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<mods:title id="3258BDC9DC13A3F7C871AD1661A1BE91">Taxonomic Revision of Tynanthus (Bignonieae, Bignoniaceae)</mods:title>
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<mods:namePart id="B670FF7CD639BFB66B1D8A396DFA3C24">Medeiros, Maria Cláudia Melo Pacheco De</mods:namePart>
<mods:affiliation id="905EB97EA2257C7DBA7CB2FD8EA9046C">Universidade de São Paulo, Instituto de Biociências, Departamento de Botânica, Rua do Matão, 277, 05508 - 090, São Paulo, SP, Brazil. &amp; Universidade de São Paulo, Instituto de Biociências, Departamento de Botânica, Rua do Matão, 277, 05508 - 090, São Paulo, SP, Brazil. &amp; Authors for correspondence: mariaclaudiamedeiros @ hotmail. com; llohmann @ usp. br</mods:affiliation>
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<mods:namePart id="027BA782E4DDBD50C4775F5357BFBF17">Lohmann, Lúcia G.</mods:namePart>
<mods:affiliation id="49671895570D2B1B51AC548ACCFD2B1B">Universidade de São Paulo, Instituto de Biociências, Departamento de Botânica, Rua do Matão, 277, 05508 - 090, São Paulo, SP, Brazil. &amp; Universidade de São Paulo, Instituto de Biociências, Departamento de Botânica, Rua do Matão, 277, 05508 - 090, São Paulo, SP, Brazil. &amp; Authors for correspondence: mariaclaudiamedeiros @ hotmail. com; llohmann @ usp. br</mods:affiliation>
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<paragraph id="A20F36ADFFB4FF99FF2B7BCB331E272D" blockId="15.[151,1437,1780,1913]" pageId="15">
<emphasis id="90C4EABFFFB4FF99FF2B7BCB31D12775" bold="true" box="[151,172,1780,1805]" pageId="15">4.</emphasis>
<taxonomicName id="65B04D2EFFB4FF99FF087BCB35722775" ID-CoL="99W59" authority="M. C. Medeiros &amp; L. G. Lohmann (2014: 82)" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityPageNumber="82" authorityYear="2014" box="[180,1039,1780,1805]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="15" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFB4FF99FF087BCB337F2775" bold="true" box="[180,514,1780,1805]" italics="true" pageId="15">Tynanthus espiritosantensis</emphasis>
<materialsCitation id="12D83CF0FFB4FF99FDB77BCB35722775" box="[523,1039,1780,1805]" collectingDate="2008-02-06" collectionCode="CVRD, A, SPF" country="Brazil" county="Type" location="Reserva Natural" municipality="Linhares" pageId="15" specimenCount="2" stateProvince="Espirito Santo" typeStatus="holotype">
M.C. Medeiros &amp; L.G. 
<bibRefCitation id="C6214B5CFFB4FF99FCA37BCB35722775" author="Lohmann, L. G. &amp; Taylor, C. M." box="[799,1039,1780,1805]" pageId="15" pagination="348 - 489" refId="ref46804" refString="Lohmann, L. G. &amp; Taylor, C. M. (2014) A new generic classification of Tribe Bignonieae (Bignoniaceae). Annals of the Missouri Botanical Garden 99: 348 - 489. http: // dx. doi. org / 10.3417 / 2003187" type="journal article" year="2014">Lohmann (2014: 82)</bibRefCitation>
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:— 
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. 
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: 
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, 
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da CVRD, 
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, km 2.3, 
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, 
<emphasis id="90C4EABFFFB4FF99FBA37A2635B02749" box="[1055,1229,1816,1841]" italics="true" pageId="15">D.A. Folli 5931</emphasis>
(
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CVRD! (11073); 
<typeStatus id="7D0B880FFFB4FF99FF407A033036272D" box="[252,331,1852,1877]" pageId="15" type="isotype">isotype</typeStatus>
SPF! (barcode 199170)).
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<figureCitation id="3A8B2A28FFB4FF99FF2B7A5F31A62701" box="[151,219,1888,1913]" captionStart="FIGURE 6" captionStartId="16.[151,247,1863,1885]" captionTargetBox="[156,1430,206,1830]" captionTargetId="figure-87@16.[151,1436,191,1842]" captionTargetPageId="16" captionText="FIGURE 6. Tynanthus espiritosantensis: A. Flowering branch; B–D. Leaflet with pubescent domatia in the abaxial surface; E. Interpetiolar region with bromeliad-like prophylls of the axillary buds; F. Detail of the inflorescence axis, showing bracts, simple and peltate trichomes; G. Open corolla showing the androecium; H. Anther; I. Open calyx showing the gynoecium (D.A. Folli 5931, SPF). Illustration by Klei Sousa. Figure from Medeiros &amp; Lohmann (2014)." figureDoi="http://doi.org/10.5281/zenodo.13634487" httpUri="https://zenodo.org/record/13634487/files/figure.png" pageId="15">Fig. 6</figureCitation>
: A–I
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<emphasis id="90C4EABFFFB4FF99FF2B751331CA2830" bold="true" box="[151,183,2092,2120]" pageId="15">16</emphasis>
• 
<emphasis id="90C4EABFFFB4FF99FF5D750D303F283F" box="[225,322,2098,2119]" italics="true" pageId="15">Phytotaxa</emphasis>
216 (1) © 2015 Magnolia Press
</paragraph>
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MEDEIROS &amp; LOHMANN TAXONOMIC REVISION OF 
<taxonomicName id="65B04D2EFFABFF86FE747509338A2833" authority="(BIGNONIEAE)" baseAuthorityName="BIGNONIEAE" box="[456,759,2102,2123]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="16" phylum="Tracheophyta" rank="genus">
<emphasis id="90C4EABFFFABFF86FE74750933302833" box="[456,589,2102,2123]" italics="true" pageId="16">TYNANTHUS</emphasis>
(BIGNONIEAE)
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<emphasis id="90C4EABFFFABFF86FF2B7A78306D2725" bold="true" box="[151,272,1863,1885]" pageId="16">FIGURE 6.</emphasis>
<taxonomicName id="65B04D2EFFABFF86FEA47A7833522724" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" box="[280,559,1863,1884]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="16" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFABFF86FEA47A7833522724" box="[280,559,1863,1884]" italics="true" pageId="16">Tynanthus espiritosantensis</emphasis>
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: A. Flowering branch; B–D. Leaflet with pubescent domatia in the abaxial surface; E. Interpetiolar region with bromeliad-like prophylls of the axillary buds; F. Detail of the inflorescence axis, showing bracts, simple and peltate trichomes; G. Open corolla showing the androecium; H. Anther; I. Open calyx showing the gynoecium (D.A. Folli 5931, SPF). Illustration by Klei Sousa. Figure from 
<bibRefCitation id="C6214B5CFFABFF86FDAD7A95325227C7" author="Medeiros, M. C. M. P. &amp; Lohmann, L. G." box="[529,815,1962,1983]" pageId="16" pagination="77 - 85" refId="ref46989" refString="Medeiros, M. C. M. P. &amp; Lohmann, L. G. (2014) Two new species of Tynanthus Miers (Bignonieae, Bignoniaceae) from Brazil. Phytokeys 42: 77 - 85. http: // dx. doi. org / 10.3897 / phytokeys. 42.8210" type="journal article" year="2014">Medeiros &amp; Lohmann (2014)</bibRefCitation>
.
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<emphasis id="90C4EABFFFABFF86FC00750835602834" box="[956,1053,2103,2124]" italics="true" pageId="16">Phytotaxa</emphasis>
216 (1) © 2015 Magnolia Press • 
<emphasis id="90C4EABFFFABFF86FA3C750D34DD2836" bold="true" box="[1408,1440,2098,2126]" pageId="16">17</emphasis>
<emphasis id="90C4EABFFFAAFF87FF2B7DA7301320C8" box="[151,366,151,176]" italics="true" pageId="17">Lianas. Branchlets</emphasis>
tetragonal to terete, with or without ritidome, finely striated, lenticeled to densely lenticeled, glabrescent (sometimes pubescent at the nodes), with peltate and patelliform trichomes (sometimes also with simple); interpetiolar ridge absent; interpetiolar patelliform glands absent; prophylls of the axillary buds 
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long, 
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wide, bromeliad-like, glabrescent (rarely puberulent) throughout, with peltate trichomes (rarely also with simple). 
<emphasis id="90C4EABFFFAAFF87FE7E7C16336C2138" box="[450,529,297,320]" italics="true" pageId="17">Leaves</emphasis>
2–3 foliolated (more commonly 3); terminal leaflets often modified into trifid tendrils, without adhesive-disks on tip; petioles and petiolules puberulent throughout surface, with simple and peltate trichomes; petioles 
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long; petiolules 
<quantity id="65489B48FFAAFF87FD687C50322C21FF" box="[724,849,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="17" unit="cm" value="2.0" valueMax="3.5" valueMin="0.5">0.5–3.5 cm</quantity>
long; leaflets (4–) 
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long, (1.5–) 
<quantity id="65489B48FFAAFF87FAA17C5034E121FF" box="[1309,1436,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="5.4" metricValueMin="1.9" pageId="17" unit="cm" value="3.65" valueMax="5.4" valueMin="1.9">1.9–5.4 cm</quantity>
wide, membranous to chartaceous, discolor, elliptic; apex acuminate or caudate, mucronate; base cuneate, symmetrical; margin entire; the abaxial surface glabrescent (sometimes pubescent) on and near the veins, with peltate and patelliform trichomes (sometimes also simple); the adaxial surface glabrescent on and near the veins, with peltate and patelliform trichomes; glandular trichomes distributed especially on the abaxial surface; second venation weak brochidodromous; pocket domatia with trichomes. 
<emphasis id="90C4EABFFFAAFF87FC357F1B355D2245" box="[905,1056,548,573]" italics="true" pageId="17">Inflorescence</emphasis>
axilar, a thyrse, lax, with conical aspect, 
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long; axis pubescent, with simple and peltate trichomes; bracts of the inflorescence predominantly caducous, pubescent throughout or only at margins, 0.7–3.9(–9) mm long; floral bracts 
<quantity id="65489B48FFAAFF87FB367F53346922FB" box="[1162,1300,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="17" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
long; floral pedicels 
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long. 
<taxonomicName id="65B04D2EFFAAFF87FE047FB0308722D0" box="[440,506,655,680]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="17" phylum="Porifera" rank="genus">
<emphasis id="90C4EABFFFAAFF87FE047FB0308722D0" box="[440,506,655,680]" italics="true" pageId="17">Calyx</emphasis>
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green, 
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long, 
<quantity id="65489B48FFAAFF87FC9D7FB032D722DF" box="[801,938,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="17" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 mm</quantity>
wide, with transversal (sometimes oblique) aperture, minutely 5-denticulate, glabrescent (sometimes pubescent at teeth) outside, with patelliform glands; lobes 
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long. 
<emphasis id="90C4EABFFFAAFF87FED07FE830B92288" box="[364,452,727,752]" italics="true" pageId="17">Corolla</emphasis>
white, 
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long, 
<quantity id="65489B48FFAAFF87FD557FE832092297" box="[745,884,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="17" unit="mm" value="2.95" valueMax="3.4" valueMin="2.5">2.5–3.4 mm</quantity>
wide at the tube opening; tube 
<quantity id="65489B48FFAAFF87FB597FE834272297" box="[1253,1370,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="17" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
long, internally tomentose to pubescent at the base or glabrescent, with simple and long and short stipitate trichomes; nectar guides absent, but with a path of long and short stipitate trichomes; lobes densely pubescent to pubescent throughout lower ones and at the margin of upper ones; upper ones 
<quantity id="65489B48FFAAFF87FC157E7B354E2323" box="[937,1075,836,861]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="17" unit="mm" value="0.75" valueMax="1.1" valueMin="0.4">0.4–1.1 mm</quantity>
long, 
<quantity id="65489B48FFAAFF87FBC27E7B358F2323" box="[1150,1266,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
wide, acute to obtuse; lower ones 
<quantity id="65489B48FFAAFF87FECA7E58337D2307" box="[374,512,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="17" unit="mm" value="2.5" valueMax="3.2" valueMin="1.8">1.8–3.2 mm</quantity>
long, 
<quantity id="65489B48FFAAFF87FDF67E5833AE2307" box="[586,723,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="17" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1–2.5 mm</quantity>
wide, obtuse to rounded (sometimes acute). 
<emphasis id="90C4EABFFFAAFF87FB6B7E58341C23F8" box="[1239,1377,871,896]" italics="true" pageId="17">Androecium</emphasis>
with fertile stamens inserted 
<quantity id="65489B48FFAAFF87FE1B7EB3336723DB" box="[423,538,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
from the base of the corolla; shorter ones 
<quantity id="65489B48FFAAFF87FC437EB335F523DB" box="[1023,1160,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="17" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
long; longer ones 
<quantity id="65489B48FFAAFF87FAE17EB331BE23BF" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="17" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
long; anthers thecae 
<quantity id="65489B48FFAAFF87FE027E90333423BF" box="[446,585,943,968]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="17" unit="mm" value="0.95" valueMax="1.1" valueMin="0.8">0.8–1.1 mm</quantity>
long, obovate to elliptic, subexserted; connective extending 
<quantity id="65489B48FFAAFF87FAAD7E9034E023BF" box="[1297,1437,943,968]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="17" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
beyond anther attachment; staminode ca. 
<quantity id="65489B48FFAAFF87FD347EEB339E2393" box="[648,739,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="17" unit="mm" value="2.4">2.4 mm</quantity>
long, glabrescent, with long and short stipitate trichomes. 
<emphasis id="90C4EABFFFAAFF87FF2B7EC730642477" box="[151,281,1016,1039]" italics="true" pageId="17">Gynoecium</emphasis>
<quantity id="65489B48FFAAFF87FE9C7EC830E82477" box="[288,405,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="17" unit="mm" value="5.25" valueMax="6.0" valueMin="4.5">4.5–6 mm</quantity>
long; ovary 
<quantity id="65489B48FFAAFF87FD947EC833E02477" box="[552,669,1015,1040]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="17" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
long, 
<quantity id="65489B48FFAAFF87FD5B7EC8320C2477" box="[743,881,1015,1040]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="17" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
wide, conical, velutinous; style 
<quantity id="65489B48FFAAFF87FB5A7EC834262477" box="[1254,1371,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="5.0" metricValueMin="3.3" pageId="17" unit="mm" value="4.15" valueMax="5.0" valueMin="3.3">3.3–5 mm</quantity>
long, tomentose at the base. 
<emphasis id="90C4EABFFFAAFF87FE29792230B3244C" box="[405,462,1053,1076]" italics="true" pageId="17">Fruit</emphasis>
not seen. 
<emphasis id="90C4EABFFFAAFF87FDFC792333FD244D" box="[576,640,1052,1077]" italics="true" pageId="17">Seeds</emphasis>
not seen.
</paragraph>
<paragraph id="A20F36ADFFAAFF87FF707900350A2420" blockId="17.[151,1437,151,1149]" box="[204,1143,1087,1112]" pageId="17">
<emphasis id="90C4EABFFFAAFF87FF707900302F2420" bold="true" box="[204,338,1087,1112]" pageId="17">Phenology:</emphasis>
—Flowers from December to February; the fruiting season is unknown.
</paragraph>
</subSubSection>
<subSubSection id="EAAA6526FFAAFF84FF70795B33EA22FD" lastPageId="18" pageId="17" type="distribution">
<paragraph id="A20F36ADFFAAFF87FF70795B34512405" blockId="17.[151,1437,151,1149]" box="[204,1324,1124,1149]" pageId="17">
<emphasis id="90C4EABFFFAAFF87FF70795B30852405" bold="true" box="[204,504,1124,1149]" pageId="17">Distribution and habitat:</emphasis>
—Occurs in moist broadleaf forests from 
<collectingCountry id="DAA7763DFFAAFF87FC71795B356C2405" box="[973,1041,1124,1149]" name="Brazil" pageId="17">Brazil</collectingCountry>
(
<collectingRegion id="6074F84FFFAAFF87FB9C795B35BF2405" box="[1056,1218,1124,1149]" country="Brazil" name="Espirito Santo" pageId="17">Espírito Santo</collectingRegion>
) (
<figureCitation id="3A8B2A28FFAAFF87FB65795B34632405" box="[1241,1310,1124,1149]" captionStart="FIGURE 7" captionStartId="17.[151,244,2003,2025]" captionTargetBox="[259,1344,1198,1972]" captionTargetId="figure-448@17.[255,1349,1193,1980]" captionTargetPageId="17" captionText="FIGURE 7. Distribution of Tynanthus espiritosantensis." figureDoi="http://doi.org/10.5281/zenodo.13634489" httpUri="https://zenodo.org/record/13634489/files/figure.png" pageId="17">Fig. 7</figureCitation>
).
</paragraph>
<caption id="F6CF6625FFAAFF87FF2B7AEC33C82791" ID-DOI="http://doi.org/10.5281/zenodo.13634489" ID-Zenodo-Dep="13634489" box="[151,693,2003,2025]" httpUri="https://zenodo.org/record/13634489/files/figure.png" pageId="17" startId="17.[151,244,2003,2025]" targetBox="[259,1344,1198,1972]" targetPageId="17" targetType="figure">
<paragraph id="A20F36ADFFAAFF87FF2B7AEC33C82791" blockId="17.[151,693,2003,2025]" box="[151,693,2003,2025]" pageId="17">
<emphasis id="90C4EABFFFAAFF87FF2B7AEC30712791" bold="true" box="[151,268,2003,2025]" pageId="17">FIGURE 7.</emphasis>
Distribution of 
<taxonomicName id="65B04D2EFFAAFF87FE1A7AEB33D22791" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" box="[422,687,2004,2025]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="17" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFAAFF87FE1A7AEB33D22791" box="[422,687,2004,2025]" italics="true" pageId="17">Tynanthus espiritosantensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A20F36ADFFAAFF87FF2B75133301283F" blockId="17.[151,636,2092,2121]" box="[151,636,2092,2121]" pageId="17">
<emphasis id="90C4EABFFFAAFF87FF2B751331CA2830" bold="true" box="[151,183,2092,2120]" pageId="17">18</emphasis>
• 
<emphasis id="90C4EABFFFAAFF87FF5D750D303F283F" box="[225,322,2098,2119]" italics="true" pageId="17">Phytotaxa</emphasis>
216 (1) © 2015 Magnolia Press
</paragraph>
<paragraph id="A20F36ADFFAAFF87FB34750D34E1283F" blockId="17.[1160,1436,2098,2119]" box="[1160,1436,2098,2119]" pageId="17">MEDEIROS &amp; LOHMANN</paragraph>
<paragraph id="A20F36ADFFA9FF84FF707DA830462165" blockId="18.[151,1440,151,645]" pageId="18">
<emphasis id="90C4EABFFFA9FF84FF707DA8332C20C8" bold="true" box="[204,593,151,176]" pageId="18">Additional specimens examined:</emphasis>
— 
<materialsCitation id="12D83CF0FFA9FF84FDD27DA834E620D7" box="[622,1435,151,176]" collectingDate="1984-12-07" collectionCode="G" country="Brazil" location="Rancho Alto" municipality="Linhares" pageId="18" specimenCount="1" stateProvince="Espirito Santo">
<collectingCountry id="DAA7763DFFA9FF84FDD27DA833AB20C8" box="[622,726,151,176]" name="Brazil" pageId="18">BRAZIL</collectingCountry>
. 
<emphasis id="90C4EABFFFA9FF84FD5D7DA832EB20C8" bold="true" box="[737,918,151,176]" pageId="18">
<collectingRegion id="6074F84FFFA9FF84FD5D7DA832EC20C8" box="[737,913,151,176]" country="Brazil" name="Espirito Santo" pageId="18">Espírito Santo</collectingRegion>
:
</emphasis>
<collectingMunicipality id="426BACD7FFA9FF84FC227DA8357E20C8" box="[926,1027,151,176]" pageId="18">Linhares</collectingMunicipality>
, 
<location id="A76F6076FFA9FF84FBB37DA835DC20C8" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FBB37DA835DC20C8" box="[1039,1185,151,176]" country="Brazil" municipality="Linhares" name="Rancho Alto" pageId="18" stateProvince="Espirito Santo">Rancho Alto</location>
, 
<date id="D60E106DFFA9FF84FB117DA7340420C8" box="[1197,1401,151,176]" pageId="18" value="1984-12-07">
<collectingDate id="C64AE985FFA9FF84FB117DA7340420C8" box="[1197,1401,151,176]" pageId="18" value="1984-12-07">7 December 1984</collectingDate>
</date>
, 
<collectionCode id="C4A1AE68FFA9FF84FA387DA734E620D7" box="[1412,1435,152,175]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="18" type="Herbarium">
<emphasis id="90C4EABFFFA9FF84FA387DA734E620D7" box="[1412,1435,152,175]" italics="true" pageId="18">G</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="90C4EABFFFA9FF84FA207DA730FC20AC" italics="true" pageId="18">
. 
<materialsCitation id="12D83CF0FFA9FF84FF2B7D83300020AC" box="[151,381,188,213]" collectionCode="J, M" country="Brazil" location="Hatschbach" pageId="18" specimenCount="1">
<location id="A76F6076FFA9FF84FF2B7D83305920AD" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FF2B7D83305920AD" box="[151,292,188,213]" country="Brazil" name="Hatschbach" pageId="18">Hatschbach</location>
&amp; 
<collectionCode id="C4A1AE68FFA9FF84FEF07D82302120AC" box="[332,348,189,212]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="18" type="Herbarium">J</collectionCode>
. 
<collectionCode id="C4A1AE68FFA9FF84FEDC7D82300020AC" box="[352,381,189,212]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="18" type="Herbarium">M</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="12D83CF0FFA9FF84FE367D83338E20AD" box="[394,755,188,213]" collectionCode="MBM, MO" country="United States of America" location="Silva" pageId="18" specimenCount="1">
<emphasis id="90C4EABFFFA9FF84FE367D83336A20AD" box="[394,535,188,213]" italics="true" pageId="18">
<location id="A76F6076FFA9FF84FE367D8330B820AD" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FE367D8330B820AD" box="[394,453,188,213]" country="United States of America" name="Silva" pageId="18">Silva</location>
48693
</emphasis>
(
<collectionCode id="C4A1AE68FFA9FF84FD947D83330E20AD" box="[552,627,188,213]" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="18">MBM</collectionCode>
, 
<collectionCode id="C4A1AE68FFA9FF84FD3E7D8333CE20AD" box="[642,691,188,213]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="18" type="Herbarium">MO</collectionCode>
, 
<collectingCountry id="DAA7763DFFA9FF84FD7D7D83339720AD" box="[705,746,188,213]" name="United States of America" pageId="18">US</collectingCountry>
)
</materialsCitation>
; 
<materialsCitation id="12D83CF0FFA9FF84FCBD7D83353E20AD" box="[769,1091,188,213]" collectionCode="CVRD" country="United States of America" location="Reserva Natural" pageId="18" specimenCount="1">
<location id="A76F6076FFA9FF84FCBD7D8332BF20AD" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FCBD7D8332BF20AD" box="[769,962,188,213]" country="United States of America" name="Reserva Natural" pageId="18">Reserva Natural</location>
da 
<collectionCode id="C4A1AE68FFA9FF84FC4C7D83353E20AD" box="[1008,1091,188,213]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13923" lsid="urn:lsid:biocol.org:col:13923" name="Reserva Natural da Vale" pageId="18" type="Herbarium">CVRD</collectionCode>
</materialsCitation>
, 
<materialsCitation id="12D83CF0FFA9FF84FBEE7D833537208F" collectingDate="2014-01-27" collectionCode="M, C" country="United States of America" elevation="53" location="Estrada Oiticica" pageId="18" specimenCount="1">
<location id="A76F6076FFA9FF84FBEE7D83346820AD" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FBEE7D83346820AD" box="[1106,1301,188,213]" country="United States of America" name="Estrada Oiticica" pageId="18">Estrada Oiticica</location>
, próximo à porteira, antes do cruzamento com a estrada municipal, 
<quantity id="65489B48FFA9FF84FCB47DE0323E208F" box="[776,835,223,248]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="18" unit="m" value="53.0">
<elevation id="299DD19EFFA9FF84FCB47DE0323E208F" box="[776,835,223,248]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="18" unit="m" value="53.0">53 m</elevation>
</quantity>
, 
<date id="D60E106DFFA9FF84FCF37DE035762080" box="[847,1035,223,248]" pageId="18" value="2014-01-27">
<collectingDate id="C64AE985FFA9FF84FCF37DE035762080" box="[847,1035,223,248]" pageId="18" value="2014-01-27">27 January 2014</collectingDate>
</date>
, 
<emphasis id="90C4EABFFFA9FF84FBAA7DDF3537208F" box="[1046,1098,224,247]" italics="true" pageId="18">
<collectionCode id="C4A1AE68FFA9FF84FBAA7DDF354E208F" box="[1046,1075,224,247]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="18" type="Herbarium">M</collectionCode>
. 
<collectionCode id="C4A1AE68FFA9FF84FB8A7DDF3537208F" box="[1078,1098,224,247]" country="Denmark" name="University of Copenhagen" pageId="18" type="Herbarium">C</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="90C4EABFFFA9FF84FBF67DDF346F2080" box="[1098,1298,223,248]" italics="true" pageId="18">
. 
<materialsCitation id="12D83CF0FFA9FF84FBE97DE034702080" box="[1109,1293,223,248]" collectionCode="R, B" country="United States of America" location="Medeiros" pageId="18" specimenCount="1">
<location id="A76F6076FFA9FF84FBE97DE035C32080" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FBE97DE035C32080" box="[1109,1214,223,248]" country="United States of America" name="Medeiros" pageId="18">Medeiros</location>
&amp; 
<collectionCode id="C4A1AE68FFA9FF84FB5D7DDE35882080" box="[1249,1269,225,248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18">R</collectionCode>
. 
<collectionCode id="C4A1AE68FFA9FF84FB457DDE34702080" box="[1273,1293,225,248]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="18" type="Herbarium">B</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="12D83CF0FFA9FF84FAA47DE0304A2165" collectionCode="CVRD, SPF" country="United States of America" location="Louzada" pageId="18" specimenCount="1">
<emphasis id="90C4EABFFFA9FF84FAA47DE034E12080" box="[1304,1436,223,248]" italics="true" pageId="18">
<location id="A76F6076FFA9FF84FAA47DE034042080" LSID="urn:lsid:plazi:treatment:2A1987BBFFB4FF84FF2B7BCB33EA22FD:A76F6076FFA9FF84FAA47DE034042080" box="[1304,1401,223,248]" country="United States of America" name="Louzada" pageId="18">Louzada</location>
41
</emphasis>
(
<collectionCode id="C4A1AE68FFA9FF84FF237C3B318C2165" box="[159,241,260,285]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13923" lsid="urn:lsid:biocol.org:col:13923" name="Reserva Natural da Vale" pageId="18" type="Herbarium">CVRD</collectionCode>
, 
<collectionCode id="C4A1AE68FFA9FF84FF407C3B30522165" box="[252,303,260,285]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="18" type="Herbarium">SPF</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A20F36ADFFA9FF84FF707C1833EA22FD" blockId="18.[151,1440,151,645]" pageId="18">
<emphasis id="90C4EABFFFA9FF84FF707C1830D62138" bold="true" box="[204,427,295,320]" pageId="18">Taxonomic notes:</emphasis>
— 
<taxonomicName id="65B04D2EFFA9FF84FE767C18326A2147" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" box="[458,791,295,320]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFA9FF84FE767C18326A2147" box="[458,791,295,320]" italics="true" pageId="18">Tynanthus espiritosantensis</emphasis>
</taxonomicName>
shares bromeliad-like prophylls of the axillary buds, glabrescent elliptic leaflets, corolla tube internally tomentose at base and lax inflorescences with 
<taxonomicName id="65B04D2EFFA9FF84FA3A7C72303721F0" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FA3A7C72303721F0" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
. Nevertheless, 
<taxonomicName id="65B04D2EFFA9FF84FE437C4E339D21FF" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" box="[511,736,368,392]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFA9FF84FE437C4E339D21FF" box="[511,736,368,392]" italics="true" pageId="18">T. espiritosantensis</emphasis>
</taxonomicName>
is easily separated by the pubescent leaflet domatia (versus glabrous domatia in 
<taxonomicName id="65B04D2EFFA9FF84FEC07CAA333B21D5" box="[380,582,404,429]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FEC07CAA333B21D5" box="[380,582,404,429]" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
), lack of patelliform glands in the petioles, petiolules and inflorescence axis (versus presence of patteliform glands in 
<taxonomicName id="65B04D2EFFA9FF84FDD07C86324521A8" box="[620,824,439,464]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FDD07C86324521A8" box="[620,824,439,464]" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
) and larger calyx, with 2.3–2.7 × 
<quantity id="65489B48FFA9FF84FB077C88343921B7" box="[1211,1348,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="18" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 mm</quantity>
(versus smaller calyx, with 1–2 × 
<quantity id="65489B48FFA9FF84FE6E7CE3331C218B" box="[466,609,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="18" unit="mm" value="1.5" valueMax="1.9" valueMin="1.1">1.1–1.9 mm</quantity>
in 
<taxonomicName id="65B04D2EFFA9FF84FD357CE23222218D" box="[649,863,476,501]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FD357CE23222218D" box="[649,863,476,501]" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="C6214B5CFFA9FF84FCC67CE335B0218D" author="Medeiros, M. C. M. P. &amp; Lohmann, L. G." box="[890,1229,476,501]" pageId="18" pagination="77 - 85" refId="ref46989" refString="Medeiros, M. C. M. P. &amp; Lohmann, L. G. (2014) Two new species of Tynanthus Miers (Bignonieae, Bignoniaceae) from Brazil. Phytokeys 42: 77 - 85. http: // dx. doi. org / 10.3897 / phytokeys. 42.8210" type="journal article" year="2014">Medeiros &amp; Lohmann 2014</bibRefCitation>
). Additionally, 
<taxonomicName id="65B04D2EFFA9FF84FA3B7CE2302E226F" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFA9FF84FA3B7CE2302E226F" italics="true" pageId="18">T. espiritosantensis</emphasis>
</taxonomicName>
occurs in the Atlantic Forest of 
<collectingRegion id="6074F84FFFA9FF84FD017CC032222260" box="[701,863,511,536]" country="Brazil" name="Espirito Santo" pageId="18">Espírito Santo</collectingRegion>
, while 
<taxonomicName id="65B04D2EFFA9FF84FC0C7F3E35042260" box="[944,1145,511,536]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FC0C7F3E35042260" box="[944,1145,511,536]" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
is an Amazonian species. Apart from their morphological similarity, 
<taxonomicName id="65B04D2EFFA9FF84FD2C7F1A320F2244" authorityName="M. C. Medeiros &amp; L. G. Lohmann" authorityYear="2014" box="[656,882,549,572]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="espiritosantensis">
<emphasis id="90C4EABFFFA9FF84FD2C7F1A320F2244" box="[656,882,549,572]" italics="true" pageId="18">T. espiritosantensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="65B04D2EFFA9FF84FC107F1A35022245" box="[940,1151,548,573]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="schumannianus">
<emphasis id="90C4EABFFFA9FF84FC107F1A35022245" box="[940,1151,548,573]" italics="true" pageId="18">T. schumannianus</emphasis>
</taxonomicName>
are also closely related, belonging to a clade that also contains 
<taxonomicName id="65B04D2EFFA9FF84FDE67F76327D2218" box="[602,768,583,608]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="fasciculatus">
<emphasis id="90C4EABFFFA9FF84FDE67F76327D2218" box="[602,768,583,608]" italics="true" pageId="18">T. fasciculatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="65B04D2EFFA9FF84FCAC7F7632F72218" box="[784,906,583,608]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="labiatus">
<emphasis id="90C4EABFFFA9FF84FCAC7F7632F72218" box="[784,906,583,608]" italics="true" pageId="18">T. labiatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="65B04D2EFFA9FF84FC7F7F76351D2218" box="[963,1120,583,608]" class="Magnoliopsida" family="Bignoniaceae" genus="Tynanthus" kingdom="Plantae" order="Lamiales" pageId="18" phylum="Tracheophyta" rank="species" species="micranthus">
<emphasis id="90C4EABFFFA9FF84FC7F7F76351D2218" box="[963,1120,583,608]" italics="true" pageId="18">T. micranthus</emphasis>
</taxonomicName>
, all distributed through the Atlantic Forest (
<bibRefCitation id="C6214B5CFFA9FF84FEF17F5333F622FD" author="Medeiros, M. C. M. P. &amp; Lohmann, L. G." box="[333,651,620,645]" pageId="18" pagination="32 - 40" refId="ref47044" refString="Medeiros, M. C. M. P. &amp; Lohmann, L. G. (2015) Phylogeny and biogeography of Tynanthus Miers (Bignonieae, Bignoniaceae). Molecular Phylogenetics and Evolution 85: 32 - 40. http: // dx. doi. org / 10.1016 / j. ympev. 2015.01.010" type="journal article" year="2015">Medeiros &amp; Lohmann 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>