197 lines
18 KiB
XML
197 lines
18 KiB
XML
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE0081035FFBFFE63D9269C486B6B" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus naviauxi Frolov et Cambefort 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="142" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="141" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>49</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>154</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331161" ID-GBIF-Taxon="167368584" ID-Zenodo-Dep="4331161" LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B" httpUri="http://treatment.plazi.org/id/F91EE0081035FFBFFE63D9269C486B6B" lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141">
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<subSubSection box="[399,984,885,909]" pageId="12" pageNumber="141" type="nomenclature">
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<paragraph blockId="12.[399,984,885,941]" box="[399,984,885,909]" pageId="12" pageNumber="141">
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<heading box="[399,984,885,909]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="141" reason="2">
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<taxonomicName ID-CoL="34H5B" authority="Frolov et Cambefort" authorityName="Frolov et Cambefort" authorityYear="2014" box="[399,881,885,909]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
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<emphasis bold="true" box="[399,633,885,909]" italics="true" pageId="12" pageNumber="141">Delopleurus naviauxi</emphasis>
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Frolov et Cambefort
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</taxonomicName>
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,
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<taxonomicNameLabel box="[894,984,886,909]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[578,806,917,941]" pageId="12" pageNumber="141" type="description">
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<paragraph blockId="12.[399,984,885,941]" box="[578,806,917,941]" pageId="12" pageNumber="141">
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(
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<figureCitation box="[586,715,917,941]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figures 1D</figureCitation>
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,
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<figureCitation box="[731,797,917,941]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">2A–C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="141" type="diagnosis">
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<paragraph blockId="12.[212,1172,982,1077]" box="[212,320,982,1005]" pageId="12" pageNumber="141">
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<heading box="[212,320,982,1005]" fontSize="10" level="3" pageId="12" pageNumber="141" reason="5">
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<emphasis box="[212,320,982,1005]" italics="true" pageId="12" pageNumber="141">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="12.[212,1172,982,1077]" pageId="12" pageNumber="141">
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This species can be separated from other
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[665,797,1022,1046]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis box="[665,797,1022,1046]" italics="true" pageId="12" pageNumber="141">Delopleurus</emphasis>
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</taxonomicName>
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species by having deep transverse fossa on the pygidium and by the fore tibia with a small acute tooth near apical spur.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141" type="description">
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<paragraph blockId="12.[212,1172,1150,1691]" box="[212,339,1150,1173]" pageId="12" pageNumber="141">
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<heading box="[212,339,1150,1173]" fontSize="10" level="3" pageId="12" pageNumber="141" reason="5">
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<emphasis box="[212,339,1150,1173]" italics="true" pageId="12" pageNumber="141">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" box="[212,1172,1189,1213]" pageId="12" pageNumber="141">
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<emphasis box="[212,410,1189,1213]" italics="true" pageId="12" pageNumber="141">
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<typeStatus box="[212,317,1189,1213]" pageId="12" pageNumber="141">Holotype</typeStatus>
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, female.
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</emphasis>
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Body strongly convex, black, glabrous, length
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<quantity box="[939,1024,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="141" unit="mm" value="4.9">4.9 mm</quantity>
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(
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<figureCitation box="[1040,1155,1189,1213]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2A</figureCitation>
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).
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</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Clypeus quadridentate. All four clypeal teeth acute, two medial ones relatively slender. Head without carinae on disc, small carinae present near inner margin of eyes. Genae right-angled, indistinctly separated from clypeus. Frontoclypeal and genal suturae indistinct. Eyes small, their dorsal parts slit-shaped, ventral parts subrectangular. Clypeus rugose in anterior part and laterally, frons densely punctate with elongate punctures.</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Pronotum trapezoidal, about two times wider than long. Anterior and lateral margins bordered, base without border. Pronotum relatively densely punctate on disc (punctures separated by one to two puncture diameters), punctation becoming denser laterally.</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Elytra trapezoidal, as wide as long, shiny. Stria distinct, with punctures larger than striae. Elytral intervals slightly convex on disc, with minute punctation.</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">
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Anterior tibiae with three outer teeth and a small acute tooth between first outer tooth and apical spur (
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<figureCitation box="[474,592,1635,1659]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1D</figureCitation>
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).
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</paragraph>
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<paragraph blockId="12.[212,1172,1150,1691]" lastBlockId="13.[144,1104,168,256]" lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141">
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Pygidium with deep transversal slit-shaped fossa with yellowish setae (
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<figureCitation box="[1043,1155,1667,1691]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2B</figureCitation>
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).
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<emphasis box="[144,252,169,192]" italics="true" pageId="13" pageNumber="142">
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<typeStatus box="[144,249,169,192]" pageId="13" pageNumber="142">Paratype</typeStatus>
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.
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</emphasis>
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female, slightly larger than the
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<typeStatus box="[649,747,168,192]" pageId="13" pageNumber="142">holotype</typeStatus>
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(6.0 mm). The specimen has malformed or worn, reduced anterior tibiae without outer teeth and tarsi, and reduced clypeal teeth. It also differs from the
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<typeStatus box="[650,748,232,256]" pageId="13" pageNumber="142">holotype</typeStatus>
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in having less shiny dorsal side.
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</paragraph>
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<paragraph blockId="13.[144,324,303,327]" box="[144,324,303,327]" pageId="13" pageNumber="142">
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<heading box="[144,324,303,327]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
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<emphasis box="[144,202,303,327]" italics="true" pageId="13" pageNumber="142">Male</emphasis>
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unknown.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="142" type="materials_examined">
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<paragraph blockId="13.[144,1104,397,557]" box="[144,300,397,421]" pageId="13" pageNumber="142">
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<heading box="[144,300,397,421]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
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<emphasis box="[144,300,397,421]" italics="true" pageId="13" pageNumber="142">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="13.[144,1104,397,557]" pageId="13" pageNumber="142">
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<materialsCitation ID-GBIF-Occurrence="3757906304" collectingDateMax="1937-05-31" collectingDateMin="1937-03-31" collectionCode="BMNH" collectorName="C. D. Knight'" country="Kenya" location="Colony Makuyu" pageId="13" pageNumber="142" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[144,253,437,461]" pageId="13" pageNumber="142">Holotype</typeStatus>
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,
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<specimenCount box="[268,342,437,461]" pageId="13" pageNumber="142" type="female">female</specimenCount>
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with the labels ‘
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<collectingCountry box="[545,620,438,462]" name="Kenya" pageId="13" pageNumber="142">Kenya</collectingCountry>
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<location LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B:746807C51034FFBFFD9BDBE69D706ED9" box="[631,822,437,461]" country="Kenya" name="Colony Makuyu" pageId="13" pageNumber="142">Colony Makuyu</location>
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iii-v 1937
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<collectorName box="[954,1096,437,461]" pageId="13" pageNumber="142">C.D.Knight’</collectorName>
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, ‘Imp. Inst. Entom.’ (
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<collectionCode box="[384,473,469,493]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="13" pageNumber="142">BMNH</collectionCode>
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) and
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3757906301" collectingDate="1978-04-30" collectionCode="MNHN" collectorName="R. Naviaux'" country="Kenya" location="Maji ya Chumvi" pageId="13" pageNumber="142" specimenCount="1" typeStatus="paratype">
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<typeStatus box="[543,645,470,494]" pageId="13" pageNumber="142">paratype</typeStatus>
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,
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<specimenCount box="[659,733,469,493]" pageId="13" pageNumber="142" type="female">female</specimenCount>
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with the labels ‘
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<collectingCountry box="[928,1002,470,494]" name="Kenya" pageId="13" pageNumber="142">Kenya</collectingCountry>
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<location LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B:746807C51034FFBFFC19DB869EAC6D19" country="Kenya" name="Maji ya Chumvi" pageId="13" pageNumber="142">Maji ya Chumvi</location>
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<date box="[247,328,501,525]" pageId="13" pageNumber="142" value="78-04-30">
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<collectingDate box="[247,328,501,525]" pageId="13" pageNumber="142" value="78-04-30">30.4.78</collectingDate>
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</date>
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R.N.’, ‘coll.
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<collectorName box="[494,638,501,525]" pageId="13" pageNumber="142">R. Naviaux’</collectorName>
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, ‘
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[666,803,501,525]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="152" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
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n. sp? Y.Cambefort det’ (
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<collectionCode box="[152,248,533,557]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="142">MNHN</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="142" type="distribution">
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<paragraph blockId="13.[144,1104,627,723]" box="[144,276,627,651]" pageId="13" pageNumber="142">
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<heading box="[144,276,627,651]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
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<emphasis box="[144,276,627,651]" italics="true" pageId="13" pageNumber="142">Distribution</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="13.[144,1104,627,723]" pageId="13" pageNumber="142">
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The species is known from two localities in Northern Acacia-Commiphora bushlands and thickets ecoregion in East Africa (
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<figureCitation box="[585,700,699,723]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="13" pageNumber="142">Figure 2C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection box="[144,859,794,858]" pageId="13" pageNumber="142" type="etymology">
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<paragraph blockId="13.[144,859,794,858]" box="[144,265,794,818]" pageId="13" pageNumber="142">
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||
<heading box="[144,265,794,818]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
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||
<emphasis box="[144,265,794,818]" italics="true" pageId="13" pageNumber="142">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="13.[144,859,794,858]" box="[144,859,833,858]" pageId="13" pageNumber="142">The species is named after Roger Naviaux, a French collector.</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="142" type="discussion">
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<paragraph blockId="13.[144,1104,928,1151]" box="[144,242,928,952]" pageId="13" pageNumber="142">
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<heading box="[144,242,928,952]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
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<emphasis box="[144,242,928,952]" italics="true" pageId="13" pageNumber="142">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="13.[144,1104,928,1151]" pageId="13" pageNumber="142">
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Yves Cambefort (MNHN) examined the
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[608,739,968,992]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">
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<emphasis box="[608,739,968,992]" italics="true" pageId="13" pageNumber="142">Delopleurus</emphasis>
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</taxonomicName>
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specimen from Maji ya Chumvi in the 1980s and noted that it was distinct from other described species of this genus. However, he did not describe it at that time. In the course of the revision of the genus, I examined the specimen from Makuyu which is housed in the BMNH. After comparison of the two specimens we concluded that they are conspecific and belong to a new species described above.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |