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</taxonomicName>
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and closely related
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<taxonomicName authorityName="Hope" authorityYear="1833" box="[580,705,721,745]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis box="[580,705,721,745]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
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</taxonomicName>
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share somewhat rectangular metepisternum, widest in its hind part, very convex epipleural margin (
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<figureCitation box="[917,1035,752,776]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="2" pageNumber="131">Figure 1A</figureCitation>
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), and other taxonomic characters of the
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[540,666,785,809]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[540,666,785,809]" italics="true" pageId="2" pageNumber="131">Sarophorus</emphasis>
|
||
</taxonomicName>
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||
group of genera (
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||
<bibRefCitation author="Frolov AV & Akhmetova LA & Scholtz CH" box="[875,1082,784,808]" pageId="2" pageNumber="131" pagination="1477 - 1508" refId="ref9962" refString="Frolov AV, Akhmetova LA, Scholtz CH. 2008. Revision of the obligate mushroom- feeding African " dung beetle " genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae). J Nat Hist. 42: 1477 - 1508." type="journal article" year="2008">Frolov et al. 2008</bibRefCitation>
|
||
). From
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[212,337,817,841]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[212,337,817,841]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
it can easily be separated by quadridentate clypeus and smaller body size (
|
||
<quantity box="[219,354,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.1" metricValueMin="3.5" pageId="2" pageNumber="131" unit="mm" value="4.8" valueMax="6.1" valueMin="3.5">3.5–6.1 mm</quantity>
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||
in
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[393,525,848,872]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[393,525,848,872]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
and 8.0–20.0 mm in
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[770,895,848,872]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[770,895,848,872]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="135" pageId="2" pageNumber="131" type="description">
|
||
<paragraph blockId="2.[212,1172,947,1137]" box="[212,339,947,970]" pageId="2" pageNumber="131">
|
||
<heading box="[212,339,947,970]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
|
||
<emphasis box="[212,339,947,970]" italics="true" pageId="2" pageNumber="131">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="2.[212,1172,947,1137]" box="[212,1129,986,1010]" pageId="2" pageNumber="131">
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Beetles are small (
|
||
<quantity box="[419,556,986,1010]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.1" metricValueMin="3.5" pageId="2" pageNumber="131" unit="mm" value="4.8" valueMax="6.1" valueMin="3.5">3.5–6.1 mm</quantity>
|
||
), strongly convex, black or dark brown, glabrous.
|
||
</paragraph>
|
||
<paragraph blockId="2.[212,1172,947,1137]" lastBlockId="3.[144,1104,1313,1688]" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131">
|
||
Clypeus distinctly quadridentate in most species. Two medial teeth are almost always acute, lateral ones are acute to right-angled. In some species, lateral clypeal teeth are almost not separated from the lateral margin (
|
||
<figureCitation box="[840,993,1082,1106]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="2" pageNumber="131">Figure 7A, D</figureCitation>
|
||
). Head without carinae on disc, and lacking horns or tubercles. Small carina present near inner margin of eye. Genae right-angled, indistinctly separated from clypeus. Frontoclypeal and genal suturae almost indistinct. Eyes small, their dorsal parts small, slit-shaped in some species, ventral parts larger, sub-rectangular. Distance between eye and gula approximately twice the width of eye in ventral view. Gula with longitudinal groove (
|
||
<figureCitation box="[152,265,1440,1464]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="3" pageNumber="132">Figure 9F</figureCitation>
|
||
). Dorsal surface of clypeus rugose in most species, frons densely punctate to rugose.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002599" ID-Zenodo-Dep="4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="2" pageNumber="131" startId="2.[212,279,1608,1630]" targetBox="[212,1172,1197,1575]" targetPageId="2">
|
||
<paragraph blockId="2.[212,1171,1608,1688]" pageId="2" pageNumber="131">
|
||
Figure 1.
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[316,434,1608,1629]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[316,434,1608,1629]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
spp. (A)
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[532,642,1608,1630]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
|
||
<emphasis box="[532,642,1608,1630]" italics="true" pageId="2" pageNumber="131">D. fossatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[649,732,1608,1629]" pageId="2" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B)
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[599,692,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[599,692,1637,1659]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
|
||
</taxonomicName>
|
||
, male; (C)
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[817,910,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[817,910,1637,1659]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
|
||
</taxonomicName>
|
||
, female; (D)
|
||
<taxonomicName authorityName="Frolov et Cambefort" authorityYear="2014" box="[1056,1166,1637,1658]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="naviauxi">
|
||
<emphasis box="[1056,1166,1637,1658]" italics="true" pageId="2" pageNumber="131">D.naviauxi</emphasis>
|
||
</taxonomicName>
|
||
, female, holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002601" ID-Zenodo-Dep="4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="3" pageNumber="132" startId="3.[144,211,1201,1223]" targetBox="[148,1104,172,1169]" targetPageId="3">
|
||
<paragraph blockId="3.[144,1104,1201,1252]" pageId="3" pageNumber="132">
|
||
Figure 2.
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[249,367,1201,1222]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[249,367,1201,1222]" italics="true" pageId="3" pageNumber="132">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C)
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[571,688,1230,1252]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
|
||
<emphasis box="[571,688,1230,1252]" italics="true" pageId="3" pageNumber="132">D. naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[696,779,1231,1252]" pageId="3" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (D–G)
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[865,995,1230,1252]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
|
||
<emphasis box="[865,995,1230,1252]" italics="true" pageId="3" pageNumber="132">D. pubescens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1003,1084,1231,1252]" pageId="3" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[144,1104,1313,1688]" pageId="3" pageNumber="132">Pronotum more or less trapezoidal, about two times wider than long. Anterior and lateral margins with distinct border, base with or without border. Pronotum is not excavated, without horns or ridges but with fine short longitudinal groove basally along the midline.</paragraph>
|
||
<paragraph blockId="3.[144,1104,1313,1688]" lastBlockId="4.[212,1172,1565,1685]" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132">
|
||
Elytra trapezoidal, as wide as long, shiny to opaque, with deep sinuation on lateral margins receiving convex metepisternum (
|
||
<figureCitation box="[703,823,1664,1688]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="3" pageNumber="132">Figure 1A</figureCitation>
|
||
). Striae fine but distinct, punctate; striae 1–7 reach base of elytron, stria 8 reaches the sinuation but not the base; striae 9–10 are very close to epipleura and mostly inseparable from each other except apically. Elytral intervals flat to convex, with minute punctation. Elytra fused along suture. Scutellum not visible from above.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002603" ID-Zenodo-Dep="4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="4" pageNumber="133" startId="4.[212,279,1464,1486]" targetBox="[212,1172,168,1432]" targetPageId="4">
|
||
<paragraph blockId="4.[212,1171,1464,1515]" pageId="4" pageNumber="133">
|
||
Figure 3.
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[315,497,1464,1485]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[315,497,1464,1485]" italics="true" pageId="4" pageNumber="133">Delopleurus pullus</emphasis>
|
||
</taxonomicName>
|
||
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002605" ID-Zenodo-Dep="4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="5" pageNumber="134" startId="5.[144,211,1307,1329]" targetBox="[165,1079,176,1275]" targetPageId="5">
|
||
<paragraph blockId="5.[144,1104,1307,1359]" pageId="5" pageNumber="134">
|
||
Figure 4.
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[253,509,1308,1329]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
|
||
<emphasis box="[253,509,1308,1329]" italics="true" pageId="5" pageNumber="134">Delopleurus darrenmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[522,608,1308,1329]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[144,1104,1419,1667]" pageId="5" pageNumber="134">
|
||
Wings well developed and feature a number of characters that
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[904,1035,1420,1444]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[904,1035,1420,1444]" italics="true" pageId="5" pageNumber="134">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
share with species of the
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[370,496,1452,1476]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[370,496,1452,1476]" italics="true" pageId="5" pageNumber="134">Sarophorus</emphasis>
|
||
</taxonomicName>
|
||
group of genera (
|
||
<bibRefCitation author="Frolov AV & Akhmetova LA & Scholtz CH" box="[710,922,1451,1475]" pageId="5" pageNumber="134" pagination="1477 - 1508" refId="ref9962" refString="Frolov AV, Akhmetova LA, Scholtz CH. 2008. Revision of the obligate mushroom- feeding African " dung beetle " genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae). J Nat Hist. 42: 1477 - 1508." type="journal article" year="2008">Frolov et al. 2008</bibRefCitation>
|
||
): brown colour except for very basal part, reduced anal area (veins J and AP
|
||
<subScript attach="left" box="[862,895,1494,1510]" fontSize="7" pageId="5" pageNumber="134">3+4</subScript>
|
||
are absent), CuA widened apically along the wing margin, RA
|
||
<subScript attach="left" box="[664,674,1525,1541]" fontSize="7" pageId="5" pageNumber="134">4</subScript>
|
||
does not reach the wing margin and becomes wide and indistinct apically.
|
||
</paragraph>
|
||
<paragraph blockId="5.[144,1104,1419,1667]" lastBlockId="6.[212,1172,168,607]" lastPageId="6" lastPageNumber="135" pageId="5" pageNumber="134">
|
||
Anterior tibiae have three outer teeth. Margin basad of third tooth feebly or not serrate. Spur of anterior tibia more or less bifurcated in males and apically acute to rounded and curved inwards in females. In one species (
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[768,897,1644,1667]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
|
||
<emphasis box="[768,897,1644,1667]" italics="true" pageId="5" pageNumber="134">D. naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[904,994,1643,1667]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), anterior tibia with one small acute tooth between first outer tooth and spur (
|
||
<figureCitation box="[968,1084,168,192]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="6" pageNumber="135">Figure 1D</figureCitation>
|
||
). Outer margins of middle and posterior tibiae without transverse carinae, serrate. Tarsi of all legs well developed, shorter than tibiae. Claws about half the length of fifth tarsal segment.
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,168,607]" pageId="6" pageNumber="135">
|
||
Pygidium about twice as wide as long, its sculpture is species-specific and, in some species, differs strongly from that in the other scarabaeine taxa. In some
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[1040,1171,328,352]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1040,1171,328,352]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
species basal and apical borders are relatively narrow and the disc occupies most of pygidium (
|
||
<figureCitation box="[334,447,391,415]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figure 2E</figureCitation>
|
||
). In others, however, the borders are much wider and the disc is almost slit-shaped (
|
||
<figureCitation box="[444,563,423,447]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figure 2B</figureCitation>
|
||
). Sculpture of pygidium is sex-dependent in some species.
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,168,607]" pageId="6" pageNumber="135">
|
||
Aedeagus of typical scarabaeine shape (
|
||
<figureCitation box="[722,849,487,511]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figures 2F</figureCitation>
|
||
,
|
||
<figureCitation box="[868,901,487,511]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="6" pageNumber="135">3C</figureCitation>
|
||
,
|
||
<figureCitation box="[919,951,487,511]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="6" pageNumber="135">4C</figureCitation>
|
||
,
|
||
<figureCitation box="[970,1002,487,511]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="6" pageNumber="135">5C</figureCitation>
|
||
,
|
||
<figureCitation box="[1021,1054,487,511]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="6" pageNumber="135">6C</figureCitation>
|
||
,
|
||
<figureCitation box="[1072,1105,487,511]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="6" pageNumber="135">7C</figureCitation>
|
||
,
|
||
<figureCitation box="[1123,1156,487,511]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="6" pageNumber="135">8C</figureCitation>
|
||
). Phallobase with two symmetrical tubercles. Parameres symmetrical, without setae apically, slightly distinctive in different species. Internal sac with armature consisting of a few sclerites of complex shape.
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,671,798]" box="[212,425,671,695]" pageId="6" pageNumber="135">
|
||
<heading box="[212,425,671,695]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
|
||
<emphasis box="[212,425,671,695]" italics="true" pageId="6" pageNumber="135">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,671,798]" pageId="6" pageNumber="135">
|
||
Males can be easily separated from females by having more or less bifurcated spurs of anterior tibiae (
|
||
<figureCitation box="[394,548,742,766]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="6" pageNumber="135">Figure 1B, C</figureCitation>
|
||
). Some species differ strongly in the sculpture of the pygidium.
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,862,1054]" box="[212,452,862,886]" pageId="6" pageNumber="135">
|
||
<heading box="[212,452,862,886]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
|
||
<emphasis box="[212,452,862,886]" italics="true" pageId="6" pageNumber="135">Diagnostic characters</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,862,1054]" pageId="6" pageNumber="135">
|
||
Unlike other scarabaeines,
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[519,650,902,926]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[519,650,902,926]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
species can be best separated by the sculpture of the pygidium. Although shape of the parameres and internal sac armature shows some species specific features, they are not considered reliable diagnostic characters. However, availability of the other characters, especially that of the pygidium, makes dissection unnecessary in routine identification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="135" type="distribution">
|
||
<paragraph blockId="6.[212,1172,1117,1532]" box="[212,344,1117,1141]" pageId="6" pageNumber="135">
|
||
<heading box="[212,344,1117,1141]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
|
||
<emphasis box="[212,344,1117,1141]" italics="true" pageId="6" pageNumber="135">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,1117,1532]" pageId="6" pageNumber="135">
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,1157,1181]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[212,343,1157,1181]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
has a disjunctive range covering most of the Afrotropical Region and Indian Peninsula. The ranges of species are rather limited (except for
|
||
<taxonomicName authority="Janssens" authorityName="Janssens" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="gilleti">
|
||
<emphasis box="[1067,1171,1189,1213]" italics="true" pageId="6" pageNumber="135">D. gilleti</emphasis>
|
||
Janssens
|
||
</taxonomicName>
|
||
) although this may be a result of undersampling. Two regional faunas, Afrotropical and Indian, share no common species, which may indicate that they have long developed in isolation. The genus range disjunction can be best explained by the contraction of a previously very large range since the late Neogene. It is possible that in the late Miocene, when huge regions of Africa and Eurasia, including the territories of the present-day Sahara and Mediterranean, were covered by the African-type savannas,
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[497,628,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[497,628,1412,1436]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
was widely distributed in the Eurasia. No palaeontological data exist on the past occurrence of
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[855,986,1444,1468]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[855,986,1444,1468]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
