139 lines
13 KiB
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139 lines
13 KiB
XML
<document id="AD57EF3CE18934ABC42A0D17A504C89A" ID-CLB-Dataset="34819" ID-DOI="http://dx.doi.org/10.3897/zookeys.113.1353" ID-GBIF-Dataset="953ba201-d189-4d18-b7cb-dfceb46900a3" ID-PMC="PMC3187627" ID-Pensoft-Pub="1313-2970-113-39" ID-PubMed="21976992" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-113-39" ModsDocOrigin="ZooKeys 113" ModsDocTitle="The dazed and confused identity of Agassiz’s land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation" checkinTime="1451250228616" checkinUser="pensoft" docAuthor="Murphy, Robert W., Berry, Kristin H., Edwards, Taylor, Leviton, Alan E., Lathrop, Amy & Riedle, J. Daren" docDate="2011" docId="6E53861C67BFC80390CBF48B0B2984EF" docLanguage="en" docName="ZooKeys 113: 39-71" docOrigin="ZooKeys 113" docSource="http://dx.doi.org/10.3897/zookeys.113.1353" docTitle="Gopherus agassizii Cooper 1861" docType="treatment" docVersion="5" lastPageNumber="49" masterDocId="FFFB155FD763FF9D2C30FF8882076558" masterDocTitle="The dazed and confused identity of Agassiz's land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation" masterLastPageNumber="71" masterPageNumber="39" pageNumber="48" updateTime="1732854536317" updateUser="ExternalLinkService">
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<mods:title id="C7DF87FAFADD1F54128B62D094782004">The dazed and confused identity of Agassiz's land tortoise, Gopherus agassizii (Testudines, Testudinidae) with the description of a new species, and its consequences for conservation</mods:title>
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<mods:namePart id="84C0CA6B3DF1557CBC2E82EDC11C5AE0">Murphy, Robert W.</mods:namePart>
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<mods:namePart id="C4356EA058C9CDDCE768250F550B681A">Berry, Kristin H.</mods:namePart>
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<mods:namePart id="45B2CCD1961F0E03B5CF5F1CCF316A8D">Edwards, Taylor</mods:namePart>
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<mods:namePart id="939725B131093E42F05B182BBEA599D7">Leviton, Alan E.</mods:namePart>
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<mods:namePart id="5861D4B73A7AAE043DBAD56C3E50725C">Lathrop, Amy</mods:namePart>
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<mods:namePart id="34DF6F6A67C58D2CE9A3440DE98A6A34">Riedle, J. Daren</mods:namePart>
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<mods:title id="975F51A94A1E5A6BBD27002AC9C0AB32">ZooKeys</mods:title>
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<mods:date id="0849567B5799AF45CFC1A36B22BC2949">2011</mods:date>
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<mods:detail id="5DE4089611C5E11B8B9A51B3037A1861" type="volume">
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<mods:number id="13E416F46F1D869D680318B5BD1651D5">113</mods:number>
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<mods:start id="29D6E25CF1FCC60454A0619E511F4497">39</mods:start>
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<mods:end id="D59B0FEA7E72B5CA1208995D9E197334">71</mods:end>
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<mods:url id="403D3944FDA2F9A7C1D79BBF19EAB610">http://dx.doi.org/10.3897/zookeys.113.1353</mods:url>
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<mods:classification id="95029CEC35A4934E62AECA28D97234E2">journal article</mods:classification>
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<mods:identifier id="6FA32993A040A136367B6E77A330D36B" type="DOI">http://dx.doi.org/10.3897/zookeys.113.1353</mods:identifier>
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<mods:identifier id="FFB7499AC763C12EF2FB1B8034031C4A" type="Pensoft-Pub">1313-2970-113-39</mods:identifier>
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<treatment id="6E53861C67BFC80390CBF48B0B2984EF" ID-GBIF-Taxon="152030064" LSID="urn:lsid:plazi:treatment:6E53861C67BFC80390CBF48B0B2984EF" httpUri="http://treatment.plazi.org/id/6E53861C67BFC80390CBF48B0B2984EF" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" scope_class="Testudines" scope_family="Testudinidae">
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<subSubSection id="4E5D3B34DFE36F07B590D3D4272878C7" pageId="9" pageNumber="48" type="nomenclature">
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<paragraph id="6F0BF851B9F771AD85DE6921DF990C7F" pageId="9" pageNumber="48">
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<normalizedToken id="FD11F63C2F4E0CFA76DE4CBB953F5D08" originalValue="Agassiz’s">Agassiz's</normalizedToken>
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Desert Tortoise
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<taxonomicName id="BEF89FD4D5298ED05E95AEF7FCF1A685" ID-CoL="3H2LD" LSID="http://species-id.net/wiki/Gopherus_agassizii" authority="Cooper, 1861" authorityName="Cooper" authorityYear="1861" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="9" pageNumber="48" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii (Cooper, 1861)</taxonomicName>
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Figs 48
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</paragraph>
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</subSubSection>
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<subSubSection id="6F5BF5AFE2E43119BC5DB01EFD52341F" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="lectotype">
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<paragraph id="C45EDF1CE48315D8CDE8308724A12AB9" pageId="9" pageNumber="48">Lectotype.</paragraph>
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<paragraph id="742CFA8EA8A699FE63F937C365192A24" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
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USNM (National Museum of Natural History, Smithsonian Institution) 7888; terra typica restricta: California, San Bernardino County; Mountains of California, near Fort Mojave; Soda Valley (very approximately
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<geoCoordinate id="AEC8A65EEA759DEC27986D78D492811A" direction="north" orientation="latitude" precision="925" value="35.1">35°6'N</geoCoordinate>
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,
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<geoCoordinate id="0762CFD05742325631B06AB8454C8BD7" direction="west" orientation="longitude" precision="925" value="-116.1">116°6'W</geoCoordinate>
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). We restrict the type locality to that published by
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<bibRefCitation id="E9D8C85734B216E313D8B8FF4D9958BF" author="Cooper, JG" journalOrPublisher="1)" pageId="21" pageNumber="60" pagination="118 - 123" title="New Californian animals. Proceedings of the California Academy of Sciences (ser." volume="2" year="1861">Cooper (1861)</bibRefCitation>
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and the parchment tag
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<pageBreakToken id="270F40CDE59759B793AD74AD8F43E9F7" pageId="10" pageNumber="49" start="start">associated</pageBreakToken>
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with the specimen. References to other localities, including Solado Valley, an apparent synonym of Soda Valley, should be considered to be in error.
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</paragraph>
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<paragraph id="FF2F302720B2CF10B8BA4B7D9584509A" pageId="10" pageNumber="49">
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The evolutionary species concept (
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<bibRefCitation id="68063672B85AEB9CF02D6833799FEABE" author="Simpson, GG" journalOrPublisher="Columbia University Press, New York" pageId="21" pageNumber="60" title="Principles of Animal Taxonomy" year="1961">Simpson 1961</bibRefCitation>
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;
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<bibRefCitation id="8329EA12D1028AB70DD18022A625237A" author="Wiley, EO" journalOrPublisher="Systematic Biology" pageId="21" pageNumber="60" pagination="17 - 26" title="The evolutionary species concept reconsidered" volume="27" year="1978">Wiley 1978</bibRefCitation>
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) suggests that the Sonoran population of the desert tortoise should be recognized as a new taxon.
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<bibRefCitation id="778597D48448880DABFC3228CC00701A" author="Frost, DR" journalOrPublisher="Herpetologica" pageId="21" pageNumber="60" pagination="87 - 104" title="Species in concept and practice: herpetological applications." volume="46" year="1990">Frost and Hillis (1990)</bibRefCitation>
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effectively argue that subspecies should not be recognized for continuously distributed species; we agree. Given these two observations, at least two species of desert tortoise should be recognized. The DNA sequence data exclude application of the available name
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<taxonomicName id="08D8741AAD3D479593E365487332321B" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus lepidocephalus" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="lepidocephalus">Gopherus lepidocephalus</taxonomicName>
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for the Sonoran Desert population of
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<taxonomicName id="522FAA1892DD68D4336FB3C97C4A5B03" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="genus">Gopherus</taxonomicName>
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that occurs west and south of the Colorado River and they confirm that the lectotype of
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<taxonomicName id="4849F62865C22287657B2FF9278FEF08" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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is from the Mojave Desert, and not Arizona. Because no names are available for the tortoise population occurring in the Sonoran Desert south and east of the Colorado River, we describe it as a new species.
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</paragraph>
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<caption id="E8FBB82CC602038A11900F7B76D8C390" pageId="10" pageNumber="49">
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<paragraph id="3C78AB0BC6441628680E0A29FF1F167B" pageId="10" pageNumber="49">
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Figure 4. Dorsal view of the lectotype of
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<taxonomicName id="8D9EAE3D2112A2B0BFCB798D7EC5DB4F" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption id="7E2F4E6D954F0E685A2985EECD89DB2E" pageId="10" pageNumber="49">
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<paragraph id="DBF9FA319EEE93CC9298EDEDFB3504A1" pageId="10" pageNumber="49">
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Figure 5. Ventral view of the holotype of
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<taxonomicName id="5BFA0FCF00D5D1283E9835BBB202883B" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption id="A07A3121D4FB6A7F43234AE231A49F10" pageId="10" pageNumber="49">
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<paragraph id="2F0C8A8317B53BD1A493E99D781A17F7" pageId="10" pageNumber="49">
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Figure 6. Anterior view of the holotype of
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<taxonomicName id="3B91F9D7E336D8A395F850CB6D2808AD" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888. Black bar is 3 cm.
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</paragraph>
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</caption>
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<caption id="6D2842A5D04A55F24CE4117025853FC5" pageId="10" pageNumber="49">
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<paragraph id="0212BC4231399B629EAB06D192C3546A" pageId="10" pageNumber="49">
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Figure 7. Posterior view of the holotype of
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<taxonomicName id="F78D27171BEFB8D7AC14C41B34E370F9" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888.
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</paragraph>
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</caption>
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<caption id="1AA295F47845FD51208404A7A2A73570" pageId="10" pageNumber="49">
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<paragraph id="26E2987A878ECC26C995EBCCFD8D4EAB" pageId="10" pageNumber="49">
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Figure 8. Left, lateral view of the holotype of
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<taxonomicName id="5821139B9A4D6D8B8C292144A6859F7D" class="Testudines" family="Testudinidae" genus="Gopherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gopherus agassizii" order="Cryptodira" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="agassizii">Gopherus agassizii</taxonomicName>
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, USNM 7888.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |