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<document id="88B628CB8E6F0FA40758476C6356BC1C" ID-DOI="10.11646/phytotaxa.260.2.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13672420" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725401489335" checkinUser="felipe" docAuthor="Jiménez-Mejías, Pedro &amp; Escudero, Marcial" docDate="2016" docId="2F0A87951A31D165FF5998237760F7D3" docLanguage="en" docName="phytotaxa.260.2.8.pdf" docOrigin="Phytotaxa 260 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.260.2.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Carex roalsoniana Jim., sp. nov." docType="treatment" docVersion="2" lastPageNumber="187" masterDocId="D333FFED1A30D167FFD19A617627FFC0" masterDocTitle="Notes on South American Carex section Schiedeanae and description of the new species Carex roalsoniana" masterLastPageNumber="192" masterPageNumber="185" pageNumber="186" updateTime="1725422461735" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FEA7704131BB34FB4D1E4AF022CDF027">Notes on South American Carex section Schiedeanae and description of the new species Carex roalsoniana</mods:title>
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<mods:namePart id="55E4BAC672F5C715F60D3E50FC4FDA04">Jiménez-Mejías, Pedro</mods:namePart>
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<mods:namePart id="B6ED4801D2AB298039B65D5A8DF56485">Escudero, Marcial</mods:namePart>
<mods:affiliation id="6FAD3BE7A1067E1C78B4743FC092DD1A">Department of Plant Biology and Ecology, University of Seville, Av. Reina Mercedes sn, 41012 Seville, Spain;</mods:affiliation>
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<mods:title id="B6ECDA8CA741CB174ACFAAD52CA99408">Phytotaxa</mods:title>
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<mods:date id="838098EBE796D5BDCFBABAFCB49E76AF">2016</mods:date>
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<treatment id="2F0A87951A31D165FF5998237760F7D3" ID-DOI="http://doi.org/10.5281/zenodo.13672454" ID-Zenodo-Dep="13672454" LSID="urn:lsid:plazi:treatment:2F0A87951A31D165FF5998237760F7D3" httpUri="http://treatment.plazi.org/id/2F0A87951A31D165FF5998237760F7D3" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
<subSubSection id="EFB965081A31D166FF5998237588FD9B" box="[136,943,575,603]" pageId="1" pageNumber="186" type="nomenclature">
<paragraph id="A71C36831A31D166FF5998237588FD9B" blockId="1.[136,943,575,603]" box="[136,943,575,603]" pageId="1" pageNumber="186">
<heading id="FC5481EF1A31D166FF5998237588FD9B" box="[136,943,575,603]" fontSize="11" level="2" pageId="1" pageNumber="186" reason="4">
<emphasis id="95D7EA911A31D166FF599823750AFD9B" bold="true" box="[136,813,575,603]" pageId="1" pageNumber="186">
1.
<taxonomicName id="60A34D001A31D166FF7598217791FD9B" ID-CoL="RBL3" authority="Jim." authorityName="Jim." box="[164,438,575,603]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="roalsoniana">
<emphasis id="95D7EA911A31D166FF759821775CFD9A" bold="true" box="[164,379,575,602]" italics="true" pageId="1" pageNumber="186">Carex roalsoniana</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; M.Escudero,
<taxonomicNameLabel id="8EE457EA1A31D166FD049821750AFD9B" box="[725,813,576,603]" pageId="1" pageNumber="186" rank="species">
<emphasis id="95D7EA911A31D166FD0498217501FD9A" bold="true" box="[725,806,576,602]" italics="true" pageId="1" pageNumber="186">sp. nov</emphasis>
.
</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3F982A061A31D166FCED98237581FD9B" box="[828,934,578,603]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1505,1526]" captionStartId-1="3.[136,229,1428,1449]" captionTargetBox-1="[283,1308,194,1400]" captionTargetId-1="figure-14@3.[280,1308,190,1403]" captionTargetPageId-1="3" captionText-0="FIGURE 1. Holotype of Carex roalsoniana Jim.-Mejías &amp; M.Escudero." captionText-1="FIGURE 2. Detailed pictures of the holotype (A; NY herbarium) and isotype (B–G; A herbarium) of Carex roalsoniana Jim.-Mejías &amp; M. Escudero. A. Inflorescences; notice in the upper inflorescence that the lowermost spike is covered by the bract; the arrow depicts the male segment at the top of an androgynous spike. B. Spike axis after most utricles are dropping; the arrow depicts the utriculiform cladoprophyll from which the spike axis protrudes; notice that the male segment at the top of the spike remains after the utricles and female glumes has dropped. C. Female glumes. D–E. Ripe utricles, abaxial side; notice in D the short style with strongly convoluted backward stigmas. F–G. Ripe achenes. Scale bar = 1 mm." figureDoi-1="http://doi.org/10.5281/zenodo.13672424" httpUri-1="https://zenodo.org/record/13672424/files/figure.png" pageId="1" pageNumber="186">Figs. 1–2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFB965081A31D166FF5998EB7264FC5F" pageId="1" pageNumber="186" type="description">
<paragraph id="A71C36831A31D166FF5998EB7264FC5F" blockId="1.[136,1452,650,927]" pageId="1" pageNumber="186">
This new species is similar to
<taxonomicName id="60A34D001A31D166FE2598EB75D9FD63" authority="G. A. Wheeler &amp; Guagl." authorityName="G. A. Wheeler &amp; Guagl." box="[500,1022,650,675]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="subandrogyna">
<emphasis id="95D7EA911A31D166FE2598EB74C5FD63" box="[500,738,650,675]" italics="true" pageId="1" pageNumber="186">Carex subandrogyna</emphasis>
G.A.Wheeler &amp; Guagl.
</taxonomicName>
, native from northern
<collectingCountry id="DFB476131A31D166FAD998EB7350FD63" box="[1288,1399,650,675]" name="Argentina" pageId="1" pageNumber="186">Argentina</collectingCountry>
and
<collectingCountry id="DFB476131A31D166FF5998CF76FAFD07" box="[136,221,686,711]" name="Bolivia" pageId="1" pageNumber="186">Bolivia</collectingCountry>
, and to
<taxonomicName id="60A34D001A31D166FEED98CF7430FD07" authority="Boott" authorityName="Boott" box="[316,535,686,711]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="lepida">
<emphasis id="95D7EA911A31D166FEED98CF77E8FD07" box="[316,463,686,711]" italics="true" pageId="1" pageNumber="186">Carex lepida</emphasis>
Boott
</taxonomicName>
from
<collectingCountry id="DFB476131A31D166FDB198CF74E7FD07" box="[608,704,686,711]" name="Ecuador" pageId="1" pageNumber="186">Ecuador</collectingCountry>
. From
<taxonomicName id="60A34D001A31D166FCC298CF75F2FD07" box="[787,981,686,711]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="subandrogyna">
<emphasis id="95D7EA911A31D166FCC298CF75F2FD07" box="[787,981,686,711]" italics="true" pageId="1" pageNumber="186">C. subandrogyna</emphasis>
</taxonomicName>
the new species clearly differs in utricle morphology (
<quantity id="605B9B661A31D166FEF198B3778FFD2B" box="[288,424,722,747]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.1" metricValueMin="3.6" pageId="1" pageNumber="186" unit="mm" value="3.8499999999999996" valueMax="4.1" valueMin="3.6">3.6–4.1 mm</quantity>
long, with a beak
<quantity id="605B9B661A31D166FDA798B374DBFD2B" box="[630,764,722,747]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="1" pageNumber="186" unit="mm" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 mm</quantity>
in
<taxonomicName id="60A34D001A31D166FCF198B375E1FD2B" box="[800,966,722,747]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="roalsoniana">
<emphasis id="95D7EA911A31D166FCF198B375E1FD2B" box="[800,966,722,747]" italics="true" pageId="1" pageNumber="186">C. roalsoniana</emphasis>
</taxonomicName>
vs.
<quantity id="605B9B661A31D166FC2598B3725CFD2B" box="[1012,1147,722,747]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="1" pageNumber="186" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4–2.9 mm</quantity>
long, with a beak up to
<quantity id="605B9B661A31D166FA5998B37693FCCF" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="186" unit="mm" value="0.3">0.3 mm</quantity>
long or beakless in
<taxonomicName id="60A34D001A31D166FE4B9897747CFCCF" box="[410,603,758,783]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="subandrogyna">
<emphasis id="95D7EA911A31D166FE4B9897747CFCCF" box="[410,603,758,783]" italics="true" pageId="1" pageNumber="186">C. subandrogyna</emphasis>
</taxonomicName>
) and leaves (up to
<quantity id="605B9B661A31D166FCE6989775B1FCCF" box="[823,918,758,783]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="186" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
wide, stiff, in
<taxonomicName id="60A34D001A31D166FBEC989772C2FCCF" box="[1085,1253,758,783]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="roalsoniana">
<emphasis id="95D7EA911A31D166FBEC989772C2FCCF" box="[1085,1253,758,783]" italics="true" pageId="1" pageNumber="186">C. roalsoniana</emphasis>
</taxonomicName>
vs. up to
<quantity id="605B9B661A31D166FA8698977693FCF3" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.5" metricValueMin="5.6" pageId="1" pageNumber="186" unit="mm" value="6.55" valueMax="7.5" valueMin="5.6">5.6–7.5 mm</quantity>
wide, very soft in
<taxonomicName id="60A34D001A31D166FE5A997B746BFCF3" box="[395,588,794,819]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="subandrogyna">
<emphasis id="95D7EA911A31D166FE5A997B746BFCF3" box="[395,588,794,819]" italics="true" pageId="1" pageNumber="186">C. subandrogyna</emphasis>
</taxonomicName>
). From
<taxonomicName id="60A34D001A31D166FD78997B7536FCF3" box="[681,785,794,819]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="lepida">
<emphasis id="95D7EA911A31D166FD78997B7536FCF3" box="[681,785,794,819]" italics="true" pageId="1" pageNumber="186">C. lepida</emphasis>
</taxonomicName>
the new species differs by its habit densely caespitose and by having trigonous achenes with three stigmas (
<taxonomicName id="60A34D001A31D166FD76995F752AFC97" box="[679,781,830,855]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="lepida">
<emphasis id="95D7EA911A31D166FD76995F752AFC97" box="[679,781,830,855]" italics="true" pageId="1" pageNumber="186">C. lepida</emphasis>
</taxonomicName>
has long rhizomes and biconvex achenes with two stigmas). The glabrous nerveless utricles of the three South American species readily distinguish them from the North American species of the section that have scabrid to hispidulous, conspicuously nerved utricles.
</paragraph>
</subSubSection>
<subSubSection id="EFB965081A31D165FF5999B175E0F9FF" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186" type="materials_examined">
<paragraph id="A71C36831A31D166FF5999B17293FBC9" blockId="1.[136,1452,976,1033]" pageId="1" pageNumber="186">
<emphasis id="95D7EA911A31D166FF5999B176E5FC25" bold="true" box="[136,194,976,997]" pageId="1" pageNumber="186">Type:</emphasis>
—
<materialsCitation id="17CB3CDE1A31D166FF0A99B17297FBC9" collectingDate="1945-08-12" collectionCode="W, H, NY, A, K, MICH" collectorName="de Oro" country="Ecuador" elevation="2591" location="Sevilla de Oro" municipality="eastern Cordillera" pageId="1" pageNumber="186" specimenCount="2" stateProvince="Azuay" typeStatus="holotype">
<collectingCountry id="DFB476131A31D166FF0A99B17775FC25" box="[219,338,976,997]" name="Ecuador" pageId="1" pageNumber="186">ECUADOR</collectingCountry>
.
<collectingRegion id="6567F8611A31D166FE8B99B177BBFC25" box="[346,412,976,997]" country="Ecuador" name="Azuay" pageId="1" pageNumber="186">Azuay</collectingRegion>
: the
<collectingMunicipality id="4778ACF91A31D166FE1A99B1745EFC25" box="[459,633,976,997]" pageId="1" pageNumber="186">eastern Cordillera</collectingMunicipality>
,
<quantity id="605B9B661A31D166FD5599B174E9FC25" box="[644,718,976,997]" metricMagnitude="3" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" pageId="1" pageNumber="186" unit="km" value="4.5" valueMax="8.0" valueMin="1.0">1–8 km</quantity>
north of the village of
<location id="A27C60581A31D166FC6099B17267FC25" LSID="urn:lsid:plazi:treatment:2F0A87951A31D165FF5998237760F7D3:A27C60581A31D166FC6099B17267FC25" box="[945,1088,976,997]" country="Ecuador" municipality="eastern Cordillera" name="Sevilla de Oro" pageId="1" pageNumber="186" stateProvince="Azuay">
Sevilla
<collectorName id="0A5653551A31D166FC2B99B17267FC25" box="[1018,1088,976,997]" pageId="1" pageNumber="186">de Oro</collectorName>
</location>
,
<quantity id="605B9B661A31D166FB9B99B172C7FC25" box="[1098,1248,976,997]" metricMagnitude="3" metricUnit="m" metricValue="2.5908" metricValueMax="2.7432" metricValueMin="2.4384" pageId="1" pageNumber="186" unit="ft" value="8500.0" valueMax="9000.0" valueMin="8000.0">
<elevation id="2C8ED1B01A31D166FB9B99B172C7FC25" box="[1098,1248,976,997]" metricMagnitude="3" metricUnit="m" metricValue="2.5908" metricValueMax="2.7432" metricValueMin="2.4384" pageId="1" pageNumber="186" unit="ft" value="8500.0" valueMax="9000.0" valueMin="8000.0">8,000–9,000 ft.</elevation>
</quantity>
, in cliffside seepage, plants in clumps,
<date id="D31D10431A31D166FEB6999577C2FBC9" box="[359,485,1012,1033]" pageId="1" pageNumber="186" value="1945-08-12">
<collectingDate id="C359E9AB1A31D166FEB6999577C2FBC9" box="[359,485,1012,1033]" pageId="1" pageNumber="186" value="1945-08-12">12 Aug 1945</collectingDate>
</date>
,
<emphasis id="95D7EA911A31D166FE3E999574B4FBC9" box="[495,659,1012,1033]" italics="true" pageId="1" pageNumber="186">
<collectionCode id="C1B2AE461A31D166FE3E99957422FBC9" box="[495,517,1012,1033]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="186" type="Herbarium">W</collectionCode>
.
