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<document id="A7C65029F469116584E2113C1693EC5B" ID-DOI="10.5281/zenodo.183029" ID-GBIF-Dataset="41d158a9-22d2-48c9-aae2-b0a6b2d09266" ID-ISSN="1175-5326" ID-Zenodo-Dep="183029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459888806107" checkinUser="plazi" docAuthor="Bruce, Niel L." docDate="2008" docId="6949084EFF8AD669868BFEE4FE7067D7" docLanguage="en" docName="zt01823p064.pdf" docOrigin="Zootaxa 1823" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Kagalana Bruce, 2008, gen. nov." docType="treatment" docVersion="8" lastPageNumber="53" masterDocId="95707036FF8BD66B861CFF83FFDB642D" masterDocTitle="New species and a new genus of Cirolanidae (Isopoda: Cymothoida: Crustacea) from groundwater in calcretes in the Pilbarra, northern Western Australia" masterLastPageNumber="64" masterPageNumber="51" pageNumber="52" updateTime="1698227922467" updateUser="plazi">
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<mods:title id="829E2AD64119DA8CDA6E8B5C52FB0DCD">New species and a new genus of Cirolanidae (Isopoda: Cymothoida: Crustacea) from groundwater in calcretes in the Pilbarra, northern Western Australia</mods:title>
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<mods:namePart id="3480F1F7DB8018CECF150C14E3E1A83F">Bruce, Niel L.</mods:namePart>
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<mods:date id="87238016CEFEB6DA1C7E3D2ABD3D94A0">2008</mods:date>
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<treatment id="6949084EFF8AD669868BFEE4FE7067D7" ID-DOI="http://doi.org/10.5281/zenodo.6228262" ID-GBIF-Taxon="119371759" ID-Zenodo-Dep="6228262" LSID="urn:lsid:plazi:treatment:6949084EFF8AD669868BFEE4FE7067D7" httpUri="http://treatment.plazi.org/id/6949084EFF8AD669868BFEE4FE7067D7" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<subSubSection id="A9FAEAD3FF8AD66A868BFEE4FE5A65AC" box="[151,385,359,385]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="E15FB958FF8AD66A868BFEE4FE5A65AC" blockId="1.[151,385,359,385]" box="[151,385,359,385]" pageId="1" pageNumber="52">
<heading id="BA170E34FF8AD66A868BFEE4FE5A65AC" bold="true" box="[151,385,359,385]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="D394654AFF8AD66A868BFEE4FE5A65AC" bold="true" box="[151,385,359,385]" pageId="1" pageNumber="52">
<taxonomicName id="26E0C2DBFF8AD66A868BFEE4FED765AC" ID-CoL="7PLYS" box="[151,268,359,385]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D394654AFF8AD66A868BFEE4FED765AC" bold="true" box="[151,268,359,385]" italics="true" pageId="1" pageNumber="52">Kagalana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8A7D831FF8AD66A870FFEE4FE5A65AC" box="[275,385,359,385]" pageId="1" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF8AD66A868BFE31FCA165E4" box="[151,890,434,458]" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="E15FB958FF8AD66A868BFE31FCA165E4" blockId="1.[151,890,434,458]" box="[151,890,434,458]" pageId="1" pageNumber="52">
<typeStatus id="3E5B07FAFF8AD66A868BFE31FF1765E4" box="[151,204,434,457]" pageId="1" pageNumber="52">Type</typeStatus>
species:
<taxonomicName id="26E0C2DBFF8AD66A8731FE31FE0365E4" box="[301,472,434,457]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="tonde" status="sp. nov.">
<emphasis id="D394654AFF8AD66A8731FE31FE0365E4" box="[301,472,434,457]" italics="true" pageId="1" pageNumber="52">Kagalana tonde</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8A7D831FF8AD66A87C3FE31FDEF65E7" box="[479,564,434,458]" pageId="1" pageNumber="52" rank="species">
<emphasis id="D394654AFF8AD66A87C3FE31FDEF65E7" bold="true" box="[479,564,434,458]" pageId="1" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
, here designated (monotypic).
</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF8AD66A868BFE79FABD6701" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="E15FB958FF8AD66A868BFE79FABD6701" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">
<emphasis id="D394654AFF8AD66A868BFE79FECC6639" bold="true" box="[151,279,506,532]" pageId="1" pageNumber="52">Diagnosis.</emphasis>
Head with rostral point. Clypeus ventrally flat, not blade-like, not projecting. Pleonites 4 and 5 narrow, laterally overlapped by margins of pleonite 3 and pleonite 5 narrower than the pleotelson anterior margin; pleonite 1 and most of pleonite 2 not visible in dorsal view. Pereopod 1 robust, propodus expanded, dactylus haptorial; pereopods 27 ambulatory, slender. Pereopods 13 with ischium superior distal margins weakly produced, setose; merus superior distal margin weakly produced, not overriding propodus; pereopods 2 and 3 merus inferior margin RS not molariform; pereopods 57 basis without long PMS, ischium and merus distally without long setae. Penial processes present on sternite 7 as mutually adjacent papillae. Pleopod 1 endopod narrow, half as wide as exopod. Pleopod 2 appendix masculina basally inserted, apex sub-acute.
</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF8AD66A86DAFCB9FD9A6311" pageId="1" pageNumber="52" type="description">
<paragraph id="E15FB958FF8AD66A86DAFCB9FD6F67D9" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">
<emphasis id="D394654AFF8AD66A86DAFCB9FE856779" bold="true" box="[198,350,826,852]" pageId="1" pageNumber="52">Description.</emphasis>
Head wide, approximately 75% as wide as pereonite 1, anterior margin evenly rounded. Body surfaces unornamented; pereonite 1 about 1.7 times as long as pereonite
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dorsal view. Pleon unornamented, about 9% BL, with 4 visible unfused segments, pleonite 1 and most of pleonite 2 largely concealed by pereonite 7; pleonite 2 epimera not produced;. Pleotelson without longitudinal carinae, ridges or without tubercles; posterior margin with PMS and RS.
</paragraph>
<paragraph id="E15FB958FF8AD66A86D9FB81FB9460B9" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">Antennule peduncle articles collinear, articles 1 and 2 fused; peduncular article 2 not at right angles to article 1; article 3 well developed, about as long as combined lengths of articles 1 and 2; flagellum shorter than peduncle; without callynophore. Antenna peduncle comprised of 5 articles, peduncular articles 13 shortest, articles 4 and 5 longest, 5 longer than 4; flagellum about as long as peduncle.</paragraph>
<paragraph id="E15FB958FF8AD66A86D9FB21FB7461A9" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">Frontal lamina lateral margins parallel; ventrally flat, without lateral angles, anteriorly rounded; posteriorly abutting clypeus; approximately 2 times as long as basal width, not projecting anteroventrally from posterior. Clypeus ventral surface not projecting relative to frontal lamina. Mandible incisors wide, right incisor tricuspidate; spine row with 45 RS. Maxillule mesial lobe with 3 CP RS. Maxilliped palp articles 3 and 4 mesial margin lobed; lateral margins of articles 25 with long setae; articles 3 and 4 distal margin width greater than proximal margin of article 4 and 5 respectively; endite with 1 coupling hook.</paragraph>
<paragraph id="E15FB958FF8AD66A86D9FA11FABF6209" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">Pereopods 17 dactylus with small secondary unguis present. Pereopod 1 dactylus longer than palm; complex RS opposing dactylus. Pereopod 7 basis not noticeably broader in distal half compared to proximal half; margins with few discontinuous setae; ischium and merus not flattened, distal margin weakly expanded, inferior margins with few setae; inferodistal angles of ischium, merus and carpus with low cuticular nodules.</paragraph>
<paragraph id="E15FB958FF8AD66A86D9F9B1FB1462E9" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">Pleopod 1 rami lamellar; endopod narrow, 3.5 times as long as wide. Pleopod 2 appendix masculina longer than endopod. Pleopods 15 with PMS present on all exopods and endopods of pleopods 1 and 2; endopods 35 without PMS, smaller (0.70.8) than exopods; endopod of pleopod 5 without proximomesial lobe. Uropod peduncle mesial margin strongly produced; exopod lateral margin not excised.</paragraph>
<paragraph id="E15FB958FF8AD66A86DAF951FD9A6311" blockId="1.[151,1437,506,2012]" pageId="1" pageNumber="52">Females. Ovigerous and non-ovigerous females similar to males but slightly larger. Brood pouch made up of 4 pairs of oostegites arising from coxae 25, the posterior pair being largest, extending over most of the marsupium (anteriorly to sternite 3).</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF8AD66986DAF8C9FB3D6647" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="discussion">
<paragraph id="E15FB958FF8AD66986DAF8C9FB72649F" blockId="1.[151,1437,506,2012]" lastBlockId="2.[151,1437,152,1018]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="D394654AFF8AD66A86DAF8C9FEE46349" bold="true" box="[198,319,1866,1892]" pageId="1" pageNumber="52">Remarks.</emphasis>
<taxonomicName id="26E0C2DBFF8AD66A8757F8C8FE646349" box="[331,447,1867,1892]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D394654AFF8AD66A8757F8C8FE646349" box="[331,447,1867,1892]" italics="true" pageId="1" pageNumber="52">Kagalana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C8A7D831FF8AD66A87D5F8C9FDE36349" box="[457,568,1866,1892]" pageId="1" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
is characterised by and can be identified by the typical cirolanid body shape (see
<bibRefCitation id="8571C4A9FF8AD66A8707F8F1FE7363A1" author="Bruce" box="[283,424,1906,1932]" pageId="1" pageNumber="64" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement, 6, 1 - 239." type="journal article" year="1986">Bruce 1986</bibRefCitation>
;
<bibRefCitation id="8571C4A9FF8AD66A87A9F8F1FD4863A1" author="Brusca" box="[437,659,1906,1932]" pageId="1" pageNumber="64" refString="Brusca, R. C., Wetzer, R. &amp; France, S. C. (1995) Cirolanidae (Crustacea: Isopoda: Flabellifera) of the tropical eastern Pacific. Proceedings of the San Diego Society of Natural History, 30, 1 - 96." type="journal article" year="1995">
Brusca
<emphasis id="D394654AFF8AD66A840CF8F0FD9E63A1" box="[528,581,1907,1932]" italics="true" pageId="1" pageNumber="52">et al</emphasis>
. 1995
</bibRefCitation>
), together with the antennule peduncle and flagellum being short (extending no further than the anterior of pereonite 1), pleonites 4 and 5 narrow, laterally overlapped by margins of pleonite 3 and pleonite 5 narrower than pleotelson anterior margin, pereopod 1 being robust and haptorial and the inferior margins of the merus of pereopods 2 and 3 lacking molariform setae.
</paragraph>
<paragraph id="E15FB958FF89D66986DAFF3CFB3D6647" blockId="2.[151,1437,152,1018]" pageId="2" pageNumber="53">
<emphasis id="D394654AFF89D66986DAFF3CFEAC64F4" bold="true" box="[198,375,191,217]" pageId="2" pageNumber="53">Relationships.</emphasis>
The most similar genus is
<taxonomicName id="26E0C2DBFF89D66984DEFF43FC6464F7" authority="Bolivar" authorityName="Bolivar" box="[706,959,192,218]" class="Malacostraca" family="Cirolanidae" genus="Speocirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66984DEFF43FC8664F4" box="[706,861,192,217]" italics="true" pageId="2" pageNumber="53">Speocirolana</emphasis>
Bolívar
</taxonomicName>
y Pieltain, 1950, a genus of 12 species endemic to
<collectingCountry id="99F7F9C8FF89D669873DFF6BFEA0652F" box="[289,379,232,258]" name="Mexico" pageId="2" pageNumber="53">Mexico</collectingCountry>
and Texas, which differs to
<taxonomicName id="26E0C2DBFF89D66984D0FF6BFC9B652C" box="[716,832,232,257]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66984D0FF6BFC9B652C" box="[716,832,232,257]" italics="true" pageId="2" pageNumber="53">Kagalana</emphasis>
</taxonomicName>
in the large size (adults
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) of the species, pleonites 4 and 5 subequal in width and both narrower than the pleotelson anterior margin, in having haptorial and robust pereopods 2 and 3 and prominent penial processes; the palm of pereopods 13 typically has several large robust setae, but there is no robust seta opposing the base of the dactylus.
<taxonomicName id="26E0C2DBFF89D669825AFEE3FB3A6554" box="[1094,1249,352,377]" class="Malacostraca" family="Cirolanidae" genus="Speocirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D669825AFEE3FB3A6554" box="[1094,1249,352,377]" italics="true" pageId="2" pageNumber="53">Speocirolana</emphasis>
</taxonomicName>
was first established as a subgenus of
<taxonomicName id="26E0C2DBFF89D66987A5FE0BFD1A658F" authority="Leach, 1818" authorityName="Leach" authorityYear="1818" box="[441,705,392,418]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66987A5FE0BFDF9658C" box="[441,546,392,417]" italics="true" pageId="2" pageNumber="53">Cirolana</emphasis>
Leach, 1818
</taxonomicName>
,
<bibRefCitation id="8571C4A9FF89D66984CCFE0BFC4A658F" author="Bowman" box="[720,913,392,418]" pageId="2" pageNumber="64" refString="Bowman, T. E. (1964) Antrolana lira, a new genus and species of troglobitic cirolanid isopod from Madison Cave, Virginia. International Journal of Speleology, 1, 229 - 236, pls 50 - 56." type="journal article" year="1964">Bowman (1964)</bibRefCitation>
later elevating it to genus on the basis that pereopods 13 were prehensile.
<bibRefCitation id="8571C4A9FF89D6698432FE33FD2B65E7" author="Bowman" box="[558,752,432,458]" pageId="2" pageNumber="64" refString="Bowman, T. E. (1964) Antrolana lira, a new genus and species of troglobitic cirolanid isopod from Madison Cave, Virginia. International Journal of Speleology, 1, 229 - 236, pls 50 - 56." type="journal article" year="1964">Bowman (1964)</bibRefCitation>
did not rediagnose the genus and, surprisingly given the number of described species, no revised diagnosis to the genus has since been given.
<bibRefCitation id="8571C4A9FF89D66982A6FE5BFF3E6637" lastPageId="2" lastPageNumber="64" pageId="2" pageNumber="63" refString="Botosaneanu, L., Iliffe, T. M. &amp; Hendrickson, D. A. (1998) On a collection of stygobitic cirolanids (Isopoda: Cirolanidae) from northern Mexico, with description of a new species. Bulletin de l'Institute Royale des Sciences Naturelles de Belgique, 68, 123 - 134." type="journal article">
Botosaneanu
<emphasis id="D394654AFF89D6698341FE5BFA4F65DC" box="[1373,1428,472,497]" italics="true" pageId="2" pageNumber="53">et al</emphasis>
. (1998)
</bibRefCitation>
offered some additional states apparently demonstrating monophyly, though the distribution and significance of the states mentioned was not discussed; some included characters such as pereopods 47 ambulatory, a state that is widespread in the genera close to
<taxonomicName id="26E0C2DBFF89D669853CFDD3FC526644" box="[800,905,592,617]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D669853CFDD3FC526644" box="[800,905,592,617]" italics="true" pageId="2" pageNumber="53">Cirolana</emphasis>
</taxonomicName>
, are of little diagnostic value.
</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF89D66986D9FDFBFBA16787" pageId="2" pageNumber="53" type="reference_group">
<paragraph id="E15FB958FF89D66986D9FDFBFCEF6777" blockId="2.[151,1437,152,1018]" pageId="2" pageNumber="53">
<taxonomicName id="26E0C2DBFF89D66986D9FDFBFEE366BC" ID-CoL="7PLYS" box="[197,312,632,657]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66986D9FDFBFEE366BC" box="[197,312,632,657]" italics="true" pageId="2" pageNumber="53">Kagalana</emphasis>
</taxonomicName>
shows few obvious affinities to other genera. The form of pleonites 4 and 5 and pereopod 1 is similar to but not the same as that of
<taxonomicName id="26E0C2DBFF89D6698457FD23FD3D6694" ID-CoL="7Q24T" box="[587,742,672,697]" class="Malacostraca" family="Cirolanidae" genus="Speocirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D6698457FD23FD3D6694" box="[587,742,672,697]" italics="true" pageId="2" pageNumber="53">Speocirolana</emphasis>
</taxonomicName>
, but convergence can be supposed. The morphology of the antennule, antenna, mouthparts and uropods are most similar to that of
<taxonomicName id="26E0C2DBFF89D66985CDFD4BFBE166CC" ID-CoL="7NSYL" box="[977,1082,712,737]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66985CDFD4BFBE166CC" box="[977,1082,712,737]" italics="true" pageId="2" pageNumber="53">Cirolana</emphasis>
</taxonomicName>
while the complex robust seta opposing the base of the dactylus of pereopod 1 could be interpreted as similar to genera related to
<taxonomicName id="26E0C2DBFF89D6698334FD73FE52671F" ID-CoL="7PH6D" authority="Kussakin, 1979" authorityName="Kussakin" authorityYear="1979" class="Malacostraca" family="Cirolanidae" genus="Metacirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D6698334FD73FF10671C" italics="true" pageId="2" pageNumber="53">Metacirolana</emphasis>
Kussakin, 1979
</taxonomicName>
(see
<bibRefCitation id="8571C4A9FF89D66987DBFC9BFD89671F" author="Bruce" box="[455,594,792,818]" pageId="2" pageNumber="64" refString="Bruce, N. L. (1986) Cirolanidae (Crustacea: Isopoda) of Australia. Records of the Australian Museum, Supplement, 6, 1 - 239." type="journal article" year="1986">Bruce 1986</bibRefCitation>
;
<bibRefCitation id="8571C4A9FF89D6698442FC9BFCE3671F" author="Brusca" box="[606,824,792,818]" pageId="2" pageNumber="64" refString="Brusca, R. C., Wetzer, R. &amp; France, S. C. (1995) Cirolanidae (Crustacea: Isopoda: Flabellifera) of the tropical eastern Pacific. Proceedings of the San Diego Society of Natural History, 30, 1 - 96." type="journal article" year="1995">
Brusca
<emphasis id="D394654AFF89D66984ABFC9BFD30671C" box="[695,747,792,817]" italics="true" pageId="2" pageNumber="53">et al</emphasis>
. 1995
</bibRefCitation>
). At present I regard the genus as of uncertain affinities, but probably derived from a
<taxonomicName id="26E0C2DBFF89D669843AFCC3FD546774" ID-CoL="7NSYL" box="[550,655,832,857]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D669843AFCC3FD546774" box="[550,655,832,857]" italics="true" pageId="2" pageNumber="53">Cirolana</emphasis>
</taxonomicName>
-like ancestor.
</paragraph>
<paragraph id="E15FB958FF89D66986D9FCEBFBA16787" blockId="2.[151,1437,152,1018]" pageId="2" pageNumber="53">
Many cave-dwelling cirolanids have long antennules and antennae, including relatively long flagellae, and
<taxonomicName id="26E0C2DBFF89D66986D4FC13FEE76784" ID-CoL="7PLYS" box="[200,316,912,937]" class="Malacostraca" family="Cirolanidae" genus="Kagalana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D394654AFF89D66986D4FC13FEE76784" box="[200,316,912,937]" italics="true" pageId="2" pageNumber="53">Kagalana</emphasis>
</taxonomicName>
is distinctive in having a very short antennule peduncle and flagellum.
</paragraph>
</subSubSection>
<subSubSection id="A9FAEAD3FF89D66986DAFC34FE7067D7" pageId="2" pageNumber="53" type="etymology">
<paragraph id="E15FB958FF89D66986DAFC34FE7067D7" blockId="2.[151,1437,152,1018]" pageId="2" pageNumber="53">
<emphasis id="D394654AFF89D66986DAFC34FE8867FC" bold="true" box="[198,339,951,977]" pageId="2" pageNumber="53">Etymology.</emphasis>
Kaga is an Aboriginal word for a water hole; combined with lana to indicate the family affinity; gender female.
</paragraph>
</subSubSection>
</treatment>
</document>