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<document id="8892D1B730EE46157F5EBC883A206F10" ID-DOI="10.3897/afrinvertebr.65.140524" ID-ZooBank="BCB49D2E-F772-49EB-A17A-47EB21194212" ID-publisher-id="140524" URI-arpha="8BBD979A-A43B-550B-853F-36895E8AFD3A" URI-zoobank="http://zoobank.org/BCB49D2E-F772-49EB-A17A-47EB21194212" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1734967240283" checkinUser="pensoft" docAuthor="Muller, Burgert S., Swart, Vaughn R. &amp; Snyman, Louwrens P." docDate="2024" docId="2FC9D5FD4A0B528482E49493789FECF8" docLanguage="en" docName="African Invertebrates 65(2): 247-327" docOrigin="African Invertebrates 65 (2)" docSource="https://africaninvertebrates.pensoft.net/article/140524/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Suragina pauliani Stuckenberg 1965" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="327" masterDocId="8BBD979AA43B550B853F36895E8AFD3A" masterDocTitle="Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)" masterLastPageNumber="327" masterPageNumber="247" pageNumber="247" updateTime="1735011578534" updateUser="admin">
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<mods:title id="D4F29447FD910460146D6E3FC908B59E">Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)</mods:title>
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<mods:namePart id="5CB27AFF91F8F6429C078DF8012F2B92">Muller, Burgert S.</mods:namePart>
<mods:nameIdentifier id="07C9B364731E5BFD9DA3049C20B47D77" type="ORCID">0000-0002-7304-4050</mods:nameIdentifier>
<mods:affiliation id="74D9677AF98303687F9582014F91AC51">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="63C359C49158ADC906CA515A0A15F573">Swart, Vaughn R.</mods:namePart>
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<mods:affiliation id="A140EEE33EDE9FCBAACE77DAEFDD1492">Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="18B7A96DA3EA9BFF43970FD7A0110FA9">Snyman, Louwrens P.</mods:namePart>
<mods:nameIdentifier id="2F28134F31511DB304BE8B5B5CCAE231" type="ORCID">0000-0002-5768-7216</mods:nameIdentifier>
<mods:affiliation id="9E69DCDB413D40DAD51F21AD8CEE1283">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada &amp; Department of Veterinary Microbiology, University of Saskatchewan, 52 Campus Drive, Saskatoon, Saskatchewan, S 7 N 5 B 4, Canada</mods:affiliation>
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<mods:title id="694D99EFC1728EFCE697E80CCD4D2E80">African Invertebrates</mods:title>
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<mods:part id="CE98ED4CBB465ED2E9062B4703AF9FF0">
<mods:date id="E23EC7838247B60D571883064BD16D35">2024</mods:date>
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<mods:number id="D7B4D5B09B34B716840F5D5776298D72">2024-12-23</mods:number>
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<mods:number id="5C39EBEAA7FD80359C303EACA7FE4805">65</mods:number>
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<treatment id="2FC9D5FD4A0B528482E49493789FECF8" ID-DOI="http://doi.org/10.5281/zenodo.14548381" ID-Zenodo-Dep="14548381" ID-arpha="2FC9D5FD-4A0B-5284-82E4-9493789FECF8" LSID="urn:lsid:plazi:treatment:2FC9D5FD4A0B528482E49493789FECF8" httpUri="http://treatment.plazi.org/id/2FC9D5FD4A0B528482E49493789FECF8">
<subSubSection id="BDCA1D522ED4ED66527848EC16F8114B" type="nomenclature">
<paragraph id="6CF2022F3BCDC42B7409AFBDF2B6027E">
<taxonomicName id="325192A2AC73A143436208978B253A4D" ID-CoL="53H79" authority="Stuckenberg, 1965" authorityName="Stuckenberg" authorityYear="1965" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="CBECD495DA59D51DB11D0DBAEADBAA91" italics="true">Suragina pauliani</emphasis>
<bibRefCitation id="DD27AA294D1AD8445CA4462E0823F929" author="Stuckenberg BR" journalOrPublisher="Annals of the Natal Museum" pagination="89-170" refId="B22" refString="Stuckenberg BR (1965) The Rhagionidae of Madagascar (Diptera). Annals of the Natal Museum 18: 89 - 170. https: // hdl. handle. net / 10520 / AJA 03040798 _ 759 [Last accessed: 2024 / 10 / 29]" title="The Rhagionidae of Madagascar (Diptera)." url="https://hdl.handle.net/10520/AJA03040798_759" volume="18" year="1965">Stuckenberg, 1965</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="E812BA9EC041727C6DD08581AA94F690">
<figureCitation id="FEF7FC74CEACFDB3A0380E9148E62195" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">Figs 32</figureCitation>
,
<figureCitation id="515CDC2D04C9789E3C6736E5FCD561BA" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">33</figureCitation>
,
<figureCitation id="88526545C43D1C857CD5685CB689C0CB" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">52</figureCitation>
</paragraph>
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<paragraph id="39EA3018EA0546400D16387FAE3E6434">
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<treatmentCitation id="B4F07ABA7685ED4F6D5909C32F6F7976" author="Stuckenberg BR" page="96" year="1965">
<taxonomicName id="19F611EBBF1FD6F3D7AF9E07D53E2A16" ID-CoL="53H79" authority="Stuckenberg, 1965: 96" authorityName="Stuckenberg" authorityPageNumber="96" authorityYear="1965" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="3B119870E38F3C92A9D17461A3A08C80" italics="true">Suragina pauliani</emphasis>
<comment id="B07362DF4C0A40F62423831B44942870">
<bibRefCitation id="8140882969C1E5E9FCF2F4BF9DA50FCD" author="Stuckenberg BR" journalOrPublisher="Annals of the Natal Museum" pagination="89-170" refId="B22" refString="Stuckenberg BR (1965) The Rhagionidae of Madagascar (Diptera). Annals of the Natal Museum 18: 89 - 170. https: // hdl. handle. net / 10520 / AJA 03040798 _ 759 [Last accessed: 2024 / 10 / 29]" title="The Rhagionidae of Madagascar (Diptera)." url="https://hdl.handle.net/10520/AJA03040798_759" volume="18" year="1965">Stuckenberg, 1965: 96</bibRefCitation>
, figs 1, 2, 5, 6, 911;
<treatmentCitation id="DF95108EF3EF311A0FAD6DF4E0123334" author="Stuckenberg BR" page="313" year="1980">
<bibRefCitation id="00E1A1FA71A3E10CCE949670AF46D807" author="Stuckenberg BR" editor="Crosskey RW" firstAuthor="Stuckenberg" journalOrPublisher="British Museum (Natural History), London" page="313" pagination="312-313" refId="B23" refString="Stuckenberg BR (1980) 23. Family Athericidae. In: Crosskey RW (Ed.) Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, 312313." title="23. Family Athericidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Stuckenberg 1980: 313</bibRefCitation>
</treatmentCitation>
.
</comment>
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</paragraph>
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<subSubSection id="SECID0EWVBK" type="type material">
<paragraph id="A755B0A860E707210A1485D750BF9AFB">
<heading id="BB31DF6331590419067DE117566FB2CF" reason="title">Type material not examined.</heading>
</paragraph>
<paragraph id="A842B313A1DA0B2B5A0934888628FB18">
<materialsCitation id="A2D9F3F7A4752CC49C5B19B81343F1C1" collectingDate="1956-01" collectionCode="PBZT" collectorName="B. R. Stuckenberg" country="Madagascar" county="Ankaratra Massif" elevation="1700" latitude="-19.35" location="Manjakatompo Forest Station" longLatPrecision="12" longitude="47.3" specimenCount="1" specimenCount-male="1" stateProvince="Vakinankaratra Region" typeStatus="holotype">
<emphasis id="C50E3045DC92BA935FAE5078A5178685" bold="true" italics="true">
<typeStatus id="9835CB4F60D32743591B65794B22ED28" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="368698C79A85E604E0D3A4A531B7F960" name="Madagascar">Madagascar</collectingCountry>
:
<specimenCount id="C4E8AC5012A831A278E7732A5BA5512E" count="1" type="male">1 ♂</specimenCount>
; [
<collectingRegion id="9974D3634358AF4B3A659AC2DD4A01A5">Vakinankaratra Region</collectingRegion>
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<location id="4DAA598BCF46634C10BF3C8D9B3ED9EC" LSID="urn:lsid:plazi:treatment:2FC9D5FD4A0B528482E49493789FECF8:4DAA598BCF46634C10BF3C8D9B3ED9EC" country="Madagascar" county="Ankaratra Massif" latitude="-19.35" longLatPrecision="12" longitude="47.3" name="Manjakatompo Forest Station" stateProvince="Vakinankaratra Region">Manjakatompo Forest Station</location>
[
<collectingCounty id="5821161E0A1B2C57B4500C4D8F6CF125">Ankaratra Massif</collectingCounty>
]; [
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<geoCoordinate id="62FD89F4A239297242BBB7ABB62588EE" degrees="19" direction="south" minutes="21.00" orientation="latitude" precision="9" value="-19.35">19°21.00'S</geoCoordinate>
,
<geoCoordinate id="1464F0EA7910F5C3C2C0D1F64997F8D1" degrees="47" direction="east" minutes="18.00" orientation="longitude" precision="9" value="47.3">47°18.00'E</geoCoordinate>
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<quantity id="6DD68079395489BD7DEC86E235EA66E5" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="634DDFA6897157C000C4C4B18A7C3C37" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 masl</elevation>
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;
<date id="57F76936F3A65DE18BA95ADF46F999F3" value="1956-01">
<collectingDate id="25D69333FF46B5E99F2BBF2FAB595BA7" value="1956-01">Jan. 1956</collectingDate>
</date>
;
<collectorName id="B1D22ECD9D96C960F8020B329C136A7F">B. R. Stuckenberg</collectorName>
leg.; (
<named-content id="3D7109316BD1F2CC5F9D18E63F62EBAE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/parc-botanique-et-zoologique-de-tsimbazaza" xlink_title="Parc Botanique et Zoologique de Tsimbazaza">
<collectionCode id="92A726363B37AC71775BA86F341978D5" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" type="">PBZT</collectionCode>
</named-content>
).
</materialsCitation>
</paragraph>
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<paragraph id="76E549C1E8323DEA52F12D3B426C7379">
<heading id="392654B5578602BBEFD50871FCCD4166" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="FEA50B3F0A512450D6C64055BD5FC06B">
<materialsCitation id="93AF14E40FA18AE6F3AE106F8419F963" collectingDate="1957-12-11" collectingDateMax="1957-12-15" collectingDateMin="1957-12-11" collectionCode="NMSA" collectorName="B. R. Stuckenberg" country="Madagascar" county="Ambatolampy District" elevation="2300" latitude="-19.3995" location="Mahafompona Pass" longLatPrecision="9" longitude="47.2503" municipality="Col de Mahafompeno" specimenCode="DIP 028172" specimenCount="1" specimenCount-male="1" stateProvince="Madagascar Centre" typeStatus="paratype">
<emphasis id="EE9E6FA1D0F90A749F425B76C453D1B7" bold="true" italics="true">
<typeStatus id="F3D3460E06AC0597B094070E0BE06270" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="900B3FE6C101C6CA54F32C613A0E0BA7" name="Madagascar">Madagascar</collectingCountry>
:
<specimenCount id="D65DEAECB4552960616E20351D09A442" count="1" type="male">1 ♂</specimenCount>
;
<collectingCounty id="77DB029E4103D2E3F03B59003494E4B5">Ambatolampy District</collectingCounty>
[Vakinankaratra];
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,
<collectingMunicipality id="89F47AA4DD31B311C9F97BD22FA7A43C">Col de Mahafompeno</collectingMunicipality>
[
<location id="529FEF75D2ED4091CE6FFD835120A262" LSID="urn:lsid:plazi:treatment:2FC9D5FD4A0B528482E49493789FECF8:529FEF75D2ED4091CE6FFD835120A262" country="Madagascar" county="Ambatolampy District" latitude="-19.3995" longLatPrecision="9" longitude="47.2503" municipality="Col de Mahafompeno" name="Mahafompona Pass" stateProvince="Madagascar Centre">Mahafompona Pass</location>
]; [
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<geoCoordinate id="7019B2B31F597F65A60DD036E0E4F33D" degrees="19" direction="south" minutes="23.97" orientation="latitude" precision="9" value="-19.3995">19°23.97'S</geoCoordinate>
,
<geoCoordinate id="F2AAA506948AC682DD63A710DDFA96A9" degrees="47" direction="east" minutes="15.018" orientation="longitude" precision="1" value="47.2503">47°15.018'E</geoCoordinate>
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<quantity id="09FD267B871D41409E3B43E6163A2069" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" unit="m" value="2300.0" valueMax="2400.0" valueMin="2200.0">
<elevation id="B16D5E92B14A201386F5912FC3E75A34" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" unit="m" value="2300.0" valueMax="2400.0" valueMin="2200.0">22002400 masl</elevation>
</quantity>
;
<date id="657A8CAE1872F47381B14B37EB6DEB5D" value="1957-12-11" valueMax="1957-12-15" valueMin="1957-12-11">
<collectingDate id="043268448D1639518F4EDB816AFA4CE1" value="1957-12-11" valueMax="1957-12-15" valueMin="1957-12-11">1115 Dec. 1957</collectingDate>
</date>
;
<collectorName id="3992CDD36214C0312CA00E96AFC76CBD">B. R. Stuckenberg</collectorName>
leg.; [on yellow bordered label: Paratype
<taxonomicName id="9EF5AE99A4B6B3AF73EDF6B7F910E279" authority="Stuckenberg, 1965" authorityName="Stuckenberg" authorityYear="1965" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pauliani">
<originalDetermination id="FE0570DB9E6DC04B7C1E8A8A168483BA">
<emphasis id="B24C5EE6F9BBE9F6ED6D59D8D71CCBA4" italics="true">Suragina pauliani</emphasis>
<bibRefCitation id="91881824BE314BE13CBDF468FC55C54F" author="Stuckenberg BR" journalOrPublisher="Annals of the Natal Museum" pagination="89-170" refId="B22" refString="Stuckenberg BR (1965) The Rhagionidae of Madagascar (Diptera). Annals of the Natal Museum 18: 89 - 170. https: // hdl. handle. net / 10520 / AJA 03040798 _ 759 [Last accessed: 2024 / 10 / 29]" title="The Rhagionidae of Madagascar (Diptera)." url="https://hdl.handle.net/10520/AJA03040798_759" volume="18" year="1965">Stuckenberg, 1965</bibRefCitation>
</originalDetermination>
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];
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<collectionCode id="3971DC411CB4BA59690298F166A4F32A" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="7AE5918AAF94BB75D5EA83944D6BBBF6">DIP 028172</specimenCode>
) (
<named-content id="F8D1A564E07791ABB2E4A6EE950F6E6C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
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)
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.
<materialsCitation id="F754785B2EC7E4465058C264F5EE036D" collectingDate="1956-01" collectionCode="NMSA" collectorName="B. R. Stuckenberg" county="Ankaratra Massif" elevation="1700" latitude="-19.35" location="Manjakatompo Forest Station" longLatPrecision="12" longitude="47.3" specimenCode="DIP 028173, DIP 028174" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Vakinankaratra">
<specimenCount id="D2453CFE1AAB16829B85F4E618F6A8AF" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="E98102C127E5D3DA41A42A602B734A65" count="2" type="female">2 ♀</specimenCount>
; [
<collectingRegion id="2B9BF98FD629BAE250C3078D0B43A241">Vakinankaratra</collectingRegion>
];
<collectingCounty id="6AA691A6F78EB86636F8E739C7F3C64A">Ankaratra Massif</collectingCounty>
,
<location id="E2D653370D9F0F6DBDE451E57E8171A9" LSID="urn:lsid:plazi:treatment:2FC9D5FD4A0B528482E49493789FECF8:E2D653370D9F0F6DBDE451E57E8171A9" county="Ankaratra Massif" latitude="-19.35" longLatPrecision="12" longitude="47.3" name="Manjakatompo Forest Station" stateProvince="Vakinankaratra">Manjakatompo Forest Station</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWXBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[47.300000,-19.350000]}">
<geoCoordinate id="5066AC6F737C9DD71302C043A9A95650" degrees="19" direction="south" minutes="21.00" orientation="latitude" precision="9" value="-19.35">19°21.00'S</geoCoordinate>
,
<geoCoordinate id="CC18878DAD24030D99CC2BDE2BFCFB86" degrees="47" direction="east" minutes="18.00" orientation="longitude" precision="9" value="47.3">47°18.00'E</geoCoordinate>
</named-content>
];
<quantity id="9951A581ACBCA95BD4BC04508F8ABDC7" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="ECB92C5512939E0A8B3FB5B45E4FD507" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 masl</elevation>
</quantity>
;
<date id="C8CE967DFF8AB283203D3CA9C4740EF5" value="1956-01">
<collectingDate id="5FE1758D7CDA328778F35779678A7DB0" value="1956-01">Jan. 1956</collectingDate>
</date>
;
<collectorName id="173834F9FA3233FA4511C5868E56249E">B. R. Stuckenberg</collectorName>
leg.; (
<specimenCount id="FBC15F715D6EF8B6DEFA3D6E42FA147F" count="1" type="male"></specimenCount>
:
<named-content id="BF745FF0B340A05BA4A5DA7656DCDC06" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="E5F6BC7E15EC43ABC0F311C454770159" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="F19DC37D731A9463D6D9B8C93A74CC99">DIP 028173</specimenCode>
;
<specimenCount id="62419B2FEF22604B3FB0C4F3B46959E1" count="2" type="female">2 ♀</specimenCount>
:
<named-content id="87CF8782E866595B7E567456FC712A09" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="E0EF69B5CC4E68B7974B2316E4B8D847" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="1B3BC853A4954D3C8B3EF5D4431476DF">DIP 028174</specimenCode>
, 158441) (
<named-content id="7F9264CB773EB2EDF59E15411A84B8C4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="47149AB4B4FEAD374F05CD620B641902" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHYBK" type="diagnosis">
<paragraph id="114E94F007580FB31E4B558768ED2093">
<heading id="9C8B55A0A02F79C450A5EF9D270C10EB" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D8423E1DB8D79609C061A6E05CC19292">
A more compact and hirsute species compared to other Afrotropical
<taxonomicName id="1F67892FD260D722EFE0FF9F769E118D" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="874A50F4F2389D692640DDE97F09EDBD" italics="true">Suragina</emphasis>
</taxonomicName>
. The males have the head narrowly dichoptic, with the frons distinctly velvety-black in appearance, much like in females.
<taxonomicName id="5B7B3CE431273B43547AADF79A87581B" authorityName="Stuckenberg" authorityYear="1965" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="3651AD57477F47E4EA47DD3AC78B135B" italics="true">Suragina pauliani</emphasis>
</taxonomicName>
has its thorax brown for the most part, the abdomen orange-yellow with dark subtriangular dorsal and elongated lateral markings throughout. It differs from the other two Madagascan endemics in colouration:
<taxonomicName id="65D49FD5A5207BEE83ECD779F6D66A3C" authorityName="CSCA" baseAuthorityName="Seguy" baseAuthorityYear="1951" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="73774FCFE243BB9592C90483BF02B0D3" italics="true">S. milloti</emphasis>
</taxonomicName>
is a mostly orange-yellow species and a comparatively more slender species, whereas
<taxonomicName id="DB3F08C1A4A8B424BF862912FB83AC16" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bilobata" status="sp. nov.">
<emphasis id="13FEEE07A88767AC1DB0693311A4AE2B" italics="true">S. bilobata</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="EC6572A1985041F0A826B5E4CDE95FC8" rank="species">sp. nov.</taxonomicNameLabel>
is a blackish species with a unique bilobed 1
<superScript id="28F1FF73E61F963598EF477DADEFA0E6">st</superScript>
flagellomere.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXZBK" type="redescription">
<paragraph id="5903423DC7B5D2CD9EEA61C9A52D91B6">
<heading id="6D515995C3B3FDDDEAC8B98BDA0ADD4D" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="BB5C7871EC78CD5B6328C77F132D8D98">
<emphasis id="8422CB5350C151A22680F94D3AAEAD18" bold="true" italics="true">Measurements</emphasis>
: Wing span:
<specimenCount id="8F4A2881B65F17BE4EC40AE83706C8C7" count="1" type="male"></specimenCount>
<quantity id="69BA78E2530CC5B0DEE7850D4C66DE38" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.5" metricValueMin="6.6" unit="mm" value="7.05" valueMax="7.5" valueMin="6.6">6.67.5 mm</quantity>
;
<specimenCount id="886B5312807C4C33CA609BBE9B413BC5" count="1" type="female"></specimenCount>
<quantity id="9279A0EBFF6403131716DCC9495BD327" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="7.9" metricValueMin="7.8" unit="mm" value="7.85" valueMax="7.9" valueMin="7.8">7.87.9 mm</quantity>
; body length:
<specimenCount id="E7F71B8BE7D83A17E015FC73488BE678" count="1" type="male"></specimenCount>
<quantity id="6767AC0FD70B5B5A31F06D2EAA6B79C8" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.4" metricValueMin="7.6" unit="mm" value="8.0" valueMax="8.4" valueMin="7.6">7.68.4 mm</quantity>
;
<specimenCount id="6D2F4AFE88F48E59E9CE9B736C1BAB91" count="1" type="female"></specimenCount>
<quantity id="F7D4A87B4A9B46333765F1EB1B9F84BC" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.2" metricValueMin="7.8" unit="mm" value="8.0" valueMax="8.2" valueMin="7.8">7.88.2 mm</quantity>
; wing span to body length ratio (avg.):
<specimenCount id="64DFA25AD0D2901765287584D38C1CC1" count="1" type="male"></specimenCount>
0.88;
<specimenCount id="4D8FDF0346902B903AB08AED5CD29D57" count="1" type="female"></specimenCount>
0.99.
</paragraph>
<paragraph id="5E24AA51D8D5F7609B04B8DC52C2E98A">
<emphasis id="AADB09322A62446C9DE6241999954C32" bold="true">Male</emphasis>
(Fig.
<figureCitation id="5CBB55E601D67FC0FAC5C68CD8CC36D9" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">32</figureCitation>
).
</paragraph>
<paragraph id="CDFF2D130542D47D5DEF9022697829AE">
<emphasis id="5B3B4FDA934E31596D529119852776BB" bold="true" italics="true">Head</emphasis>
: Brown colour, with grey pruinosity on majority of head; eye bare; narrowly dichoptic, frons at narrowest as wide as anterior ocellus, widening towards antennal bases; ommatidia of similar size; lateral edge of eye without indentation; ocellar tubercle slightly more elevated than frons, dark brown to blackish with some short dark setulae; vertex grey pruinose, dark brown when viewed posterodorsally, with long dark setulae; anterior ocellus larger than posterior pair; ocellar tubercle in front of dorsal margin of eye (margin less indented than in
<specimenCount id="E79EA70DE2CF3E9CFB920DA02A555604" count="1" type="female"></specimenCount>
); dorsal inner edge of eye without discernible paired dark markings; occiput with same greyish pruinosity as rest of head, except for paired narrow subrectangular dark brown almost black markings with short dark setulae on upper occiput, abutting posterior margin of eyes, flanking vertex; upper occiput with short dark setulae; lateral margins of occiput with pale setulae; lower occiput with long pale setulae, these continue ventrally on head to base of mouthparts that have similar long pale ventral setulae; frons dark velvety-brown from ocellar triangle down to a small area above antennal bases that is greyish; dark part of frons with conspicuous dark proclinate setulae, lower grey part of frons with some pale setulae; face and gena silvery-grey with pale setulae; clypeus brown with some greyish pruinosity, bare; face separated from clypeus by a prominent, deep suture on anterior and lateral edges; face not appearing to bulge laterally when viewed in profile; clypeus visible in profile; antennal bases separated ca 0.5 × width of scape, with slight longitudinal groove; scape, pedicel, 1
<superScript id="963840DBF3C65B8379910E318A489E39">st</superScript>
flagellomere brown with some whitish pruinosity, 2
<superScript id="9F928CAAB49F2D93EB798BEE28D8AD6F">nd</superScript>
flagellomere brown; 1
<superScript id="7D55C11E92CFF6C550987B4CAB3C83B1">st</superScript>
flagellomere reniform, only slightly larger than pedicel and scape; 2
<superScript id="995D9BF7F04302D06335984F850267C2">nd</superScript>
flagellomere arista-like; scape and pedicel with dark dorsal setulae, pedicel with setulae dorsally and ventrally, scape with setulae only on dorsal surface; palpus blackish with some sparse greyish pruinosity, well-developed, ca 0.5 × length of proboscis, with dark setulae; proboscis blackish-brown with long dark setulae and some scattered shorter dark setulae.
</paragraph>
<paragraph id="DC1807E37EF684D16BA811CB44851F7A">
<emphasis id="F8B123D058F5FD3ECC7709B881BCC923" bold="true" italics="true">Thorax</emphasis>
: Brown ground colour; median dorsal surface of scutum with short dark setulae, postsutural surface with long pale setulae; postsutural setulae longer than presutural setulae; postpronotal lobe orange-yellow with long dark setulae; scutum mostly dark brown with two dorsocentral brownish-grey pruinose vittae running to middle of scutum in dorsal view; postalar wall and callus appearing orange-yellow; scutellum almost entirely orange-yellow except for brown basal margin, median surface with short dark setulae, apical margin with long pale setulae; pleura mostly blackish-brown with greyish pruinosity, except for posterior surface of anepimeron and entire katatergite that are yellowish-brown with similar pruinosity; notopleuron greyish pruinose with long dark setulae; area surrounding anterior and posterior spiracle orange-yellow, postspiracular scale orange-brown; anterior spiracle bare posteriorly; proepisternum, pronotum similar in appearance to aforementioned pleura; proepimeron, proepisternum, anepisternum, parts of anepimeron and entire katatergite with pale setulae; rest of pleura bare; postscutellum dark brown.
</paragraph>
<paragraph id="A76D48E9323083120FE44CEC8ECD2367">
<emphasis id="16979FDE3B1DE1518542142674140697" bold="true" italics="true">Legs</emphasis>
: Fore and hind coxae brown on anterior surface and orange-yellow posteriorly; mid coxa brown on majority of surface, orange-yellow on apical posterior surface; all coxae with long pale setulae on anterior surface, in addition fore coxa has long pale setulae on posterior surface and hind coxa with long pale setulae on lateral apical edges, and with well-developed anterior apical point; all trochanters brown with short dark setulae; fore and mid femora entirely orange-yellow, hind femur orange-yellow with dark brown band on middle of hind femur; fore and mid tibiae orange-yellow and apically darker, hind tibia mostly dark brown, yellowish-brown at base and apex; fore and mid tarsi brownish, hind tarsi darker brown with more yellowish-brown terminal tarsal segments; fore and mid femora covered with pale setulae on almost all surfaces except for dorso-apical and posteroventral surface with some short dark setulae, hind femur with mixed long pale and dark setulae on dorsal and ventral surfaces, basally with long pale setulae, anteriorly with short setulae and posteriorly with longer setulae; hind leg overall stouter than remaining legs; fore tarsi covered with long sensory setulae along antero- and posteroventral surfaces, sensory setulae ca 2 × as long as tarsal segment is wide; fore and mid tibiae covered in short dark setulae, hind tibia with longer dark setulae; hind basal tarsal segment with long dark posterior setulae; fore tarsal claws asymmetrical, outer claw much larger than inner claw, foreleg empodium ca 2 × size of inner pulvillus, outer pulvillus ca 2 × length of inner, approaching size of outer claw; hind tarsal segments 1.1 × as long as hind tibia; tibial spur formula 0: 2: 2.
</paragraph>
<paragraph id="A9E7CCA26B528269F7CF867D1D814F00">
<emphasis id="2A581AAA8DBE3A9C5D9A652787EBEBDE" bold="true" italics="true">Wing</emphasis>
: Overall slight light brown suffused appearance; with a dark brown stigma over area of veins
<emphasis id="D3547739AFD66DF59AFC483E8185FA7B" italics="true">
R
<subScript id="3C1D059082DF975FBC917D8EDF09BABC">1</subScript>
</emphasis>
and
<emphasis id="9D0188629837A817C4CD6201B43267A9" italics="true">
R
<subScript id="F3E631B3137AC30751C7892E57F0EFA8">2 + 3</subScript>
</emphasis>
and cell
<emphasis id="59114D09696BB5833CD6F74F76D1B9F1" italics="true">
r
<subScript id="16005B9885679BA7A3C680E09985EE15">1</subScript>
</emphasis>
; veins brown; substigmal marking extends down over base of cell
<emphasis id="2514DF2BF7C589FB86DC435592AD216F" italics="true">
r
<subScript id="50B94208C387A3523263953120DFC832">2 + 3</subScript>
</emphasis>
, apex of cells
<emphasis id="3C7AC72B8319E8C5B4AD83AF0622DEFB" italics="true">br</emphasis>
and
<emphasis id="52E9BB5A3D9DB2CF7249100FF1634E57" italics="true">bm</emphasis>
, base of cells
<emphasis id="0A18E686BC928DB724634266542F9698" italics="true">
r
<subScript id="CAB75BCF5574E4B9389FA19D9C39C704">5</subScript>
</emphasis>
and
<emphasis id="DD708E12D67BF779EE2D57C49DEB2C3C" italics="true">
m
<subScript id="4B60938B8FDDC4295930A7628FE4EE49">2</subScript>
</emphasis>
, crossvein
<emphasis id="9F2C2E0686AD211D616E677B66A165AC" italics="true">r m</emphasis>
and basal third of discal cell; additional darker suffusion over majority of cell
<emphasis id="D68B3BE99B90E2CCBBC131C25F0ED17F" italics="true">br</emphasis>
; hyaline appearance in cell
<emphasis id="C32F12DBB5CB8F365C0BE5D4D14FB54D" italics="true">cua</emphasis>
, apical two-thirds of discal cell, base of cell
<emphasis id="E15CB842385B90721D6B329493482496" italics="true">
r
<subScript id="71BF8D5A775CC5823FBC581CD99FEF32">4</subScript>
</emphasis>
, post basal half of cell
<emphasis id="72EF8A9F9FF677DFF82D20EC42B42CAC" italics="true">
r
<subScript id="53881E5C3248FB52D173EE9CDEFDA06D">5</subScript>
</emphasis>
; costa without distinct downward flexure over stigma; cell
<emphasis id="B01B45CA0C837E48D0926841C03FCF19" italics="true">cua</emphasis>
closed a short distance from wing margin; cell
<emphasis id="A1571A0A982E6D1AC1B5B4AA2EEFEB19" italics="true">
m
<subScript id="27A334233C346F83420C5AEA2C1765C9">3</subScript>
</emphasis>
open, veins
<emphasis id="3A305C6525C47D86BD74E840DDC19EAD" italics="true">
M
<subScript id="18053E26111E8D3C27275B6FB0FDA4F1">1</subScript>
</emphasis>
,
<emphasis id="32DD4B3F5634682B190D3F598FE86E94" italics="true">
M
<subScript id="367C32F649F0C7A158813900B6948B9C">2</subScript>
</emphasis>
,
<emphasis id="BA569B35503C1293E9CD6F3E649D5122" italics="true">
M
<subScript id="B9950DBDDFC0D0262B44A102656FBEF2">3</subScript>
</emphasis>
present; haltere orange-yellow with a brownish knob and very short dark setulae.
</paragraph>
<paragraph id="F434990B6257EEDE49EB0F2EAD0DFB41">
<emphasis id="CA9F3E143117355D33D3284DBAC2D68C" bold="true" italics="true">Abdomen</emphasis>
: Overall orange-yellow colour; tergites 15 with broad dark brown subtriangular median vittae, remaining tergites with only a slight median marking; tergites 13 with long pale setulae, remaining tergites with shorter dark setulae; tergites 17 with dark lateral markings and especially on tergites 15 with prominent long pale setulae; sternites orange-yellow with pale setulae; tergite 1 without median longitudinal suture.
</paragraph>
<paragraph id="0C32F73EDD7029B8C1C92CECF0E440FB">
<emphasis id="AE2928C83FC6A8A05D9729B63961EE87" bold="true" italics="true">Terminalia</emphasis>
: Epandrium orange with dark setulae, hypandrium orange-brown, cercus blackish, both with dark setulae, terminalia not dissected.
</paragraph>
<paragraph id="DDBF14D6A2AB65F0E6C049AFFB5D59A5">
<emphasis id="BEF1A6D791B50E9B975DB2D1725DC453" bold="true">Female</emphasis>
(Fig.
<figureCitation id="29B52BB0AA12F4D60415626053074459" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">33</figureCitation>
): Similar to
<specimenCount id="B102908EF762998EBCF9F81ABB204D71" count="1" type="male"></specimenCount>
except for the following:
</paragraph>
<paragraph id="44B58BDBAE6AA82D516EF9AB0BE70FD7">
<emphasis id="32950B0713E9814F85EA6962E084464F" bold="true" italics="true">Head</emphasis>
: Widely dichoptic; dorsal margin of eye more indented than in
<specimenCount id="88795E02FBE8BBAA6B71950003E46DB0" count="1" type="male"></specimenCount>
; vertex wider than in
<specimenCount id="182A92BF788520C8D6F970ECF3A8C6D6" count="1" type="male"></specimenCount>
, dorsal inner edge of eye brownish-grey pruinose, appearing as an extension of vertex surrounding ocellar tubercle; occiput with paired dark brown almost black subtriangular (more narrow and subrectangular in
<specimenCount id="55A49CEA66BEDD27996A772EA31ED7B6" count="1" type="male"></specimenCount>
) markings on upper occiput; frons similar in colour appearance to that of
<specimenCount id="A1A889E419BFFDA6B139D2563A150BA7" count="1" type="male"></specimenCount>
but at narrowest ca 1.6 × width of ocellar tubercle, widening slightly towards antennal base, gradually fading at edges into bronzy brown colour; face separated anteriorly from clypeus by shallow transverse suture, deeper sutures laterally; face slightly visible in profile; 1
<superScript id="40314CDDBC36EFDB7B01474962B9B17B">st</superScript>
flagellomere comparatively much larger than in
<specimenCount id="AA75E543BB8A3EEFFE6777B503CAE859" count="1" type="male"></specimenCount>
; scape ca 2 × as long as pedicel; 1
<superScript id="433DD18C883A04C32B36EB8731CDC396">st</superScript>
flagellomere ca 2 × as long as pedicel.
</paragraph>
<paragraph id="9E8FCB8E9A8157085FA35C8812E4A3DA">
<emphasis id="64F5F3F51DAF9E552AFB7C54C07D36EA" bold="true" italics="true">Thorax</emphasis>
: Scutum slightly lighter than in
<specimenCount id="8779629ED28B2557020814D52745E3FE" count="1" type="male"></specimenCount>
, otherwise with similar appearance; postpronotal lobes with similar dark setulae as in
<specimenCount id="0A5252B8FD7BEDDC5349FDD3E0BC6C59" count="1" type="male"></specimenCount>
but lobe with long pale setulae as well; pleura reddish-brown with grey pruinosity, with similar patterning as in
<specimenCount id="E3F086CCA783B5F5BAF221270FEA0CA7" count="1" type="male"></specimenCount>
just lighter; postspiracular scale orange in colour; scutellum with only pale setulae.
</paragraph>
<paragraph id="0ED9B4FC6A86B83EC98B9D0A795643FE">
<emphasis id="7C0F3C52B76A863D9CE63F81932BC32D" bold="true" italics="true">Legs</emphasis>
: Coxae similar to
<specimenCount id="9C3492FD2815E9183905405517195B0E" count="1" type="male"></specimenCount>
, just overall lighter in colour; hind fermur with median band light brown; fore and mid legs yellow with tarsi brownish, but basal two-thirds of basitarsi yellowish; fore tarsal claws symmetrical; hind leg with general orange-yellow colour on segments, including tarsi; hind femur with mix of short pale and dark setulae compared to
<specimenCount id="99C3DB95471135AB3B4C957CB1A31B64" count="1" type="male"></specimenCount>
; hind tarsal segments 0.9 × as long as hind tibia.
</paragraph>
<paragraph id="2322BD3DD6BD809FC96C8288B83F1A65">
<emphasis id="38B71ECABAD79F614F380282B8FD007E" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="E6D98054974FC44A5B4B4368EB2ED565" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">52</figureCitation>
): Overall brownish suffusion, with upper third of wing somewhat darker suffused, stigma also markedly lighter compared to
<specimenCount id="932A17CA906F2B293344F99B8E863033" count="1" type="male"></specimenCount>
; no prominent dark suffused substigmal markings running longitudinally on wing as in
<specimenCount id="B03AC3223D52B718BC7659E413205280" count="1" type="male"></specimenCount>
.
</paragraph>
<paragraph id="EA19A1C2EBB7DB09C455549B1C5E2A27">
<emphasis id="797E11ED9C4A88A76B63159C6D6DF186" bold="true" italics="true">Abdomen</emphasis>
: Similar pattering and colour as
<specimenCount id="6D7F9D8888B0FB1D158EBB4285D8E6F1" count="1" type="male"></specimenCount>
; lateral dark markings on tergites appearing as more uniform and continuous dark marginal line.
</paragraph>
<paragraph id="9829102967EC5BF628AA31E2704EDCF4">
<emphasis id="181975D00471BD03591C49792E165D19" bold="true" italics="true">Terminalia</emphasis>
: Cercus orange-brown with pale setulae; terminalia not dissected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES6BK" type="distribution">
<paragraph id="3D7CD5B3A4F37F8646E6B37859FCE64D">
<heading id="0695527C9C6EE9A6E009B08DB3512AF5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="E1DB164B99A903954D21FF19823BBAD6">
<collectingCountry id="0DE4B08931A584D4D6428DE290295F53" name="Madagascar">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>