<collectionCode id="C1B2AE461A31D166FDD69995743CFBC9" box="[519,539,1012,1033]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="186" type="Herbarium">H</collectionCode>
. Camp 4662
</emphasis>
(
<typeStatus id="781888211A31D166FD71999574D1FBC9" box="[672,758,1012,1033]" pageId="1" pageNumber="186" type="holotype">holotype</typeStatus>
<collectionCode id="C1B2AE461A31D166FD2A99957504FBC9" box="[763,803,1012,1033]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="1" pageNumber="186" type="Herbarium">NY</collectionCode>
!
<figureCitation id="3F982A061A31D166FCFD999575AFFBC9" box="[812,904,1012,1033]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1505,1526]" captionStartId-1="3.[136,229,1428,1449]" captionTargetBox-1="[283,1308,194,1400]" captionTargetId-1="figure-14@3.[280,1308,190,1403]" captionTargetPageId-1="3" captionText-0="FIGURE 1. Holotype of Carex roalsoniana Jim.-Mejías &amp; M.Escudero." captionText-1="FIGURE 2. Detailed pictures of the holotype (A; NY herbarium) and isotype (B–G; A herbarium) of Carex roalsoniana Jim.-Mejías &amp; M. Escudero. A. Inflorescences; notice in the upper inflorescence that the lowermost spike is covered by the bract; the arrow depicts the male segment at the top of an androgynous spike. B. Spike axis after most utricles are dropping; the arrow depicts the utriculiform cladoprophyll from which the spike axis protrudes; notice that the male segment at the top of the spike remains after the utricles and female glumes has dropped. C. Female glumes. D–E. Ripe utricles, abaxial side; notice in D the short style with strongly convoluted backward stigmas. F–G. Ripe achenes. Scale bar = 1 mm." figureDoi-1="http://doi.org/10.5281/zenodo.13672424" httpUri-1="https://zenodo.org/record/13672424/files/figure.png" pageId="1" pageNumber="186">Figs. 1–2</figureCitation>
;
<typeStatus id="781888211A31D166FC43999575C6FBC9" box="[914,993,1012,1033]" pageId="1" pageNumber="186" type="isotype">isotypes</typeStatus>
<collectionCode id="C1B2AE461A31D166FC34999575DCFBC9" box="[997,1019,1012,1033]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="186" type="Herbarium">A</collectionCode>
!
<figureCitation id="3F982A061A31D166FBD499957218FBC9" box="[1029,1087,1012,1033]" captionStart="FIGURE 2" captionStartId="3.[136,229,1428,1449]" captionTargetBox="[283,1308,194,1400]" captionTargetId="figure-14@3.[280,1308,190,1403]" captionTargetPageId="3" captionText="FIGURE 2. Detailed pictures of the holotype (A; NY herbarium) and isotype (B–G; A herbarium) of Carex roalsoniana Jim.-Mejías &amp; M. Escudero. A. Inflorescences; notice in the upper inflorescence that the lowermost spike is covered by the bract; the arrow depicts the male segment at the top of an androgynous spike. B. Spike axis after most utricles are dropping; the arrow depicts the utriculiform cladoprophyll from which the spike axis protrudes; notice that the male segment at the top of the spike remains after the utricles and female glumes has dropped. C. Female glumes. D–E. Ripe utricles, abaxial side; notice in D the short style with strongly convoluted backward stigmas. F–G. Ripe achenes. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13672424" httpUri="https://zenodo.org/record/13672424/files/figure.png" pageId="1" pageNumber="186">Fig. 2</figureCitation>
,
<collectionCode id="C1B2AE461A31D166FB989995727BFBC9" box="[1097,1116,1012,1033]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="186" type="Herbarium">K</collectionCode>
,
<collectionCode id="C1B2AE461A31D166FBB79995728EFBC9" box="[1126,1193,1012,1033]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="1" pageNumber="186" type="Herbarium">MICH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A71C36831A31D166FF599E5B72F2F8E3" blockId="1.[136,1452,1082,2079]" pageId="1" pageNumber="186">
Perennial; caespitose. Stems
<quantity id="605B9B661A31D166FE059E5B7462FB93" box="[468,581,1082,1107]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" metricValueMax="5.2" metricValueMin="1.0" pageId="1" pageNumber="186" unit="cm" value="31.0" valueMax="52.0" valueMin="10.0">10–52 cm</quantity>
long,
<quantity id="605B9B661A31D166FD409E5B753DFB93" box="[657,794,1082,1107]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="186" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.8–1.2 mm</quantity>
width in its middle length, shorter than or as long as the leaves, trigonous, with narrowly winged margins, faintly scabrid in most its length, with pickles in both antrorse and retrorse directions; basal sheaths up to
<quantity id="605B9B661A31D166FD919EE3748EFB5B" box="[576,681,1154,1179]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="186" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
high, soft, ± entire, dark brown. Leaves flat or slightly
<collectionCode id="C1B2AE461A31D166FAF19EE3731EFB5B" box="[1312,1337,1154,1179]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="186" type="Herbarium">M</collectionCode>
shaped in cross section, hypostomatic, the uppermost of the flowering stems with a blade
<quantity id="605B9B661A31D166FC289EC7724FFB7F" box="[1017,1128,1190,1215]" metricMagnitude="-1" metricUnit="m" metricValue="3.85" metricValueMax="5.4" metricValueMin="2.3" pageId="1" pageNumber="186" unit="cm" value="38.5" valueMax="54.0" valueMin="23.0">23–54 cm</quantity>
long, the widest with a blade
<quantity id="605B9B661A31D166FF599EAB76C2FB23" box="[136,229,1226,1251]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="186" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
width; ligule
<quantity id="605B9B661A31D166FE519EAB77F9FB23" box="[384,478,1226,1251]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="1" pageNumber="186" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
, longer than wide, acute, hyaline; lowermost leaves reduced, with a short blade and the side opposite to the blade brownish. Inflorescence
<quantity id="605B9B661A31D166FD149E8F7564FAC7" box="[709,835,1262,1287]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.6" metricValueMin="1.9" pageId="1" pageNumber="186" unit="cm" value="2.75" valueMax="3.6" valueMin="1.9">1.9–3.6 cm</quantity>
long, with 2–3 androgynous spikes, sometimes one of them reduced, the lowest one ± covered at the base by the lowermost bract; lowermost bract
<quantity id="605B9B661A31D166FB4C9F7372D8FAEB" box="[1181,1279,1298,1323]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="1" pageNumber="186" unit="cm" value="9.5" valueMax="13.0" valueMin="6.0">6–13 cm</quantity>
×
<quantity id="605B9B661A31D166FACF9F737380FAEB" box="[1310,1447,1298,1323]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.5" metricValueMin="1.7" pageId="1" pageNumber="186" unit="mm" value="2.6" valueMax="3.5" valueMin="1.7">1.7–3.5 mm</quantity>
, much longer than the inflorescence, sheathless; second lowermost bract
<quantity id="605B9B661A31D166FC709F57723BFA8F" box="[929,1052,1334,1359]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="5.4" metricValueMin="2.5" pageId="1" pageNumber="186" unit="cm" value="3.95" valueMax="5.4" valueMin="2.5">2.5–5.4 cm</quantity>
×
<quantity id="605B9B661A31D166FBE69F577299FA8F" box="[1079,1214,1334,1359]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.2" metricValueMin="0.8" pageId="1" pageNumber="186" unit="mm" value="1.5" valueMax="2.2" valueMin="0.8">0.8–2.2 mm</quantity>
, longer than its spike. Spikes androgynous, densely flowered, with up to 40–50 utricles, being the female part much more developed than the male one; uppermost spike 20–22.5 ×
<quantity id="605B9B661A31D166FDE59F1F748FFA57" box="[564,680,1406,1431]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="1" pageNumber="186" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
, generally larger than the lowermost one, with a male part
<quantity id="605B9B661A31D166FAEB9F1F738BFA57" box="[1338,1452,1406,1431]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="1" pageNumber="186" unit="mm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 mm</quantity>
long; lowermost spike 11.7–20 ×
<quantity id="605B9B661A31D166FDD39FC37451FA7B" box="[514,630,1442,1467]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="1" pageNumber="186" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 mm</quantity>
, with a male part
<quantity id="605B9B661A31D166FC919FC375EFFA7B" box="[832,968,1442,1467]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.5" metricValueMin="1.7" pageId="1" pageNumber="186" unit="mm" value="2.6" valueMax="3.5" valueMin="1.7">1.7–3.5 mm</quantity>
long; cladoprophylls present at the base of each spike, ±
<quantity id="605B9B661A31D166FEF19FA77751FA1F" box="[288,374,1478,1503]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="186" unit="mm" value="2.2">2.2 mm</quantity>
long, utriculiform, hyaline to reddish, with a funnel-shaped beak with ciliate borders, containing a female flower although frequently aborting it, if developing then the achene body protrudes from the cladoprophyll beak. Male glumes 2 ×
<quantity id="605B9B661A31D166FE5E9C6F743FF9E7" box="[399,536,1550,1575]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="186" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
, entirely hyaline or with the central nerve green, acute, scabrid above, crowded at the top of the spike. Anthers
<quantity id="605B9B661A31D166FE169C53746AF98B" box="[455,589,1586,1611]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="186" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
long, filaments
<quantity id="605B9B661A31D166FCD09C537554F98B" box="[769,883,1586,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="1" pageNumber="186" unit="mm" value="1.3" valueMax="1.6" valueMin="1.0">1–1.6 mm</quantity>
long, reddish. Female glumes deciduous, dropping before utricles, with a body 2.5–4 ×
<quantity id="605B9B661A31D166FDF99C377495F9AF" box="[552,690,1622,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="1" pageNumber="186" unit="mm" value="1.0" valueMax="1.4" valueMin="0.6">0.6–1.4 mm</quantity>
, hyaline, narrowly obovate to oblong, with a central band greenish with 1–3 nerves, acute, mucronate, the ones from the middle part of the spike with the mucro 0.5–1.1 long, sparsely scabrid. Utricle 3.6–4.1 ×
<quantity id="605B9B661A31D166FE6B9CFF7461F977" box="[442,582,1694,1719]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="1" pageNumber="186" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
, elliptical, soft, membranaceous, hyaline to straw-colored, nerveless except for the two lateral nerves, attenuated into a
<quantity id="605B9B661A31D166FDBE9CA374D1F91B" box="[623,758,1730,1755]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="1" pageNumber="186" unit="mm" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 mm</quantity>
smooth beak. Stigmas 3, very short, conspicuously convolute backward, reddish-brown and densely papillose; style base bulbous, deciduous, leaving a flattish lignified remnant at the top of the achene. Achene 2.1–2.9 ×
<quantity id="605B9B661A31D166FD9D9D6B7483F8E3" box="[588,676,1802,1827]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="186" unit="mm" value="1.2">1.2 mm</quantity>
, trigonous; rachilla vestigial (~
<quantity id="605B9B661A31D166FBD19D6B727EF8E3" box="[1024,1113,1802,1827]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="186" unit="mm" value="0.1">0.1 mm</quantity>
) or absent.
</paragraph>
<paragraph id="A71C36831A31D166FF6C9D4F743BF7DF" blockId="1.[136,1452,1082,2079]" pageId="1" pageNumber="186">
<emphasis id="95D7EA911A31D166FF6C9D4F77CBF887" bold="true" box="[189,492,1838,1863]" pageId="1" pageNumber="186">Habitat and distribution:</emphasis>
—Known only from two populations, one in
<collectingCountry id="DFB476131A31D166FC379D4F7264F887" box="[998,1091,1838,1863]" name="Ecuador" pageId="1" pageNumber="186">Ecuador</collectingCountry>
and another one in
<collectingCountry id="DFB476131A31D166FAF09D4F7370F887" box="[1313,1367,1838,1863]" name="Peru" pageId="1" pageNumber="186">Peru</collectingCountry>
, placed at a straight-line distance of
<quantity id="605B9B661A31D166FE009D33740FF8AB" box="[465,552,1874,1899]" metricMagnitude="5" metricUnit="m" metricValue="2.33" pageId="1" pageNumber="186" unit="km" value="233.0">233 km</quantity>
one from another. Both populations are in western facing slopes of the Andes foothills, at an altitude of
<quantity id="605B9B661A31D166FE699D17747EF84F" box="[440,601,1910,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="1" pageNumber="186" unit="m" value="2600.0" valueMax="2800.0" valueMin="2400.0">2400–2800 m</quantity>
. The indications provided in the herbarium labels associate the habitat of this new species to moist environments (forest, seepages on cliffs). Such ecological preferences are more similar to the requirements of
<taxonomicName id="60A34D001A31D166FE439DDF7472F817" box="[402,597,1982,2007]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="species" species="subandrogyna">
<emphasis id="95D7EA911A31D166FE439DDF7472F817" box="[402,597,1982,2007]" italics="true" pageId="1" pageNumber="186">C. subandrogyna</emphasis>
</taxonomicName>
, which grows in shady subtropical montane forest (“yungas”; Wheeler &amp; Guaglianone, 2003, 2006), than to the North American species, which are found in dry, subdesertic habitats (Reznicek, 2003;
<bibRefCitation id="C3324B721A31D166FF1E92677437F7DF" author="Gomez-Sanchez, M. &amp; Cabrera-Luna, A. &amp; Gonzalez-Elizondo, M. S. &amp; Reznicek, A. A." box="[207,528,2054,2079]" pageId="1" pageNumber="186" pagination="926 - 928" refId="ref3856" refString="Gomez-Sanchez, M., Cabrera-Luna, A., Gonzalez-Elizondo, M. S. &amp; Reznicek, A. A. (2012) Carex vizarronensis (Cyperaceae), a New Species of Carex Section Schiedeanae from Central Mexico. Systematic Botany 37: 926 - 928. http: // dx. doi. org / 10.1600 / 036364412 X 656527" type="journal article" year="2012">
Gómez-Sánchez
<emphasis id="95D7EA911A31D166FE5C926777E1F7DF" box="[397,454,2054,2079]" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
, 2012
</bibRefCitation>
).
</paragraph>
<caption id="F3DC660B1A32D165FF599F807565FA36" box="[136,834,1505,1526]" pageId="2" pageNumber="187" startId="2.[136,229,1505,1526]" targetType="figure">
<paragraph id="A71C36831A32D165FF599F807565FA36" blockId="2.[136,834,1505,1526]" box="[136,834,1505,1526]" pageId="2" pageNumber="187">
<emphasis id="95D7EA911A32D165FF599F8076D9FA36" bold="true" box="[136,254,1505,1526]" pageId="2" pageNumber="187">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="60A34D001A32D165FEAC9F807479FA36" authority="Jim." authorityName="Jim." box="[381,606,1505,1526]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="roalsoniana">
<emphasis id="95D7EA911A32D165FEAC9F807408FA36" box="[381,559,1505,1526]" italics="true" pageId="2" pageNumber="187">Carex roalsoniana</emphasis>
Jim.
</taxonomicName>
-Mejías &amp; M.Escudero.
</paragraph>
</caption>
<paragraph id="A71C36831A32D165FF6C9C4775E0F9FF" blockId="2.[136,1452,1574,2067]" box="[189,967,1574,1599]" pageId="2" pageNumber="187">
<emphasis id="95D7EA911A32D165FF6C9C477764F9FF" bold="true" box="[189,323,1574,1599]" pageId="2" pageNumber="187">Phenology:</emphasis>
—Flowers and fruits were collected from May to August.
</paragraph>
</subSubSection>
<subSubSection id="EFB965081A32D165FF6C9C2B7760F7D3" pageId="2" pageNumber="187" type="etymology">
<paragraph id="A71C36831A32D165FF6C9C2B7430F96B" blockId="2.[136,1452,1574,2067]" pageId="2" pageNumber="187">
<emphasis id="95D7EA911A32D165FF6C9C2B776FF9A3" bold="true" box="[189,328,1610,1635]" pageId="2" pageNumber="187">Etymology:</emphasis>
—Dedicated to Eric H. Roalson, North American botanist and professor of the
<collectingRegion id="6567F8611A32D165FB359C2B738BF9A3" box="[1252,1452,1610,1635]" country="United States of America" name="Washington" pageId="2" pageNumber="187">Washington State</collectingRegion>
University, expert in systematics, taxonomy and evolution of
<taxonomicName id="60A34D001A32D165FCFE9C0F7592F947" box="[815,949,1646,1671]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
, who continuously promoted and encouraged to develop the knowledge in
<taxonomicName id="60A34D001A32D165FE1D9CF37428F96B" box="[460,527,1682,1707]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="95D7EA911A32D165FE1D9CF37428F96B" box="[460,527,1682,1707]" italics="true" pageId="2" pageNumber="187">Carex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A71C36831A32D165FF6C9CD77295F933" blockId="2.[136,1452,1574,2067]" pageId="2" pageNumber="187">
<emphasis id="95D7EA911A32D165FF6C9CD77478F90F" bold="true" box="[189,607,1718,1743]" pageId="2" pageNumber="187">
Additional collections (
<typeStatus id="781888211A32D165FE049CD77475F90F" box="[469,594,1718,1743]" pageId="2" pageNumber="187" type="paratype">paratypes</typeStatus>
):
</emphasis>
—
<collectingCountry id="DFB476131A32D165FDAD9CD774E1F90F" box="[636,710,1718,1743]" name="Peru" pageId="2" pageNumber="187">PERU</collectingCountry>
. Ayabaca:
<collectingRegion id="6567F8611A32D165FC929CD775A6F90F" box="[835,897,1718,1743]" country="Peru" name="Piura" pageId="2" pageNumber="187">Piura</collectingRegion>
, ruinas de Aypate,
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–
<geoCoordinate id="C29750441A32D165FB2F9CD77380F90F" box="[1278,1447,1718,1743]" degrees="79" direction="west" minutes="34.252" orientation="longitude" pageId="2" pageNumber="187" precision="1" value="-79.57087">79° 34.252’ W</geoCoordinate>
,
<quantity id="605B9B661A32D165FF599CBB7702F933" box="[136,293,1754,1779]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="2" pageNumber="187" unit="m" value="2750.0" valueMax="2800.0" valueMin="2700.0">2700–2800 m</quantity>
, bosque secundario intervenido,
<date id="D31D10431A32D165FD449CBB750AF933" box="[661,813,1754,1779]" pageId="2" pageNumber="187" value="1996-05-22">22 May 1996</date>
,
<emphasis id="95D7EA911A32D165FCE99CBB721AF933" box="[824,1085,1754,1779]" italics="true" pageId="2" pageNumber="187">V. Quipuscoa et al. 569</emphasis>
(F!, MO).
</paragraph>
<paragraph id="A71C36831A32D165FF6C9C9F7760F7D3" blockId="2.[136,1452,1574,2067]" pageId="2" pageNumber="187">
<emphasis id="95D7EA911A32D165FF6C9C9F7742F8D7" bold="true" box="[189,357,1790,1815]" pageId="2" pageNumber="187">Observations:</emphasis>
—The name “
<taxonomicName id="60A34D001A32D165FDD49C9F753FF8D7" authority="Nees" authorityName="Nees" box="[517,792,1790,1815]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="species" species="erythrogyne">
<emphasis id="95D7EA911A32D165FDD49C9F74F1F8D7" box="[517,726,1790,1815]" italics="true" pageId="2" pageNumber="187">Carex erythrogyne</emphasis>
Nees
</taxonomicName>
” found in the vouchers from the collections from
<collectingCountry id="DFB476131A32D165FA9F9C9F738CF8D7" box="[1358,1451,1790,1815]" name="Ecuador" pageId="2" pageNumber="187">Ecuador</collectingCountry>
seems just to be a labeling error. The name is not listed in the International Plant Names Index (www.ipni.org), neither in the World Checklist of
<taxonomicName id="60A34D001A32D165FDD59D2775B5F89F" authority="(Govaerts et al., 2016)" baseAuthorityName="Govaerts" baseAuthorityYear="2016" box="[516,914,1862,1887]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="family">
Cyperaceae (
<bibRefCitation id="C3324B721A32D165FD479D2775ADF89F" author="Govaerts, R. &amp; Jimenez-Mejias, P. &amp; Koopman, J. &amp; Simpson, D. A. &amp; Goetghebeur, P. &amp; Wilson, K. L. &amp; Egorova, T. &amp; Bruhl, J. J." box="[662,906,1862,1887]" pageId="2" pageNumber="187" refId="ref3921" refString="Govaerts, R., Jimenez-Mejias, P., Koopman, J., Simpson, D. A., Goetghebeur, P., Wilson, K. L., Egorova, T. &amp; Bruhl, J. J. (2016) World checklist of Cyperaceae. Facilitated by the Royal Botanic Gardens, Kew. Available from: http: // apps. kew. org / wcsp / (accessed 17 December 2015)" type="url" year="2016">
Govaerts
<emphasis id="95D7EA911A32D165FCD49D277519F89F" box="[773,830,1862,1887]" italics="true" pageId="2" pageNumber="187">et al.</emphasis>
, 2016
</bibRefCitation>
)
</taxonomicName>
. We also unsuccessfully searched for this name in Steudel’s Synopsis Plantarum Glumacearum (1853–1855) and Boott’s Illustrations of genus
<taxonomicName id="60A34D001A32D165FB169D0B732DF843" box="[1223,1290,1898,1923]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="187" phylum="Tracheophyta" rank="genus">
<emphasis id="95D7EA911A32D165FB169D0B732DF843" box="[1223,1290,1898,1923]" italics="true" pageId="2" pageNumber="187">Carex</emphasis>
</taxonomicName>
(1858–1867), where some name authorities are assigned to Nees. In addition, we requested for the existence of possible
<typeStatus id="781888211A32D165FACB9DEF736DF867" box="[1306,1354,1934,1959]" pageId="2" pageNumber="187">type</typeStatus>
material under this name in herbaria that could harbor Nees’ collections (B, BR, CGE, GE, GZU, HBG, K, M, MSB, NHV, STR, UPS; Stafleu &amp; Cowan (1976–1997), abbreviations according Thiers (2015)) and received a negative response from all of them.
</paragraph>
</subSubSection>
</treatment>
</document>
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