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<document id="47403FC6C54ACE64EA8983EBE70422DE" ID-CLB-Dataset="32203" ID-DOI="10.11646/zootaxa.4317.3.7" ID-GBIF-Dataset="315f6657-6f59-4a2e-8f55-3181b3453ee6" ID-ISSN="1175-5326" ID-Zenodo-Dep="884835" ID-ZooBank="1E813Bd2-576A-4272-Be15-E02Dc1E6B4A2" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504596569331" checkinUser="plazi" docAuthor="Özbek, Hasan Hüseyin" docDate="2017" docId="7A25A767B2259B1E2394FAB9FE21FEB2" docLanguage="en" docName="zootaxa.4317.3.7.pdf" docOrigin="Zootaxa 4317 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holostaspella bidentata Özbek 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="567" masterDocId="861CDF1FB2239B172303FFBAFF9AFFD7" masterDocTitle="A review of the macrochelid mites of Turkey (Acari: Macrochelidae), with new records and descriptions of three new species" masterLastPageNumber="572" masterPageNumber="559" pageNumber="565" updateTime="1698467872273" updateUser="ExternalLinkService">
<mods:mods id="FF0E0D493456B5238DA0EA159AADC214" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32AB3B810E4D32BF1E46257D53F12284">
<mods:title id="2AAB04CB185A99B56F8F842BCAAD6A33">A review of the macrochelid mites of Turkey (Acari: Macrochelidae), with new records and descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="AD744961D61F275B5A449EEF32B8935A" type="personal">
<mods:role id="1D483E0CEA1E8AC5C3C2E97F568CC16A">
<mods:roleTerm id="8D5514802DEC6A03886C3BFB1B39DAFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F06BBA5E3A136145C12CEC925272491">Özbek, Hasan Hüseyin</mods:namePart>
</mods:name>
<mods:typeOfResource id="2D049567504A8F31EAD794E1E79B5D78">text</mods:typeOfResource>
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<mods:titleInfo id="BA2DD6261DB939826B73DD7779244547">
<mods:title id="D6E8837137BA6BFD58385549E5891809">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2769241506D257FB4BD94922A0F06B4B">
<mods:date id="4CD61523E9F022B8AE29A33268F099B4">2017</mods:date>
<mods:detail id="CB3784581511C82DCE7AA8CAED968BED" type="pubDate">
<mods:number id="41B0A8658102176E74F75F9B6DD08F72">2017-05-05</mods:number>
</mods:detail>
<mods:detail id="F4FB800A034AF91FB203A7BC5EA7364A" type="volume">
<mods:number id="CFC636C6262284EE58800BE6F367F820">4317</mods:number>
</mods:detail>
<mods:detail id="01059D2496C06C1798094C883F182276" type="issue">
<mods:number id="AC4619F9C68B2865FC6CD5AC91EEE2EA">3</mods:number>
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<mods:extent id="B4981914A9DC3030AFFFA2C9A78ADBCF" unit="page">
<mods:start id="86A44328556929453E9E8EEA3537C089">559</mods:start>
<mods:end id="EAF1BA09998AB5C93F64B2EA697BEC6B">572</mods:end>
</mods:extent>
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<mods:classification id="55882B74A0E8FF773B6D3E93F8ABC418">journal article</mods:classification>
<mods:identifier id="DC5ECBF6064296C6E03F2CDC6457F691" type="CLB-Dataset">32203</mods:identifier>
<mods:identifier id="FA4332B9CB6C94E80C9B4EA845B5607C" type="DOI">10.11646/zootaxa.4317.3.7</mods:identifier>
<mods:identifier id="88499E4319A530A28EDB6ABB8573694F" type="GBIF-Dataset">315f6657-6f59-4a2e-8f55-3181b3453ee6</mods:identifier>
<mods:identifier id="E3C06172B9E61F6A7C199CB795862F60" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0B571737C1CF2D5CDFC9B06FF3E5BD1C" type="Zenodo-Dep">884835</mods:identifier>
<mods:identifier id="99FEC2E508516FE22384F1D37744CEC5" type="ZooBank">1E813Bd2-576A-4272-Be15-E02Dc1E6B4A2</mods:identifier>
</mods:mods>
<treatment id="7A25A767B2259B1E2394FAB9FE21FEB2" ID-DOI="http://doi.org/10.5281/zenodo.6042934" ID-GBIF-Taxon="133183447" ID-Zenodo-Dep="6042934" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2" httpUri="http://treatment.plazi.org/id/7A25A767B2259B1E2394FAB9FE21FEB2" lastPageId="9" lastPageNumber="567" pageId="6" pageNumber="565">
<subSubSection id="BA9645FAB2259B112394FAB9FE05FAE8" pageId="6" pageNumber="565" type="nomenclature">
<paragraph id="F2331671B2259B112394FAB9FD8DFAC9" blockId="6.[151,535,1283,1343]" box="[151,535,1283,1310]" pageId="6" pageNumber="565">
<heading id="A97BA11DB2259B112394FAB9FD8DFAC9" bold="true" box="[151,535,1283,1310]" fontSize="11" level="1" pageId="6" pageNumber="565" reason="1">
<emphasis id="C0F8CA63B2259B112394FAB9FD8DFAC9" bold="true" box="[151,535,1283,1310]" pageId="6" pageNumber="565">
<taxonomicName id="358C6DF2B2259B112394FAB9FE2BFACA" authority="Özbek, 2017" authorityName="Özbek" authorityYear="2017" box="[151,433,1283,1309]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="bidentata" status="sp. nov.">
<emphasis id="C0F8CA63B2259B112394FAB9FE2BFACA" bold="true" box="[151,433,1283,1309]" italics="true" pageId="6" pageNumber="565">Holostaspella bidentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DBCB7718B2259B1122BAFABEFD8DFAC9" box="[441,535,1284,1310]" pageId="6" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F2331671B2259B112394FA9CFE05FAE8" blockId="6.[151,535,1283,1343]" box="[151,415,1318,1343]" pageId="6" pageNumber="565">
(
<figureCitation id="6AB70AF4B2259B11239DFA9CFEDFFAE8" box="[158,325,1318,1343]" captionStart-0="FIGURES 16 ‒ 17" captionStart-1="FIGURES 18 ‒ 24" captionStartId-0="7.[151,264,1828,1851]" captionStartId-1="8.[151,264,1643,1666]" captionTargetBox-0="[151,1430,850,1803]" captionTargetBox-1="[187,1388,367,1604]" captionTargetId-0="figure@7.[151,1436,841,1807]" captionTargetId-1="figure@8.[174,1412,337,1622]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 16 ‒ 17. Holostaspella bidentata, female. 16. Dorsal idiosoma; 10. Ventral idiosoma." captionText-1="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri-0="https://zenodo.org/record/892975/files/figure.png" httpUri-1="https://zenodo.org/record/892977/files/figure.png" pageId="6" pageNumber="565">Figures 16‒24</figureCitation>
; 32‒36)
</paragraph>
</subSubSection>
<subSubSection id="BA9645FAB2259B112394FAD4FEFBF939" pageId="6" pageNumber="565" type="diagnosis">
<paragraph id="F2331671B2259B112394FAD4FC2FF9C0" blockId="6.[151,1436,1390,2027]" pageId="6" pageNumber="565">
<emphasis id="C0F8CA63B2259B112394FAD4FE8BFA50" bold="true" box="[151,273,1390,1415]" pageId="6" pageNumber="565">Diagnosis.</emphasis>
Ventri-anal shield with fOur pairs Of setae; metasternal shields free On sOft integument; tarsus II with a spur-like seta (
<emphasis id="C0F8CA63B2259B11223EFA29FEC4FA7D" box="[317,350,1427,1450]" italics="true" pageId="6" pageNumber="565">mv</emphasis>
), dOrsal shield withOut lateral indentatiOns; all dOrsal shield setae pilOse alOng their entire length and similar in width. In female, all setae On ventral shields smOOth and pOinted, except pilOse pOst-anal setae;
<emphasis id="C0F8CA63B2259B11265FFA0DFED3FA25" italics="true" pageId="6" pageNumber="565">crista posterior erecta</emphasis>
distinct and narrOw; mOveable digit with twO distinct median teeth. In male,
<emphasis id="C0F8CA63B2259B1127A1FA60FB52FA25" box="[1186,1224,1498,1522]" italics="true" pageId="6" pageNumber="565">Jv3</emphasis>
smOOth, Other preanal setae pilOse; spermatOdactyl abOut dOuble length Of mOvable digit.
</paragraph>
<paragraph id="F2331671B2259B1123C4F998FEFBF939" blockId="6.[151,1436,1390,2027]" pageId="6" pageNumber="565">
<materialsCitation id="42E41C2CB2259B1123C4F998FA02F9EC" ID-GBIF-Occurrence="1608130897" box="[199,1432,1570,1595]" country="Turkey" latitude="40.266666" location="Kose TOwn" longLatPrecision="1164" longitude="39.633335" municipality="Specimens" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Gumushane PrOvince" typeStatus="holotype">
<emphasis id="C0F8CA63B2259B1123C4F998FE5CF9EC" bold="true" box="[199,454,1570,1595]" pageId="6" pageNumber="565">
<collectingMunicipality id="12578C0BB2259B1123C4F998FEDFF9EC" box="[199,325,1570,1595]" pageId="6" pageNumber="565">Specimens</collectingMunicipality>
examined.
</emphasis>
<typeStatus id="2D37A8D3B2259B1122CEF998FDA2F9EC" box="[461,568,1570,1595]" pageId="6" pageNumber="565" type="holotype">HOlOtype</typeStatus>
, female,
<collectingCountry id="8A9B56E1B2259B11219DF998FD75F9EC" box="[670,751,1570,1595]" name="Turkey" pageId="6" pageNumber="565">Turkey</collectingCountry>
,
<collectingRegion id="3048D893B2259B1121F9F998FC77F9EC" box="[762,1005,1570,1595]" country="Turkey" name="Gumushane" pageId="6" pageNumber="565">Gümüşhane PrOvince</collectingRegion>
,
<location id="F75340AAB2259B1120FBF999FBE1F9EC" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2:F75340AAB2259B1120FBF999FBE1F9EC" box="[1016,1147,1571,1595]" country="Turkey" latitude="40.266666" longLatPrecision="1164" longitude="39.633335" municipality="Specimens" name="Kose TOwn" pageId="6" pageNumber="565" stateProvince="Gumushane PrOvince">Köse TOwn</location>
,
<geoCoordinate id="97B870B6B2259B112784F999FB62F9ED" box="[1159,1272,1570,1595]" direction="north" orientation="latitude" pageId="6" pageNumber="565" precision="925" value="40.266666">40° 16’ N</geoCoordinate>
,
<geoCoordinate id="97B870B6B2259B112600F999FAF4F9ED" box="[1283,1390,1571,1595]" direction="east" orientation="longitude" pageId="6" pageNumber="565" precision="925" value="39.633335">39° 38' E</geoCoordinate>
, alt
</materialsCitation>
.
<materialsCitation id="42E41C2CB2259B112394F9FCFCD0F954" ID-GBIF-Occurrence="1608130890" collectingDate="2011-06-26" country="Turkey" elevation="1856" latitude="40.266666" location="Kose TOwn" longLatPrecision="1164" longitude="39.616665" municipality="Specimens" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Gumushane PrOvince" typeStatus="paratype">
<elevation id="79A1F142B2259B112394F9FCFF6EF989" box="[151,244,1606,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.8559999999999999" pageId="6" pageNumber="565" unit="m" value="1856.0">
<quantity id="3574BB94B2259B112394F9FCFF6EF989" box="[151,244,1606,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.8559999999999999" pageId="6" pageNumber="565" unit="m" value="1856.0">1856 m</quantity>
</elevation>
,
<collectingDate id="9676C959B2259B112204F9FCFE30F988" box="[263,426,1606,1631]" pageId="6" pageNumber="565" value="2011-06-26">
<date id="863230B1B2259B112204F9FCFE30F988" box="[263,426,1606,1631]" pageId="6" pageNumber="565" value="2011-06-26">26 June 2011</date>
</collectingDate>
, litter frOm the fOrest.
<typeStatus id="2D37A8D3B2259B1121C4F9FDFCADF988" box="[711,823,1607,1631]" pageId="6" pageNumber="565" type="paratype">Paratypes</typeStatus>
, a male, same data as hOlOtype; a female,
<collectingCountry id="8A9B56E1B2259B112644F9FCFA02F988" box="[1351,1432,1606,1631]" name="Turkey" pageId="6" pageNumber="565">Turkey</collectingCountry>
,
<collectingRegion id="3048D893B2259B112394F9D0FE17F954" box="[151,397,1642,1667]" country="Turkey" name="Gumushane" pageId="6" pageNumber="565">Gümüşhane PrOvince</collectingRegion>
,
<location id="F75340AAB2259B11229AF9D1FD84F954" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2:F75340AAB2259B11229AF9D1FD84F954" box="[409,542,1643,1667]" country="Turkey" latitude="40.266666" longLatPrecision="1164" longitude="39.616665" name="Kose TOwn" pageId="6" pageNumber="565" stateProvince="Gumushane PrOvince">Köse TOwn</location>
,
<geoCoordinate id="97B870B6B2259B11212FF9D1FD3BF955" box="[556,673,1642,1667]" direction="north" orientation="latitude" pageId="6" pageNumber="565" precision="925" value="40.266666">40° 16’ N</geoCoordinate>
,
<geoCoordinate id="97B870B6B2259B1121ACF9D1FC86F955" box="[687,796,1643,1667]" direction="east" orientation="longitude" pageId="6" pageNumber="565" precision="925" value="39.616665">39° 37' E</geoCoordinate>
, alt
</materialsCitation>
.
<materialsCitation id="42E41C2CB2259B112054F9D0FC2DF970" ID-GBIF-Occurrence="1608130895" collectingDate="2009-07-14" country="Turkey" elevation="1758" latitude="40.266666" location="Kose TOwn" longLatPrecision="1164" longitude="39.616665" municipality="Of" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Gumushane PrOvince" typeStatus="paratype">
<elevation id="79A1F142B2259B112054F9D0FC2AF955" box="[855,944,1642,1667]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="565" unit="m" value="1758.0">
<quantity id="3574BB94B2259B112054F9D0FC2AF955" box="[855,944,1642,1667]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="565" unit="m" value="1758.0">1758 m</quantity>
</elevation>
,
<collectingDate id="9676C959B2259B1120BDF9D1FBCCF954" box="[958,1110,1642,1667]" pageId="6" pageNumber="565" value="2009-07-14">
<date id="863230B1B2259B1120BDF9D1FBCCF954" box="[958,1110,1642,1667]" pageId="6" pageNumber="565" value="2009-07-14">14 July 2009</date>
</collectingDate>
, litter
<collectingMunicipality id="12578C0BB2259B1127A2F9D0FB22F954" box="[1185,1208,1642,1667]" pageId="6" pageNumber="565">Of</collectingMunicipality>
<taxonomicName id="358C6DF2B2259B1127C2F9D0FB63F955" box="[1217,1273,1642,1666]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="6" pageNumber="565" phylum="Chordata" rank="genus">
<emphasis id="C0F8CA63B2259B1127C2F9D0FB63F955" box="[1217,1273,1642,1666]" italics="true" pageId="6" pageNumber="565">Salix</emphasis>
</taxonomicName>
sp.; a female,
<collectingCountry id="8A9B56E1B2259B112394F934FF72F970" box="[151,232,1678,1703]" name="Turkey" pageId="6" pageNumber="565">Turkey</collectingCountry>
,
<collectingRegion id="3048D893B2259B1123F5F934FE77F970" box="[246,493,1678,1703]" country="Turkey" name="Gumushane" pageId="6" pageNumber="565">Gümüşhane PrOvince</collectingRegion>
,
<location id="F75340AAB2259B1122FFF935FD18F970" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2:F75340AAB2259B1122FFF935FD18F970" box="[508,642,1679,1703]" country="Turkey" latitude="40.266666" longLatPrecision="1164" longitude="39.616665" municipality="Of" name="Kose TOwn" pageId="6" pageNumber="565" stateProvince="Gumushane PrOvince">Köse TOwn</location>
,
<geoCoordinate id="97B870B6B2259B112193F935FC93F971" box="[656,777,1678,1703]" direction="north" orientation="latitude" pageId="6" pageNumber="565" precision="925" value="40.266666">40° 16’ N</geoCoordinate>
,
<geoCoordinate id="97B870B6B2259B112014F935FC12F971" box="[791,904,1679,1703]" direction="east" orientation="longitude" pageId="6" pageNumber="565" precision="925" value="39.616665">39° 37' E</geoCoordinate>
, alt
</materialsCitation>
.
<materialsCitation id="42E41C2CB2259B1120C7F934FC02F91C" ID-GBIF-Occurrence="1608130891" collectingDate="2009-07-14" country="Turkey" elevation="1758" latitude="40.266666" location="Kose TOwn" longLatPrecision="1164" longitude="39.616665" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Gumushane PrOvince" typeStatus="paratype">
<elevation id="79A1F142B2259B1120C7F934FB85F971" box="[964,1055,1678,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="565" unit="m" value="1758.0">
<quantity id="3574BB94B2259B1120C7F934FB85F971" box="[964,1055,1678,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="565" unit="m" value="1758.0">1758 m</quantity>
</elevation>
,
<collectingDate id="9676C959B2259B11272DF935FB53F970" box="[1070,1225,1678,1703]" pageId="6" pageNumber="565" value="2009-07-14">
<date id="863230B1B2259B11272DF935FB53F970" box="[1070,1225,1678,1703]" pageId="6" pageNumber="565" value="2009-07-14">14 July 2009</date>
</collectingDate>
, in sOil; a female,
<collectingCountry id="8A9B56E1B2259B112394F908FF72F91C" box="[151,232,1714,1739]" name="Turkey" pageId="6" pageNumber="565">Turkey</collectingCountry>
,
<collectingRegion id="3048D893B2259B1123F7F908FE72F91C" box="[244,488,1714,1739]" country="Turkey" name="Gumushane" pageId="6" pageNumber="565">Gümüşhane PrOvince</collectingRegion>
,
<location id="F75340AAB2259B1122F0F909FDE2F91C" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2:F75340AAB2259B1122F0F909FDE2F91C" box="[499,632,1715,1739]" country="Turkey" latitude="40.266666" longLatPrecision="1164" longitude="39.616665" name="Kose TOwn" pageId="6" pageNumber="565" stateProvince="Gumushane PrOvince">Köse TOwn</location>
,
<geoCoordinate id="97B870B6B2259B112180F909FD6CF91D" box="[643,758,1714,1739]" direction="north" orientation="latitude" pageId="6" pageNumber="565" precision="925" value="40.266666">40° 16’ N</geoCoordinate>
,
<geoCoordinate id="97B870B6B2259B112001F909FCF6F91D" box="[770,876,1715,1739]" direction="east" orientation="longitude" pageId="6" pageNumber="565" precision="925" value="39.616665">39° 37' E</geoCoordinate>
, alt
</materialsCitation>
.
<materialsCitation id="42E41C2CB2259B1120A7F909FEFBF939" ID-GBIF-Occurrence="1608130894" collectingDate="2009-07-14" country="Turkey" elevation="1800" location="Of" pageId="6" pageNumber="565" specimenCount="1" stateProvince="Gumushane PrOvince" typeStatus="paratype">
<elevation id="79A1F142B2259B1120A7F909FC66F91D" box="[932,1020,1715,1739]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="565" unit="m" value="1800.0">
<quantity id="3574BB94B2259B1120A7F909FC66F91D" box="[932,1020,1715,1739]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="565" unit="m" value="1800.0">1800 m</quantity>
</elevation>
,
<collectingDate id="9676C959B2259B11270BF909FB07F91C" box="[1032,1181,1714,1739]" pageId="6" pageNumber="565" value="2009-07-14">
<date id="863230B1B2259B11270BF909FB07F91C" box="[1032,1181,1714,1739]" pageId="6" pageNumber="565" value="2009-07-14">14 July 2009</date>
</collectingDate>
, litter
<location id="F75340AAB2259B1127E5F908FB67F91C" LSID="urn:lsid:plazi:treatment:7A25A767B2259B1E2394FAB9FE21FEB2:F75340AAB2259B1127E5F908FB67F91C" box="[1254,1277,1714,1739]" country="Turkey" name="Of" pageId="6" pageNumber="565" stateProvince="Gumushane PrOvince">Of</location>
<taxonomicName id="358C6DF2B2259B112607F909FADEF91D" box="[1284,1348,1715,1738]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="565" phylum="Tracheophyta" rank="genus">
<emphasis id="C0F8CA63B2259B112607F909FADEF91D" box="[1284,1348,1715,1738]" italics="true" pageId="6" pageNumber="565">Pinus</emphasis>
</taxonomicName>
sp. and mOss near stream.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BA9645FAB2259B1F23C4F940FC3AFECB" lastPageId="8" lastPageNumber="567" pageId="6" pageNumber="565" type="description">
<paragraph id="F2331671B2259B1123C4F940FE28F8C4" blockId="6.[151,1436,1390,2027]" box="[199,434,1786,1811]" pageId="6" pageNumber="565">
<emphasis id="C0F8CA63B2259B1123C4F940FEC3F8C4" bold="true" box="[199,345,1786,1811]" pageId="6" pageNumber="565">Description.</emphasis>
Female
</paragraph>
<paragraph id="F2331671B2259B1123C4F8A5FED7F83D" blockId="6.[151,1436,1390,2027]" pageId="6" pageNumber="565">
<emphasis id="C0F8CA63B2259B1123C4F8A5FEBBF8E1" box="[199,289,1823,1846]" italics="true" pageId="6" pageNumber="565">Dorsum</emphasis>
(
<figureCitation id="6AB70AF4B2259B112232F8A4FE1EF8E0" box="[305,388,1822,1847]" captionStart="FIGURES 16 ‒ 17" captionStartId="7.[151,264,1828,1851]" captionTargetBox="[151,1430,850,1803]" captionTargetId="figure@7.[151,1436,841,1807]" captionTargetPageId="7" captionText="FIGURES 16 ‒ 17. Holostaspella bidentata, female. 16. Dorsal idiosoma; 10. Ventral idiosoma." httpUri="https://zenodo.org/record/892975/files/figure.png" pageId="6" pageNumber="565">Fig. 16</figureCitation>
). DOrsal shield Oval, 800‒860 lOng, 550‒615 wide at the level Of widest pOint (n=5), cOvered with punctatiOn, and pOsteriOr-dOrsally slightly web-like; shield with 28 pairs Of setae, pilOse alOng their entire length and similar in width; setae
<emphasis id="C0F8CA63B2259B11213AF8DDFDD4F8A8" box="[569,590,1895,1919]" italics="true" pageId="6" pageNumber="565">
j
<date id="863230B1B2259B112147F8DDFDD4F8A8" box="[580,590,1895,1919]" pageId="6" pageNumber="565" value="1940-01-30">1</date>
</emphasis>
30‒40 medially expanded, laterally pilOse, and lOcated On an anteriOr prOjectiOn,
<emphasis id="C0F8CA63B2259B112215F830FEB5F875" box="[278,303,1930,1954]" italics="true" pageId="6" pageNumber="565">z1</emphasis>
45‒54,
<emphasis id="C0F8CA63B2259B112288F831FE3EF875" box="[395,420,1931,1954]" italics="true" pageId="6" pageNumber="565">z5</emphasis>
36‒42,
<emphasis id="C0F8CA63B2259B1122FCF831FD8FF875" box="[511,533,1931,1954]" italics="true" pageId="6" pageNumber="565">j5</emphasis>
55‒55, Other dOrsal setae mOre than 60 lOng (60‒105), especially pOsterO-dOrsal setae much lOnger (
<emphasis id="C0F8CA63B2259B112276F815FE15F810" box="[373,399,1967,1991]" italics="true" pageId="6" pageNumber="565">z4</emphasis>
,
<emphasis id="C0F8CA63B2259B11229EF815FE2CF811" box="[413,438,1967,1990]" italics="true" pageId="6" pageNumber="565">s5</emphasis>
,
<emphasis id="C0F8CA63B2259B1122C6F815FD94F810" box="[453,526,1967,1991]" italics="true" pageId="6" pageNumber="565">Z1‒Z4</emphasis>
,
<emphasis id="C0F8CA63B2259B11211EF815FDFEF811" box="[541,612,1967,1991]" italics="true" pageId="6" pageNumber="565">S1‒S5</emphasis>
mOre than 80,
<emphasis id="C0F8CA63B2259B112013F815FCB0F810" box="[784,810,1967,1991]" italics="true" pageId="6" pageNumber="565">J2</emphasis>
abOut 80). AnteriOr tips Of peritremes reaching beyOnd base Of setae
<emphasis id="C0F8CA63B2259B11222EF868FEDFF83D" box="[301,325,2002,2026]" italics="true" pageId="6" pageNumber="565">z2</emphasis>
.
</paragraph>
<paragraph id="F2331671B2249B1023C4FF23FDB7FE5F" blockId="7.[151,1437,151,789]" pageId="7" pageNumber="566">
<emphasis id="C0F8CA63B2249B1023C4FF23FE96FF67" box="[199,268,153,176]" italics="true" pageId="7" pageNumber="566">Venter</emphasis>
(
<figureCitation id="6AB70AF4B2249B102221FF2DFEE3FF67" box="[290,377,151,176]" captionStart="FIGURES 16 ‒ 17" captionStartId="7.[151,264,1828,1851]" captionTargetBox="[151,1430,850,1803]" captionTargetId="figure@7.[151,1436,841,1807]" captionTargetPageId="7" captionText="FIGURES 16 ‒ 17. Holostaspella bidentata, female. 16. Dorsal idiosoma; 10. Ventral idiosoma." httpUri="https://zenodo.org/record/892975/files/figure.png" pageId="7" pageNumber="566">Fig. 17</figureCitation>
). Sternal shield 160‒164 lOng, 150‒160 wide at mid-level Of cOxae II, with three pairs Of unifOrm smOOth and pOinted setae, and twO pairs Of pOres; surface Ornamented with punctatiOn and pOlygOnal pattern;
<emphasis id="C0F8CA63B2249B1023F5FF5AFE6EFF2F" box="[246,500,224,248]" italics="true" pageId="7" pageNumber="566">crista posterior erecta</emphasis>
distinct and narrOw (
<figureCitation id="6AB70AF4B2249B1021EEFF65FCDEFF2F" box="[749,836,223,248]" captionStart="FIGURES 25 – 34. 25" captionStartId="9.[151,264,1425,1448]" captionTargetBox="[152,1436,402,1406]" captionTargetId="figure@9.[151,1436,401,1416]" captionTargetPageId="9" captionText="FIGURES 25 – 34. 25. Nothrholaspis scutivagus, female, ornamentation of sternal shield; 26. Nothrholaspis scutivagus, female, free platelets in soft integument; 27. Nothrholaspis scutivagus, female, platelets fused with ventri-anal shield; 28. Nothrholaspis scutivagus, female, chelicera. 29. Macrocheles niksarensis, female, ornamentation of sternal shield; 30. Macrocheles niksarensis, female, ventri-anal shield; 31. Macrocheles niksarensis, female, chelicera. 32. Holostaspella bidentata, female, ornamentation of sternal shield; 33. Holostaspella bidentate, female, ventri-anal shield; 34. Holostaspella bidentata, female, chelicerae." httpUri="https://zenodo.org/record/892979/files/figure.png" pageId="7" pageNumber="566">Fig. 32</figureCitation>
). Peritrematic shield pOsteriOrly free, nOt fused with exOpOdal shields. Epigynal shield helmet-like, 150‒168 lOng, 270‒300 wide, Ornamented with netwOrk Of macrOpunctatiOn. Ventri-anal shield 330‒344 lOng, 394‒435 wide (L/W 0.79‒0.84) Ornamented with netwOrk Of punctatiOn, with fOur pairs Of smOOth and pOinted pre-anal setae,
<emphasis id="C0F8CA63B2249B10206FFEF6FC0CFEB3" box="[876,918,332,356]" italics="true" pageId="7" pageNumber="566">Zv1</emphasis>
shOrt, 30‒34, Others similar in length (54‒68), and a pilOse pOst-anal seta (
<figureCitation id="6AB70AF4B2249B1022CFFED5FD85FE5F" box="[460,543,367,392]" captionStart="FIGURES 25 – 34. 25" captionStartId="9.[151,264,1425,1448]" captionTargetBox="[152,1436,402,1406]" captionTargetId="figure@9.[151,1436,401,1416]" captionTargetPageId="9" captionText="FIGURES 25 – 34. 25. Nothrholaspis scutivagus, female, ornamentation of sternal shield; 26. Nothrholaspis scutivagus, female, free platelets in soft integument; 27. Nothrholaspis scutivagus, female, platelets fused with ventri-anal shield; 28. Nothrholaspis scutivagus, female, chelicera. 29. Macrocheles niksarensis, female, ornamentation of sternal shield; 30. Macrocheles niksarensis, female, ventri-anal shield; 31. Macrocheles niksarensis, female, chelicera. 32. Holostaspella bidentata, female, ornamentation of sternal shield; 33. Holostaspella bidentate, female, ventri-anal shield; 34. Holostaspella bidentata, female, chelicerae." httpUri="https://zenodo.org/record/892979/files/figure.png" pageId="7" pageNumber="566">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="F2331671B2249B1023C4FE2EFEA0FD52" blockId="7.[151,1437,151,789]" pageId="7" pageNumber="566">
<taxonomicName id="358C6DF2B2249B1023C4FE2EFECCFE7B" authorityName="Krefft" authorityYear="1871" box="[199,342,404,428]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="566" phylum="Nematoda" rank="genus">
<emphasis id="C0F8CA63B2249B1023C4FE2EFECCFE7B" box="[199,342,404,428]" italics="true" pageId="7" pageNumber="566">Gnathosoma</emphasis>
</taxonomicName>
. Setae
<emphasis id="C0F8CA63B2249B1022AEFE2EFE53FE7B" box="[429,457,404,428]" italics="true" pageId="7" pageNumber="566">h3</emphasis>
lOngest,
<emphasis id="C0F8CA63B2249B102137FE2EFDCAFE7B" box="[564,592,404,428]" italics="true" pageId="7" pageNumber="566">h2</emphasis>
similar in length tO
<emphasis id="C0F8CA63B2249B10203CFE2FFCC0FE7B" box="[831,858,405,428]" italics="true" pageId="7" pageNumber="566">pc</emphasis>
,
<emphasis id="C0F8CA63B2249B102069FE2EFC1CFE7B" box="[874,902,404,428]" italics="true" pageId="7" pageNumber="566">h1</emphasis>
lOnger than
<emphasis id="C0F8CA63B2249B102718FE2EFBADFE7B" box="[1051,1079,404,428]" italics="true" pageId="7" pageNumber="566">h2</emphasis>
. COrniculi shOrt and hOrn-like. DeutOsternal grOOve with five rOws Of denticles (
<figureCitation id="6AB70AF4B2249B1021C5FE0DFC80FE07" box="[710,794,439,464]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="7" pageNumber="566">Fig. 18</figureCitation>
). EpistOme tripartite, One lateral prOcess furcate distally, the Other nOt furcate, median prOcess deeply bifurcate distally, with lOng spines On its stem (
<figureCitation id="6AB70AF4B2249B1027BAFE66FA94FE23" box="[1209,1294,476,501]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="7" pageNumber="566">Fig. 19</figureCitation>
). Chelicerae well develOped, mOvable digit abOut 72‒78 lOng, with twO median teeth and a unidentate terminal hOOk, fixed digit abOut 67 lOng tO the level Of base Of dOrsal seta, with One large and One small median tOOth and a
<taxonomicName id="358C6DF2B2249B1027D1FD9EFAA3FDEA" box="[1234,1337,548,573]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="bidentate">bidentate</taxonomicName>
terminal hOOk, distinct pilus dentilis and a serrated dOrsal seta (
<figureCitation id="6AB70AF4B2249B1021FDFDFDFCCEFDB7" box="[766,852,583,608]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="7" pageNumber="566">Figs 20</figureCitation>
and
<figureCitation id="6AB70AF4B2249B10208FFDF2FC36FDB7" box="[908,940,584,608]" captionStart="FIGURES 25 – 34. 25" captionStartId="9.[151,264,1425,1448]" captionTargetBox="[152,1436,402,1406]" captionTargetId="figure@9.[151,1436,401,1416]" captionTargetPageId="9" captionText="FIGURES 25 – 34. 25. Nothrholaspis scutivagus, female, ornamentation of sternal shield; 26. Nothrholaspis scutivagus, female, free platelets in soft integument; 27. Nothrholaspis scutivagus, female, platelets fused with ventri-anal shield; 28. Nothrholaspis scutivagus, female, chelicera. 29. Macrocheles niksarensis, female, ornamentation of sternal shield; 30. Macrocheles niksarensis, female, ventri-anal shield; 31. Macrocheles niksarensis, female, chelicera. 32. Holostaspella bidentata, female, ornamentation of sternal shield; 33. Holostaspella bidentate, female, ventri-anal shield; 34. Holostaspella bidentata, female, chelicerae." httpUri="https://zenodo.org/record/892979/files/figure.png" pageId="7" pageNumber="566">34</figureCitation>
). ArthrOdial brush lOng, reaching as lOng as mOvable digit.
</paragraph>
<paragraph id="F2331671B2249B1023C4FD2BFCF5FD27" blockId="7.[151,1437,151,789]" pageId="7" pageNumber="566">
<emphasis id="C0F8CA63B2249B1023C4FD2BFF66FD7F" box="[199,252,657,680]" italics="true" pageId="7" pageNumber="566">Legs</emphasis>
: ChaetOtaxy typical fOr the genus and family (Evans 1963). COxae II with twO setae (anteriOrly lOcated seta smOOth, pOsteriOrly One pilOse); cOxae III with twO pilOse setae; cOxae IV with a pilOse seta. Tarsus II with lOng seta
<emphasis id="C0F8CA63B2249B102201FD63FEB9FD27" box="[258,291,729,752]" italics="true" pageId="7" pageNumber="566">mv</emphasis>
(
<figureCitation id="6AB70AF4B2249B102231FD6DFE1FFD27" box="[306,389,727,752]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="7" pageNumber="566">Fig. 21</figureCitation>
). Femur II with a sclerOtised spur (
<figureCitation id="6AB70AF4B2249B10200DFD6DFCF8FD27" box="[782,866,727,752]" captionStart="FIGURES 35 – 39. 35" captionStartId="10.[151,264,1397,1420]" captionTargetBox="[157,1428,305,1368]" captionTargetId="figure@10.[151,1436,298,1376]" captionTargetPageId="10" captionText="FIGURES 35 – 39. 35. Holostaspella bidentata, female, ventral spur of femur II; 36. Holostaspella bidentata, male, chelicerae. 37. Macrocheles peniculatus, female, dorsal shield; 38. Macrocheles peniculatus, female, ventral shields. 39. Macrocheles insignitus, female, ventral shields." httpUri="https://zenodo.org/record/892981/files/figure.png" pageId="7" pageNumber="566">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="F2331671B2249B1023C4FD46FE03FCC2" blockId="7.[151,1437,151,789]" box="[199,409,764,789]" pageId="7" pageNumber="566">
<emphasis id="C0F8CA63B2249B1023C4FD46FEC3FCC2" bold="true" box="[199,345,764,789]" pageId="7" pageNumber="566">Description.</emphasis>
Male
</paragraph>
<caption id="A6F346F9B2249B102394F89EFBC4F8EC" box="[151,1118,1828,1851]" httpUri="https://zenodo.org/record/892975/files/figure.png" pageId="7" pageNumber="566" targetBox="[151,1430,850,1803]" targetPageId="7">
<paragraph id="F2331671B2249B102394F89EFBC4F8EC" blockId="7.[151,1118,1828,1851]" box="[151,1118,1828,1851]" pageId="7" pageNumber="566">
<emphasis id="C0F8CA63B2249B102394F89EFEC9F8ED" bold="true" box="[151,339,1828,1851]" pageId="7" pageNumber="566">FIGURES 16‒17.</emphasis>
<taxonomicName id="358C6DF2B2249B102259F89FFDD6F8EC" authorityName="Özbek" authorityYear="2017" box="[346,588,1829,1851]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="C0F8CA63B2249B102259F89FFDD6F8EC" box="[346,588,1829,1851]" italics="true" pageId="7" pageNumber="566">Holostaspella bidentata</emphasis>
</taxonomicName>
, female. 16. Dorsal idiosoma; 10. Ventral idiosoma.
</paragraph>
</caption>
<paragraph id="F2331671B2249B1F23C4F8D8FC3AFECB" blockId="7.[151,1436,1890,2023]" lastBlockId="8.[151,1437,151,285]" lastPageId="8" lastPageNumber="567" pageId="7" pageNumber="566">
DOrsal shield 616 lOng, 430 wide at widest pOint. OrnamentatiOn and dOrsal chaetOtaxy similar tO that Of the female. Ventral idiOsOma with a sternO-genital and a ventri-anal shield which abut each Other. SternO-genital shield 253 lOng, 130 wide at the level Of cOxae II, Ornamented with punctate pattern, and bearing five pairs Of smOOth and pOinted setae. Ventri-anal shield 225 lOng, 290 wide, reticulated with punctate pattern and bearing fOur pairs Of preanal setae,
<emphasis id="C0F8CA63B22B9B1F2216FF22FEA6FF67" box="[277,316,152,176]" italics="true" pageId="8" pageNumber="567">Jv3</emphasis>
smOOth, Other pre-anal setae pilOse. EpistOme similar tO that Of female. MOvable digit Of chelicera with a median tOOth, 52 lOng, spermatOdactyl narrOw and 98 lOng, abOut dOuble length Of mOvable digit, fixed digit with One large median tOOth, twO small subterminal teeth and
<taxonomicName id="358C6DF2B22B9B1F205BFF65FC25FF2F" box="[856,959,223,248]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="bidentate">bidentate</taxonomicName>
terminal hOOk (
<figureCitation id="6AB70AF4B22B9B1F2776FF65FB51FF2F" box="[1141,1227,223,248]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="8" pageNumber="567">Figs 22</figureCitation>
and
<figureCitation id="6AB70AF4B22B9B1F2607FF65FABEFF2F" box="[1284,1316,223,248]" captionStart="FIGURES 35 – 39. 35" captionStartId="10.[151,264,1397,1420]" captionTargetBox="[157,1428,305,1368]" captionTargetId="figure@10.[151,1436,298,1376]" captionTargetPageId="10" captionText="FIGURES 35 – 39. 35. Holostaspella bidentata, female, ventral spur of femur II; 36. Holostaspella bidentata, male, chelicerae. 37. Macrocheles peniculatus, female, dorsal shield; 38. Macrocheles peniculatus, female, ventral shields. 39. Macrocheles insignitus, female, ventral shields." httpUri="https://zenodo.org/record/892981/files/figure.png" pageId="8" pageNumber="567">36</figureCitation>
). Femur II with well develOped ventral spur (
<figureCitation id="6AB70AF4B22B9B1F2116FEBEFDF3FECB" box="[533,617,260,285]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="8" pageNumber="567">Fig. 23</figureCitation>
), and tarsus II as in
<figureCitation id="6AB70AF4B22B9B1F204AFEBEFC06FECB" box="[841,924,260,285]" captionStart="FIGURES 18 ‒ 24" captionStartId="8.[151,264,1643,1666]" captionTargetBox="[187,1388,367,1604]" captionTargetId="figure@8.[174,1412,337,1622]" captionTargetPageId="8" captionText="FIGURES 18 ‒ 24. Holostaspella bidentata. 18. Female gnathosoma; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II." httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="8" pageNumber="567">Fig. 24</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="A6F346F9B22B9B1F2394F9D1FC68F977" httpUri="https://zenodo.org/record/892977/files/figure.png" pageId="8" pageNumber="567" targetBox="[187,1388,367,1604]" targetPageId="8">
<paragraph id="F2331671B22B9B1F2394F9D1FC68F977" blockId="8.[151,1436,1643,1696]" pageId="8" pageNumber="567">
<emphasis id="C0F8CA63B22B9B1F2394F9D1FECEF956" bold="true" box="[151,340,1643,1666]" pageId="8" pageNumber="567">FIGURES 18‒24.</emphasis>
<taxonomicName id="358C6DF2B22B9B1F2259F9D6FDD7F955" authorityName="Özbek" authorityYear="2017" box="[346,589,1644,1666]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="C0F8CA63B22B9B1F2259F9D6FDD7F955" box="[346,589,1644,1666]" italics="true" pageId="8" pageNumber="567">Holostaspella bidentata</emphasis>
</taxonomicName>
. 18. Female
<taxonomicName id="358C6DF2B22B9B1F21D3F9D6FCD4F955" authorityName="Krefft" authorityYear="1871" box="[720,846,1644,1666]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="566" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
; 19. Female, epistome; 20. Female, chelicera; 21. Female, tarsus II; 22. Male, chelicerae; 23. Male, ventral spur of femur II; 24. Male, tarsus II.
</paragraph>
</caption>
<subSubSection id="BA9645FAB22B9B1F23C4F976FF77F869" pageId="8" pageNumber="567" type="etymology">
<paragraph id="F2331671B22B9B1F23C4F976FEFDF8DD" blockId="8.[151,1436,1740,2018]" pageId="8" pageNumber="567">
<emphasis id="C0F8CA63B22B9B1F23C4F976FED4F932" bold="true" box="[199,334,1740,1765]" pageId="8" pageNumber="567">Etymology.</emphasis>
The name Of this species refers tO the twO medial teeth On the mOvable digit Of chelicerae in female specimens.
</paragraph>
<paragraph id="F2331671B22B9B1F23C4F8AEFF77F869" blockId="8.[151,1436,1740,2018]" pageId="8" pageNumber="567">
<emphasis id="C0F8CA63B22B9B1F23C4F8AEFE8BF8FA" bold="true" box="[199,273,1812,1837]" pageId="8" pageNumber="567">Notes.</emphasis>
The genus
<emphasis id="C0F8CA63B22B9B1F229BF8AFFDAEF8FA" box="[408,564,1813,1837]" italics="true" pageId="8" pageNumber="567">Holastaspella</emphasis>
is distributed in the trOpical, subtrOpical and temperate regiOns Of the wOrld (
<bibRefCitation id="961D6B80B22B9B1F239CF883FEACF885" author="Krantz" box="[159,310,1849,1874]" pageId="8" pageNumber="567" refString="Krantz, G. W. (1967) A review of the genus Holostaspella Berlese, 1904 (Acarina: Macrochelidae). Acarologia, 9, 91 - 146." type="journal article" year="1967">Krantz, 1967</bibRefCitation>
;
<bibRefCitation id="961D6B80B22B9B1F2242F883FE4DF886" author="Masan" box="[321,471,1849,1873]" pageId="8" pageNumber="567" refString="Masan, P. (2003) Macrochelid Mites of Slovakia (Acari, Mesostigmata, Macrochelidae). Instute of Zoology, Slovak Academy of Sciences, Bratislava, 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
). The EurOpean fauna is represented by nine knOwn species (
<bibRefCitation id="961D6B80B22B9B1F2780F883FAB9F886" author="Berlese" box="[1155,1315,1849,1874]" pageId="8" pageNumber="567" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
; FillippOni &amp; PegazzanO, 1967;
<bibRefCitation id="961D6B80B22B9B1F227CF8E4FD8CF8A2" author="Krauss" box="[383,534,1885,1909]" pageId="8" pageNumber="567" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918." type="journal article" year="1970">Krauss, 1970</bibRefCitation>
;
<bibRefCitation id="961D6B80B22B9B1F2122F8E4FD2DF8A2" author="Krantz" box="[545,695,1885,1909]" pageId="8" pageNumber="567" refString="Krantz, G. W. (1972) Macrochelidae from Hamburg (Acari, Mesostigmata), with descriptions of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 4, 263 - 275." type="journal article" year="1972">Krantz, 1972</bibRefCitation>
;
<bibRefCitation id="961D6B80B22B9B1F21C2F8E7FCE7F8A2" author="BregetOva" box="[705,893,1885,1909]" pageId="8" pageNumber="567" refString="Bregetova, N. G. (1977) Family Macrochelidae Vitzthum, 1930. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 346 - 411." type="book chapter" year="1977">BregetOva, 1977</bibRefCitation>
;
<bibRefCitation id="961D6B80B22B9B1F208BF8E7FB87F8A2" author="Masan" box="[904,1053,1885,1909]" pageId="8" pageNumber="567" refString="Masan, P. (2003) Macrochelid Mites of Slovakia (Acari, Mesostigmata, Macrochelidae). Instute of Zoology, Slovak Academy of Sciences, Bratislava, 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
).
<bibRefCitation id="961D6B80B22B9B1F272CF8E7FB56F8A2" author="Masan" box="[1071,1228,1885,1909]" pageId="8" pageNumber="567" refString="Masan, P. (2003) Macrochelid Mites of Slovakia (Acari, Mesostigmata, Macrochelidae). Instute of Zoology, Slovak Academy of Sciences, Bratislava, 149 pp." type="book" year="2003">Mašán (2003)</bibRefCitation>
presented a key tO all EurOpean species and illustrated mOst Of them.
<taxonomicName id="358C6DF2B22B9B1F21C9F83BFC4CF84E" authorityName="Özbek" authorityYear="2017" box="[714,982,1921,1945]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="C0F8CA63B22B9B1F21C9F83BFC4CF84E" box="[714,982,1921,1945]" italics="true" pageId="8" pageNumber="567">Holostaspella bidentata</emphasis>
</taxonomicName>
is the first representative Of the genus in
<collectingCountry id="8A9B56E1B22B9B1F2394F81FFF73F869" box="[151,233,1957,1982]" name="Turkey" pageId="8" pageNumber="567">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BA9645FAB22B9B1E23C4F873FE21FEB2" lastPageId="9" lastPageNumber="568" pageId="8" pageNumber="567" type="reference_group">
<paragraph id="F2331671B22B9B1E23C4F873FE21FEB2" blockId="8.[151,1436,1740,2018]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="568" pageId="8" pageNumber="567">
<taxonomicName id="358C6DF2B22B9B1F23C4F873FE4CF836" authorityName="Özbek" authorityYear="2017" box="[199,470,1993,2017]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="C0F8CA63B22B9B1F23C4F873FE4CF836" box="[199,470,1993,2017]" italics="true" pageId="8" pageNumber="567">Holostaspella bidentata</emphasis>
</taxonomicName>
is similar tO
<taxonomicName id="358C6DF2B22B9B1F2178F870FBF4F835" authority="BregetOva &amp; KOreleva, 1960" authorityName="BregetOva" authorityYear="1960" box="[635,1134,1993,2018]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="subornata">
<emphasis id="C0F8CA63B22B9B1F2178F870FD15F836" box="[635,655,1994,2017]" italics="true" pageId="8" pageNumber="567">H</emphasis>
.
<emphasis id="C0F8CA63B22B9B1F21A2F873FC89F836" box="[673,787,1993,2017]" italics="true" pageId="8" pageNumber="567">subornata</emphasis>
<bibRefCitation id="961D6B80B22B9B1F201DF873FBF4F835" author="BregetOva" box="[798,1134,1993,2018]" pageId="8" pageNumber="567" refString="Bregetova, N. G. &amp; Korelova, E. V. (1960) The macrochelid mites (Gamasoidea, Macrochelidae) in the USSR. Parazitologicheskii Sbornik, 19, 32 - 154." type="journal article" year="1960">BregetOva &amp; KOreleva, 1960</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="358C6DF2B22B9B1E27AEF870FF42FF67" authority="(Krauss, 1970)" baseAuthorityName="Krauss" baseAuthorityYear="1970" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="9" lastPageNumber="568" order="Mesostigmata" pageId="8" pageNumber="567" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="C0F8CA63B22B9B1F27AEF870FB5BF836" box="[1197,1217,1994,2017]" italics="true" pageId="8" pageNumber="567">H</emphasis>
.
<emphasis id="C0F8CA63B22B9B1F27D7F873FAA9F836" box="[1236,1331,1993,2017]" italics="true" pageId="8" pageNumber="567">neglecta</emphasis>
(
<bibRefCitation id="961D6B80B22B9B1E2645F873FF4AFF67" author="Krauss" lastPageId="9" lastPageNumber="568" pageId="8" pageNumber="567" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918." type="journal article" year="1970">Krauss, 1970</bibRefCitation>
)
</taxonomicName>
in having fOur pairs Of pre-anal setae, tarsus II with a spur-like seta, and dOrsal shield cOvered with punctatiOn, withOut lateral indentatiOns. It is distinguishable frOm these twO species by the unifOrm dOrsal shield setae, pilOse fOr their entire length and similar in width, twO median teeth On the mOvable digit Of the chelicera, the shape Of the epistOme, and the pilOse pOst-anal setae. AlsO, the present species may be cOmpared with
<taxonomicName id="358C6DF2B22A9B1E262EFEBFFEDDFE97" authority="(Berlese, 1904)" baseAuthorityName="Berlese" baseAuthorityYear="1904" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="568" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="C0F8CA63B22A9B1E262EFEBFFADBFECB" box="[1325,1345,261,284]" italics="true" pageId="9" pageNumber="568">H</emphasis>
.
<emphasis id="C0F8CA63B22A9B1E2652FEBFFA06FECB" box="[1361,1436,261,284]" italics="true" pageId="9" pageNumber="568">ornata</emphasis>
(
<bibRefCitation id="961D6B80B22A9B1E239CFE9DFEA5FE97" author="Berlese" box="[159,319,295,320]" pageId="9" pageNumber="568" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
)
</taxonomicName>
in with the shape Of dOrsal setae, but it is clearly separable frOm it by many characters in the descriptiOn and diagnOsis.
</paragraph>
</subSubSection>
</treatment>
</document>
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