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<document id="6508738BEA20C2F9E6523BA11ED13EF9" ID-CLB-Dataset="27456" ID-DOI="10.5852/ejt.2019.511" ID-GBIF-Dataset="6109d429-ee5b-4857-9b13-a580bbf39fce" ID-Zenodo-Dep="2626854" ID-ZooBank="8FDAD45D-1B7D-446F-8B34-026EDF192210" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554321381323" checkinUser="plazi" docAuthor="Kolicka, Małgorzata" docDate="2019" docId="892F87BC731E45791A6DCD09FCA75449" docLanguage="en" docName="ejt-511_kolicka.pdf.imf" docOrigin="European Journal of Taxonomy 511" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaetonotus (Chaetonotus) invitatus Kolicka 2019, sp. nov." docType="treatment" docUuid="5350662C-4565-4EB6-96B0-4742ED038286" docUuidSource="ZooBank" docVersion="10" lastPageNumber="17" masterDocId="7516FFC4731B45681867CC02FFF55677" masterDocTitle="Gastrotricha - not only in sediments: new epiphytic species of Chaetonotida from the Jubilee Greenhouse of the Botanical Garden in Kraków" masterLastPageNumber="100" masterPageNumber="1" pageNumber="6" updateTime="1698717331031" updateUser="ExternalLinkService">
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<mods:title id="C2CF07F541039826305DDE3725CEF79A">Gastrotricha - not only in sediments: new epiphytic species of Chaetonotida from the Jubilee Greenhouse of the Botanical Garden in Kraków</mods:title>
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<mods:namePart id="E697B7FE8BB3D53089DE0CED8B180C14">Kolicka, Małgorzata</mods:namePart>
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<mods:title id="AF6C9F073A32E4CFE2C2A15F379ED0A9">European Journal of Taxonomy</mods:title>
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<mods:date id="6F4609E0859997C6FC89AAFC34E9CD70">2019</mods:date>
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<mods:number id="51927336471FA0B83655F5FF1CDDC5E6">2019-04-02</mods:number>
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<mods:number id="CFAB64E852B3848EF6FE729ECA6CA1E0">511</mods:number>
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<treatment id="892F87BC731E45791A6DCD09FCA75449" ID-DOI="http://doi.org/10.5281/zenodo.5620059" ID-GBIF-Taxon="154533728" ID-Zenodo-Dep="5620059" LSID="urn:lsid:zoobank.org:act:5350662C-4565-4EB6-96B0-4742ED038286" httpUri="http://treatment.plazi.org/id/892F87BC731E45791A6DCD09FCA75449" lastPageId="17" lastPageNumber="17" pageId="5" pageNumber="6">
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<paragraph id="013936AA731E456D1A6DCD09FBDC5751" blockId="5.[379,1208,267,364]" box="[522,1065,267,294]" pageId="5" pageNumber="6">
<heading id="5A7181C6731E456D1A6DCD09FBDC5751" box="[522,1065,267,294]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="C6864D29731E456D1A6DCD09FC325752" ID-CoL="7P6C4" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[522,967,267,294]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="6" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731E456D1A6DCD09FC325752" bold="true" box="[522,967,267,294]" pageId="5" pageNumber="6">
<emphasis id="33F2EAB8731E456D1A6DCD09FD565751" bold="true" box="[522,675,267,294]" italics="true" pageId="5" pageNumber="6">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8731E456D1AD2CD09FCBB5751" bold="true" box="[693,846,267,294]" italics="true" pageId="5" pageNumber="6">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731E456D1B38CD09FC325752" bold="true" box="[863,967,267,293]" italics="true" pageId="5" pageNumber="6">invitatus</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731E456D1BA8CD0EFBDC5751" box="[975,1065,268,294]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="499C6521731E456D1A23CD50FC05571B" box="[580,1008,338,364]" pageId="5" pageNumber="6" type="description">
<paragraph id="013936AA731E456D1A23CD50FC05571B" blockId="5.[379,1208,267,364]" box="[580,1008,338,364]" pageId="5" pageNumber="6">
<figureCitation id="99BD2A2F731E456D1A23CD50FD4F571B" box="[580,698,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="6.[179,222,531,559]" captionStartId-1="7.[189,232,1647,1673]" captionStartId-2="8.[189,232,1929,1955]" captionStartId-3="9.[189,232,1944,1970]" captionStartId-4="10.[189,232,1894,1920]" captionStartId-5="11.[189,232,1894,1920]" captionStartId-6="12.[189,232,1888,1914]" captionStartId-7="13.[189,232,1929,1955]" captionStartId-8="14.[189,232,1964,1990]" captionStartId-9="15.[189,232,1964,1990]" captionTargetBox-0="[236,1862,635,1741]" captionTargetBox-1="[225,1359,275,1607]" captionTargetBox-2="[197,1390,1228,1889]" captionTargetBox-3="[189,1398,1052,1906]" captionTargetBox-4="[231,1356,266,1858]" captionTargetBox-5="[189,1398,1009,1864]" captionTargetBox-6="[189,1398,1005,1860]" captionTargetBox-7="[189,1398,1041,1893]" captionTargetBox-8="[189,1398,1077,1929]" captionTargetBox-9="[189,1398,1080,1934]" captionTargetId-0="figure@6.[231,1880,625,1751]" captionTargetId-1="figure@7.[219,1368,265,1613]" captionTargetId-2="figure@8.[189,1398,1218,1901]" captionTargetId-3="figure@9.[189,1398,1052,1907]" captionTargetId-4="figure@10.[231,1356,266,1858]" captionTargetId-5="figure@11.[189,1398,1009,1864]" captionTargetId-6="figure@12.[189,1398,1005,1860]" captionTargetId-7="figure@13.[189,1398,1041,1893]" captionTargetId-8="figure@14.[189,1398,1077,1929]" captionTargetId-9="figure@15.[189,1398,1080,1934]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." captionText-1="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." captionText-2="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." captionText-3="Fig. 5. Chaetonotus (Chaetonotus) invitatus sp. nov. Habitus. Phase contrast microphotographs. A. Juvenile specimen. B. Adult specimen. C. Specimen with mature egg." captionText-4="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." captionText-5="Fig. 7. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Detail of dorsal head view of adult specimen. B. Detail of dorsal head view of juvenile specimen. C. Dorsal view of posterior trunk region of specimen with two scales 17. D. Dorsal view of posterior trunk region of specimen with three scales 17." captionText-6="Fig. 8. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Dorsal view of the posterior trunk region, furca base and furcal appendage scales. B. Ventral view of the posterior trunk region, furca base and furcal appendage scales." captionText-7="Fig. 9. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotographs. A. Detail of the mouth ring, dorsal view. B. Detail of the mouth ring, internal view. C. Detail of the pharynx." captionText-8="Fig. 10. Chaetonotus (Chaetonotus) invitatus sp. nov. Detail of internal morphology. Bright field microphotography. X-organ." captionText-9="Fig. 11. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotography. Living euglenids inside intestine." httpUri-0="https://zenodo.org/record/2626858/files/figure.png" httpUri-1="https://zenodo.org/record/2626860/files/figure.png" httpUri-2="https://zenodo.org/record/2626862/files/figure.png" httpUri-3="https://zenodo.org/record/2626864/files/figure.png" httpUri-4="https://zenodo.org/record/2626866/files/figure.png" httpUri-5="https://zenodo.org/record/2626868/files/figure.png" httpUri-6="https://zenodo.org/record/2626870/files/figure.png" httpUri-7="https://zenodo.org/record/2626872/files/figure.png" httpUri-8="https://zenodo.org/record/2626874/files/figure.png" httpUri-9="https://zenodo.org/record/2626876/files/figure.png" pageId="5" pageNumber="6">Figs 211</figureCitation>
, A
<figureCitation id="99BD2A2F731E456D1ABBCD50FCFA571B" box="[732,783,338,364]" captionStart="Fig" captionStartId="2.[189,232,1271,1297]" captionTargetBox="[189,1398,780,1232]" captionTargetId="figure@2.[189,1398,780,1232]" captionTargetPageId="2" captionText="Fig. 1. Study area. A. The inside of Jubilee Greenhouse at the Botanical Garden in Kraków. B. Sampling site." httpUri="https://zenodo.org/record/2626856/files/figure.png" pageId="5" pageNumber="6">1A</figureCitation>
<figureCitation id="99BD2A2F731E456D1B68CD50FCEB571B" box="[783,798,338,364]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="5" pageNumber="6">2</figureCitation>
;
<tableCitation id="4C040311731E456D1B4DCD50FC7B571B" box="[810,910,338,364]" captionStart="Table 2" captionStartId="9.[189,258,267,293]" captionText="Table 2. Main morphometric characters of Chaetonotus (Chaetonotus) invitatus sp. nov. N = number of specimens analysed; Range = the smallest and the largest structure found among all specimens measured. All measurements in micrometers (μm)." pageId="5" pageNumber="6">Tables 2</tableCitation>
, A
<tableCitation id="4C040311731E456D1BD7CD50FC14571B" box="[944,993,338,364]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Sampling sites with specified aquatic plant species." pageId="5" pageNumber="6">1A</tableCitation>
<tableCitation id="4C040311731E456D1B86CD50FC05571B" box="[993,1008,338,364]" captionStart="Table 2" captionStartId="9.[189,258,267,293]" captionText="Table 2. Main morphometric characters of Chaetonotus (Chaetonotus) invitatus sp. nov. N = number of specimens analysed; Range = the smallest and the largest structure found among all specimens measured. All measurements in micrometers (μm)." pageId="5" pageNumber="6">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="499C6521731E456D18DACD91FDA5520F" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="013936AA731E456D18DACD91FEC357DA" blockId="5.[189,310,403,429]" box="[189,310,403,429]" pageId="5" pageNumber="6">
<heading id="5A7181C6731E456D18DACD91FEC357DA" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="33F2EAB8731E456D18DACD91FEC357DA" bold="true" box="[189,310,403,429]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA731E456D18DACDC0FDA5520F" blockId="5.[189,1399,450,1144]" pageId="5" pageNumber="6">
Slender, bottle-shaped, body measuring from 174.6 to 194.7 μm in length. Head five-lobed, cephalion, epipleurae and hypopleurae clearly demarcated in head outline. Hypostomium kidney-shaped with horn-like protuberances anterolaterally and pair of reinforcements near anterior edge. Two additional plates beyond posterior hypostomium edge. Ocellar granules absent. In mouth ring, two weak cuticular teeth arising from anterior pharynx region. All scales one-lobed, with shallow to deep posterior notches, keeled and spined. Scales distributed in 2325 single longitudinal rows (57D +4DL+6L+4LV+
<date id="7538106A731E456D1D2ECE70FA9B54FB" box="[1353,1390,626,652]" pageId="5" pageNumber="6">4V</date>
) with 2527 scales in central row. Scales strongly differing morphologically in body areas. Scales located close to one another and with overlapping edges on furcal base and furcal appendages. Spines basally bent and thick, subsequently strongly and gradually tapering towards hair-like end, ventral spines hairlike along entire length. Lateral spines more strongly curved than dorsal and dorsolateral ones. Length of spines gradually increases from dorsal, dorsolateral, lateral and ventrolateral surfaces towards ventral surface and from head towards widest trunk region. Dorsally on posterior trunk region, two or three scales with longer and thicker spines. Dorsally and dorsolaterally on posterior trunk region, on furcal base and furcal appendages, scales with shorter spines, with rudimentary spines or without any spine. Dorsolaterally on furcal appendages, one pair of scales with long, rigid and spike-like spines reaching to furca inner indentation. Furcal appendages with two pairs of scales with long, thick parafurcal spines tapering to their ends. Ventral interciliary field covered with semirectangular plates on pharyngeal region and one-lobed, keeled scales on intestine region. Four pairs of ventral interciliary field terminal scales. Pharynx narrow, with anterior and posterior dilatations. Intestine straight without anterior section differing in form and morphology.
</paragraph>
</subSubSection>
<subSubSection id="499C6521731E456D18DAC8AAFAD35287" box="[189,1318,1192,1265]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="013936AA731E456D18DAC8AAFEB152B5" blockId="5.[189,324,1192,1218]" box="[189,324,1192,1218]" pageId="5" pageNumber="6">
<heading id="5A7181C6731E456D18DAC8AAFEB152B5" bold="true" box="[189,324,1192,1218]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="33F2EAB8731E456D18DAC8AAFEB152B5" bold="true" box="[189,324,1192,1218]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA731E456D18DAC8D5FAD35287" blockId="5.[189,1318,1238,1265]" box="[189,1318,1238,1265]" pageId="5" pageNumber="6">
From the Latin ʻ
<taxonomicName id="C6864D29731E456D1919C8D5FE165287" authorityName="Kolicka" authorityYear="2019" box="[382,483,1239,1264]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="6" phylum="Gastrotricha" rank="species" species="invitatus">
<emphasis id="33F2EAB8731E456D1919C8D5FE165287" box="[382,483,1239,1264]" italics="true" pageId="5" pageNumber="6">invitatus</emphasis>
</taxonomicName>
ʼ = ʻinvitedʼ, referring to the artificial place to which it was transferred.
</paragraph>
</subSubSection>
<subSubSection id="499C6521731E456D18DAC91DFC945016" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="013936AA731E456D18DAC91DFE5F534E" blockId="5.[189,426,1311,1337]" box="[189,426,1311,1337]" pageId="5" pageNumber="6">
<heading id="5A7181C6731E456D18DAC91DFE5F534E" bold="true" box="[189,426,1311,1337]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="33F2EAB8731E456D18DAC91DFE5F534E" bold="true" box="[189,426,1311,1337]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA731E456D188DC94CFEA9531F" blockId="5.[189,1399,1358,1490]" box="[234,348,1358,1384]" pageId="5" pageNumber="6">
<materialsCitation id="B1EE3CF7731E456D188DC94CFEA9531F" ID-GBIF-Occurrence="2234225218" box="[234,348,1358,1384]" collectingDate="2013-11-15" collectorName="M. Kolicka" country="Poland" latitude="50.060555" location="POLAND" longLatPrecision="18" longitude="19.958334" pageId="5" pageNumber="6" specimenCode="NHC-GCCI-20-1-25/h" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<emphasis id="33F2EAB8731E456D188DC94CFEA9531F" bold="true" box="[234,348,1358,1384]" pageId="5" pageNumber="6">
<typeStatus id="DE3D8808731E456D188DC94CFEA9531F" box="[234,348,1358,1384]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="013936AA731E456D18DAC973FEB753A5" blockId="5.[189,1399,1358,1490]" pageId="5" pageNumber="6">
<collectingCountry id="7991763A731E456D18DAC973FEC153FC" box="[189,308,1393,1419]" name="Poland" pageId="5" pageNumber="6">POLAND</collectingCountry>
<specimenCount id="1780FD23731E456D1936C973FE7A53FC" box="[337,399,1393,1419]" pageId="5" pageNumber="6" type="adult">adult</specimenCount>
; Kraków, Botanical Garden, Jubilee Greenhouse, site 1;
<geoCoordinate id="64B2506D731E456D1C5EC96DFB2553FC" box="[1081,1232,1391,1419]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="15" value="50.060555">50°03'38&quot; N</geoCoordinate>
,
<geoCoordinate id="64B2506D731E456D1CB9C96DFA8453FC" box="[1246,1393,1391,1419]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="15" value="19.958334">19°57'30&quot; E</geoCoordinate>
;
<date id="7538106A731E456D18DAC996FE8453D9" box="[189,369,1428,1455]" pageId="5" pageNumber="6" value="2013-11-15">
<collectingDate id="657CE982731E456D18DAC996FE8453D9" box="[189,369,1428,1455]" pageId="5" pageNumber="6" value="2013-11-15">15 Nov. 2013</collectingDate>
</date>
;
<collectorName id="AC73537C731E456D19E1C996FDE053D9" box="[390,533,1428,1454]" pageId="5" pageNumber="6">M. Kolicka</collectorName>
leg.;
<specimenCode id="51209ED1731E456D1A0EC996FC8153D8" box="[617,884,1428,1455]" pageId="5" pageNumber="6">NHC-GCCI-20-1-25/h</specimenCode>
(photomicrographs, also in the author's collection).
</paragraph>
<paragraph id="013936AA731E456D188DCA02FC945016" blockId="5.[189,1399,1536,1633]" pageId="5" pageNumber="6">
<materialsCitation id="B1EE3CF7731E456D188DCA02FCA85016" ID-GBIF-Occurrence="2234225212" country="Poland" location="POLAND" pageId="5" pageNumber="6" specimenCode="NHC-GCCI-20-26- 100/p" specimenCount="34" specimenCount-adult="24" specimenCount-juvenile="5" specimenCount-subadult="5" typeStatus="paratype">
<emphasis id="33F2EAB8731E456D188DCA02FE92506D" bold="true" box="[234,359,1536,1562]" pageId="5" pageNumber="6">
<typeStatus id="DE3D8808731E456D188DCA02FE92506D" box="[234,359,1536,1562]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7991763A731E456D18DACA21FEC1504A" box="[189,308,1571,1597]" name="Poland" pageId="5" pageNumber="6">POLAND</collectingCountry>
<specimenCount id="1780FD23731E456D192CCA21FE42504A" box="[331,439,1571,1597]" pageId="5" pageNumber="6" type="adult">24 adults</specimenCount>
,
<specimenCount id="1780FD23731E456D19A5CA21FDBC504A" box="[450,585,1571,1598]" pageId="5" pageNumber="6" type="subadult">5 subadults</specimenCount>
,
<specimenCount id="1780FD23731E456D1A34CA21FD23504A" box="[595,726,1571,1598]" pageId="5" pageNumber="6" type="juvenile">5 juveniles</specimenCount>
; same collection data as for holotype;
<specimenCode id="51209ED1731E456D1CF6CA21FEF55016" pageId="5" pageNumber="6">NHC-GCCI-20-26- 100/p</specimenCode>
(photomicrographs, also in the authors collection)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="499C6521731E457818DACA8DFE0D5500" lastPageId="16" lastPageNumber="17" pageId="5" pageNumber="6" type="description">
<paragraph id="013936AA731E456D18DACA8DFEBB50DE" blockId="5.[189,334,1679,1705]" box="[189,334,1679,1705]" pageId="5" pageNumber="6">
<heading id="5A7181C6731E456D18DACA8DFEBB50DE" bold="true" box="[189,334,1679,1705]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="33F2EAB8731E456D18DACA8DFEBB50DE" bold="true" box="[189,334,1679,1705]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA731E456D18DACABDFE2D51B8" blockId="5.[189,1399,1726,1999]" pageId="5" pageNumber="6">
HABITUS.
<taxonomicName id="C6864D29731E456D194DCABCFD2D50AF" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[298,728,1726,1753]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="5" pageNumber="6" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731E456D194DCABCFE4B50AF" box="[298,446,1726,1752]" italics="true" pageId="5" pageNumber="6">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8731E456D19A9CABCFD9750AF" box="[462,610,1726,1752]" italics="true" pageId="5" pageNumber="6">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731E456D1A14CABDFD2D50AF" box="[627,728,1727,1752]" italics="true" pageId="5" pageNumber="6">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731E456D1AB9CABDFCCD50AE" box="[734,824,1727,1753]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
has a slender, bottle-shaped body (
<figureCitation id="99BD2A2F731E456D1CAACABCFAED50AF" box="[1229,1304,1726,1752]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731E456D1D44CABCFAC150AE" box="[1315,1332,1726,1753]" captionStart="Fig" captionStartId="9.[189,232,1944,1970]" captionTargetBox="[189,1398,1052,1906]" captionTargetId="figure@9.[189,1398,1052,1907]" captionTargetPageId="9" captionText="Fig. 5. Chaetonotus (Chaetonotus) invitatus sp. nov. Habitus. Phase contrast microphotographs. A. Juvenile specimen. B. Adult specimen. C. Specimen with mature egg." httpUri="https://zenodo.org/record/2626864/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). The head is wide and the neck constriction is strongly marked. The neck gradually tapers to the beginning of the trunk (ca U38) (
<figureCitation id="99BD2A2F731E456D19AACB06FDEE5168" box="[461,539,1796,1823]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731E456D1A4ECB06FDCD5168" box="[553,568,1796,1823]" captionStart="Fig" captionStartId="9.[189,232,1944,1970]" captionTargetBox="[189,1398,1052,1906]" captionTargetId="figure@9.[189,1398,1052,1907]" captionTargetPageId="9" captionText="Fig. 5. Chaetonotus (Chaetonotus) invitatus sp. nov. Habitus. Phase contrast microphotographs. A. Juvenile specimen. B. Adult specimen. C. Specimen with mature egg." httpUri="https://zenodo.org/record/2626864/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). The trunk is only slightly wider than the head and gradually dilates from ca U39 to ca U59, where it is at its maximum width. Then it gradually tapers slightly towards a weakly demarcated furcal base (from U85) (
<figureCitation id="99BD2A2F731E456D1AB7CB49FCEF5112" box="[720,794,1867,1893]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
). The furcal indentation is parabolic in shape. The furcal branches are set apart and point slightly outwards (
<figureCitation id="99BD2A2F731E456D1B00CB6CFC4751FF" box="[871,946,1902,1928]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="5" pageNumber="6">Figs 6</figureCitation>
GH, 8). The adhesive tubes are long, slightly bent, narrow and very gradually taper slightly towards their ends. The adhesive tube ends are blunt (
<figureCitation id="99BD2A2F731E456D196ECBB6FEA051B8" box="[265,341,1972,1999]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731E456D1905CBB6FE8551B9" box="[354,368,1972,1998]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
GH, 8).
</paragraph>
<caption id="55F96622731D456E18D4CE11FC365425" httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="6" pageNumber="7" startId="6.[179,222,531,559]" targetBox="[236,1862,635,1741]" targetPageId="6">
<paragraph id="013936AA731D456E18D4CE11FC365425" blockId="6.[179,1932,531,594]" pageId="6" pageNumber="7">
<emphasis id="33F2EAB8731D456E18D4CE11FF0A5458" bold="true" box="[179,255,531,559]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
<taxonomicName id="C6864D29731D456E1961CE11FD465458" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[262,691,531,559]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="6" pageNumber="7" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731D456E1961CE11FE6F5458" box="[262,410,531,559]" italics="true" pageId="6" pageNumber="7">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8731D456E19CDCE11FDCB5458" box="[426,574,531,559]" italics="true" pageId="6" pageNumber="7">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731D456E1A29CE11FD465458" box="[590,691,531,559]" italics="true" pageId="6" pageNumber="7">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731D456E1ADDCE11FCE65458" box="[698,787,531,559]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
Schematic drawings.
<emphasis id="33F2EAB8731D456E1C74CE11FBDD5458" bold="true" box="[1043,1064,531,559]" pageId="6" pageNumber="7">A</emphasis>
. Dorsal body view.
<emphasis id="33F2EAB8731D456E1D77CE11FAD15458" bold="true" box="[1296,1316,531,559]" pageId="6" pageNumber="7">B</emphasis>
. View of internal morphology.
<emphasis id="33F2EAB8731D456E1EE8CE11F9515458" bold="true" box="[1679,1700,531,559]" pageId="6" pageNumber="7">C</emphasis>
. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands.
</paragraph>
</caption>
<caption id="55F96622731C456F18DACA6DFD59519D" httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1647,1673]" targetBox="[225,1359,275,1607]" targetPageId="7">
<paragraph id="013936AA731C456F18DACA6DFD59519D" blockId="7.[189,1399,1647,2026]" pageId="7" pageNumber="8">
<emphasis id="33F2EAB8731C456F18DACA6DFEE450FE" bold="true" box="[189,273,1647,1673]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
<taxonomicName id="C6864D29731C456F1978CA6DFD2950FE" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[287,732,1647,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="7" pageNumber="8" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731C456F1978CA6DFE4650FE" box="[287,435,1647,1673]" italics="true" pageId="7" pageNumber="8">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8731C456F19ACCA6DFDAA50FE" box="[459,607,1647,1673]" italics="true" pageId="7" pageNumber="8">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731C456F1A10CA72FD2950FE" box="[631,732,1648,1673]" italics="true" pageId="7" pageNumber="8">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731C456F1A8DCA72FCBE50FD" box="[746,843,1648,1674]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
Schematic drawings of the scales.
<emphasis id="33F2EAB8731C456F1D69CA6DFAD650FE" bold="true" box="[1294,1315,1647,1673]" pageId="7" pageNumber="8">A</emphasis>
. Head anteriormost dorsal scale (scale 1).
<emphasis id="33F2EAB8731C456F1A39CA91FD8750DA" bold="true" box="[606,626,1683,1709]" pageId="7" pageNumber="8">B</emphasis>
. Head dorsal scale (scale 2).
<emphasis id="33F2EAB8731C456F1BABCA91FC1450DA" bold="true" box="[972,993,1683,1709]" pageId="7" pageNumber="8">C</emphasis>
. Head dorsolateral scale (scale 3).
<emphasis id="33F2EAB8731C456F18DACAB4FF2750A7" bold="true" box="[189,210,1718,1744]" pageId="7" pageNumber="8">D</emphasis>
. Head lateral and ventrolateral scale (scale 4).
<emphasis id="33F2EAB8731C456F1A92CAB4FCFC50A7" bold="true" box="[757,777,1718,1744]" pageId="7" pageNumber="8">E</emphasis>
. Head ventral scale (scale 5).
<emphasis id="33F2EAB8731C456F1C0FCAB4FB8F50A7" bold="true" box="[1128,1146,1718,1744]" pageId="7" pageNumber="8">F</emphasis>
. Anterior neck dorsal scale (scale 6).
<emphasis id="33F2EAB8731C456F1913CADBFE7E5084" bold="true" box="[372,395,1753,1779]" pageId="7" pageNumber="8">G</emphasis>
. Anterior neck dorsolateral scale (scale 7).
<emphasis id="33F2EAB8731C456F1BE9CADBFC505084" bold="true" box="[910,933,1753,1779]" pageId="7" pageNumber="8">H</emphasis>
. Anterior neck lateral and ventrolateral scale (scale 8).
<emphasis id="33F2EAB8731C456F19E4CAFFFE7B5160" bold="true" box="[387,398,1789,1815]" pageId="7" pageNumber="8">I</emphasis>
. Anterior neck ventral scale (scale 9).
<emphasis id="33F2EAB8731C456F1B1CCAFFFC7F5160" bold="true" box="[891,906,1789,1815]" pageId="7" pageNumber="8">J</emphasis>
. Posterior neck dorsal scale (scale 10).
<emphasis id="33F2EAB8731C456F18DACB22FF21514D" bold="true" box="[189,212,1824,1850]" pageId="7" pageNumber="8">K</emphasis>
. Posterior neck dorsolateral scale (scale 11).
<emphasis id="33F2EAB8731C456F1A94CB22FCF2514D" bold="true" box="[755,775,1824,1850]" pageId="7" pageNumber="8">L</emphasis>
. Posterior neck lateral and ventrolateral scale (scale 12).
<emphasis id="33F2EAB8731C456F1894CB41FEFA512A" bold="true" box="[243,271,1859,1885]" pageId="7" pageNumber="8">M</emphasis>
. Posterior neck ventral scale (scale 13).
<emphasis id="33F2EAB8731C456F1A88CB41FCF1512A" bold="true" box="[751,772,1859,1885]" pageId="7" pageNumber="8">N</emphasis>
. Trunk dorsal scale (scale 14).
<emphasis id="33F2EAB8731C456F1C1FCB41FB7A512A" bold="true" box="[1144,1167,1859,1885]" pageId="7" pageNumber="8">O</emphasis>
. Trunk dorsolateral scale (scale 15).
<emphasis id="33F2EAB8731C456F19E1CB64FE6D51F7" bold="true" box="[390,408,1894,1920]" pageId="7" pageNumber="8">P</emphasis>
. Trunk lateral and ventrolateral scale (scale 16).
<emphasis id="33F2EAB8731C456F1B82CB64FC0951F7" bold="true" box="[997,1020,1894,1920]" pageId="7" pageNumber="8">Q</emphasis>
. Trunk ventral scale (scale 17).
<emphasis id="33F2EAB8731C456F18DACB8BFF2751D4" bold="true" box="[189,210,1929,1955]" pageId="7" pageNumber="8">R</emphasis>
. Scale 18.
<emphasis id="33F2EAB8731C456F1932CB8BFE9051D4" bold="true" box="[341,357,1929,1955]" pageId="7" pageNumber="8">S</emphasis>
. Scale 19.
<emphasis id="33F2EAB8731C456F198FCB8BFE0951D4" bold="true" box="[488,508,1929,1955]" pageId="7" pageNumber="8">T</emphasis>
. Scale 20.
<emphasis id="33F2EAB8731C456F1A19CB8BFD6651D4" bold="true" box="[638,659,1929,1955]" pageId="7" pageNumber="8">U</emphasis>
. Scale 21.
<emphasis id="33F2EAB8731C456F1B71CB8BFCDE51D4" bold="true" box="[790,811,1929,1955]" pageId="7" pageNumber="8">V</emphasis>
. Scale 22.
<emphasis id="33F2EAB8731C456F1BC9CB8BFC3E51D4" bold="true" box="[942,971,1929,1955]" pageId="7" pageNumber="8">W</emphasis>
. Scale 23.
<emphasis id="33F2EAB8731C456F1C29CB8BFB9651D4" bold="true" box="[1102,1123,1929,1955]" pageId="7" pageNumber="8">X</emphasis>
. Scale 24.
<emphasis id="33F2EAB8731C456F1C81CB8BFB0E51D4" bold="true" box="[1254,1275,1929,1955]" pageId="7" pageNumber="8">Y</emphasis>
. Scale 25.
<emphasis id="33F2EAB8731C456F18DACBAFFF2451B0" bold="true" box="[189,209,1965,1991]" pageId="7" pageNumber="8">Z</emphasis>
. Scale 26.
<emphasis id="33F2EAB8731C456F193ECBAEFE7651B1" bold="true" box="[345,387,1964,1990]" pageId="7" pageNumber="8">AA</emphasis>
. Scale 27.
<emphasis id="33F2EAB8731C456F1A6ACBAFFDC151B0" bold="true" box="[525,564,1965,1991]" pageId="7" pageNumber="8">BB</emphasis>
. Scale 28.
<emphasis id="33F2EAB8731C456F1ADACBAEFD1251B1" bold="true" box="[701,743,1964,1990]" pageId="7" pageNumber="8">CC</emphasis>
. Scale 29.
<emphasis id="33F2EAB8731C456F1B17CBAFFC6F51B0" bold="true" box="[880,922,1965,1991]" pageId="7" pageNumber="8">DD</emphasis>
. Scale 30.
<emphasis id="33F2EAB8731C456F1C44CBAFFBBF51B0" bold="true" box="[1059,1098,1965,1991]" pageId="7" pageNumber="8">EE</emphasis>
. Scale 31.
<emphasis id="33F2EAB8731C456F1CB4CBAFFB0251B0" bold="true" box="[1235,1271,1965,1991]" pageId="7" pageNumber="8">FF</emphasis>
. Scale 32.
<emphasis id="33F2EAB8731C456F18DACBCDFF1E519E" bold="true" box="[189,235,1999,2025]" pageId="7" pageNumber="8">GG</emphasis>
. Scale 33.
<emphasis id="33F2EAB8731C456F190BCBD2FE6F519D" bold="true" box="[364,410,2000,2026]" pageId="7" pageNumber="8">HH</emphasis>
. Scale 34.
<emphasis id="33F2EAB8731C456F1A7CCBD2FDC7519D" bold="true" box="[539,562,2000,2026]" pageId="7" pageNumber="8">II</emphasis>
. Scale 35.
</paragraph>
</caption>
<paragraph id="013936AA7313456018DACD0EFE89527E" blockId="8.[189,1399,267,1033]" pageId="8" pageNumber="9">
HEAD. The head is five-lobed, long and parabolic. The cephalion (U1U7) is long and wide, it is visible on the dorsal head surface as an inverted, rounded triangle. It is clearly demarcated in the body outline and its dorsal edge is free and does not adhere to the head surface (
<figureCitation id="99BD2A2F731345601BEACD50FC24571B" box="[909,977,338,364]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1398,1009,1864]" captionTargetId="figure@11.[189,1398,1009,1864]" captionTargetPageId="11" captionText="Fig. 7. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Detail of dorsal head view of adult specimen. B. Detail of dorsal head view of juvenile specimen. C. Dorsal view of posterior trunk region of specimen with two scales 17. D. Dorsal view of posterior trunk region of specimen with three scales 17." httpUri="https://zenodo.org/record/2626868/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
AB). The pleurae are large, convex and clearly demarcated in the head outline, the hypopleurae are only slightly larger than the epipleurae (
<figureCitation id="99BD2A2F7313456018A1CD9AFEFA57C5" box="[198,271,408,434]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). Between the epipleurae and hypopleurae, deep notches are present. The epipleurae (U3U7) are located on the dorsal, dorsolateral, lateral and ventrolateral surfaces, whereas the hypopleurae (U8 U12) cover the dorsal, dorsolateral, lateral, ventrolateral and ventral surfaces in such a manner that their greatest portion is located on the ventral side. The hypostomium is kidney-shaped and has two horn-like protuberances placed anterolaterally and a pair of strong reinforcements near its anterior edge (U3U6) (
<figureCitation id="99BD2A2F7313456018A2CE4AFED35415" box="[197,294,584,610]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="8" pageNumber="9">Figs 2C</figureCitation>
,
<figureCitation id="99BD2A2F731345601954CE4AFEAD5414" box="[307,344,584,611]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="8" pageNumber="9">6B</figureCitation>
). One pair of additional plates is located posterolaterally to the hypostomium, at U6U7. These plates are wide and short and their lateral edges extend to the edges of the ventral epipleurae (
<figureCitation id="99BD2A2F7313456018A1CE8CFED454DE" box="[198,289,654,681]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="8" pageNumber="9">Fig. 6B</figureCitation>
). Two pairs of cephalic ciliary tufts are present. The anterior tufts emerge laterally between the cephalion edge and the dorsal edge of the epipleurae (at U3). The anterior tufts consist of 5 cilia which are bent and short, and their length increases from the first to the last cilium (
<tableCitation id="4C040311731345601C35CED7FB375498" box="[1106,1218,725,751]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Sampling sites with specified aquatic plant species." pageId="8" pageNumber="9">Table A1</tableCitation>
). The posterior tufts have 5 cilia each and emerge laterally between the epipleurae and the hypopleurae (at U7U8). Cilia in the posterior tufts are long and nearly straight and their length increases from the first cilium to the last, long cilium (
<tableCitation id="4C0403117313456019B8CF3CFDA5552E" box="[479,592,830,857]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Sampling sites with specified aquatic plant species." pageId="8" pageNumber="9">Table A1</tableCitation>
). Ocellar granules are absent. The mouth ring is located terminally at U1U3, is wide and has long, strong finger-like reinforcements as well as very long cuticular inner hairs (
<figureCitation id="99BD2A2F731345601962CF87FEA555E8" box="[261,336,901,927]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F73134560193BCF87FE9F55E8" box="[348,362,901,927]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[189,1398,1041,1893]" captionTargetId="figure@13.[189,1398,1041,1893]" captionTargetPageId="13" captionText="Fig. 9. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotographs. A. Detail of the mouth ring, dorsal view. B. Detail of the mouth ring, internal view. C. Detail of the pharynx." httpUri="https://zenodo.org/record/2626872/files/figure.png" pageId="8" pageNumber="9">9</figureCitation>
AB). Inside the mouth ring, two small and weak cuticular teeth arising from the anterior pharynx region are present (
<figureCitation id="99BD2A2F731345601A6BCFAAFDA755B5" box="[524,594,936,962]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[189,1398,1041,1893]" captionTargetId="figure@13.[189,1398,1041,1893]" captionTargetPageId="13" captionText="Fig. 9. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotographs. A. Detail of the mouth ring, dorsal view. B. Detail of the mouth ring, internal view. C. Detail of the pharynx." httpUri="https://zenodo.org/record/2626872/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
BC). Suboral bristles emerge in three pairs of tufts, laterally to the mouth ring, near the lateral edges of the hypostomium and in lines below the additional hypostomium plates (
<figureCitation id="99BD2A2F731345601976CFECFE9B527E" box="[273,366,1006,1033]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="013936AA7313456518DAC832FDF6552E" blockId="8.[189,1399,1072,1169]" lastBlockId="13.[189,1399,267,857]" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9">
SCALES. The body is covered with one-lobed scales that adhere along their entire surface to the cuticle (
<figureCitation id="99BD2A2F7313456018A2C851FEE1521A" box="[197,276,1107,1133]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="8" pageNumber="9">Figs 3</figureCitation>
,
<figureCitation id="99BD2A2F731345601942C851FEA05219" box="[293,341,1107,1134]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,232,1894,1920]" captionStartId-1="11.[189,232,1894,1920]" captionTargetBox-0="[231,1356,266,1858]" captionTargetBox-1="[189,1398,1009,1864]" captionTargetId-0="figure@10.[231,1356,266,1858]" captionTargetId-1="figure@11.[189,1398,1009,1864]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." captionText-1="Fig. 7. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Detail of dorsal head view of adult specimen. B. Detail of dorsal head view of juvenile specimen. C. Dorsal view of posterior trunk region of specimen with two scales 17. D. Dorsal view of posterior trunk region of specimen with three scales 17." httpUri-0="https://zenodo.org/record/2626866/files/figure.png" httpUri-1="https://zenodo.org/record/2626868/files/figure.png" pageId="8" pageNumber="9">67</figureCitation>
). The scales have a straight keel and are egg-shaped with very weak posterior notches to very deep posterior notches. The scales are distributed in 2325 longitudinal alternating rows (57D +4DL+6L+4LV+
<date id="7538106A731045631989CD09FDE65752" box="[494,531,267,293]" pageId="11" pageNumber="12">4V</date>
) with 2527 scales in the central row. The longitudinal rows of scales run straight and are arranged in parallel to one another from the top of the head to the widest body region. Dorsally and dorsolaterally on the posterior trunk region, furca base and furcal appendages, the scales are arranged in rounded arcs and in a rosette (
<figureCitation id="99BD2A2F731045631B41CD77FC8657F8" box="[806,883,373,399]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731045631B18CD77FC7857F8" box="[895,909,373,399]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1398,1009,1864]" captionTargetId="figure@11.[189,1398,1009,1864]" captionTargetPageId="11" captionText="Fig. 7. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Detail of dorsal head view of adult specimen. B. Detail of dorsal head view of juvenile specimen. C. Dorsal view of posterior trunk region of specimen with two scales 17. D. Dorsal view of posterior trunk region of specimen with three scales 17." httpUri="https://zenodo.org/record/2626868/files/figure.png" pageId="11" pageNumber="12">7</figureCitation>
CD, 8A). The scales are located close to one another, but on the head, neck and anterior and central trunk part they do not overlap. The edges of the scales overlap only on the furcal base and furcal appendages (
<figureCitation id="99BD2A2F731045631C6ACDB9FBAB57A2" box="[1037,1118,443,469]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731045631C09CDB9FB6357A2" box="[1134,1174,443,469]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,1005,1860]" captionTargetId="figure@12.[189,1398,1005,1860]" captionTargetPageId="12" captionText="Fig. 8. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Dorsal view of the posterior trunk region, furca base and furcal appendage scales. B. Ventral view of the posterior trunk region, furca base and furcal appendage scales." httpUri="https://zenodo.org/record/2626870/files/figure.png" pageId="11" pageNumber="12">8A</figureCitation>
). The scales show morphological diversity throughout the particular body regions in terms of shape and size (
<figureCitation id="99BD2A2F731045631CBDCDDCFAC1578E" box="[1242,1332,478,505]" captionStart="Fig" captionStartId="2.[189,232,1271,1297]" captionTargetBox="[189,1398,780,1232]" captionTargetId="figure@2.[189,1398,780,1232]" captionTargetPageId="2" captionText="Fig. 1. Study area. A. The inside of Jubilee Greenhouse at the Botanical Garden in Kraków. B. Sampling site." httpUri="https://zenodo.org/record/2626856/files/figure.png" pageId="11" pageNumber="12">Fig. A1</figureCitation>
). The head anteriormost scales are situated near the posterior edge of the cephalion. These scales are oval with shallow posterior notches (scale 1;
<figureCitation id="99BD2A2F731045631A36CE27FD585448" box="[593,685,549,575]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3A</figureCitation>
). The remaining head dorsal scales (scale 2;
<figureCitation id="99BD2A2F731045631CCCCE27FAF75448" box="[1195,1282,549,575]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3B</figureCitation>
) are more elongated and have shallower posterior notches. The head dorsolateral scales (scale 3;
<figureCitation id="99BD2A2F731045631CD5CE4AFAFE5415" box="[1202,1291,584,610]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
) become shorter than the dorsal scales and have deeper posterior notches. The head lateral and ventrolateral scales (scale 4;
<figureCitation id="99BD2A2F731045631910CE8CFE2754DE" box="[375,466,654,681]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3D</figureCitation>
) become more semi-rounded, shorter and wider than the dorsolateral ones and possess deep posterior notches, whereas the ventral scales are smaller, wide and have more pointed anterior edges (scale 5;
<figureCitation id="99BD2A2F7310456319BCCED7FDC25498" box="[475,567,725,751]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3E</figureCitation>
). On the neck, the scales are shorter and wider than the scales on the head area (scales 69;
<figureCitation id="99BD2A2F7310456319B4CEFAFDEC5565" box="[467,537,760,786]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
FI) and their size gradually increases towards the trunk region (scales 1013;
<figureCitation id="99BD2A2F731045631970CF19FE9E5542" box="[279,363,795,821]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3J</figureCitation>
). The neck dorsal scales (scales 6 and 10;
<figureCitation id="99BD2A2F731045631B0FCF19FC2D5541" box="[872,984,795,822]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3F, J</figureCitation>
) are egg-shaped and possess weak posterior notches. The neck dorsolateral scales (scales 7 and 11;
<figureCitation id="99BD2A2F731045631BA4CF3CFBB0552E" box="[963,1093,830,857]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3G, K</figureCitation>
) become gradually larger than the dorsal scales and have deeper posterior notches. The neck lateral and ventrolateral scales (scales 8 and 12;
<figureCitation id="99BD2A2F731045631953CF87FE4455E8" box="[308,433,901,927]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">Fig. 3H, L</figureCitation>
) become more semi-rounded and deeply posteriorly notched. They are wider and gradually larger than the dorsolateral ones, whereas the ventral scales (scales 9 and 13;
<figureCitation id="99BD2A2F731045631CAFCFAAFAB655B4" box="[1224,1347,936,963]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="11" pageNumber="12">
Fig.
<date id="7538106A731045631C98CFAAFAEC55B5" box="[1279,1305,936,962]" pageId="11" pageNumber="12">3I</date>
, M
</figureCitation>
) are smaller, wide and have more pointed anterior edges (
<figureCitation id="99BD2A2F731745641B78CD09FC8E5752" box="[799,891,267,293]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="12" pageNumber="13">Fig. 2C</figureCitation>
). The trunk dorsal scales (scale 14;
<figureCitation id="99BD2A2F731745641D73CD09FA9B5752" box="[1300,1390,267,293]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3N</figureCitation>
) from the anterior to posterior trunk part are egg-shaped, but with a more tapered anterior edge than the neck scales. These scales have weak posterior notches. The trunk dorsolateral scales (scale 15;
<figureCitation id="99BD2A2F7317456418DACD77FEEC57F8" box="[189,281,373,399]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3O</figureCitation>
) become gradually larger than the dorsal scales and have deeper posterior notches. The trunk lateral and ventrolateral scales (scale 16;
<figureCitation id="99BD2A2F731745641ACCCD9AFCF757C5" box="[683,770,408,434]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3P</figureCitation>
) become more oval, gradually larger and wider than the dorsolateral ones, have a more pointed anterior edge and possess deep posterior notches. The trunk ventral scales (scale 17;
<figureCitation id="99BD2A2F7317456419B8CDDCFDCF578E" box="[479,570,478,505]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3Q</figureCitation>
) are smaller, shorter and wider, have more pointed anterior edges and a very deep posterior notch (
<figureCitation id="99BD2A2F731745641A77CE00FD87546B" box="[528,626,514,540]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="12" pageNumber="13">Figs 2A</figureCitation>
,
<figureCitation id="99BD2A2F731745641A18CE00FD57546A" box="[639,674,514,541]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="12" pageNumber="13">6F</figureCitation>
). The scales in the ventral longitudinal row located closest to the ciliary band are distinctly smaller than the other scales and their anterior edge is oriented towards the bands at an angle of ca 40° (
<figureCitation id="99BD2A2F731745641A62CE4AFD915415" box="[517,612,584,610]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="12" pageNumber="13">Figs 2C</figureCitation>
,
<figureCitation id="99BD2A2F731745641A17CE4AFD0D5414" box="[624,760,584,611]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="12" pageNumber="13">6B, D, F, H</figureCitation>
). Dorsally on the trunk posterior region (at U73U77), two or three slightly larger scales with deeper posterior notches and much longer and stronger spines are present (scale 18;
<figureCitation id="99BD2A2F7317456419F3CE8CFE0454DE" box="[404,497,654,681]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3R</figureCitation>
). Near the scales with longer and stronger spines and posteriorly to them (at U75U80), smaller, trifle pear-shaped scales are located with weak posterior notches, straight keels and without spines (scale 19;
<figureCitation id="99BD2A2F73174564198ACED7FDB05498" box="[493,581,725,751]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3S</figureCitation>
). Subsequently, on U80U85, dorsally on the posterior trunk region, 8 scales, that are triangle-shaped with strongly rounded edges, with clear posterior notches and a straight keel and spine, are situated (scale 20;
<figureCitation id="99BD2A2F731745641A18CF19FD2C5542" box="[639,729,795,821]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3T</figureCitation>
). On the dorsal posterior trunk region, one pair of small, semirounded scales with a straight keel and a short, straight spine is located in the notch of a pair of double keeled scales with trunk sensory bristles (at U82U83) (scale 21;
<figureCitation id="99BD2A2F731745641C68CF60FB99550B" box="[1039,1132,866,892]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="12" pageNumber="13">Fig. 3U</figureCitation>
). Dorsolaterally on the posterior trunk region and furcal base, two pairs of small semi-triangular scales with strongly rounded edges and shallow posterior notches, straight keels and straight rudimental spines are present (at U84 U87) (scale 22;
<figureCitation id="99BD2A2F731645651919CD09FE285752" box="[382,477,267,293]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">
Fig.
<date id="7538106A7316456519D2CD09FE285752" box="[437,477,267,293]" pageId="13" pageNumber="14">3V</date>
</figureCitation>
). Laterally on the posterior trunk region and on the furcal base, at U82U88, two pairs of narrow, elongated pear-shaped scales with very weak posterior notches are present. These scales have long straight keels but are not spined (scale 23;
<figureCitation id="99BD2A2F731645651BE0CD50FC18571B" box="[903,1005,338,364]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">Fig. 3W</figureCitation>
). Dorsally on the furcal base and furcal appendages at U85U88, three semi-rounded scales with weak posterior notches, straight keels and short spines are present (scale 24;
<figureCitation id="99BD2A2F731645651AE8CD9AFD1B57C5" box="[655,750,408,434]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">
Fig.
<date id="7538106A731645651AA0CD9AFD1B57C5" box="[711,750,408,434]" pageId="13" pageNumber="14">3X</date>
</figureCitation>
). Laterally to them (to scales 24), on the dorsal furcal base, one pair of large (5.49.1 ×3.25.3 μm), oval scales with a shallow posterior notch, straight keel and a short, straight spine is situated (at U84U87) (scale 25;
<figureCitation id="99BD2A2F731645651BD5CDDCFBE7578E" box="[946,1042,478,505]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">Fig. 3Y</figureCitation>
). Dorsolaterally on the furcal appendages at U87U89, one pair of large, rounded scales with prominent posterior notches is present (scale 26;
<figureCitation id="99BD2A2F73164565195ACE27FE6F5448" box="[317,410,549,575]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">Fig. 3Z</figureCitation>
). These scales have straight keels and long, rigid spines. Laterally to this pair of scales (to scales 26) at U88U89, two pairs of small (2.66.1×1.84.1 μm), oval-shaped scales with a shallow indentation and straight keels and spines are present (scale 27;
<figureCitation id="99BD2A2F731645651B9FCE69FBCF54F2" box="[1016,1082,619,645]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
AA). On the lateral surface of the furcal appendage, two pairs of scales with parafurcal spines are present. The first pair is located at U88U89 and the second one, i.e. the last pair of furcal lateral scales, is located at U90U91. Both pairs are similar in shape to the posterior trunk scales (
<figureCitation id="99BD2A2F731645651B27CED7FC785498" box="[832,909,725,751]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="13" pageNumber="14">Figs 2</figureCitation>
,
<figureCitation id="99BD2A2F731645651BFDCED7FC345487" box="[922,961,725,752]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="13" pageNumber="14">6G</figureCitation>
). Ventrolaterally and ventrally on the furcal appendages and furcal base, two pairs of small (2.25.3 × 1.42.8 μm) oval scales without any posterior notches are present (at U85U88). These scales have straight keels and straight rudimentary spines (scale 28;
<figureCitation id="99BD2A2F7316456519E1CF3CFE3F552E" box="[390,458,830,857]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
BB).
</paragraph>
<caption id="55F966227313456018DACB8BFB80519C" httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1929,1955]" targetBox="[197,1390,1228,1889]" targetPageId="8">
<paragraph id="013936AA7313456018DACB8BFB80519C" blockId="8.[189,1398,1929,2027]" pageId="8" pageNumber="9">
<emphasis id="33F2EAB87313456018DACB8BFEFE51D3" bold="true" box="[189,267,1929,1956]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
<taxonomicName id="C6864D29731345601974CB88FD3151D3" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[275,708,1930,1956]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="8" pageNumber="9" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731345601974CB88FE5251D3" box="[275,423,1930,1956]" italics="true" pageId="8" pageNumber="9">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87313456019DECB88FDB851D3" box="[441,589,1930,1956]" italics="true" pageId="8" pageNumber="9">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731345601A38CB89FD3151D3" box="[607,708,1931,1956]" italics="true" pageId="8" pageNumber="9">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731345601AABCB88FCD251D3" box="[716,807,1930,1956]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
Schematic drawings of the spine types.
<emphasis id="33F2EAB8731345601D63CB8BFAEC51D4" bold="true" box="[1284,1305,1929,1955]" pageId="8" pageNumber="9">A</emphasis>
. Dorsal and dorsolateral spine type.
<emphasis id="33F2EAB8731345601A6CCBAFFDEA51B0" bold="true" box="[523,543,1965,1991]" pageId="8" pageNumber="9">B</emphasis>
. Lateral and ventrolateral spine type.
<emphasis id="33F2EAB8731345601B85CBAFFC0251B0" bold="true" box="[994,1015,1965,1991]" pageId="8" pageNumber="9">C</emphasis>
. Ventral spine type.
<emphasis id="33F2EAB8731345601C8BCBAFFAF451B0" bold="true" box="[1260,1281,1965,1991]" pageId="8" pageNumber="9">D</emphasis>
. Scale 18 spine type.
<emphasis id="33F2EAB8731345601924CBD2FEA2519D" bold="true" box="[323,343,2000,2026]" pageId="8" pageNumber="9">E</emphasis>
. Dorsolateral furcal appendage spine type.
<emphasis id="33F2EAB8731345601B32CBD2FC92519D" bold="true" box="[853,871,2000,2026]" pageId="8" pageNumber="9">F</emphasis>
. Parafurcal spine type.
</paragraph>
</caption>
<caption id="55F966227312456118DACD09FD66571C" pageId="9" pageNumber="10">
<paragraph id="013936AA7312456118DACD09FD66571C" blockId="9.[189,1399,266,364]" pageId="9" pageNumber="10">
<emphasis id="33F2EAB87312456118DACD09FEEA5753" bold="true" box="[189,287,266,293]" pageId="9" pageNumber="10">Table 2.</emphasis>
Main morphometric characters of
<taxonomicName id="C6864D29731245611AD3CD09FB965753" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[692,1123,267,293]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="9" pageNumber="10" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731245611AD3CD09FCBD5752" box="[692,840,267,293]" italics="true" pageId="9" pageNumber="10">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8731245611B3ECD09FC185752" box="[857,1005,267,293]" italics="true" pageId="9" pageNumber="10">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731245611B99CD09FB965753" box="[1022,1123,267,292]" italics="true" pageId="9" pageNumber="10">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731245611C0DCD09FB315752" box="[1130,1220,267,293]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
N = number of specimens analysed; Range = the smallest and the largest structure found among all specimens measured. All measurements in micrometers (μm).
</paragraph>
</caption>
<paragraph id="013936AA7312456119F5CD91FAEF55A4" pageId="9" pageNumber="10">
<table id="7386C40A7312BA971884CD91FA9255A3" box="[227,1383,403,980]" colsContinueIn="30.[206,1388,437,1299]" gridcols="4" gridrows="14" pageId="9" pageNumber="10">
<tr id="BFB634E87312BA971884CD91FA9257DC" box="[227,1383,403,427]" gridrow="0" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CD91FD3557DC" box="[227,704,403,427]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="33F2EAB87312456119F5CD91FDE457DC" bold="true" box="[402,529,403,427]" pageId="9" pageNumber="10">Characters</emphasis>
</th>
<th id="FC675D947312BA971A9ACD91FCED57DC" box="[765,792,403,427]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="33F2EAB8731245611B66CD91FCE157DC" bold="true" box="[769,788,403,427]" pageId="9" pageNumber="10">N</emphasis>
</th>
<th id="FC675D947312BA971B5BCD91FBD857DC" box="[828,1069,403,427]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="33F2EAB8731245611BE6CD91FC1C57DC" bold="true" box="[897,1001,403,427]" pageId="9" pageNumber="10">Holotype</emphasis>
</th>
<th id="FC675D947312BA971C2FCD91FA9257DC" box="[1096,1383,403,427]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">
<emphasis id="33F2EAB8731245611C2FCD91FA9257DC" bold="true" box="[1096,1383,403,427]" pageId="9" pageNumber="10">Range of adult paratypes</emphasis>
</th>
</tr>
<tr id="BFB634E87312BA971884CDC1FA9257AC" box="[227,1383,451,475]" gridrow="1" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CDC1FD3557AC" box="[227,704,451,475]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Body length</th>
<td id="FC675D947312BA971A9ACDC1FCED57AC" box="[765,792,451,475]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">23</td>
<td id="FC675D947312BA971B5BCDC1FBD857AC" box="[828,1069,451,475]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">193.18</td>
<td id="FC675D947312BA971C2FCDC1FA9257AC" box="[1096,1383,451,475]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">174.64194.66</td>
</tr>
<tr id="BFB634E87312BA971884CDE9FA925474" box="[227,1383,491,515]" gridrow="2" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CDE9FD355474" box="[227,704,491,515]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">Furca length</th>
<td id="FC675D947312BA971A9ACDE9FCED5474" box="[765,792,491,515]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCDE9FBD85474" box="[828,1069,491,515]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">33.17</td>
<td id="FC675D947312BA971C2FCDE9FA925474" box="[1096,1383,491,515]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">31.1434.45</td>
</tr>
<tr id="BFB634E87312BA971884CE11FA92545C" box="[227,1383,531,555]" gridrow="3" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CE11FD35545C" box="[227,704,531,555]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">Adhesive tube length</th>
<td id="FC675D947312BA971A9ACE11FCED545C" box="[765,792,531,555]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCE11FBD8545C" box="[828,1069,531,555]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">18.97</td>
<td id="FC675D947312BA971C2FCE11FA92545C" box="[1096,1383,531,555]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">16.3919.20</td>
</tr>
<tr id="BFB634E87312BA971884CE38FA925424" box="[227,1383,570,595]" gridrow="4" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CE38FD355424" box="[227,704,570,595]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">Number of scales in central longitudinal row</th>
<td id="FC675D947312BA971A9ACE38FCED5424" box="[765,792,570,595]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">25</td>
<td id="FC675D947312BA971B5BCE38FBD85424" box="[828,1069,570,595]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">27</td>
<td id="FC675D947312BA971C2FCE38FA925424" box="[1096,1383,570,595]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">2527</td>
</tr>
<tr id="BFB634E87312BA971884CE60FA9254E2" box="[227,1383,610,661]" gridrow="5" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CE60FD3554E2" box="[227,704,610,661]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">Total noumber of longitudinal alternating rows of scales</th>
<td id="FC675D947312BA971A9ACE60FCED54E2" box="[765,792,610,661]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">25</td>
<td id="FC675D947312BA971B5BCE60FBD854E2" box="[828,1069,610,661]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">25</td>
<td id="FC675D947312BA971C2FCE60FA9254E2" box="[1096,1383,610,661]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">2325</td>
</tr>
<tr id="BFB634E87312BA971884CEA7FA9254CA" box="[227,1383,677,701]" gridrow="6" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CEA7FD3554CA" box="[227,704,677,701]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">Head scales length × width</th>
<td id="FC675D947312BA971A9ACEA7FCED54CA" box="[765,792,677,701]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCEA7FBD854CA" box="[828,1069,677,701]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">1.944.94 × 1.914.23</td>
<td id="FC675D947312BA971C2FCEA7FA9254CA" box="[1096,1383,677,701]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">1.585.19 × 1.604.91</td>
</tr>
<tr id="BFB634E87312BA971884CECEFA925492" box="[227,1383,716,741]" gridrow="7" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CECEFD355492" box="[227,704,716,741]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">Neck scales length × width</th>
<td id="FC675D947312BA971A9ACECEFCED5492" box="[765,792,716,741]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCECEFBD85492" box="[828,1069,716,741]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">2.556.29 × 3.025.71</td>
<td id="FC675D947312BA971C2FCECEFA925492" box="[1096,1383,716,741]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">2.087.09 × 2.466.44</td>
</tr>
<tr id="BFB634E87312BA971884CEF6FA92557A" box="[227,1383,756,781]" gridrow="8" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CEF6FD35557A" box="[227,704,756,781]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">Trunk scales length × width</th>
<td id="FC675D947312BA971A9ACEF6FCED557A" box="[765,792,756,781]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCEF6FBD8557A" box="[828,1069,756,781]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">3.578.78 × 2.795.97</td>
<td id="FC675D947312BA971C2FCEF6FA92557A" box="[1096,1383,756,781]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">2.849.38 × 2.227.39</td>
</tr>
<tr id="BFB634E87312BA971884CF1EFA925543" box="[227,1383,796,820]" gridrow="9" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CF1EFD355543" box="[227,704,796,820]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">Head spines length</th>
<td id="FC675D947312BA971A9ACF1EFCED5543" box="[765,792,796,820]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCF1EFBD85543" box="[828,1069,796,820]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">3.8824.11</td>
<td id="FC675D947312BA971C2FCF1EFA925543" box="[1096,1383,796,820]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">3.3126.39</td>
</tr>
<tr id="BFB634E87312BA971884CF46FA92552B" box="[227,1383,836,860]" gridrow="10" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CF46FD35552B" box="[227,704,836,860]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">Neck spines length</th>
<td id="FC675D947312BA971A9ACF46FCED552B" box="[765,792,836,860]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCF46FBD8552B" box="[828,1069,836,860]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">6.3727.41</td>
<td id="FC675D947312BA971C2FCF46FA92552B" box="[1096,1383,836,860]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">5.0331.86</td>
</tr>
<tr id="BFB634E87312BA971884CF6EFA9255F3" box="[227,1383,876,900]" gridrow="11" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CF6EFD3555F3" box="[227,704,876,900]" gridcol="0" gridrow="11" pageId="9" pageNumber="10">Trunk spines length</th>
<td id="FC675D947312BA971A9ACF6EFCED55F3" box="[765,792,876,900]" gridcol="1" gridrow="11" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCF6EFBD855F3" box="[828,1069,876,900]" gridcol="2" gridrow="11" pageId="9" pageNumber="10">10.0931.06</td>
<td id="FC675D947312BA971C2FCF6EFA9255F3" box="[1096,1383,876,900]" gridcol="3" gridrow="11" pageId="9" pageNumber="10">7.8434.26</td>
</tr>
<tr id="BFB634E87312BA971884CF96FA9255DB" box="[227,1383,916,940]" gridrow="12" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CF96FD3555DB" box="[227,704,916,940]" gridcol="0" gridrow="12" pageId="9" pageNumber="10">Mouth ring diameter</th>
<td id="FC675D947312BA971A9ACF96FCED55DB" box="[765,792,916,940]" gridcol="1" gridrow="12" pageId="9" pageNumber="10">22</td>
<td id="FC675D947312BA971B5BCF96FBD855DB" box="[828,1069,916,940]" gridcol="2" gridrow="12" pageId="9" pageNumber="10">8.43</td>
<td id="FC675D947312BA971C2FCF96FA9255DB" box="[1096,1383,916,940]" gridcol="3" gridrow="12" pageId="9" pageNumber="10">7.589.47</td>
</tr>
<tr id="BFB634E87312BA971884CFB9FA9255A3" box="[227,1383,955,980]" gridrow="13" pageId="9" pageNumber="10">
<th id="FC675D947312BA971884CFB9FD3555A3" box="[227,704,955,980]" gridcol="0" gridrow="13" pageId="9" pageNumber="10">Pharynx length</th>
<td id="FC675D947312BA971A9ACFB9FCED55A3" box="[765,792,955,980]" gridcol="1" gridrow="13" pageId="9" pageNumber="10">23</td>
<td id="FC675D947312BA971B5BCFB9FBD855A3" box="[828,1069,955,980]" gridcol="2" gridrow="13" pageId="9" pageNumber="10">56.84</td>
<td id="FC675D947312BA971C2FCFB9FA9255A3" box="[1096,1383,955,980]" gridcol="3" gridrow="13" pageId="9" pageNumber="10">51.0259.64</td>
</tr>
</table>
</paragraph>
<caption id="55F966227312456118DACB9AFBE051A1" httpUri="https://zenodo.org/record/2626864/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1944,1970]" targetBox="[189,1398,1052,1906]" targetPageId="9">
<paragraph id="013936AA7312456118DACB9AFBE051A1" blockId="9.[189,1399,1944,2006]" pageId="9" pageNumber="10">
<emphasis id="33F2EAB87312456118DACB9AFEE651C5" bold="true" box="[189,275,1944,1970]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
<taxonomicName id="C6864D29731245611944CB9AFD1151C5" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[291,740,1944,1970]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="9" pageNumber="10" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731245611944CB9AFE4251C5" box="[291,439,1944,1970]" italics="true" pageId="9" pageNumber="10">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87312456119B6CB9AFD9051C5" box="[465,613,1944,1970]" italics="true" pageId="9" pageNumber="10">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731245611A18CB9BFD1151C5" box="[639,740,1945,1970]" italics="true" pageId="9" pageNumber="10">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731245611A93CB9AFCA251C5" box="[756,855,1944,1970]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
Habitus. Phase contrast microphotographs.
<emphasis id="33F2EAB87312456118DACBB9FF2751A2" bold="true" box="[189,210,1979,2005]" pageId="9" pageNumber="10">A</emphasis>
. Juvenile specimen.
<emphasis id="33F2EAB87312456119A0CBB9FE2E51A2" bold="true" box="[455,475,1979,2005]" pageId="9" pageNumber="10">B</emphasis>
. Adult specimen.
<emphasis id="33F2EAB8731245611AD7CBB9FD3051A2" bold="true" box="[688,709,1979,2005]" pageId="9" pageNumber="10">C</emphasis>
. Specimen with mature egg.
</paragraph>
</caption>
<caption id="55F966227311456218DACB64FE19519E" httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1894,1920]" targetBox="[231,1356,266,1858]" targetPageId="10">
<paragraph id="013936AA7311456218DACB64FE19519E" blockId="10.[189,1399,1894,2025]" pageId="10" pageNumber="11">
<emphasis id="33F2EAB87311456218DACB64FEF851F7" bold="true" box="[189,269,1894,1920]" pageId="10" pageNumber="11">Fig. 6.</emphasis>
<taxonomicName id="C6864D29731145621970CB64FD3851F7" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[279,717,1894,1920]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="10" pageNumber="11" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731145621970CB64FE5E51F7" box="[279,427,1894,1920]" italics="true" pageId="10" pageNumber="11">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87311456219A7CB64FDA151F7" box="[448,596,1894,1920]" italics="true" pageId="10" pageNumber="11">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731145621A0FCB65FD3851F7" box="[616,717,1895,1920]" italics="true" pageId="10" pageNumber="11">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731145621AB0CB64FCC051F7" box="[727,821,1894,1920]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
Bright field microphotographs.
<emphasis id="33F2EAB8731145621CDECB64FB3B51F7" bold="true" box="[1209,1230,1894,1920]" pageId="10" pageNumber="11">A</emphasis>
. Head, dorsal view.
<emphasis id="33F2EAB873114562196FCB8BFEE951D4" bold="true" box="[264,284,1929,1955]" pageId="10" pageNumber="11">B</emphasis>
. Head, ventral view.
<emphasis id="33F2EAB8731145621A4FCB8BFDC851D4" bold="true" box="[552,573,1929,1955]" pageId="10" pageNumber="11">C</emphasis>
. Neck, dorsal view.
<emphasis id="33F2EAB8731145621B26CB8BFCA351D4" bold="true" box="[833,854,1929,1955]" pageId="10" pageNumber="11">D</emphasis>
. Neck, ventral view.
<emphasis id="33F2EAB8731145621C04CB8BFB8251D4" bold="true" box="[1123,1143,1929,1955]" pageId="10" pageNumber="11">E</emphasis>
. Trunk, dorsal view.
<emphasis id="33F2EAB87311456218DACBAEFF3A51B1" bold="true" box="[189,207,1964,1990]" pageId="10" pageNumber="11">F</emphasis>
. Trunk, ventral view.
<emphasis id="33F2EAB87311456219BACBAEFE0151B1" bold="true" box="[477,500,1964,1990]" pageId="10" pageNumber="11">G</emphasis>
. Furca base and furcal appendages, dorsal view.
<emphasis id="33F2EAB8731145621C2ECBAEFB9551B1" bold="true" box="[1097,1120,1964,1990]" pageId="10" pageNumber="11">H</emphasis>
. Furca base and furcal appendages, ventral view.
</paragraph>
</caption>
<caption id="55F966227310456318DACB64FDD7519D" httpUri="https://zenodo.org/record/2626868/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1894,1920]" targetBox="[189,1398,1009,1864]" targetPageId="11">
<paragraph id="013936AA7310456318DACB64FDD7519D" blockId="11.[189,1399,1894,2026]" pageId="11" pageNumber="12">
<emphasis id="33F2EAB87310456318DACB64FEFF51F7" bold="true" box="[189,266,1894,1920]" pageId="11" pageNumber="12">Fig. 7.</emphasis>
<taxonomicName id="C6864D29731045631975CB64FD3451F7" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[274,705,1894,1921]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="11" pageNumber="12" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731045631975CB64FE5351F7" box="[274,422,1894,1920]" italics="true" pageId="11" pageNumber="12">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87310456319D0CB64FDBE51F7" box="[439,587,1894,1920]" italics="true" pageId="11" pageNumber="12">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731045631A3BCB65FD3451F7" box="[604,705,1895,1920]" italics="true" pageId="11" pageNumber="12">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731045631AAECB65FCD651F6" box="[713,803,1895,1921]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
Bright field microphotographs.
<emphasis id="33F2EAB8731045631CFBCB64FB4451F7" bold="true" box="[1180,1201,1894,1920]" pageId="11" pageNumber="12">A</emphasis>
. Detail of dorsal head view of adult specimen.
<emphasis id="33F2EAB8731045631A40CB8BFDCE51D4" bold="true" box="[551,571,1929,1955]" pageId="11" pageNumber="12">B</emphasis>
. Detail of dorsal head view of juvenile specimen.
<emphasis id="33F2EAB8731045631CFACB8BFB4751D4" bold="true" box="[1181,1202,1929,1955]" pageId="11" pageNumber="12">C</emphasis>
. Dorsal view of posterior trunk region of specimen with two scales 17.
<emphasis id="33F2EAB8731045631B04CBAFFC8D51B0" bold="true" box="[867,888,1965,1991]" pageId="11" pageNumber="12">D</emphasis>
. Dorsal view of posterior trunk region of specimen with three scales 17.
</paragraph>
</caption>
<caption id="55F966227317456418DACB62FD1351B7" httpUri="https://zenodo.org/record/2626870/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1888,1914]" targetBox="[189,1398,1005,1860]" targetPageId="12">
<paragraph id="013936AA7317456418DACB62FD1351B7" blockId="12.[189,1399,1888,1984]" pageId="12" pageNumber="13">
<emphasis id="33F2EAB87317456418DACB62FEFE510D" bold="true" box="[189,267,1888,1914]" pageId="12" pageNumber="13">Fig. 8.</emphasis>
<taxonomicName id="C6864D29731745641974CB62FD31510D" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[275,708,1888,1915]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="12" pageNumber="13" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731745641974CB62FE52510D" box="[275,423,1888,1914]" italics="true" pageId="12" pageNumber="13">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87317456419DECB62FDB8510D" box="[441,589,1888,1914]" italics="true" pageId="12" pageNumber="13">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731745641A38CB63FD31510D" box="[607,708,1889,1914]" italics="true" pageId="12" pageNumber="13">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731745641AABCB63FCD2510C" box="[716,807,1889,1915]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
Bright field microphotographs.
<emphasis id="33F2EAB8731745641CC5CB62FB42510D" bold="true" box="[1186,1207,1888,1914]" pageId="12" pageNumber="13">A</emphasis>
. Dorsal view of the posterior trunk region, furca base and furcal appendage scales.
<emphasis id="33F2EAB8731745641BA2CB86FC2C51E9" bold="true" box="[965,985,1924,1950]" pageId="12" pageNumber="13">B</emphasis>
. Ventral view of the posterior trunk region, furca base and furcal appendage scales.
</paragraph>
</caption>
<paragraph id="013936AA7316456618DACF82FDF1550B" blockId="13.[189,1398,896,993]" lastBlockId="14.[189,1399,267,892]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
SPINES. All spines emerging from the posterior scale region have no lateral denticles. The dorsal, dorsolateral (
<figureCitation id="99BD2A2F731645651933CFA1FE5B55CA" box="[340,430,931,957]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="13" pageNumber="14">Fig.4A</figureCitation>
),lateral and ventrolateral (
<figureCitation id="99BD2A2F731645651ABDCFA1FCC555CA" box="[730,816,931,957]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="13" pageNumber="14">Fig. 4B</figureCitation>
)spines are basally bent and thick,and subsequently strongly and gradually taper towards the slightly bent hair-like ends. The spines arising from the lateral, ventrolateral and ventral scales are much more strongly bent than the dorsal and dorsolateral spines. The ventral scale spines are hair-like along their entire length (
<figureCitation id="99BD2A2F731545661B08CD2CFC38573E" box="[879,973,302,329]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="14" pageNumber="15">Fig. 4C</figureCitation>
). The length of the spines gradually increases from the head towards the widest body region (from 3.924.1 to 10.131.1 μm). Then the length of the dorsal spines gradually shortens to the posterior trunk region. Moreover, the length of the spines gradually increases from the dorsal surface, along the dorsolateral, lateral, ventrolateral and ventral surfaces towards the ciliary bands (head: from 1.67.4 to 13.326.4 μm; neck: from 5.010.7 to 15.931.9 μm; trunk: from 7.816.5 to 19.634.3 μm) (
<tableCitation id="4C040311731545661B22CDDCFC41578E" box="[837,948,478,505]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Sampling sites with specified aquatic plant species." pageId="14" pageNumber="15">Table A1</tableCitation>
). On the dorsal posterior trunk region, two or three spines of different
<typeStatus id="DE3D8808731545661A4ACE00FD9E546B" box="[557,619,514,540]" pageId="14" pageNumber="15">types</typeStatus>
are present (arising from scales 18). These spines are much longer (16.620.0 μm) and stronger than the other dorsal spines and taper to blunt ends (
<figureCitation id="99BD2A2F731545661C1BCE27FB2B5448" box="[1148,1246,549,575]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="14" pageNumber="15">Figs 2A</figureCitation>
,
<figureCitation id="99BD2A2F731545661C8CCE27FB0C5437" box="[1259,1273,549,576]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="14" pageNumber="15">6</figureCitation>
CD). The spines arising from the dorsal and dorsolateral posterior trunk and furcal base scales (namely from scales 20, 22, 24, 27) are thick and straight, tapering to blunt ends (
<figureCitation id="99BD2A2F731545661BF5CE69FC0754F2" box="[914,1010,619,645]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="14" pageNumber="15">Fig. 4D</figureCitation>
). The posteriormost lateral trunk spines (16.125.1 μm) do not vary in length from the other lateral spines in the posterior trunk region. From the dorsolateral furcal appendage scales (scale 26) arise very long (14.518.4 μm), thick, rigid, basally bent and spike-like spines that extend to the inner furcal indentation (
<figureCitation id="99BD2A2F731545661C01CED7FB3E5498" box="[1126,1227,725,751]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="14" pageNumber="15">Figs 2A</figureCitation>
,
<figureCitation id="99BD2A2F731545661CBDCED7FB085498" box="[1242,1277,725,751]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="14" pageNumber="15">4E</figureCitation>
,
<figureCitation id="99BD2A2F731545661D6BCED7FAC75498" box="[1292,1330,725,751]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="14" pageNumber="15">6G</figureCitation>
,
<figureCitation id="99BD2A2F731545661D26CED7FA9C5498" box="[1345,1385,725,751]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,1005,1860]" captionTargetId="figure@12.[189,1398,1005,1860]" captionTargetPageId="12" captionText="Fig. 8. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Dorsal view of the posterior trunk region, furca base and furcal appendage scales. B. Ventral view of the posterior trunk region, furca base and furcal appendage scales." httpUri="https://zenodo.org/record/2626870/files/figure.png" pageId="14" pageNumber="15">8A</figureCitation>
). These spines taper slightly to pointed ends. Two pairs of parafurcal spines, emerging from the lateral scales of the furcal appendages (at U88U89 and at U90U91), are thicker and longer (11.313.5 μm) than the other lateral body spines (
<figureCitation id="99BD2A2F731545661A2CCF3CFD51552E" box="[587,676,830,857]" captionStart="Fig" captionStartId="8.[189,232,1929,1955]" captionTargetBox="[197,1390,1228,1889]" captionTargetId="figure@8.[189,1398,1218,1901]" captionTargetPageId="8" captionText="Fig. 4. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the spine types. A. Dorsal and dorsolateral spine type. B. Lateral and ventrolateral spine type. C. Ventral spine type. D. Scale 18 spine type. E. Dorsolateral furcal appendage spine type. F. Parafurcal spine type." httpUri="https://zenodo.org/record/2626862/files/figure.png" pageId="14" pageNumber="15">Fig. 4F</figureCitation>
). These spines are basally bent and taper slightly towards their pointed ends (
<figureCitation id="99BD2A2F731545661905CF60FE36550B" box="[354,451,866,892]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="14" pageNumber="15">Figs 6G</figureCitation>
,
<figureCitation id="99BD2A2F7315456619B7CF60FE02550B" box="[464,503,866,892]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,1005,1860]" captionTargetId="figure@12.[189,1398,1005,1860]" captionTargetPageId="12" captionText="Fig. 8. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Dorsal view of the posterior trunk region, furca base and furcal appendage scales. B. Ventral view of the posterior trunk region, furca base and furcal appendage scales." httpUri="https://zenodo.org/record/2626870/files/figure.png" pageId="14" pageNumber="15">8A</figureCitation>
).
</paragraph>
<caption id="55F966227316456518DACB8BFED0519E" httpUri="https://zenodo.org/record/2626872/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1929,1955]" targetBox="[189,1398,1041,1893]" targetPageId="13">
<paragraph id="013936AA7316456518DACB8BFED0519E" blockId="13.[189,1399,1929,2025]" pageId="13" pageNumber="14">
<emphasis id="33F2EAB87316456518DACB8BFEF851D4" bold="true" box="[189,269,1929,1955]" pageId="13" pageNumber="14">Fig. 9.</emphasis>
<taxonomicName id="C6864D29731645651970CB8BFD3951D4" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[279,716,1929,1956]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="13" pageNumber="14" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731645651970CB8BFE5E51D4" box="[279,427,1929,1955]" italics="true" pageId="13" pageNumber="14">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87316456519D8CB8BFDA651D4" box="[447,595,1929,1955]" italics="true" pageId="13" pageNumber="14">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731645651A00CB88FD3951D4" box="[615,716,1930,1955]" italics="true" pageId="13" pageNumber="14">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731645651AB2CB88FCC651D3" box="[725,819,1930,1956]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
Adult specimen. Bright field microphotographs.
<emphasis id="33F2EAB87316456518DACBAEFF2751B1" bold="true" box="[189,210,1964,1990]" pageId="13" pageNumber="14">A</emphasis>
. Detail of the mouth ring, dorsal view.
<emphasis id="33F2EAB8731645651AD4CBAEFD3251B1" bold="true" box="[691,711,1964,1990]" pageId="13" pageNumber="14">B</emphasis>
. Detail of the mouth ring, internal view.
<emphasis id="33F2EAB8731645651CDECBAEFB3B51B1" bold="true" box="[1209,1230,1964,1990]" pageId="13" pageNumber="14">C</emphasis>
. Detail of the pharynx.
</paragraph>
</caption>
<paragraph id="013936AA7315456718DACFA6FC9B57F8" blockId="14.[189,1399,931,1028]" lastBlockId="15.[189,1399,267,399]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
DORSAL SENSORY BRISTLES. This species has three pairs of dorsal sensory bristles. The first pair is located on the head, directly between the lateral edge of the cephalion and the epipleurae (at U4), and emerges from small, round papillae. The second pair of sensory bristles is located on the neck (U19) and emerges from small, rounded papillae. The third posterior pair of sensory bristles emerges from the doublekeeled scales located on the posterior trunk region at U81U83. These scales are shaped like triangles with strongly rounded edges and a deep posterior notch. The keels of these scales are slightly bent, not connected and pass into rudimentary spines (
<figureCitation id="99BD2A2F731445671AACCD77FCD857F8" box="[715,813,373,399]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="15" pageNumber="16">Figs 2A</figureCitation>
,
<figureCitation id="99BD2A2F731445671B5ECD77FC9457F8" box="[825,865,373,399]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,1005,1860]" captionTargetId="figure@12.[189,1398,1005,1860]" captionTargetPageId="12" captionText="Fig. 8. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Dorsal view of the posterior trunk region, furca base and furcal appendage scales. B. Ventral view of the posterior trunk region, furca base and furcal appendage scales." httpUri="https://zenodo.org/record/2626870/files/figure.png" pageId="15" pageNumber="16">8A</figureCitation>
).
</paragraph>
<caption id="55F966227315456618DACBAEFDF9519D" httpUri="https://zenodo.org/record/2626874/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1964,1990]" targetBox="[189,1398,1077,1929]" targetPageId="14">
<paragraph id="013936AA7315456618DACBAEFDF9519D" blockId="14.[189,1398,1964,2026]" pageId="14" pageNumber="15">
<emphasis id="33F2EAB87315456618DACBAEFED551B1" bold="true" box="[189,288,1964,1990]" pageId="14" pageNumber="15">Fig. 10.</emphasis>
<taxonomicName id="C6864D2973154566194ACBAEFD1C51B1" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[301,745,1964,1991]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="14" pageNumber="15" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB873154566194ACBAEFE3451B1" box="[301,449,1964,1990]" italics="true" pageId="14" pageNumber="15">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87315456619BECBAEFD9851B1" box="[473,621,1964,1990]" italics="true" pageId="14" pageNumber="15">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731545661AE3CBAFFD1C51B1" box="[644,745,1965,1990]" italics="true" pageId="14" pageNumber="15">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731545661A90CBAFFCAD51B0" box="[759,856,1965,1991]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
Detail of internal morphology. Bright field microphotography. X-organ.
</paragraph>
</caption>
<paragraph id="013936AA7314457818DACDB5FD7357A2" blockId="15.[189,1399,438,1028]" lastBlockId="16.[189,1399,267,470]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
VENTRAL CILIARY BANDS AND VENTRAL INTERCILIARY FIELD. Longitudinal ventral ciliary bands begin at U8 and run back to U86 (
<figureCitation id="99BD2A2F7314456719A6CDDBFDD75783" box="[449,546,473,500]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="15" pageNumber="16">Figs 2C</figureCitation>
,
<figureCitation id="99BD2A2F731445671A48CDDBFD485783" box="[559,701,473,500]" captionStart="Fig" captionStartId="10.[189,232,1894,1920]" captionTargetBox="[231,1356,266,1858]" captionTargetId="figure@10.[231,1356,266,1858]" captionTargetPageId="10" captionText="Fig. 6. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Head, dorsal view. B. Head, ventral view. C. Neck, dorsal view. D. Neck, ventral view. E. Trunk, dorsal view. F. Trunk, ventral view. G. Furca base and furcal appendages, dorsal view. H. Furca base and furcal appendages, ventral view." httpUri="https://zenodo.org/record/2626866/files/figure.png" pageId="15" pageNumber="16">6B, D, F, H</figureCitation>
). On the head, at ca U8U10, the ciliary bands are merged and lie close to the hypopleurae. The ventral interciliary field scales are distributed in single (on the pharyngeal region) and 57 (on the intestinal region) longitudinal alternating rows, with 2933 scales in the central row, including 1518 plate-like scales and 1417 one-lobed scales. The entire pharyngeal region (from U10 to U31) is covered with short, wide plates (scale 29;
<figureCitation id="99BD2A2F731445671B8ACE64FBDA54F7" box="[1005,1071,614,640]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
CC) and their size gradually increases from the anterior to the posterior end (from 1.01.7 ×1.93.5 μm to 2.65.2 ×7.111.4 μm). On the intestinal region, one-lobed scales with overlapped edges are present (from U32 to U84). On the anterior part of the intestinal region the scales are semi-rectangular in shape, have weak keels and their posterior edges are straight, without posterior notches (scale 30,
<figureCitation id="99BD2A2F731445671BC8CEF1FC07557A" box="[943,1010,755,781]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
DD). Towards the furcal base the keels become more pronounced and on the edges of the anterior scales small, horn-like perturbances begin to appear; the posterior edges become notched (scale 31;
<figureCitation id="99BD2A2F731445671BD1CF3BFC0E5523" box="[950,1019,825,852]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
EE). The last alternating row of interciliary field scales is of a different
<typeStatus id="DE3D8808731445671AEDCF5FFD4A5500" box="[650,703,861,887]" pageId="15" pageNumber="16">type</typeStatus>
, i.e., they are oval in shape without posterior notches, have straight keels and present rudimentary spines (scale 32;
<figureCitation id="99BD2A2F731445671B29CF82FC6455ED" box="[846,913,896,922]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
FF). The longitudinal row of ventral field scales closest to the ventral ciliary bands has significantly smaller and narrower scales than in the central rows. The first pair of terminal scales on the ventral interciliary field is large (14.817.1 × 6.68.1) and located at U84U87. They are oval, asymmetrical in shape, possess a straight keel and a straight, thick spine (scale 33;
<figureCitation id="99BD2A2F730B457819A4CD09FDF25752" box="[451,519,267,293]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
GG). The second pair of terminal scales on the ventral interciliary field is located on the furcal appendages (at U86U88). These scales are oval in shape, have long, straight keels and thick, curved spines reaching to the internal furcal indentation (scale 34;
<figureCitation id="99BD2A2F730B45781C31CD50FB6F571B" box="[1110,1178,338,364]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
HH). The first and second pairs slightly overlap. The third (at U87U88) and fourth (at 88U89) pairs are definitely small (2.04.8× 1.33.0 μm), egg-shaped, are located on the furcal appendages, and have straight keels and rudimentary spines (scale 35;
<figureCitation id="99BD2A2F730B45781A7ACDB9FD9457A2" box="[541,609,443,469]" captionStart="Fig" captionStartId="7.[189,232,1647,1673]" captionTargetBox="[225,1359,275,1607]" captionTargetId="figure@7.[219,1368,265,1613]" captionTargetPageId="7" captionText="Fig. 3. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings of the scales. A. Head anteriormost dorsal scale (scale 1). B. Head dorsal scale (scale 2). C. Head dorsolateral scale (scale 3). D. Head lateral and ventrolateral scale (scale 4). E. Head ventral scale (scale 5). F. Anterior neck dorsal scale (scale 6). G. Anterior neck dorsolateral scale (scale 7). H. Anterior neck lateral and ventrolateral scale (scale 8). I. Anterior neck ventral scale (scale 9). J. Posterior neck dorsal scale (scale 10). K. Posterior neck dorsolateral scale (scale 11). L. Posterior neck lateral and ventrolateral scale (scale 12). M. Posterior neck ventral scale (scale 13). N. Trunk dorsal scale (scale 14). O. Trunk dorsolateral scale (scale 15). P. Trunk lateral and ventrolateral scale (scale 16). Q. Trunk ventral scale (scale 17). R. Scale 18. S. Scale 19. T. Scale 20. U. Scale 21. V. Scale 22. W. Scale 23. X. Scale 24. Y. Scale 25. Z. Scale 26. AA. Scale 27. BB. Scale 28. CC. Scale 29. DD. Scale 30. EE. Scale 31. FF. Scale 32. GG. Scale 33. HH. Scale 34. II. Scale 35." httpUri="https://zenodo.org/record/2626860/files/figure.png" pageId="16" pageNumber="17">
Fig.
<date id="7538106A730B45781A34CDB9FD9457A2" box="[595,609,443,469]" pageId="16" pageNumber="17">3</date>
</figureCitation>
II).
</paragraph>
<caption id="55F966227314456718DACBAEFDB6519E" httpUri="https://zenodo.org/record/2626876/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1964,1990]" targetBox="[189,1398,1080,1934]" targetPageId="15">
<paragraph id="013936AA7314456718DACBAEFDB6519E" blockId="15.[189,1399,1964,2025]" pageId="15" pageNumber="16">
<emphasis id="33F2EAB87314456718DACBAEFEEC51B1" bold="true" box="[189,281,1964,1990]" pageId="15" pageNumber="16">Fig. 11.</emphasis>
<taxonomicName id="C6864D29731445671946CBAEFD2651B1" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[289,723,1964,1991]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="15" pageNumber="16" phylum="Gastrotricha" rank="subGenus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8731445671946CBAEFE4051B1" box="[289,437,1964,1990]" italics="true" pageId="15" pageNumber="16">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB87314456719AFCBAEFDA951B1" box="[456,604,1964,1990]" italics="true" pageId="15" pageNumber="16">Chaetonotus</emphasis>
)
<emphasis id="33F2EAB8731445671A09CBAFFD2651B1" box="[622,723,1965,1990]" italics="true" pageId="15" pageNumber="16">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3731445671ABBCBAFFCCD51B0" box="[732,824,1965,1991]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
Adult specimen. Bright field microphotography. Living euglenids inside intestine.
</paragraph>
</caption>
<paragraph id="013936AA730B457818DACDFFFE0D5500" blockId="16.[189,1399,509,887]" pageId="16" pageNumber="17">
INTERNAL MORPHOLOGY. The pharynx (from U2 to U31) is narrow and has marked anterior and posterior dilatations. The posterior dilatation is stronger than the anterior one (
<figureCitation id="99BD2A2F730B45781B85CE22FBCA544D" box="[994,1087,544,570]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[189,1398,1041,1893]" captionTargetId="figure@13.[189,1398,1041,1893]" captionTargetPageId="13" captionText="Fig. 9. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotographs. A. Detail of the mouth ring, dorsal view. B. Detail of the mouth ring, internal view. C. Detail of the pharynx." httpUri="https://zenodo.org/record/2626872/files/figure.png" pageId="16" pageNumber="17">Fig. 9C</figureCitation>
). In the anterior dilatation, weak cuticular reinforcements are located in the form of two straight, connected rods (at U4). The pharynx is connected through the pharyngealintestinal junction by the straight intestine (running from U32 to U86). The pharyngealintestinal junction is clearly demarcated, short and wide (U32) (
<figureCitation id="99BD2A2F730B45781D6ACE8BFA9C54D3" box="[1293,1385,649,676]" captionStart="Fig" captionStartId="6.[179,222,531,559]" captionTargetBox="[236,1862,635,1741]" captionTargetId="figure@6.[231,1880,625,1751]" captionTargetPageId="6" captionText="Fig. 2. Chaetonotus (Chaetonotus) invitatus sp. nov. Schematic drawings. A. Dorsal body view. B. View of internal morphology. C. Ventral body view. Light grey areas indicate the areas of the insertion of ciliary bands." httpUri="https://zenodo.org/record/2626858/files/figure.png" pageId="16" pageNumber="17">Fig. 2B</figureCitation>
). The intestine does not have a separate, different anterior section. The X-organ of this species (observed in one specimen) is located at U84U86 near the terminal part of the intestine. It is bilobed, built from two extensions enveloped in a thin coat and connected by a thinner band located below the intestine end, at the ventral surface (
<figureCitation id="99BD2A2F730B457819DACF14FDE15547" box="[445,532,790,816]" captionStart="Fig" captionStartId="14.[189,232,1964,1990]" captionTargetBox="[189,1398,1077,1929]" captionTargetId="figure@14.[189,1398,1077,1929]" captionTargetPageId="14" captionText="Fig. 10. Chaetonotus (Chaetonotus) invitatus sp. nov. Detail of internal morphology. Bright field microphotography. X-organ." httpUri="https://zenodo.org/record/2626874/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
). The extensions have a granular appearance. The thin coat and the cellular bridge connecting the extensions have a smooth and homogeneous structure. The sperm packets of this species were not observed.
</paragraph>
</subSubSection>
<subSubSection id="499C6521730B457818DACF9CFEFC5376" pageId="16" pageNumber="17" type="discussion">
<paragraph id="013936AA730B457818DACF9CFEDA55CF" blockId="16.[189,303,926,952]" box="[189,303,926,952]" pageId="16" pageNumber="17">
<heading id="5A7181C6730B457818DACF9CFEDA55CF" bold="true" box="[189,303,926,952]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="33F2EAB8730B457818DACF9CFEDA55CF" bold="true" box="[189,303,926,952]" pageId="16" pageNumber="17">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA730B457818DACFCFFEFC5376" blockId="16.[189,1399,973,1281]" pageId="16" pageNumber="17">
Because of the long pharynx in relation to the length of the body and intestine, the pharynxintestine ratio (I) has high values in this species. Juvenile and subadult specimens have a pharynxintestine ratio (I) higher than 65%, rather than higher than 55%, as is usually observed in other species. In juvenile specimens, the first head scales from each dorsal longitudinal row are located under the free dorsal cephalion edges, whereas in adults they are nto located under the cephalion but below their dorsal edges (
<figureCitation id="99BD2A2F730B457818A1C87FFEFF52E0" box="[198,266,1149,1175]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1398,1009,1864]" captionTargetId="figure@11.[189,1398,1009,1864]" captionTargetPageId="11" captionText="Fig. 7. Chaetonotus (Chaetonotus) invitatus sp. nov. Bright field microphotographs. A. Detail of dorsal head view of adult specimen. B. Detail of dorsal head view of juvenile specimen. C. Dorsal view of posterior trunk region of specimen with two scales 17. D. Dorsal view of posterior trunk region of specimen with three scales 17." httpUri="https://zenodo.org/record/2626868/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
AB). Of the 25 reported adults, one individual had a large, developing egg (
<figureCitation id="99BD2A2F730B45781CE2C87FFB2B52EF" box="[1157,1246,1149,1176]" captionStart="Fig" captionStartId="9.[189,232,1944,1970]" captionTargetBox="[189,1398,1052,1906]" captionTargetId="figure@9.[189,1398,1052,1907]" captionTargetPageId="9" captionText="Fig. 5. Chaetonotus (Chaetonotus) invitatus sp. nov. Habitus. Phase contrast microphotographs. A. Juvenile specimen. B. Adult specimen. C. Specimen with mature egg." httpUri="https://zenodo.org/record/2626864/files/figure.png" pageId="16" pageNumber="17">Fig. 5C</figureCitation>
) and another possessed a distinct X-organ (
<figureCitation id="99BD2A2F730B45781A79C8A2FD8252CD" box="[542,631,1184,1210]" captionStart="Fig" captionStartId="14.[189,232,1964,1990]" captionTargetBox="[189,1398,1077,1929]" captionTargetId="figure@14.[189,1398,1077,1929]" captionTargetPageId="14" captionText="Fig. 10. Chaetonotus (Chaetonotus) invitatus sp. nov. Detail of internal morphology. Bright field microphotography. X-organ." httpUri="https://zenodo.org/record/2626874/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
). Four specimens had live, motile euglenids inside their intestine (
<figureCitation id="99BD2A2F730B457818A1C8C1FEE852AA" box="[198,285,1219,1245]" captionStart="Fig" captionStartId="15.[189,232,1964,1990]" captionTargetBox="[189,1398,1080,1934]" captionTargetId="figure@15.[189,1398,1080,1934]" captionTargetPageId="15" captionText="Fig. 11. Chaetonotus (Chaetonotus) invitatus sp. nov. Adult specimen. Bright field microphotography. Living euglenids inside intestine." httpUri="https://zenodo.org/record/2626876/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
) and were listed in an earlier paper as
<taxonomicName id="C6864D29730B45781A98C8C1FBB052A9" authority="(Chaetonotus)" authorityName="Ehrenberg" authorityYear="1830" baseAuthorityName="Chaetonotus" box="[767,1093,1219,1246]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="subGenus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781A98C8C1FC6652AA" box="[767,915,1219,1245]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8730B45781BC0C8C1FBCE52AA" box="[935,1083,1219,1245]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
)
</taxonomicName>
sp. 1 (
<bibRefCitation id="65174B5B730B45781CFAC8C1FF085376" author="Kisielewska G. &amp; Kolicka M. &amp; Zawierucha K." journalOrPublisher="European Journal of Protistology" pageId="16" pageNumber="17" pagination="138 - 141" part="51" refId="ref21719" refString="Kisielewska G., Kolicka M. &amp; Zawierucha K. 2015. Prey or parasite? The first observations of live Euglenida in the intestine of Gastrotricha. European Journal of Protistology 51: 138 - 141. https: // doi. org / 10.1016 / j. ejop. 2014.12.003" title="Prey or parasite? The first observations of live Euglenida in the intestine of Gastrotricha" type="journal article" year="2015">
Kisielewska
<emphasis id="33F2EAB8730B45781D5EC8C6FA8252AA" box="[1337,1399,1219,1245]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="499C6521730B457918DAC92AFCA75449" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="013936AA730B457818DAC92AFE3C5335" blockId="16.[189,457,1320,1346]" box="[189,457,1320,1346]" pageId="16" pageNumber="17">
<heading id="5A7181C6730B457818DAC92AFE3C5335" bold="true" box="[189,457,1320,1346]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="33F2EAB8730B457818DAC92AFE3C5335" bold="true" box="[189,457,1320,1346]" pageId="16" pageNumber="17">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="013936AA730B457918DAC954FCA75449" blockId="16.[189,1399,1366,2026]" lastBlockId="17.[189,1399,267,575]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Of all the 92 currently known nominal representatives of the subgenus
<taxonomicName id="C6864D29730B45781C48C954FE7153E3" authority="(Chaetonotus) Ehrenberg, 1830" authorityName="Ehrenberg" authorityYear="1830" baseAuthorityName="Chaetonotus" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="subGenus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781C48C954FB365307" box="[1071,1219,1366,1392]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8730B45781CBFC954FA995307" box="[1240,1388,1366,1392]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
) Ehrenberg, 1830
</taxonomicName>
,
<taxonomicName id="C6864D29730B457819F5C978FDA653E4" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[402,595,1402,1428]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="invitatus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B457819F5C978FE5853E3" box="[402,429,1402,1428]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B457819A7C978FE2E53E3" box="[448,475,1402,1428]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781989C978FDA653E4" box="[494,595,1402,1427]" italics="true" pageId="16" pageNumber="17">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3730B45781A3BC978FD4D53E3" box="[604,696,1402,1428]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
most closely resembles
<taxonomicName id="C6864D29730B45781BBAC978FA8453E3" authority="Ehrenberg, 1830" authorityName="Ehrenberg" authorityYear="1830" box="[989,1393,1401,1428]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="maximus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781BBAC978FC0D53E3" box="[989,1016,1402,1428]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781C6CC978FBD353E3" box="[1035,1062,1402,1428]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781C5EC978FB5453E4" box="[1081,1185,1402,1427]" italics="true" pageId="16" pageNumber="17">maximus</emphasis>
Ehrenberg, 1830
</taxonomicName>
,
<taxonomicName id="C6864D29730B457818DAC99FFD2253C0" authority="Preobrajenskaja, 1926" authorityName="Preobrajenskaja" authorityYear="1926" box="[189,727,1437,1463]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="microchaetus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B457818DAC99FFF2D53C0" box="[189,216,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781888C99FFEFF53C0" box="[239,266,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781946C99FFE4B53C0" box="[289,446,1437,1463]" italics="true" pageId="16" pageNumber="17">microchaetus</emphasis>
Preobrajenskaja, 1926
</taxonomicName>
,
<taxonomicName id="C6864D29730B45781A8DC99FFBA753C0" authority="Zelinka, 1889" authorityName="Zelinka" authorityYear="1889" box="[746,1106,1437,1463]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="similis" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781A8DC99FFCF153C0" box="[746,772,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781B7CC99FFCC353C0" box="[795,822,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781B2AC99FFC6F53C0" box="[845,922,1437,1463]" italics="true" pageId="16" pageNumber="17">similis</emphasis>
Zelinka, 1889
</taxonomicName>
,
<taxonomicName id="C6864D29730B45781C03C99FFE8453AD" authority="Balsamo, 1978" authorityName="Balsamo" authorityYear="1978" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="heterospinosus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781C03C99FFB8A53C0" box="[1124,1151,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781CF1C99FFB4453C0" box="[1174,1201,1437,1463]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781CAFC99FFA8253C0" box="[1224,1399,1437,1463]" italics="true" pageId="16" pageNumber="17">heterospinosus</emphasis>
Balsamo, 1978
</taxonomicName>
,
<taxonomicName id="C6864D29730B45781918C9C2FCEB53AD" authority="Marcolongo, 1910" authorityName="Marcolongo" authorityYear="1910" box="[383,798,1472,1498]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="laroides" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781918C9C2FE6F53AD" box="[383,410,1472,1498]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B457819CAC9C2FE3D53AD" box="[429,456,1472,1498]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B457819BCC9C2FDCF53AD" box="[475,570,1472,1498]" italics="true" pageId="16" pageNumber="17">laroides</emphasis>
Marcolongo, 1910
</taxonomicName>
and
<taxonomicName id="C6864D29730B45781B3DC9C2FB0853AD" authority="Greuter, 1917" authorityName="Greuter" authorityYear="1917" box="[858,1277,1472,1498]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="polyspinosus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781B3DC9C2FC8053AD" box="[858,885,1472,1498]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781BEFC9C2FC5653AD" box="[904,931,1472,1498]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781BD1C9C2FBBB53AD" box="[950,1102,1472,1498]" italics="true" pageId="16" pageNumber="17">polyspinosus</emphasis>
<bibRefCitation id="65174B5B730B45781C30C9C2FB0853AD" accessDate="2019-01-28" author="Greuter A." box="[1111,1277,1472,1498]" journalOrPublisher="Revue suisse de Zoologie" pageId="16" pageNumber="17" pagination="35 - 76" part="25" publicationUrl="https://biodiversitylibrary.org/page/10710334" refId="ref21072" refString="Greuter A. 1917. Beitrage zur Systematik der Gastrotrichen in der Schweiz. Revue suisse de Zoologie 25: 35 - 76. Available from https: // biodiversitylibrary. org / page / 10710334 [accessed 28 Jan. 2019]." title="Beitrage zur Systematik der Gastrotrichen in der Schweiz" type="journal article" year="1917">Greuter, 1917</bibRefCitation>
</taxonomicName>
. Of all 32 species belonging to the subgenus
<taxonomicName id="C6864D29730B45781A3AC9E1FBE6538A" authority="Schwank, 1990" authorityName="Schwank" authorityYear="1990" box="[605,1043,1507,1533]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="subGenus" subGenus="Hystricochaetonotus">
<emphasis id="33F2EAB8730B45781A3AC9E1FCA5538A" box="[605,848,1507,1533]" italics="true" pageId="16" pageNumber="17">Hystricochaetonotus</emphasis>
<bibRefCitation id="65174B5B730B45781B3EC9E1FBE6538A" author="Schwank P." box="[857,1043,1507,1533]" editor="Schwoerbel J." journalOrPublisher="Susswasserfauna von Mitteleuropa, 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York" pageId="16" pageNumber="17" pagination="3" refId="ref22912" refString="Schwank P. 1990. Gastrotricha. In: Schwoerbel J. &amp; Zwick P. (eds) Susswasserfauna von Mitteleuropa, 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York." type="book chapter" volumeTitle="Gastrotricha" year="1990">Schwank, 1990</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="C6864D29730B45781C46C9E6FF0D5057" authority="Balsamo, 1990" authorityName="Balsamo" authorityYear="1990" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="trispinosus" subGenus="Hystricochaetonotus">
<emphasis id="33F2EAB8730B45781C46C9E6FBC95389" box="[1057,1084,1508,1534]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781C28C9E6FB99538A" box="[1103,1132,1508,1533]" italics="true" pageId="16" pageNumber="17">H.</emphasis>
)
<emphasis id="33F2EAB8730B45781C19C9E6FB0A538A" box="[1150,1279,1508,1533]" italics="true" pageId="16" pageNumber="17">trispinosus</emphasis>
<bibRefCitation id="65174B5B730B45781D6FC9E1FF0D5057" author="Balsamo M." journalOrPublisher="Bolletino di Zoologia" pageId="16" pageNumber="17" pagination="165 - 178" part="57" refId="ref20637" refString="Balsamo M. 1990. Gastrotrichs from Lakes Bolsena, Chiusi and Montepulciano (central Italy), with the description of four new species. Bolletino di Zoologia 57 (2): 165 - 178. https: // doi. org / 10.1080 / 11250009009355693" title="Gastrotrichs from Lakes Bolsena, Chiusi and Montepulciano (central Italy), with the description of four new species" type="journal article" year="1990">Balsamo, 1990</bibRefCitation>
</taxonomicName>
seems to be the most similar to the newly described taxon, whereas of all the currently known 22 nominal representatives of the subgenus
<emphasis id="33F2EAB8730B45781AC5CA28FD485033" box="[674,701,1578,1604]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781AB7CA2BFC865034" box="[720,883,1577,1603]" italics="true" pageId="16" pageNumber="17">Primochaetus</emphasis>
)
<bibRefCitation id="65174B5B730B45781BE2CA2BFB995034" author="Kisielewski J." box="[901,1132,1577,1603]" journalOrPublisher="Annales Zoologici" pageId="16" pageNumber="17" pagination="145 - 151" part="46" refId="ref21921" refString="Kisielewski J. 1997 b. On the subgeneric division of the genus Chaetonotus Ehrenberg (Gastrotricha). Annales Zoologici 46: 145 - 151." title="On the subgeneric division of the genus Chaetonotus Ehrenberg (Gastrotricha)" type="journal article" year="1997">Kisielewski, 1997b</bibRefCitation>
, the new species most closely resembles
<taxonomicName id="C6864D29730B457819F2CA4FFCD65010" authority="Balsamo, 1978" authorityName="Balsamo" authorityYear="1978" box="[405,803,1613,1639]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="mutinensis" subGenus="P.">
<emphasis id="33F2EAB8730B457819F2CA4FFE455010" box="[405,432,1613,1639]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B457819A5CA4CFE205010" box="[450,469,1614,1639]" italics="true" pageId="16" pageNumber="17">P.</emphasis>
)
<emphasis id="33F2EAB8730B4578198ECA4CFD935010" box="[489,614,1614,1639]" italics="true" pageId="16" pageNumber="17">mutinensis</emphasis>
Balsamo, 1978
</taxonomicName>
.
<taxonomicName id="C6864D29730B45781B48CA4FFB995010" authorityName="Ehrenberg" authorityYear="1830" box="[815,1132,1613,1639]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="maximus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781B48CA4FFC365010" box="[815,963,1613,1639]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8730B45781BB2CA4FFC1A5010" box="[981,1007,1613,1639]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781C66CA4CFB995010" box="[1025,1132,1614,1639]" italics="true" pageId="16" pageNumber="17">maximus</emphasis>
</taxonomicName>
<emphasis id="33F2EAB8730B45781C0ACA4CFB845010" box="[1133,1137,1614,1639]" italics="true" pageId="16" pageNumber="17">,</emphasis>
<taxonomicName id="C6864D29730B45781C1ECA4FFA875010" authorityName="Preobrajenskaja" authorityYear="1926" box="[1145,1394,1613,1639]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="microchaetus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781C1ECA4FFB615010" box="[1145,1172,1613,1639]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781CC2CA4FFB355010" box="[1189,1216,1613,1639]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781CB5CA4FFA875010" box="[1234,1394,1613,1639]" italics="true" pageId="16" pageNumber="17">microchaetus</emphasis>
</taxonomicName>
<emphasis id="33F2EAB8730B45781D15CA4FFA835010" box="[1394,1398,1613,1639]" italics="true" pageId="16" pageNumber="17">,</emphasis>
<taxonomicName id="C6864D29730B457818DACA72FE9E50FD" authorityName="Zelinka" authorityYear="1889" box="[189,363,1648,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="similis" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B457818DACA72FF2D50FD" box="[189,216,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B4578188BCA72FEF250FD" box="[236,263,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B4578197BCA72FE9E50FD" box="[284,363,1648,1674]" italics="true" pageId="16" pageNumber="17">similis</emphasis>
</taxonomicName>
<emphasis id="33F2EAB8730B4578190CCA72FE8550FD" box="[363,368,1648,1674]" italics="true" pageId="16" pageNumber="17">,</emphasis>
<taxonomicName id="C6864D29730B4578191DCA72FD7E50FD" authorityName="Balsamo" authorityYear="1978" box="[378,651,1648,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="heterospinosus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B4578191DCA72FE6050FD" box="[378,405,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B457819CDCA72FE3150FD" box="[426,452,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B457819BECA72FD7E50FD" box="[473,651,1648,1674]" italics="true" pageId="16" pageNumber="17">heterospinosus</emphasis>
</taxonomicName>
<emphasis id="33F2EAB8730B45781AECCA72FD7A50FD" box="[651,655,1648,1674]" italics="true" pageId="16" pageNumber="17">,</emphasis>
<taxonomicName id="C6864D29730B45781AFDCA72FCAE50FD" authorityName="Marcolongo" authorityYear="1910" box="[666,859,1648,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="laroides" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781AFDCA72FD4050FD" box="[666,693,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781AAECA72FD1150FD" box="[713,740,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781A9ECA72FCAE50FD" box="[761,859,1648,1674]" italics="true" pageId="16" pageNumber="17">laroides</emphasis>
</taxonomicName>
<emphasis id="33F2EAB8730B45781B3CCA72FCAA50FD" box="[859,863,1648,1674]" italics="true" pageId="16" pageNumber="17">,</emphasis>
<taxonomicName id="C6864D29730B45781B0DCA72FB9550FD" authorityName="Greuter" authorityYear="1917" box="[874,1120,1648,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="polyspinosus" subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781B0DCA72FC7050FD" box="[874,901,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781BFECA72FC4150FD" box="[921,948,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781BAFCA72FB9550FD" box="[968,1120,1648,1674]" italics="true" pageId="16" pageNumber="17">polyspinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="C6864D29730B45781CC7CA72FA8350FD" authorityName="Balsamo" authorityYear="1978" box="[1184,1398,1648,1674]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="mutinensis" subGenus="P.">
<emphasis id="33F2EAB8730B45781CC7CA72FB4E50FD" box="[1184,1211,1648,1674]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781CA8CA73FB1750FD" box="[1231,1250,1649,1674]" italics="true" pageId="16" pageNumber="17">P.</emphasis>
)
<emphasis id="33F2EAB8730B45781C9ECA73FA8350FD" box="[1273,1398,1649,1674]" italics="true" pageId="16" pageNumber="17">mutinensis</emphasis>
</taxonomicName>
are the most similar to the newly described species in terms of: body shape similarity, range of body length, range of pharynx length, range of adhesive tube length, presence of a scale of a different
<typeStatus id="DE3D8808730B45781D23CAB5FA8250A6" box="[1348,1399,1719,1745]" pageId="16" pageNumber="17">type</typeStatus>
on the dorsal and dorsolateral surfaces on the furcal base, presence of rigid, long spines on the furcal appendages and a similar
<typeStatus id="DE3D8808730B4578199ACAFFFDC55160" box="[509,560,1789,1815]" pageId="16" pageNumber="17">type</typeStatus>
of terminal scales on the ventral interciliary field.
<taxonomicName id="C6864D29730B45781CC0CAFFFECB514D" authorityName="Balsamo" authorityYear="1990" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="trispinosus" subGenus="Hystricochaetonotus">
<emphasis id="33F2EAB8730B45781CC0CAFFFACE5160" box="[1191,1339,1789,1815]" italics="true" pageId="16" pageNumber="17">Chaetonotus</emphasis>
(
<emphasis id="33F2EAB8730B45781D37CAFCFA985160" box="[1360,1389,1790,1815]" italics="true" pageId="16" pageNumber="17">H.</emphasis>
)
<emphasis id="33F2EAB8730B457818DACB23FECB514D" box="[189,318,1825,1850]" italics="true" pageId="16" pageNumber="17">trispinosus</emphasis>
</taxonomicName>
was selected for comparison with
<taxonomicName id="C6864D29730B45781ABCCB22FC6F514D" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[731,922,1824,1850]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="invitatus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B45781ABCCB22FD03514D" box="[731,758,1824,1850]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B45781B6FCB22FCD6514D" box="[776,803,1824,1850]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B45781B52CB23FC6F514D" box="[821,922,1825,1850]" italics="true" pageId="16" pageNumber="17">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3730B45781BC4CB22FC0B514D" box="[931,1022,1824,1850]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
due to its similarity in terms of having three larger scales with longer and stronger spines present on the dorsal posterior trunk region, length variation of large spines on different body sides and by the presence of scales of a different
<typeStatus id="DE3D8808730B457818DACB88FF0551D3" box="[189,240,1930,1956]" pageId="16" pageNumber="17">type</typeStatus>
on the dorsal and dorsolateral surfaces of the furcal base. Despite the fact that these eight species have the highest number of common features with the newly described species, they are significantly different from
<taxonomicName id="C6864D29730B4578190ECBD2FDD2519D" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[361,551,2000,2026]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="16" pageNumber="17" phylum="Gastrotricha" rank="species" species="invitatus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8730B4578190ECBD2FE71519D" box="[361,388,2000,2026]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
(
<emphasis id="33F2EAB8730B457819F1CBD2FE45519D" box="[406,432,2000,2026]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
)
<emphasis id="33F2EAB8730B457819A5CBD3FDD2519D" box="[450,551,2001,2026]" italics="true" pageId="16" pageNumber="17">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3730B45781A49CBD2FD7C519D" box="[558,649,2000,2026]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
most strikingly by scale
<typeStatus id="DE3D8808730B45781BA1CBD2FC0C519D" box="[966,1017,2000,2026]" pageId="16" pageNumber="17">type</typeStatus>
and shape variation, spine
<typeStatus id="DE3D8808730B45781D5ECBD2FA82519D" box="[1337,1399,2000,2026]" pageId="16" pageNumber="17">types</typeStatus>
and length variation. Comparisons between the new species and the most morphologically similar taxa are summarised in
<tableCitation id="4C040311730A457919FACD2CFDFE573F" box="[413,523,302,328]" captionStart="Table 2" captionStartId="9.[189,258,267,293]" captionText="Table 2. Main morphometric characters of Chaetonotus (Chaetonotus) invitatus sp. nov. N = number of specimens analysed; Range = the smallest and the largest structure found among all specimens measured. All measurements in micrometers (μm)." pageId="17" pageNumber="18">Table A2</tableCitation>
. Taxa chosen for the differential diagnosis often occur in different regions of the world and are presented very differently by various researchers. Thus, a different description/ specification or taxonomic classification of theoretically the same species may suggest that we are dealing with a very large range of plasticity of these taxa or with a larger number of undiagnosed, separate species. To avoid any doubts in comparisons with
<taxonomicName id="C6864D29730A45791B19CDB9FBCB57A2" authority="Kolicka, 2019" authorityName="Kolicka" authorityYear="2019" box="[894,1086,443,469]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="17" pageNumber="18" phylum="Gastrotricha" rank="species" species="invitatus" status="sp. nov." subGenus="Chaetonotus">
<emphasis id="33F2EAB8730A45791B19CDB9FC6C57A2" box="[894,921,443,469]" italics="true" pageId="17" pageNumber="18">C.</emphasis>
(
<emphasis id="33F2EAB8730A45791BCCCDB9FC3357A2" box="[939,966,443,469]" italics="true" pageId="17" pageNumber="18">C.</emphasis>
)
<emphasis id="33F2EAB8730A45791BBECDBEFBCB57A2" box="[985,1086,444,469]" italics="true" pageId="17" pageNumber="18">invitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28C157C3730A45791C21CDB9FB5057A2" box="[1094,1189,443,469]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, only the original descriptions of these species and three of the most detailed papers with morphological data were used (
<bibRefCitation id="65174B5B730A457918A2CE03FE87546C" author="Balsamo M." box="[197,370,513,539]" journalOrPublisher="Guide per il Riconoscimento delle Specie animali delle Acque interne italiane" pageId="17" pageNumber="18" pagination="1 - 92" part="20" refId="ref20603" refString="Balsamo M. 1983. Gastrotrichi. In: Consiglio Nazionale delle Ricerche (ed.) Guide per il Riconoscimento delle Specie animali delle Acque interne italiane 20: 1 - 92." title="Gastrotrichi" type="journal article" volumeTitle="Consiglio Nazionale delle Ricerche" year="1983">Balsamo 1983</bibRefCitation>
;
<bibRefCitation id="65174B5B730A45791919CE03FDDA546C" author="Schwank P." box="[382,559,513,539]" editor="Schwoerbel J." journalOrPublisher="Susswasserfauna von Mitteleuropa, 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York" pageId="17" pageNumber="18" pagination="3" refId="ref22912" refString="Schwank P. 1990. Gastrotricha. In: Schwoerbel J. &amp; Zwick P. (eds) Susswasserfauna von Mitteleuropa, 3. Gastrotricha und Nemertini. Gustav Fischer Verlag, Stuttgart, Jena, New York." type="book chapter" volumeTitle="Gastrotricha" year="1990">Schwank 1990</bibRefCitation>
;
<bibRefCitation id="65174B5B730A45791A5CCE03FCEC546C" author="Kisielewski J." box="[571,793,513,539]" journalOrPublisher="Fauna Slodkowodna Polski, Zeszyt" pageId="17" pageNumber="18" pagination="1 - 157" part="31" refId="ref21899" refString="Kisielewski J. 1997 a. Brzuchorzeski (Gastrotricha). Fauna Slodkowodna Polski, Zeszyt 31: 1 - 157." title="Brzuchorzeski (Gastrotricha)" type="journal article" year="1997">Kisielewski 1997a</bibRefCitation>
). For information regarding
<taxonomicName id="C6864D29730A45791C02CE00FAC6546C" authorityName="Balsamo" authorityYear="1978" box="[1125,1331,513,540]" family="Chaetonotidae" genus="Chaetonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chaetonotida" pageId="17" pageNumber="18" phylum="Gastrotricha" rank="species" species="mutinensis" subGenus="P.">
<emphasis id="33F2EAB8730A45791C02CE00FB8A546B" box="[1125,1151,514,540]" italics="true" pageId="17" pageNumber="18">C.</emphasis>
(
<emphasis id="33F2EAB8730A45791CF7CE00FB56546C" box="[1168,1187,514,539]" italics="true" pageId="17" pageNumber="18">P.</emphasis>
)
<emphasis id="33F2EAB8730A45791CD1CE00FAC6546C" box="[1206,1331,514,539]" italics="true" pageId="17" pageNumber="18">mutinensis</emphasis>
</taxonomicName>
, only original data are considered in the differential diagnosis.
</paragraph>
</subSubSection>
</treatment>
</document>