in Europe and extratropical Asia, but considering very low taphonomic potential of these soil dwellers, discovery of any fossil
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[522,653,1508,1532]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[522,653,1508,1532]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
seems highly improbable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="137" pageId="6" pageNumber="135" type="biology_ecology">
|
||
<paragraph blockId="6.[212,1172,1596,1692]" box="[212,297,1596,1620]" pageId="6" pageNumber="135">
|
||
<heading box="[212,297,1596,1620]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
|
||
<emphasis box="[212,297,1596,1620]" italics="true" pageId="6" pageNumber="135">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[212,1172,1596,1692]" lastBlockId="8.[212,1172,850,1384]" lastPageId="8" lastPageNumber="137" pageId="6" pageNumber="135">
|
||
Little information is available about the biology of
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[821,952,1636,1660]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[821,952,1636,1660]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
and the immature stages of the genus are unknown. In
|
||
<collectingCountry box="[635,773,1667,1692]" name="Ivory Coast" pageId="6" pageNumber="135">Ivory Coast</collectingCountry>
|
||
, a few specimens of
|
||
<taxonomicName box="[1011,1110,1667,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="gilleti">
|
||
<emphasis box="[1011,1110,1667,1691]" italics="true" pageId="6" pageNumber="135">D. gilleti</emphasis>
|
||
</taxonomicName>
|
||
were collected on mushrooms of
|
||
<taxonomicName box="[469,625,1535,1558]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="136" phylum="Basidiomycota" rank="genus">
|
||
<emphasis box="[469,625,1535,1558]" italics="true" pageId="7" pageNumber="136">Termitomyces</emphasis>
|
||
</taxonomicName>
|
||
cf.
|
||
<emphasis box="[674,779,1534,1558]" italics="true" pageId="7" pageNumber="136">schimperi</emphasis>
|
||
along with more numerous
|
||
<taxonomicName authority="Boheman" authorityName="Boheman" authorityYear="1857" box="[144,537,1566,1590]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="nitidipennis">
|
||
<emphasis box="[144,412,1566,1590]" italics="true" pageId="7" pageNumber="136">Coptorhina nitidipennis</emphasis>
|
||
Boheman
|
||
</taxonomicName>
|
||
(Yves Cambefort, pers. comm.). In
|
||
<collectingCountry box="[1004,1103,1566,1590]" name="Namibia" pageId="7" pageNumber="136">Namibia</collectingCountry>
|
||
(
|
||
<collectingRegion box="[151,239,1598,1622]" country="Namibia" name="Zambezi" pageId="7" pageNumber="136">Caprivi</collectingRegion>
|
||
, Darren Mann, pers. comm.), beetles of
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[746,927,1598,1622]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
|
||
<emphasis box="[746,927,1598,1622]" italics="true" pageId="7" pageNumber="136">D. darrenmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[942,1037,1599,1622]" pageId="7" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were collected in mushrooms (
|
||
<taxonomicName box="[433,589,1631,1654]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="136" phylum="Basidiomycota" rank="genus">
|
||
<emphasis box="[433,589,1631,1654]" italics="true" pageId="7" pageNumber="136">Termitomyces</emphasis>
|
||
</taxonomicName>
|
||
sp.) on and around termite mounds, also in association with
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[342,467,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[342,467,1662,1686]" italics="true" pageId="7" pageNumber="136">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[488,653,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="nitidipennis">
|
||
<emphasis box="[488,653,1662,1686]" italics="true" pageId="7" pageNumber="136">C. nitidipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Arrow" authorityName="Arrow" authorityYear="1906" box="[720,949,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="angolensis">
|
||
<emphasis box="[720,858,1662,1686]" italics="true" pageId="7" pageNumber="136">C.angolensis</emphasis>
|
||
Arrow
|
||
</taxonomicName>
|
||
). There were apparently
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[340,465,850,874]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[340,465,850,874]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
burrows under the fungal umbrella or a few centimetres away. No smaller burrows of
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[475,606,882,906]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[475,606,882,906]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
were seen however. It is possible that
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[1040,1171,882,906]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1040,1171,882,906]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
can use mushroom pieces buried by
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[628,753,914,938]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[628,753,914,938]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002607" ID-Zenodo-Dep="4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="7" pageNumber="136" startId="7.[144,211,1422,1444]" targetBox="[145,1103,172,1390]" targetPageId="7">
|
||
<paragraph blockId="7.[144,1103,1422,1473]" pageId="7" pageNumber="136">
|
||
Figure 5.
|
||
<taxonomicName box="[247,428,1422,1443]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="gilleti">
|
||
<emphasis box="[247,428,1422,1443]" italics="true" pageId="7" pageNumber="136">Delopleurus gilleti</emphasis>
|
||
</taxonomicName>
|
||
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002609" ID-Zenodo-Dep="4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="8" pageNumber="137" startId="8.[212,279,738,760]" targetBox="[214,1170,168,697]" targetPageId="8">
|
||
<paragraph blockId="8.[212,1171,738,789]" pageId="8" pageNumber="137">
|
||
Figure 6.
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[317,525,738,759]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
|
||
<emphasis box="[317,525,738,759]" italics="true" pageId="8" pageNumber="137">Delopleurus krikkeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[533,617,738,759]" pageId="8" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[254,385,945,969]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[254,385,945,969]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
beetles have never been found in or under dung pads, and they have never been collected by pitfalls baited with dung or carrion [Philippe Walter collected a few specimens in traps baited with fish (Yves Cambefort, pers. comm.) but these were most probably chance captures].
|
||
</paragraph>
|
||
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
|
||
Available data suggest that
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[589,721,1073,1097]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[589,721,1073,1097]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
are specialist feeders of agaric mushrooms (
|
||
<taxonomicName authorityName="Underwood" authorityYear="1899" box="[306,430,1105,1129]" class="Agaricomycetes" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="137" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
|
||
). They seem to be associated with the termitophilous genus
|
||
<taxonomicName box="[212,368,1138,1161]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="137" phylum="Basidiomycota" rank="genus">
|
||
<emphasis box="[212,368,1138,1161]" italics="true" pageId="8" pageNumber="137">Termitomyces</emphasis>
|
||
</taxonomicName>
|
||
, a widespread fungal genus with large fruit bodies. In the case of
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,1169,1193]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[212,343,1169,1193]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Hope" authorityYear="1833" box="[405,530,1169,1193]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[405,530,1169,1193]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
|
||
</taxonomicName>
|
||
, feeding on mushrooms can be a secondary adaptation from the putatively generalist saprophagy of their ancestors. Feeding on mushrooms as an adaptation to arid biotopes in beetles was hypothesized by
|
||
<bibRefCitation author="Striganova BR" box="[957,1161,1232,1256]" pageId="8" pageNumber="137" refId="ref10319" refString="Striganova BR. 1980. Pitanie pochvennykh saprofagov [Feeding of soil saprophages]. Moscow: Nauka." type="book" year="1980">Striganova (1980)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
|
||
All
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[299,430,1264,1288]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[299,430,1264,1288]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
specimens examined have well developed wings and apparently are good fliers. But no specimens have been collected at light, according to the labels and data from collectors. This, along with small eyes (typical for all members of the
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,338,1360,1384]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[212,338,1360,1384]" italics="true" pageId="8" pageNumber="137">Sarophorus</emphasis>
|
||
</taxonomicName>
|
||
group genera), suggests that the beetles have diurnal flight activity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="141" pageId="8" pageNumber="137" type="key">
|
||
<paragraph blockId="8.[212,510,1459,1524]" box="[212,510,1459,1484]" pageId="8" pageNumber="137">
|
||
<heading box="[212,510,1459,1484]" fontSize="10" level="3" pageId="8" pageNumber="137" reason="5">
|
||
<emphasis bold="true" box="[212,510,1459,1484]" pageId="8" pageNumber="137">
|
||
Key to
|
||
<emphasis bold="true" box="[295,510,1459,1483]" italics="true" pageId="8" pageNumber="137">
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[295,426,1459,1483]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
|
||
species
|
||
</emphasis>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[212,510,1459,1524]" box="[212,280,1500,1524]" pageId="8" pageNumber="137">
|
||
<emphasis box="[212,280,1500,1524]" italics="true" pageId="8" pageNumber="137">Males</emphasis>
|
||
</paragraph>
|
||
<key lastPageId="11" lastPageNumber="140" pageId="8" pageNumber="137">
|
||
<paragraph blockId="8.[212,1172,1571,1691]" pageId="8" pageNumber="137">
|
||
<keyLead pageId="8" pageNumber="137">
|
||
<keyStep pageId="8" pageNumber="137">
|
||
1. Disc of pygidium more or less densely punctate (
|
||
<figureCitation box="[801,925,1571,1595]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="8" pageNumber="137">Figures 3B</figureCitation>
|
||
,
|
||
<figureCitation box="[939,970,1571,1595]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="8" pageNumber="137">4B</figureCitation>
|
||
,
|
||
<figureCitation box="[986,1016,1571,1595]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="8" pageNumber="137">5B</figureCitation>
|
||
) or granulate with short brown setae (
|
||
<figureCitation box="[526,641,1603,1627]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="8" pageNumber="137">Figure 2E</figureCitation>
|
||
). Species from Africa .................................. 2 Disc of pygidium almost smooth, without setae (
|
||
<figureCitation box="[801,914,1635,1659]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="8" pageNumber="137">Figure 8B</figureCitation>
|
||
). Species from Asia ... ...............................................................................................................
|
||
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[1064,1171,1668,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis box="[1064,1171,1668,1691]" italics="true" pageId="8" pageNumber="137">D. parvus</emphasis>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002611" ID-Zenodo-Dep="4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="9" pageNumber="138" startId="9.[144,211,1410,1432]" targetBox="[145,1103,169,1378]" targetPageId="9">
|
||
<paragraph blockId="9.[144,1103,1410,1461]" pageId="9" pageNumber="138">
|
||
Figure 7.
