217 lines
24 KiB
XML
217 lines
24 KiB
XML
<document id="054B331CA7EF938388F0DA331B6FB800" ID-DOI="10.5281/zenodo.157661" ID-GBIF-Dataset="65483b24-e197-4508-9c49-50fb2a6ec291" ID-ISSN="11755326" ID-Zenodo-Dep="157661" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461014164758" checkinUser="plazi" docAuthor="Julka, J. M., Blanchart, Eric & Chapuis-Lardy, Lydie" docDate="2004" docId="C067F90D257B9B53FEEC98A4C67BFDAC" docLanguage="en" docName="zt00486.pdf" docOrigin="Zootaxa 486" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Shimodrilus Julka, Blanchart & Chapuis-Lardy, 2004, gen. nov." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="3C5E817525739B59FFE49E33C252FFDA" masterDocTitle="New genera and new species of earthworms (Oligochaeta: Octochaetidae) from Western Ghats, South India" masterLastPageNumber="27" masterPageNumber="1" pageNumber="9" updateTime="1698568502420" updateUser="plazi">
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<mods:titleInfo id="5B6FCFF8A17F9265347A82764CBA68A1">
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<mods:title id="AC53D0D5BEC247456AA2DCFF2DDF8BCA">New genera and new species of earthworms (Oligochaeta: Octochaetidae) from Western Ghats, South India</mods:title>
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<mods:name id="70D30C1B8429460C33EC16DE9F79EF1F" type="personal">
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<mods:roleTerm id="EF82B69C95D4F040F500DDAD146D1179">Author</mods:roleTerm>
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<mods:namePart id="1FDC334414EEA088E0951AA9380E494A">Julka, J. M.</mods:namePart>
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<mods:roleTerm id="831B4C7503567EFF2C8B4D0A4B54330A">Author</mods:roleTerm>
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<mods:namePart id="5F2D05B19DB0F1977D6024D21A058AA3">Blanchart, Eric</mods:namePart>
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<mods:name id="3726E535189DAA277724057179799E43" type="personal">
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<mods:roleTerm id="A8B94E979DCC05CC22A3E4AA215DEE26">Author</mods:roleTerm>
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<mods:namePart id="97A4796A21A8620951D1315439A30070">Chapuis-Lardy, Lydie</mods:namePart>
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<mods:title id="0C309359880D78B29D4F753DA7E179AA">Zootaxa</mods:title>
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<mods:part id="6E1D9B07D7D5910E53C22B6D46162024">
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<mods:date id="1BE2D51D2BAC4B588763EF31AE823CBB">2004</mods:date>
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<mods:number id="0D40F6A5A9B28C181A67B78BE968A93E">486</mods:number>
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<mods:identifier id="A60BEB0DB06C99D7DF2B7CF0B018530C" type="DOI">10.5281/zenodo.157661</mods:identifier>
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<treatment id="C067F90D257B9B53FEEC98A4C67BFDAC" ID-DOI="http://doi.org/10.5281/zenodo.6269560" ID-GBIF-Taxon="119601044" ID-Zenodo-Dep="6269560" LSID="urn:lsid:plazi:treatment:C067F90D257B9B53FEEC98A4C67BFDAC" httpUri="http://treatment.plazi.org/id/C067F90D257B9B53FEEC98A4C67BFDAC" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="00D41B90257B9B51FEEC98A4C03AF96B" box="[264,616,1687,1713]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="4871481B257B9B51FEEC98A4C03AF96B" blockId="8.[264,616,1687,1713]" box="[264,616,1687,1713]" pageId="8" pageNumber="9">
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<heading id="1339FF77257B9B51FEEC98A4C03AF96B" bold="true" box="[264,616,1687,1713]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="7ABA9409257B9B51FEEC98A4C03AF96B" bold="true" box="[264,616,1687,1713]" pageId="8" pageNumber="9">
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Genus
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<taxonomicName id="8FCE3398257B9B51FEBB98A4C3A1F96B" box="[351,499,1687,1713]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="7ABA9409257B9B51FEBB98A4C3A1F96B" bold="true" box="[351,499,1687,1713]" italics="true" pageId="8" pageNumber="9">Shimodrilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="61892972257B9B51FE1F98A4C03AF96B" box="[507,616,1687,1713]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="00D41B90257B9B50FEEC98D2C74FFE0C" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="4871481B257B9B50FEEC98D2C74FFE0C" blockId="8.[264,1324,1761,1868]" lastBlockId="9.[264,1324,284,670]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<emphasis id="7ABA9409257B9B51FEEC98D2C3DAF921" bold="true" box="[264,392,1761,1787]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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Prostomium prolobic to proepilobic. Setae lumbricine. Male pores paired, in seminal grooves, on 18; prostatic pores paired, at ends of seminal grooves, on 17 and 19. Oesophagus with a single gizzard in 5; much enlarged in 16 with calciferous ridges extending to 18; discrete calciferous glands, intestinal caeca and supraintestinal glands absent; typhlosole lamelliform, simple. Penial setae present. Micromeronephridia astomate, exonephric, small paired tufts on body wall in 2–4, on septa in 5–12, in clusters of
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<quantity id="8F36E5FE257A9B50FEEC9FA7C301FE74" box="[264,339,404,430]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="9" pageNumber="10" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
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13 and posteriad segments; paired, stomate, exonephric megameronephridia lateral to micromeronephridia in 17 and posteriad segments, funnels at or lateral to
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<emphasis id="7ABA9409257A9B50FB6C9F8FC6C5FE0F" box="[1160,1175,444,469]" italics="true" pageId="9" pageNumber="10">b</emphasis>
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lines in
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<emphasis id="7ABA9409257A9B50FB1F9F8FC745FE0F" box="[1275,1303,444,469]" italics="true" pageId="9" pageNumber="10">bc</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="00D41B90257A9B50FEDC9FD7C0C8FD44" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="4871481B257A9B50FEDC9FD7C0C8FD44" blockId="9.[264,1324,284,670]" pageId="9" pageNumber="10">
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<taxonomicName id="8FCE3398257A9B50FEDC9FD7C394FE27" box="[312,454,484,509]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="10" phylum="Annelida" rank="genus" status="gen. nov.">
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<emphasis id="7ABA9409257A9B50FEDC9FD7C394FE27" box="[312,454,484,509]" italics="true" pageId="9" pageNumber="10">Shimodrilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="61892972257A9B50FE299FD7C06BFE24" box="[461,569,484,510]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
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belongs to a group of Oriental octochaetid genera with a single oesophageal gizzard and stomate exonephric megameronephridia but without discrete calciferous glands. It can be distinguished form other members of the group,
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<taxonomicName id="8FCE3398257A9B50FB259C07C38FFDAC" authority="Stephenson, 1921" authorityName="Stephenson" authorityYear="1921" class="Clitellata" family="Acanthodrilidae" genus="Ramiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
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<emphasis id="7ABA9409257A9B50FB259C07C779FD97" box="[1217,1323,564,589]" italics="true" pageId="9" pageNumber="10">Ramiella</emphasis>
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<bibRefCitation id="2C5F35EA257A9B50FEEC9C6FC38FFDAC" author="Stephenson" box="[264,477,604,630]" pageId="9" pageNumber="27" refString="Stephenson, J. (1921) Contributions to the morphology, classification, and zoogeography of India Oligochaeta. I. The affinities and systematic position of the genus Eudichogaster Michaelsen, and some related questions. II. On polyphyly in the Oligochaeta. III. Some general considerations in the geographical distribution of Indian Oligochaeta. Proceedings of the Zoological Society of London, 1921, 103 - 142." type="proceedings paper" year="1921">Stephenson, 1921</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="7ABA9409257A9B50FE099C6FC03CFDAF" box="[493,622,604,629]" italics="true" pageId="9" pageNumber="10">Mallehulla</emphasis>
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<bibRefCitation id="2C5F35EA257A9B50FD9C9C6FC123FDAC" author="Julka" box="[632,881,604,630]" pageId="9" pageNumber="26" refString="Julka, J. M. & Rao, B. R. (1982) A new earthworm Mallehulla indica gen. et sp. nov. (Oligochaeta: Octochaetidae) from Karnataka, India. Journal of Soil Biology and Ecology, 2 (2), 67 - 72." type="journal article" year="1982">Julka and Rao, 1982</bibRefCitation>
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and
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<emphasis id="7ABA9409257A9B50FC4B9C6FC612FDAF" box="[943,1088,604,629]" italics="true" pageId="9" pageNumber="10">Konkadrilus</emphasis>
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<bibRefCitation id="2C5F35EA257A9B50FBAF9C6FC687FDAC" author="Julka" box="[1099,1237,604,630]" pageId="9" pageNumber="26" refString="Julka, J. M. (1988) The Fauna of India and the adjacent countries. Megadrile Oligochaeta (Haplotaxida: Lumbricina: Megascolecoidea: Octochaetidae). Zoological Survey of India, Calcutta, xiv + 400 pp." type="book" year="1988">Julka, 1988</bibRefCitation>
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by the characteristics as given in
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<tableCitation id="054C7DA0257A9B50FDD99CB7C0C4FD44" box="[573,662,644,670]" captionStart="TABLE 2" captionStartId="9.[264,352,725,749]" captionTargetBox="[267,1271,788,926]" captionTargetPageId="9" captionText="TABLE 2. Comparison of Shimodrilus with related genera." httpUri="http://table.plazi.org/id/1CB11893257A9B50FEEC9CE6C1D1FD34" pageId="9" pageNumber="10" tableUuid="1CB11893257A9B50FEEC9CE6C1D1FD34">Table 2</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<caption id="1CB11893257A9B50FEEC9CE6C1D1FD34" ID-Table-UUID="1CB11893257A9B50FEEC9CE6C1D1FD34" box="[264,899,724,750]" httpUri="http://table.plazi.org/id/1CB11893257A9B50FEEC9CE6C1D1FD34" pageId="9" pageNumber="10" targetBox="[267,1271,788,926]" targetIsTable="true" targetPageId="9">
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<paragraph id="4871481B257A9B50FEEC9CE6C1D1FD34" blockId="9.[264,899,724,750]" box="[264,899,724,750]" pageId="9" pageNumber="10">
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<emphasis id="7ABA9409257A9B50FEEC9CE6C329FD37" bold="true" box="[264,379,725,749]" pageId="9" pageNumber="10">TABLE 2.</emphasis>
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Comparison of
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<taxonomicName id="8FCE3398257A9B50FDCC9CE5C0FBFD37" box="[552,681,726,749]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
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<emphasis id="7ABA9409257A9B50FDCC9CE5C0FBFD37" box="[552,681,726,749]" italics="true" pageId="9" pageNumber="10">Shimodrilus</emphasis>
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</taxonomicName>
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with related genera.
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</paragraph>
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</caption>
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<paragraph id="4871481B257A9B50FEEF9D27C6A0FCA4" pageId="9" pageNumber="10">
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<table id="3ACEBABB257A64A6FEEF9D27C6A5FC44" box="[267,1271,788,926]" gridcols="4" gridrows="3" pageId="9" pageNumber="10">
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<tr id="F6FE4A59257A64A6FEEF9D27C6A5FCF3" box="[267,1271,788,809]" gridrow="0" pageId="9" pageNumber="10">
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<th id="B52F2325257A64A6FEEF9D27C0DAFCF3" box="[267,648,788,809]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">
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Characteristics
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<taxonomicName id="8FCE3398257A9B50FDF09D27C0DAFCF3" box="[532,648,788,809]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
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<emphasis id="7ABA9409257A9B50FDF09D27C0DAFCF3" box="[532,648,788,809]" italics="true" pageId="9" pageNumber="10">Shimodrilus</emphasis>
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</taxonomicName>
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</th>
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<th id="B52F2325257A64A6FD1F9D27C131FCF3" box="[763,867,788,809]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">
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<taxonomicName id="8FCE3398257A9B50FD1F9D27C100FCF3" box="[763,850,788,809]" class="Clitellata" family="Acanthodrilidae" genus="Ramiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
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<emphasis id="7ABA9409257A9B50FD1F9D27C100FCF3" box="[763,850,788,809]" italics="true" pageId="9" pageNumber="10">Ramiella</emphasis>
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</taxonomicName>
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</th>
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<th id="B52F2325257A64A6FC4B9D27C676FCF3" box="[943,1060,788,809]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">
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<emphasis id="7ABA9409257A9B50FC4B9D27C64AFCF3" box="[943,1048,788,809]" italics="true" pageId="9" pageNumber="10">Mallehulla</emphasis>
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</th>
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<th id="B52F2325257A64A6FB659D27C6A5FCF3" box="[1153,1271,788,809]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">
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<emphasis id="7ABA9409257A9B50FB659D27C6A5FCF3" box="[1153,1271,788,809]" italics="true" pageId="9" pageNumber="10">Konkadrilus</emphasis>
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</th>
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</tr>
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<tr id="F6FE4A59257A64A6FEEF9D0DC6A5FC89" box="[267,1271,830,851]" gridrow="1" pageId="9" pageNumber="10">
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<th id="B52F2325257A64A6FEEF9D0DC0DAFC89" box="[267,648,830,851]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Setae lumbricine</th>
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<td id="B52F2325257A64A6FD1F9D0DC131FC89" box="[763,867,830,851]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">lumbricine</td>
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<td id="B52F2325257A64A6FC4B9D0DC676FC89" box="[943,1060,830,851]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">perichaetine</td>
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<td id="B52F2325257A64A6FB659D0DC6A5FC89" box="[1153,1271,830,851]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">lumbricine</td>
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</tr>
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<tr id="F6FE4A59257A64A6FEEF9D5AC6A5FC44" box="[267,1271,873,926]" gridrow="2" pageId="9" pageNumber="10">
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<th id="B52F2325257A64A6FEEF9D5AC0DAFC44" box="[267,648,873,926]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">Oesophageal calciferous 16–18 lamellae</th>
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<td id="B52F2325257A64A6FD1F9D5AC131FC44" box="[763,867,873,926]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">8–11</td>
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<td id="B52F2325257A64A6FC4B9D5AC676FC44" box="[943,1060,873,926]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">14–15</td>
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<td id="B52F2325257A64A6FB659D5AC6A5FC44" box="[1153,1271,873,926]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">16, 16–½17</td>
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</tr>
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</table>
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</paragraph>
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<paragraph id="4871481B257A9B50FEEF9D80C6ECFB67" pageId="9" pageNumber="10">
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<table id="3ACEBABB257A64A6FEEF9D80C74AFB06" box="[267,1304,947,1244]" gridcols="5" gridrows="3" pageId="9" pageNumber="10">
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<tr id="F6FE4A59257A64A6FEEF9D80C74AFC32" box="[267,1304,947,1000]" gridrow="0" pageId="9" pageNumber="10">
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<th id="B52F2325257A64A6FEEF9D80C399FC32" box="[267,459,947,1000]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">Intestine begins</th>
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<th id="B52F2325257A64A6FDF79D80C0B7FC32" box="[531,741,947,1000]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">in 19</th>
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<th id="B52F2325257A64A6FD1F9D80C1CEFC32" box="[763,924,947,1000]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">in or anterior to 16</th>
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<th id="B52F2325257A64A6FC4A9D80C63FFC32" box="[942,1133,947,1000]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">in 17</th>
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<th id="B52F2325257A64A6FB659D80C74AFC32" box="[1153,1304,947,1000]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">in 19</th>
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</tr>
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<tr id="F6FE4A59257A64A6FEEF9DCDC74AFB48" box="[267,1304,1022,1170]" gridrow="1" pageId="9" pageNumber="10">
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<th id="B52F2325257A64A6FEEF9DCDC399FB48" box="[267,459,1022,1170]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Exonephric stomate megameronephridia</th>
|
||
<td id="B52F2325257A64A6FDF79DCDC0B7FB48" box="[531,741,1022,1170]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">
|
||
lateral to micromero nephridia in 17 and posteriad segments, funnels at or lateral to
|
||
<emphasis id="7ABA9409257A9B50FDF09A4EC072FB48" box="[532,544,1149,1170]" italics="true" pageId="9" pageNumber="10">b</emphasis>
|
||
lines in
|
||
<emphasis id="7ABA9409257A9B50FD959A4EC0DAFB48" box="[625,648,1149,1170]" italics="true" pageId="9" pageNumber="10">bc</emphasis>
|
||
</td>
|
||
<td id="B52F2325257A64A6FD1F9DCDC1CEFB48" box="[763,924,1022,1170]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">median most in caudalsegments, funnels close to nerve cord</td>
|
||
<td id="B52F2325257A64A6FC4A9DCDC63FFB48" box="[942,1133,1022,1170]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">median most in 6 and posteriad seg ments, funnels close to nerve cord</td>
|
||
<td id="B52F2325257A64A6FB659DCDC74AFB48" box="[1153,1304,1022,1170]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">median most in caudal segments, funnels close to nerve cord</td>
|
||
</tr>
|
||
<tr id="F6FE4A59257A64A6FEEF9A9BC74AFB06" box="[267,1304,1192,1244]" gridrow="2" pageId="9" pageNumber="10">
|
||
<th id="B52F2325257A64A6FEEF9A9BC399FB06" box="[267,459,1192,1244]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">Tufted micromero nephridia</th>
|
||
<td id="B52F2325257A64A6FDF79A9BC0B7FB06" box="[531,741,1192,1244]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">exonephric in 2–12</td>
|
||
<td id="B52F2325257A64A6FD1F9A9BC1CEFB06" box="[763,924,1192,1244]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">absent</td>
|
||
<td id="B52F2325257A64A6FC4A9A9BC63FFB06" box="[942,1133,1192,1244]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">exonephric in 3, enteronephric in 4</td>
|
||
<td id="B52F2325257A64A6FB659A9BC74AFB06" box="[1153,1304,1192,1244]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">absent</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="00D41B90257A9B50FEDC9B0CC138FA80" box="[312,874,1343,1370]" pageId="9" pageNumber="10" type="materials_examined">
|
||
<paragraph id="4871481B257A9B50FEDC9B0CC138FA80" blockId="9.[264,1324,1343,1890]" box="[312,874,1343,1370]" pageId="9" pageNumber="10">
|
||
<emphasis id="7ABA9409257A9B50FEDC9B0CC388FA83" bold="true" box="[312,474,1343,1369]" pageId="9" pageNumber="10">
|
||
<typeStatus id="9775F6B9257A9B50FEDC9B0CC327FA83" box="[312,373,1343,1369]" pageId="9" pageNumber="10">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="8FCE3398257A9B50FE059B73C155FA83" box="[481,775,1344,1369]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="bhatkalensis" status="sp. nov.">
|
||
<emphasis id="7ABA9409257A9B50FE059B73C155FA83" box="[481,775,1344,1369]" italics="true" pageId="9" pageNumber="10">Shimodrilus bhatkalensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="61892972257A9B50FCF49B73C138FA80" box="[784,874,1344,1370]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00D41B90257A9B53FEDC9B54C01AFE0C" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="4871481B257A9B50FEDC9B54C11AF9F8" blockId="9.[264,1324,1343,1890]" pageId="9" pageNumber="10">
|
||
<emphasis id="7ABA9409257A9B50FEDC9B54C382FA5B" bold="true" box="[312,464,1383,1409]" pageId="9" pageNumber="10">Description.</emphasis>
|
||
Prostomium prolobic to proepilobic. Setae lumbricine through out body. Clitellum annular. Male pores paired, minute, in seminal grooves on 18; prostatic pores paired, minute, at ends of seminal grooves on 17 and 19. Female pores paired, minute, presetal, within
|
||
<emphasis id="7ABA9409257A9B50FE4C9BD3C3E5FA23" box="[424,439,1504,1529]" italics="true" pageId="9" pageNumber="10">a</emphasis>
|
||
lines on 14. Spermathecal pores paired, tiny or transverse slits. Genital markings absent. Nephridiopores not recognised.
|
||
</paragraph>
|
||
<paragraph id="4871481B257A9B53FEDC9803C01AFE0C" blockId="9.[264,1324,1343,1890]" lastBlockId="10.[264,1323,284,630]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Unpigmented. Gizzard single, in 5. Discrete calciferous glands absent; oesophagus enlarged in 16 and with internal longitudinal calciferous ridges extending to 18. Supraintestinal glands and intestinal caeca absent; typhlosole simple, lamelliform. Dorsal blood vessel single, complete; supraoesophageal vessel single, 7–13; subneural vessel absent; extraoesophageal vessels paired, recognised from 5, joining supraoesophageal vessel in 8; lateroparietal vessels paired, passing to supraoesophageal vessel in 13; lateral hearts originating from supraoesophageal vessel with delicate connectives to dorsal vessel in 10–13. Holandric. Prostates tubular, paired, in 17 and 19. Penial setae present. Spermathecae paired, diverticulate. Ovaries paired, fanshaped, in 13. Micromeronephridia astomate; exonephric, small paired tufts on body wall in 2–4 and anterior faces of septa
|
||
<date id="3C706EDB25799B53FB6B9F77C75BFE84" box="[1167,1289,324,350]" pageId="10" pageNumber="11" value="1913-06-05" valueMax="1913-12-05" valueMin="1913-06-05">5/6–12/13</date>
|
||
, in clusters of 4–5 on body wall in 13 and posteriad segments; paired stomate exonephric megameronephridia lateral to micromeronephridia in 17 and posteriad segments, funnels at or lateral to
|
||
<emphasis id="7ABA940925799B53FE579F8FC390FE0F" box="[435,450,444,469]" italics="true" pageId="10" pageNumber="11">b</emphasis>
|
||
lines in
|
||
<emphasis id="7ABA940925799B53FDC29F8FC010FE0F" box="[550,578,444,469]" italics="true" pageId="10" pageNumber="11">bc</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00D41B9025799B53FEDC9FD0C306FD94" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="4871481B25799B53FEDC9FD0C306FD94" blockId="10.[264,1323,284,630]" pageId="10" pageNumber="11">
|
||
<emphasis id="7ABA940925799B53FEDC9FD0C396FE27" bold="true" box="[312,452,483,509]" pageId="10" pageNumber="11">Etymology.</emphasis>
|
||
<taxonomicName id="8FCE339825799B53FE359FD7C00DFE27" box="[465,607,484,509]" class="Clitellata" family="Octochaetidae" genus="Shimodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="11" phylum="Annelida" rank="genus">
|
||
<emphasis id="7ABA940925799B53FE359FD7C00DFE27" box="[465,607,484,509]" italics="true" pageId="10" pageNumber="11">Shimodrilus</emphasis>
|
||
</taxonomicName>
|
||
, gender masculine, derived from the name of the district ‘Shimoga’ in which it is commonly found, and the Greek word ‘drilos’, meaning the worm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="00D41B9025799B53FEDC9C68C67BFDAC" box="[312,1065,603,630]" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="4871481B25799B53FEDC9C68C67BFDAC" blockId="10.[264,1323,284,630]" box="[312,1065,603,630]" pageId="10" pageNumber="11">
|
||
<emphasis id="7ABA940925799B53FEDC9C68C3E3FDAF" bold="true" box="[312,433,603,629]" pageId="10" pageNumber="11">Remarks.</emphasis>
|
||
Referred as ‘Genus B’ in
|
||
<bibRefCitation id="2C5F35EA25799B53FD009C6FC677FDAC" author="Blanchart" box="[740,1061,604,630]" pageId="10" pageNumber="26" refString="Blanchart, E. and Julka, J. M. (1997) Influence of forest disturbance on earthworm communities in the Western Ghats, South India. Soil Biology and Biochemistry, 29, 303 - 306." type="journal article" year="1997">Blanchart and Julka (1997)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |