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<document id="066F5EE8B42B3D9BC23656F15AD8245B" ID-DOI="10.5281/zenodo.190886" ID-GBIF-Dataset="d5d829fb-863d-4ae1-97b4-b03b5649ad1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="190886" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461121335042" checkinUser="plazi" docAuthor="Jacobus, Luke M. &amp; Zhou, - F." docDate="2009" docId="C05E1C6BFF9EFFEE3CD9FAF4FD8B060D" docLanguage="en" docName="zt02268p058.pdf" docOrigin="Zootaxa 2268" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Serratella elissa Jacobus &amp; Zhou, 2009, new species" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="3C676413FF9DFFE83C4EFFF4FFE8077E" masterDocTitle="Two new species of Asian Serratella Edmunds (Ephemeroptera: Ephemerellidae)" masterLastPageNumber="58" masterPageNumber="52" pageNumber="55" updateTime="1698596008563" updateUser="plazi">
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<mods:title id="B9A649F88AC10D599F29108096B76AF6">Two new species of Asian Serratella Edmunds (Ephemeroptera: Ephemerellidae)</mods:title>
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<mods:namePart id="8525FA3AA703B2A4FD76BD26FE32D1B3">Jacobus, Luke M.</mods:namePart>
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<mods:namePart id="9EB5BA0860DC7AD62025CD2240996432">Zhou, - F.</mods:namePart>
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<mods:date id="BFCE4C5E07AA00F22FADF738ED5BD6C4">2009</mods:date>
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<treatment id="C05E1C6BFF9EFFEE3CD9FAF4FD8B060D" ID-DOI="http://doi.org/10.5281/zenodo.6213162" ID-GBIF-Taxon="119625630" ID-Zenodo-Dep="6213162" LSID="urn:lsid:plazi:treatment:C05E1C6BFF9EFFEE3CD9FAF4FD8B060D" httpUri="http://treatment.plazi.org/id/C05E1C6BFF9EFFEE3CD9FAF4FD8B060D" lastPageId="6" lastPageNumber="58" pageId="3" pageNumber="55">
<subSubSection id="00EDFEF6FF9EFFEB3CD9FAF4FEFF0243" pageId="3" pageNumber="55" type="nomenclature">
<paragraph id="4848AD7DFF9EFFEB3CD9FAF4FE1E0264" blockId="3.[151,502,1280,1341]" box="[151,502,1280,1306]" pageId="3" pageNumber="55">
<heading id="13001A11FF9EFFEB3CD9FAF4FE1E0264" bold="true" box="[151,502,1280,1306]" fontSize="11" level="1" pageId="3" pageNumber="55" reason="1">
<emphasis id="7A83716FFF9EFFEB3CD9FAF4FE1E0264" bold="true" box="[151,502,1280,1306]" pageId="3" pageNumber="55">
<taxonomicName id="8FF7D6FEFF9EFFEB3CD9FAF4FEB00264" ID-CoL="4WXFF" box="[151,344,1280,1306]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="elissa" status="sp. nov.">
<emphasis id="7A83716FFF9EFFEB3CD9FAF4FEB00264" bold="true" box="[151,344,1280,1306]" italics="true" pageId="3" pageNumber="55">Serratella elissa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="61B0CC14FF9EFFEB3D28FAF4FE1E0264" box="[358,502,1280,1306]" pageId="3" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4848AD7DFF9EFFEB3CD9FAD7FEFF0243" blockId="3.[151,502,1280,1341]" box="[151,279,1315,1341]" pageId="3" pageNumber="55">
(
<figureCitation id="D0CCB1F8FF9EFFEB3CD1FAD7FEE70243" box="[159,271,1315,1341]" captionStart-0="FIGURES 4, 5" captionStart-1="FIGURES 6" captionStartId-0="4.[151,269,1931,1955]" captionStartId-1="5.[151,269,1965,1989]" captionText-0="FIGURES 4, 5. Serratella elissa, new species. Fig. 4. Maxilla. Fig. 5. Claw." captionText-1="FIGURES 6. Serratella elissa, new species. Dorsal habitus." pageId="3" pageNumber="55">Figs. 46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="00EDFEF6FF9EFFEC3CD9FA84FAE905B1" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55" type="materials_examined">
<paragraph id="4848AD7DFF9EFFEB3CD9FA84FD0E02A6" blockId="3.[151,1437,1392,2037]" pageId="3" pageNumber="55">
<emphasis id="7A83716FFF9EFFEB3CD9FA84FEA602F4" bold="true" box="[151,334,1392,1418]" pageId="3" pageNumber="55">
<typeStatus id="974C13DFFF9EFFEB3CD9FA84FF3B02F4" box="[151,211,1392,1418]" pageId="3" pageNumber="55">Type</typeStatus>
material.
</emphasis>
<typeStatus id="974C13DFFF9EFFEB3D17FA84FE1302F4" box="[345,507,1392,1418]" pageId="3" pageNumber="55" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="30E0EDEDFF9EFFEB3E48FA84FDD102F4" box="[518,569,1392,1418]" name="Iran" pageId="3" pageNumber="55">Iran</collectingCountry>
, Guilan River at Lanak Waterfall,
<geoCoordinate id="2DC3CBBAFF9EFFEB3F97FA84FBA902F4" box="[985,1089,1392,1418]" direction="north" orientation="latitude" pageId="3" pageNumber="55" precision="925" value="37.0">37º00N</geoCoordinate>
,
<geoCoordinate id="2DC3CBBAFF9EFFEB3800FA84FB5902F4" box="[1102,1201,1392,1418]" direction="east" orientation="longitude" pageId="3" pageNumber="55" precision="925" value="49.866665">49º52E</geoCoordinate>
, no collection date, larva [
<collectionCode id="2EE635B8FF9EFFEB3CAEFA63FEDB02CF" box="[224,307,1431,1457]" pageId="3" pageNumber="55">PERC</collectionCode>
].
<typeStatus id="974C13DFFF9EFFEB3D08FA63FE0502CF" box="[326,493,1431,1457]" pageId="3" pageNumber="55" type="paratype">PARATYPES</typeStatus>
: same data, four larvae [
<collectionCode id="2EE635B8FF9EFFEB3F42FA63FCB702CF" box="[780,863,1431,1457]" pageId="3" pageNumber="55">PERC</collectionCode>
].
<collectingCountry id="30E0EDEDFF9EFFEB3F3FFA63FC4B02CF" box="[881,931,1431,1457]" name="Iran" pageId="3" pageNumber="55">Iran</collectingCountry>
, Havigh River,
<quantity id="8F0F0098FF9EFFEB3810FA63FB4102CF" box="[1118,1193,1431,1457]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="55" unit="km" value="20.0">20 km</quantity>
south of Astara,
<date id="3C498BBDFF9EFFEB393BFA63FF1202A6" pageId="3" pageNumber="55" value="1973-04-30">30- IV-1973</date>
, SF&amp;KD Kimball, seven larvae [
<collectionCode id="2EE635B8FF9EFFEB3E30FA4AFD3402A6" LSID="urn:lsid:biocol.org:col:34871" box="[638,732,1470,1496]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="55">USNM</collectionCode>
].
</paragraph>
<paragraph id="4848AD7DFF9EFFEC3C88FA10FC6D06E4" blockId="3.[151,1437,1392,2037]" lastBlockId="4.[151,1438,152,1183]" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
Larva (preserved in ethanol). Length: body
<quantity id="8F0F0098FF9EFFEB3E82FA10FCB50280" box="[716,861,1508,1534]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.3" metricValueMin="6.5" pageId="3" pageNumber="55" unit="mm" value="6.9" valueMax="7.3" valueMin="6.5">6.57.3 mm</quantity>
; antennae
<quantity id="8F0F0098FF9EFFEB3F95FA10FB840280" box="[987,1132,1508,1534]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="3" pageNumber="55" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.41.7 mm</quantity>
; caudal filaments 3.74.0 mm. Body color yellow to light brown, sometimes with dark brown shading. Head: Color light brown with variable, dark brown maculae. Antennal scape and pedicel brown; flagellar segments light brown, darker distally, with whorl of fine, hairlike setae distally on each segment; hairlike setae one-half length of respective segment. Vertex rough. One to two pairs of spiculate, suboccipital bumps present. Clypeus with short setae. Labrum with irregular, transverse row of long, fine setae. Mandible not extending beyong margin of head capsule, with few fine, sublateral setae. Maxilla (
<figureCitation id="D0CCB1F8FF9EFFEB3E9AF938FCF20198" box="[724,794,1740,1766]" captionStart="FIGURES 4, 5" captionStartId="4.[151,269,1931,1955]" captionText="FIGURES 4, 5. Serratella elissa, new species. Fig. 4. Maxilla. Fig. 5. Claw." pageId="3" pageNumber="55">Fig. 4</figureCitation>
) with few subdistal setae, three fimbriate medial setae, and palp with three segments. Labium with palp segment 3 length about one-half that of segment 2. Thorax: Nota without prominent protuberances and with few fine setae, sometimes with brown maculation; pronotum with no anterolateral projections. Coxae and trochanters dark brown. Femora with subproximal and subdistal dark brown bands; forefemur with broad subdistal band of few long, sharp setae; mid- and hindfemora each with scattered hairlike setae and with sparse row of few, spatulate setae recumbent on posterior margin. Tibiae dark brown basally, grading to pale distally and with scattered hairlike setae. Tarsi brown with dark brown basal band and with scattered hairlike setae. Claws (
<figureCitation id="D0CCB1F8FF9EFFEB3F41F82FFCBE008B" box="[783,854,2011,2037]" captionStart="FIGURES 4, 5" captionStartId="4.[151,269,1931,1955]" captionText="FIGURES 4, 5. Serratella elissa, new species. Fig. 4. Maxilla. Fig. 5. Claw." pageId="3" pageNumber="55">Fig. 5</figureCitation>
) with seven to nine denticles and with subdistal setae; denticles nearly subequal in size; subdistal setae sometimes difficult to discern. Abdomen: Terga without paired, submedian spines; paired tufts of fewer than four spatulate setae present on at least some posterior margins. Terga 6, 7, 9, and 10 usually dark brown to black. Dorsal lamellae of gills falcate, with brown trilobed pattern; ventral lamella of gill 6 deeply cleft; gill 7 inserted near posterolateral corner. Sterna with dark brown to black sublateral maculation; sterna 9 and 10 dark brown to black. Caudal filaments uniformly yellow to light brown, few basal segments dark brown; segments with distal whorls of stout setae and fine setae, sometimes with few long, hairlike setae laterally.
</paragraph>
<paragraph id="4848AD7DFF99FFEC3C88FE52FD8F06BE" blockId="4.[151,1438,152,1183]" box="[198,615,422,448]" pageId="4" pageNumber="56">Subimagoes and Adults. Unknown.</paragraph>
<paragraph id="4848AD7DFF99FFEC3C88FE39FE680699" blockId="4.[151,1438,152,1183]" box="[198,384,461,487]" pageId="4" pageNumber="56">Egg. Unknown.</paragraph>
<paragraph id="4848AD7DFF99FFEC3C88FE00FEA2054A" blockId="4.[151,1438,152,1183]" pageId="4" pageNumber="56">
Addition to data matrix.
<taxonomicName id="8FF7D6FEFF99FFEC3DA8FE00FDCF0570" box="[486,551,500,526]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">elissa</taxonomicName>
?? 1 0 0 0 0 0 0 0 0 0 0 - 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0 0 0 0 0 1 0???????
</paragraph>
<paragraph id="4848AD7DFF99FFEC3C88FDB5FAE905B1" blockId="4.[151,1438,152,1183]" pageId="4" pageNumber="56">Larval Variability. Body color of the new species varies from nearly uniform yellow with few or no markings to brown with distinct dark and pale markings. The caudal filaments are variable with respect to the presence or absence of lateral hairlike setae. Larvae tend to differ in body shape, depending on the instar examined; earlier instars tend to be more compact and dorsoventrally flattened than later instars.</paragraph>
</subSubSection>
<subSubSection id="00EDFEF6FF99FFEE3C88FD2FFE6E0658" lastPageId="6" lastPageNumber="58" pageId="4" pageNumber="56" type="diagnosis">
<paragraph id="4848AD7DFF99FFEC3C88FD2FFD7103E1" blockId="4.[151,1438,152,1183]" pageId="4" pageNumber="56">
<emphasis id="7A83716FFF99FFEC3C88FD2FFEAE058B" bold="true" box="[198,326,731,757]" pageId="4" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="8FF7D6FEFF99FFEC3D03FD28FDE5058B" box="[333,525,732,757]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC3D03FD28FDE5058B" box="[333,525,732,757]" italics="true" pageId="4" pageNumber="56">Serratella elissa</emphasis>
</taxonomicName>
is distinguished from other Hyrtanellini by the combination of having at least some abdominal terga with distinctive paired tufts of spatulate setae on the posterior margins (
<figureCitation id="D0CCB1F8FF99FFEC38BDFCF6FAD40462" box="[1267,1340,770,796]" captionStart="FIGURES 6" captionStartId="5.[151,269,1965,1989]" captionText="FIGURES 6. Serratella elissa, new species. Dorsal habitus." pageId="4" pageNumber="56">Fig. 6</figureCitation>
), gills 3 that are not operculate and claws with denticles nearly subequal in size (
<figureCitation id="D0CCB1F8FF99FFEC3FAEFCDDFBCF043D" box="[992,1063,809,835]" captionStart="FIGURES 4, 5" captionStartId="4.[151,269,1931,1955]" captionText="FIGURES 4, 5. Serratella elissa, new species. Fig. 4. Maxilla. Fig. 5. Claw." pageId="4" pageNumber="56">Fig. 5</figureCitation>
). This new species is somewhat similar in general appearance to
<taxonomicName id="8FF7D6FEFF99FFEC3E55FCA4FC960414" authority="Kazanci" authorityName="Kazanci" box="[539,894,848,874]" class="Insecta" family="Ephemerellidae" genus="Quatica" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="euphratica">
<emphasis id="7A83716FFF99FFEC3E55FCA4FCEA0417" box="[539,770,848,873]" italics="true" pageId="4" pageNumber="56">Quatica euphratica</emphasis>
(Kazanci)
</taxonomicName>
and
<taxonomicName id="8FF7D6FEFF99FFEC3FF7FCA4FB1C0414" authority="Kazanci" authorityName="Kazanci" box="[953,1268,848,874]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="karia">
<emphasis id="7A83716FFF99FFEC3FF7FCA4FB9D0417" box="[953,1141,848,873]" italics="true" pageId="4" pageNumber="56">Serratella karia</emphasis>
(Kazanci)
</taxonomicName>
, from
<collectingCountry id="30E0EDEDFF99FFEC390CFCA4FA700414" box="[1346,1432,848,874]" name="Turkey" pageId="4" pageNumber="56">Turkey</collectingCountry>
. These species are distinguished easily from
<taxonomicName id="8FF7D6FEFF99FFEC3ED2FC83FD1504EE" box="[668,765,887,912]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC3ED2FC83FD1504EE" box="[668,765,887,912]" italics="true" pageId="4" pageNumber="56">S. elissa</emphasis>
</taxonomicName>
by the dorsal armature of thorax and abdomen (
<bibRefCitation id="2C66D08CFF99FFEC397DFC82FF3F04C9" author="Kazanci" pageId="4" pageNumber="58" refString="Kazanci, N. (1990) Drunella karia n. sp. A second species of the genus Drunella (Ephemeroptera: Ephemerellidae) from Turkey. Hydrobiologia, 199, 35 - 42." type="journal article" year="1990">Kazanci, 1990</bibRefCitation>
).
<taxonomicName id="8FF7D6FEFF99FFEC3CA2FC69FE4704C8" box="[236,431,925,950]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC3CA2FC69FE4704C8" box="[236,431,925,950]" italics="true" pageId="4" pageNumber="56">Serratella elissa</emphasis>
</taxonomicName>
also resembles species of the
<emphasis id="7A83716FFF99FFEC3F57FC69FBC304C8" box="[793,1067,925,950]" italics="true" pageId="4" pageNumber="56">Teloganpsis mesoleuca</emphasis>
(Brauer) species group, which includes also
<taxonomicName id="8FF7D6FEFF99FFEC3D7BFC30FCED04A0" authority="Thomas, Marie &amp; Dia" authorityName="Thomas, Marie &amp; Dia" box="[309,773,964,990]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="bauernfeindi">
<emphasis id="7A83716FFF99FFEC3D7BFC30FE0004A3" box="[309,488,964,989]" italics="true" pageId="4" pageNumber="56">T. bauernfeindi</emphasis>
(Thomas, Marie &amp; Dia)
</taxonomicName>
,
<taxonomicName id="8FF7D6FEFF99FFEC3F5FFC30FB9D04A0" authority="Ikonomov" authorityName="Ikonomov" box="[785,1141,964,990]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="maculocaudata">
<emphasis id="7A83716FFF99FFEC3F5FFC30FC0B04A3" box="[785,995,964,989]" italics="true" pageId="4" pageNumber="56">T. maculocaudata</emphasis>
(Ikonomov)
</taxonomicName>
and
<taxonomicName id="8FF7D6FEFF99FFEC38E3FC30FA7404A0" authority="Allen" authorityName="Allen" box="[1197,1436,964,990]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="subsolana">
<emphasis id="7A83716FFF99FFEC38E3FC30FAA904A3" box="[1197,1345,964,989]" italics="true" pageId="4" pageNumber="56">T. subsolana</emphasis>
(Allen)
</taxonomicName>
(
<bibRefCitation id="2C66D08CFF99FFEC3CEEFC1EFDC9037A" author="Jacobus" box="[160,545,1002,1028]" pageId="4" pageNumber="58" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2008) Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society, 134, 185 - 274." type="journal article" year="2008">Jacobus &amp; McCafferty, 2008</bibRefCitation>
; Jacobus, 2009;
<bibRefCitation id="2C66D08CFF99FFEC3EB0FC1EFB64037A" author="Ubero-Pascal" box="[766,1164,1002,1028]" pageId="4" pageNumber="58" refString="Ubero-Pascal, N. &amp; Sartori, M. (2009) Phylogeny of the genus Teloganopsis Ulmer, 1939 with a redescription of Teloganopsis media Ulmer, 1939 and the description of a new Oriental species (Ephemeroptera: Ephemerellidae). Aquatic Insects, IN PRESS." type="journal article" year="2009">Ubero-Pascal &amp; Sartori, 2009</bibRefCitation>
).
<taxonomicName id="8FF7D6FEFF99FFEC38EAFC1EFA96037D" box="[1188,1406,1002,1027]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC38EAFC1EFA96037D" box="[1188,1406,1002,1027]" italics="true" pageId="4" pageNumber="56">Serratella elissa</emphasis>
</taxonomicName>
is distinguishable from these by its abdominal armature and lack of elongate apical denticles on the tarsal claws.
<taxonomicName id="8FF7D6FEFF99FFEC3CD9FBCCFEBF032F" box="[151,343,1080,1105]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC3CD9FBCCFEBF032F" box="[151,343,1080,1105]" italics="true" pageId="4" pageNumber="56">Serratella elissa</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="8FF7D6FEFF99FFEC3E39FBCCFBC2032C" authority="Bengtsson" authorityName="Bengtsson" box="[631,1066,1080,1106]" class="Insecta" family="Ephemerellidae" genus="Ephemerella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="mucronata">
<emphasis id="7A83716FFF99FFEC3E39FBCCFC79032F" box="[631,913,1080,1105]" italics="true" pageId="4" pageNumber="56">Ephemerella mucronata</emphasis>
(Bengtsson)
</taxonomicName>
, of the tribe Ephemerellini, but
<taxonomicName id="8FF7D6FEFF99FFEC3CD9FBABFF130306" box="[151,251,1119,1144]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF99FFEC3CD9FBABFF130306" box="[151,251,1119,1144]" italics="true" pageId="4" pageNumber="56">S. elissa</emphasis>
</taxonomicName>
is distinguishable by its having deeply cleft ventral lamellae of gills 6, a character shared by all Hyrtanellini (
<bibRefCitation id="2C66D08CFF99FFEC3D7AFB71FD6503E1" author="Jacobus" box="[308,653,1157,1183]" pageId="4" pageNumber="58" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2008) Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society, 134, 185 - 274." type="journal article" year="2008">Jacobus &amp; McCafferty, 2008</bibRefCitation>
).
</paragraph>
<caption id="1C88FDF5FF99FFEC3CD9F87FFC2200DD" box="[151,970,1931,1955]" pageId="4" pageNumber="56">
<paragraph id="4848AD7DFF99FFEC3CD9F87FFC2200DD" blockId="4.[151,970,1931,1955]" box="[151,970,1931,1955]" pageId="4" pageNumber="56">
<emphasis id="7A83716FFF99FFEC3CD9F87FFEC900DD" bold="true" box="[151,289,1931,1955]" pageId="4" pageNumber="56">FIGURES 4</emphasis>
, 5.
<taxonomicName id="8FF7D6FEFF99FFEC3D07F878FE1000DD" box="[329,504,1932,1955]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="elissa" status="sp. nov.">
<emphasis id="7A83716FFF99FFEC3D07F878FE1000DD" box="[329,504,1932,1955]" italics="true" pageId="4" pageNumber="56">Serratella elissa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="61B0CC14FF99FFEC3E4AF878FD6F00DD" box="[516,647,1932,1955]" pageId="4" pageNumber="56" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="7A83716FFF99FFEC3EDCF87FFD3000DD" bold="true" box="[658,728,1931,1955]" pageId="4" pageNumber="56">Fig. 4.</emphasis>
Maxilla.
<emphasis id="7A83716FFF99FFEC3F71F87FFC6D00DD" bold="true" box="[831,901,1931,1955]" pageId="4" pageNumber="56">Fig. 5.</emphasis>
Claw.
</paragraph>
</caption>
<caption id="1C88FDF5FF98FFED3CD9F859FCF100BA" box="[151,793,1965,1989]" pageId="5" pageNumber="57">
<paragraph id="4848AD7DFF98FFED3CD9F859FCF100BA" blockId="5.[151,793,1965,1989]" box="[151,793,1965,1989]" pageId="5" pageNumber="57">
<emphasis id="7A83716FFF98FFED3CD9F859FEC100BB" bold="true" box="[151,297,1965,1989]" pageId="5" pageNumber="57">FIGURES 6.</emphasis>
<taxonomicName id="8FF7D6FEFF98FFED3D60F859FE3500BA" box="[302,477,1965,1988]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="57" phylum="Arthropoda" rank="species" species="elissa" status="sp. nov.">
<emphasis id="7A83716FFF98FFED3D60F859FE3500BA" box="[302,477,1965,1988]" italics="true" pageId="5" pageNumber="57">Serratella elissa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="61B0CC14FF98FFED3DA4F859FD8500BA" box="[490,621,1965,1988]" pageId="5" pageNumber="57" rank="species">new species</taxonomicNameLabel>
. Dorsal habitus.
</paragraph>
</caption>
<paragraph id="4848AD7DFF9BFFEE3C8BFF6CFE6E0658" blockId="6.[151,1436,152,371]" pageId="6" pageNumber="58">
Couplet 20 of the recent key to ephemerellid larvae (
<bibRefCitation id="2C66D08CFF9BFFEE3F2CFF6CFAFA07CC" author="Jacobus" box="[866,1298,152,178]" pageId="6" pageNumber="58" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2008) Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society, 134, 185 - 274." type="journal article" year="2008">Jacobus &amp; McCafferty, 2008: 247</bibRefCitation>
) should be modified so that
<taxonomicName id="8FF7D6FEFF9BFFEE3D2BFF4BFE2207A6" box="[357,458,191,216]" class="Insecta" family="Ephemerellidae" genus="Serratella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="elissa">
<emphasis id="7A83716FFF9BFFEE3D2BFF4BFE2207A6" box="[357,458,191,216]" italics="true" pageId="6" pageNumber="58">S. elissa</emphasis>
</taxonomicName>
will pass through and not be misidentified as
<taxonomicName id="8FF7D6FEFF9BFFEE3FB1FF4BFAD007A6" authority="Klapalek" authorityName="Klapalek" box="[1023,1336,190,216]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="major">
<emphasis id="7A83716FFF9BFFEE3FB1FF4BFB4307A6" box="[1023,1195,191,216]" italics="true" pageId="6" pageNumber="58">Torleya major</emphasis>
(Klapálek)
</taxonomicName>
. Part 20 should include the following:
<emphasis id="7A83716FFF9BFFEE3DB2FF11FCA80780" box="[508,832,229,254]" italics="true" pageId="6" pageNumber="58">and gills 3 semi-operculate</emphasis>
. Part 20 should include the following:
<emphasis id="7A83716FFF9BFFEE3953FF11FE68065B" italics="true" pageId="6" pageNumber="58">and gills 3 not semi-operculate</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="00EDFEF6FF9BFFEE3C88FEC6FD8B060D" pageId="6" pageNumber="58" type="etymology">
<paragraph id="4848AD7DFF9BFFEE3C88FEC6FD8B060D" blockId="6.[151,1436,152,371]" pageId="6" pageNumber="58">
<emphasis id="7A83716FFF9BFFEE3C88FEC6FEBB0632" bold="true" box="[198,339,306,332]" pageId="6" pageNumber="58">Etymology.</emphasis>
The specific epithet is a noun in apposition, referring to one of three fictitious stepsisters in Edmund Spensers
<emphasis id="7A83716FFF9BFFEE3D34FEADFDB5060C" box="[378,605,345,370]" italics="true" pageId="6" pageNumber="58">The Faerie Queene</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>