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[249,453,1410,1431]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
|
||
<emphasis box="[249,453,1410,1431]" italics="true" pageId="9" pageNumber="138">Delopleurus fossatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[462,544,1411,1432]" pageId="9" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="9.[144,1109,1511,1599]" pageId="9" pageNumber="138">
|
||
<keyLead pageId="9" pageNumber="138">
|
||
<keyStep pageId="9" pageNumber="138">
|
||
2. Disc of pygidium rugose and with sparse brown setae (
|
||
<figureCitation box="[810,924,1511,1535]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="9" pageNumber="138">Figure 2E</figureCitation>
|
||
) ...................... .............................................................................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[861,1005,1543,1567]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
|
||
<emphasis box="[861,1005,1543,1567]" italics="true" pageId="9" pageNumber="138">D. pubescens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1014,1104,1544,1567]" pageId="9" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Disc of pygidium with different sculpture, without setae. ................................. 3
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep lastPageId="10" lastPageNumber="139" pageId="9" pageNumber="138">
|
||
<paragraph blockId="9.[144,1103,1623,1679]" pageId="9" pageNumber="138">
|
||
<keyLead pageId="9" pageNumber="138">
|
||
3. Border of pygidium relatively narrow; punctation of pygidium disc sparser (
|
||
<figureCitation box="[189,303,1655,1679]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="9" pageNumber="138">Figure 3B</figureCitation>
|
||
) .............................................................................................
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[1003,1103,1655,1679]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[1003,1103,1655,1679]" italics="true" pageId="9" pageNumber="138">D. pullus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002613" ID-Zenodo-Dep="4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="10" pageNumber="139" startId="10.[212,279,1391,1413]" targetBox="[212,1171,171,1359]" targetPageId="10">
|
||
<paragraph blockId="10.[212,1172,1391,1442]" pageId="10" pageNumber="139">
|
||
Figure 8.
|
||
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[317,507,1391,1413]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis box="[317,507,1391,1413]" italics="true" pageId="10" pageNumber="139">Delopleurus parvus</emphasis>
|
||
</taxonomicName>
|
||
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="10.[248,1172,1503,1559]" pageId="10" pageNumber="139">
|
||
<keyLead pageId="10" pageNumber="139">
|
||
Border of pygidium thicker (
|
||
<figureCitation box="[593,720,1503,1527]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="10" pageNumber="139">Figures 4B</figureCitation>
|
||
,
|
||
<figureCitation box="[741,772,1503,1527]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="10" pageNumber="139">6B</figureCitation>
|
||
,
|
||
<figureCitation box="[792,823,1504,1527]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="10" pageNumber="139">7B</figureCitation>
|
||
), if similar to
|
||
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[1005,1110,1503,1527]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[1005,1110,1503,1527]" italics="true" pageId="10" pageNumber="139">D. pullus</emphasis>
|
||
</taxonomicName>
|
||
then punctation of pygidium disc coarser (
|
||
<figureCitation box="[679,793,1535,1559]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="10" pageNumber="139">Figure 5B</figureCitation>
|
||
) .............................................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph blockId="10.[212,1178,1583,1671]" pageId="10" pageNumber="139">
|
||
<keyLead pageId="10" pageNumber="139">
|
||
<keyStep pageId="10" pageNumber="139">
|
||
4. Disc of pygidium almost as long as apical border (
|
||
<figureCitation box="[829,943,1583,1607]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="10" pageNumber="139">Figure 4B</figureCitation>
|
||
) ............................. .........................................................................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[897,1073,1615,1639]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
|
||
<emphasis box="[897,1073,1615,1639]" italics="true" pageId="10" pageNumber="139">D. darrenmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1082,1172,1615,1638]" pageId="10" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Disc of pygidium 1.5–2 times longer than apical border ................................... 5
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002615" ID-Zenodo-Dep="4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="11" pageNumber="140" startId="11.[144,211,939,961]" targetBox="[147,1104,168,906]" targetPageId="11">
|
||
<paragraph blockId="11.[144,1104,939,1049]" pageId="11" pageNumber="140">
|
||
Figure 9.
|
||
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[252,370,939,960]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[252,370,939,960]" italics="true" pageId="11" pageNumber="140">Delopleurus</emphasis>
|
||
</taxonomicName>
|
||
spp. and
|
||
<taxonomicName authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[482,732,939,960]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="janssensi">
|
||
<emphasis box="[482,732,939,960]" italics="true" pageId="11" pageNumber="140">Metacatharsius janssensi</emphasis>
|
||
</taxonomicName>
|
||
. (A, E) habitus (A,
|
||
<taxonomicName box="[955,1064,939,961]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis box="[955,1064,939,961]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
|
||
</taxonomicName>
|
||
, E,
|
||
<taxonomicName authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[144,266,969,990]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="janssensi">
|
||
<emphasis box="[144,266,969,990]" italics="true" pageId="11" pageNumber="140">M. janssensi</emphasis>
|
||
</taxonomicName>
|
||
); (B)
|
||
<taxonomicName box="[325,430,969,990]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis box="[325,430,969,990]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
|
||
</taxonomicName>
|
||
, head and pronotum in dorsal view; (C)
|
||
<taxonomicName box="[843,947,969,990]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis box="[843,947,969,990]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
|
||
</taxonomicName>
|
||
, pygidium; (D)
|
||
<taxonomicName box="[144,252,998,1020]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis box="[144,252,998,1020]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
|
||
</taxonomicName>
|
||
, locality map; (F)
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[458,572,998,1020]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
|
||
<emphasis box="[458,572,998,1020]" italics="true" pageId="11" pageNumber="140">D. fossatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[582,669,998,1019]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ventral view of head showing gula with distinct longitudinal groove.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[144,1103,1113,1233]" pageId="11" pageNumber="140">
|
||
<keyLead pageId="11" pageNumber="140">
|
||
<keyStep pageId="11" pageNumber="140">
|
||
5. Disc of pygidium feebly convex, almost in plane with basal border (in lateral view,
|
||
<figureCitation box="[180,294,1145,1169]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="11" pageNumber="140">Figure 5B</figureCitation>
|
||
) ..............................................................................................
|
||
<taxonomicName box="[1005,1103,1145,1169]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="gilleti">
|
||
<emphasis box="[1005,1103,1145,1169]" italics="true" pageId="11" pageNumber="140">D. gilleti</emphasis>
|
||
</taxonomicName>
|
||
Disc of pygidium strongly convex, distinctly separated from basal border by a deep hollow (in lateral view,
|
||
<figureCitation box="[516,631,1209,1233]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6B</figureCitation>
|
||
,
|
||
<figureCitation box="[646,677,1210,1233]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">7B</figureCitation>
|
||
) ..................................................... 6
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="11.[144,1104,1257,1440]" pageId="11" pageNumber="140">
|
||
<keyLead pageId="11" pageNumber="140">
|
||
<keyStep pageId="11" pageNumber="140">
|
||
6. Elytra with well-developed humeral umbones, sinuate behind umbones in dorsal view (
|
||
<figureCitation box="[251,370,1289,1313]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6A</figureCitation>
|
||
). Basal border of pygidium strongly widened medially (
|
||
<figureCitation captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6B</figureCitation>
|
||
) ........................................................................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[878,1005,1321,1345]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
|
||
<emphasis box="[878,1005,1321,1345]" italics="true" pageId="11" pageNumber="140">D. krikkeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1014,1104,1321,1344]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Elytra with feebly developed humeral umbones, feebly or not sinuate behind umbones in dorsal view (
|
||
<figureCitation box="[473,591,1384,1408]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">Figure 7A</figureCitation>
|
||
). Basal border of pygidium slightly widened medially (
|
||
<figureCitation box="[297,411,1416,1440]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">Figure 7B</figureCitation>
|
||
) ..............................................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[881,1005,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
|
||
<emphasis box="[881,1005,1416,1440]" italics="true" pageId="11" pageNumber="140">D. fossatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1014,1104,1417,1440]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
<paragraph blockId="11.[144,234,1500,1524]" box="[144,234,1500,1524]" pageId="11" pageNumber="140">
|
||
<heading box="[144,234,1500,1524]" fontSize="10" level="3" pageId="11" pageNumber="140" reason="5">
|
||
<emphasis box="[144,234,1500,1524]" italics="true" pageId="11" pageNumber="140">Females</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key lastPageId="12" lastPageNumber="141" pageId="11" pageNumber="140">
|
||
<paragraph blockId="11.[144,1103,1571,1691]" pageId="11" pageNumber="140">
|
||
<keyLead pageId="11" pageNumber="140">
|
||
<keyStep pageId="11" pageNumber="140">
|
||
1. Disc of pygidium more or less densely punctate, or slit-shaped, or granulate (with short brown setae). Species from Africa ............................................................ 2 Disc of pygidium almost smooth (
|
||
<figureCitation box="[619,742,1635,1659]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="11" pageNumber="140">Figure 8E</figureCitation>
|
||
) or granulate, without setae (
|
||
<figureCitation box="[192,308,1667,1691]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="11" pageNumber="140">Figure 9C</figureCitation>
|
||
). Species from Asia .......................................................................... 7
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="12.[212,1172,168,288]" pageId="12" pageNumber="141">
|
||
<keyLead pageId="12" pageNumber="141">
|
||
<keyStep pageId="12" pageNumber="141">
|
||
2. Fore tibia with small acute tooth near apical spur (
|
||
<figureCitation box="[819,936,168,192]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1D</figureCitation>
|
||
). Pygidium with slitshaped pubescent disc (
|
||
<figureCitation box="[516,629,200,224]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2B</figureCitation>
|
||
) .......................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[943,1072,201,224]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
|
||
<emphasis box="[943,1072,201,224]" italics="true" pageId="12" pageNumber="141">D. naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1082,1172,201,224]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Fore tibia without tooth near apical spur (
|
||
<figureCitation box="[762,882,232,256]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1C</figureCitation>
|
||
). Sculpture of pygidium different; if disc of pygidium slit-shaped then it lacks setation (
|
||
<figureCitation box="[968,1081,264,288]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="12" pageNumber="141">Figure 4E</figureCitation>
|
||
) .... .. 3
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="12.[212,1172,312,399]" pageId="12" pageNumber="141">
|
||
<keyLead pageId="12" pageNumber="141">
|
||
<keyStep pageId="12" pageNumber="141">
|
||
3. Borders of pygidium relatively narrow; disc feebly convex, granulate and pubescent (
|
||
<figureCitation box="[333,448,344,368]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="12" pageNumber="141">Figure 3E</figureCitation>
|
||
) ...................................................................................
|
||
<taxonomicName authority="Borders" authorityName="Borders" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="pullus">
|
||
<emphasis box="[1071,1171,344,368]" italics="true" pageId="12" pageNumber="141">D. pullus</emphasis>
|
||
Borders
|
||
</taxonomicName>
|
||
of pygidium relatively wide, disc without setation ............................... 4
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="12.[212,1177,423,511]" pageId="12" pageNumber="141">
|
||
<keyLead pageId="12" pageNumber="141">
|
||
<keyStep pageId="12" pageNumber="141">
|
||
4. Disc of pygidium slit-shaped, with transverse carina (
|
||
<figureCitation box="[857,971,423,447]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="12" pageNumber="141">Figure 4E</figureCitation>
|
||
) ......................... .........................................................................................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[897,1073,455,479]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
|
||
<emphasis box="[897,1073,455,479]" italics="true" pageId="12" pageNumber="141">D. darrenmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1082,1172,456,479]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Disc of pygidium 1.5–2 times longer than apical border thickness ................... 5
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="12.[212,1172,535,655]" pageId="12" pageNumber="141">
|
||
<keyLead pageId="12" pageNumber="141">
|
||
<keyStep pageId="12" pageNumber="141">
|
||
5. Disc of pygidium feebly convex, almost in plane with basal border (in lateral view,
|
||
<figureCitation box="[248,362,567,591]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="12" pageNumber="141">Figure 5E</figureCitation>
|
||
) ..............................................................................................
|
||
<taxonomicName box="[1073,1171,567,591]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="gilleti">
|
||
<emphasis box="[1073,1171,567,591]" italics="true" pageId="12" pageNumber="141">D. gilleti</emphasis>
|
||
</taxonomicName>
|
||
Disc of pygidium strongly convex, distinctly separated from basal border by a deep hollow (in lateral view,
|
||
<figureCitation box="[584,698,630,654]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="12" pageNumber="141">Figure 7E</figureCitation>
|
||
) ...............................
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[949,1073,631,655]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
|
||
<emphasis box="[949,1073,631,655]" italics="true" pageId="12" pageNumber="141">D. fossatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1082,1172,631,654]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="12.[212,1172,678,798]" pageId="12" pageNumber="141">
|
||
<keyLead pageId="12" pageNumber="141">
|
||
<keyStep pageId="12" pageNumber="141">
|
||
6. Elytral striae with larger punctures (
|
||
<figureCitation box="[683,805,678,702]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="12" pageNumber="141">Figure 8D</figureCitation>
|
||
). Disc of pygidium smooth to indistinctly rugose (
|
||
<figureCitation box="[472,587,710,734]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="12" pageNumber="141">Figure 8E</figureCitation>
|
||
) ..............................................................
|
||
<taxonomicName authority="Elytral" authorityName="Elytral" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis box="[1063,1171,711,734]" italics="true" pageId="12" pageNumber="141">D. parvus</emphasis>
|
||
Elytral
|
||
</taxonomicName>
|
||
striae with smaller punctures (
|
||
<figureCitation box="[698,819,742,766]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="12" pageNumber="141">Figure 9A</figureCitation>
|
||
). Disc of pygidium granulate (
|
||
<figureCitation box="[260,376,774,798]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="12" pageNumber="141">Figure 9C</figureCitation>
|
||
) ..........................................................................................
|
||
<taxonomicName box="[1054,1171,775,798]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis box="[1054,1171,775,798]" italics="true" pageId="12" pageNumber="141">D. striatus</emphasis>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |