treatments-xml/data/0C/2B/6F/0C2B6F42E458B43019E0FDDC0E32D934.xml
2024-06-21 12:29:13 +02:00

915 lines
113 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="AFE56E191643AF1192A8D4FA5AF38D9A" ID-DOI="10.5281/zenodo.293380" ID-GBIF-Dataset="6df5e2ab-4290-4cf9-8ec9-fa8af2c28c4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="293380" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460247565093" checkinUser="plazi" docAuthor="Coad, Brian W. &amp; Bogutskaya, Nina G." docDate="2010" docId="0C2B6F42E458B43019E0FDDC0E32D934" docLanguage="en" docName="zt02534p047.pdf" docOrigin="Zootaxa 2534" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Petroleuciscus esfahani Coad &amp; Bogutskaya, 2010, new species" docType="treatment" docVersion="7" lastPageNumber="45" masterDocId="F012173AE459B4381977FF890A01DF3E" masterDocTitle="Petroleuciscus esfahani, a new species of fish from central Iran (Actinopterygii: Cyprinidae)" masterLastPageNumber="47" masterPageNumber="37" pageNumber="38" updateTime="1698242085118" updateUser="plazi">
<mods:mods id="07E98ECB64711B5BC72FB38CB4A08123" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="13B645C6AB7FCE5E2B6342929297EB3F">
<mods:title id="C8B2308B29862A9134DC7869A71F4D79">Petroleuciscus esfahani, a new species of fish from central Iran (Actinopterygii: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="716D05655B193E9245D98874B72BF026" type="personal">
<mods:role id="5BE3AE380EE6081907453EFC1BA33D31">
<mods:roleTerm id="3B3D6271A8A33F5EC09C4AB112015D2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="422E0B4E031D9EABD887CE2B1D49EED3">Coad, Brian W.</mods:namePart>
</mods:name>
<mods:name id="938468F299ACFC1B1B714737382B6AD5" type="personal">
<mods:role id="FFC1EF3DBC63A58B128E1DD128534472">
<mods:roleTerm id="47D69958D5B29137CB0AB9017A32DD22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4666036433601F92EA59CC158BD4F035">Bogutskaya, Nina G.</mods:namePart>
</mods:name>
<mods:typeOfResource id="721CF1BF58005D4DD1EEAAD694773162">text</mods:typeOfResource>
<mods:relatedItem id="24F60DD69560EE9BE1C2D7C8C60A7785" type="host">
<mods:titleInfo id="F59C0870937C0F279E71AB857CC6F30D">
<mods:title id="392081E173025C529CE5E8C4F0729DE3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="74C010A96D71719E9AC34CA5022648C3">
<mods:date id="20817F553056B96C181BAF4C90876856">2010</mods:date>
<mods:detail id="9CC0E2AFB44336651C1214DE33D20A78" type="volume">
<mods:number id="A6167998FB6E74408B812379623C2A8D">2534</mods:number>
</mods:detail>
<mods:extent id="E41D6017F72240B19DBFCEAFC46FF1C0" unit="page">
<mods:start id="6900FF73CE6947C08919ADA1F12E4E99">37</mods:start>
<mods:end id="18E6CD62E3A630C975A75CBEDE70B53E">47</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F9955CAD41ED63E1227F80C9157AC84F">journal article</mods:classification>
<mods:identifier id="5BE2F9CE8E6931786E827CA91EB32038" type="DOI">10.5281/zenodo.293380</mods:identifier>
<mods:identifier id="A9D20E623514B0592332B019A95B6551" type="GBIF-Dataset">6df5e2ab-4290-4cf9-8ec9-fa8af2c28c4c</mods:identifier>
<mods:identifier id="657D4E794B8D4867F5572159AA4B774F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8E1243248FD48ACBAC6BC8510C6C6AB5" type="Zenodo-Dep">293380</mods:identifier>
</mods:mods>
<treatment id="0C2B6F42E458B43019E0FDDC0E32D934" ID-DOI="http://doi.org/10.5281/zenodo.5612490" ID-GBIF-Taxon="119390677" ID-Zenodo-Dep="5612490" LSID="urn:lsid:plazi:treatment:0C2B6F42E458B43019E0FDDC0E32D934" httpUri="http://treatment.plazi.org/id/0C2B6F42E458B43019E0FDDC0E32D934" lastPageId="8" lastPageNumber="45" pageId="1" pageNumber="38">
<subSubSection id="CC988DDFE458B43919E0FDDC0B37DDAC" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="843DDE54E458B43919E0FDDC0854DD51" blockId="1.[151,597,597,658]" box="[151,597,597,623]" pageId="1" pageNumber="38">
<heading id="DF756938E458B43919E0FDDC0854DD51" bold="true" box="[151,597,597,623]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B6F60246E458B43919E0FDDC0854DD51" bold="true" box="[151,597,597,623]" pageId="1" pageNumber="38">
<taxonomicName id="4382A5D7E458B43919E0FDDC0BB7DD51" ID-CoL="776C7" box="[151,438,597,623]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E458B43919E0FDDC0BB7DD51" bold="true" box="[151,438,597,623]" italics="true" pageId="1" pageNumber="38">Petroleuciscus esfahani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADC5BF3DE458B43918B2FDDC0854DD51" box="[453,597,597,623]" pageId="1" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="843DDE54E458B43919E0FDF10B37DDAC" blockId="1.[151,597,597,658]" box="[151,310,632,658]" pageId="1" pageNumber="38">
(
<figureCitation id="1CB9C2D1E458B43919E8FDF10AE4DDAC" box="[159,229,632,658]" captionStart="FIGURE 1" captionStartId="1.[151,255,1481,1505]" captionTargetBox="[171,1414,938,1457]" captionTargetId="figure@1.[171,1416,938,1457]" captionTargetPageId="1" captionText="FIGURE 1. Petroleuciscus esfahani sp. n., female holotype, CMNFI 1979 - 0249, 106.7 mm SL." httpUri="https://zenodo.org/record/293381/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
,
<figureCitation id="1CB9C2D1E458B4391984FDF10B05DDAC" box="[243,260,632,658]" captionStart="FIGURE 2" captionStartId="1.[151,255,1942,1966]" captionTargetBox="[188,1398,1556,1921]" captionTargetId="figure@1.[188,1399,1556,1921]" captionTargetPageId="1" captionText="FIGURE 2. Petroleuciscus esfahani sp. n., male paratype, CMNFI 1979 - 0251, 83.7 mm SL." httpUri="https://zenodo.org/record/294009/files/figure.png" pageId="1" pageNumber="38">2</figureCitation>
,
<figureCitation id="1CB9C2D1E458B4391866FDF10B1EDDAC" box="[273,287,632,658]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="1" pageNumber="38">3</figureCitation>
a)
</paragraph>
</subSubSection>
<subSubSection id="CC988DDFE458B43919E0FD4B081EDC4C" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="843DDE54E458B43919E0FD4B0EC4DC3C" blockId="1.[151,1437,706,882]" pageId="1" pageNumber="38">
<typeStatus id="5B3960F6E458B43919E0FD4B0B08DDE2" box="[151,265,706,732]" pageId="1" pageNumber="38" type="holotype">
<emphasis id="B6F60246E458B43919E0FD4B0B08DDE2" bold="true" box="[151,265,706,732]" pageId="1" pageNumber="38">Holotype</emphasis>
</typeStatus>
(
<figureCitation id="1CB9C2D1E458B439186FFD4B0B5CDDE2" box="[280,349,706,732]" captionStart="FIGURE 1" captionStartId="1.[151,255,1481,1505]" captionTargetBox="[171,1414,938,1457]" captionTargetId="figure@1.[171,1416,938,1457]" captionTargetPageId="1" captionText="FIGURE 1. Petroleuciscus esfahani sp. n., female holotype, CMNFI 1979 - 0249, 106.7 mm SL." httpUri="https://zenodo.org/record/293381/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
):
<collectionCode id="E2934691E458B4391801FD4B0BD2DDE2" box="[374,467,706,732]" pageId="1" pageNumber="38">CMNFI</collectionCode>
1979-0249, female,
<quantity id="437A73B1E458B4391BB1FD4B093CDDE2" box="[710,829,706,732]" metricMagnitude="-1" metricUnit="m" metricValue="1.067" pageId="1" pageNumber="38" unit="mm" value="106.7">106.7 mm</quantity>
SL,
<collectingCountry id="FC959EC4E458B4391A03FD4B09C1DDE2" box="[884,960,706,732]" name="Iran" pageId="1" pageNumber="38">IRAN</collectingCountry>
: Esfahan, stream at Dizaj in the southern Zayandeh River basin,
<geoCoordinate id="E1B6B893E458B43918D1FD61080FDC3C" box="[422,526,744,770]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="925" value="31.916666">31°55N</geoCoordinate>
,
<geoCoordinate id="E1B6B893E458B4391B6DFD61087EDC3C" box="[538,639,744,770]" direction="east" orientation="longitude" pageId="1" pageNumber="38" precision="925" value="51.5">51°30E</geoCoordinate>
, B. W. Coad and Sh. Mansoorabadi,
<date id="F03CF894E458B4391D58FD610EC0DC3C" box="[1071,1217,744,770]" pageId="1" pageNumber="38" value="1977-06-09">9 June 1977</date>
.
</paragraph>
<paragraph id="843DDE54E458B43919B2FC84081EDC4C" blockId="1.[151,1437,706,882]" pageId="1" pageNumber="38">
<emphasis id="B6F60246E458B43919B2FC840B53DC19" bold="true" box="[197,338,781,807]" pageId="1" pageNumber="38">
<typeStatus id="5B3960F6E458B43919B2FC840B4CDC19" box="[197,333,781,807]" pageId="1" pageNumber="38" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="E2934691E458B439182CFC840BBDDC19" box="[347,444,781,807]" pageId="1" pageNumber="38">CMNFI</collectionCode>
1979-0251, 134,
<quantity id="437A73B1E458B4391BEBFC840950DC19" box="[668,849,781,807]" metricMagnitude="-2" metricUnit="m" metricValue="5.29" metricValueMax="8.37" metricValueMin="2.21" pageId="1" pageNumber="38" unit="mm" value="52.9" valueMax="83.7" valueMin="22.1">22.183.7 mm</quantity>
SL,
<collectingCountry id="FC959EC4E458B4391AF8FC8409C4DC19" box="[911,965,781,807]" name="Iran" pageId="1" pageNumber="38">Iran</collectingCountry>
, Esfahan, stream
<quantity id="437A73B1E458B4391DD4FC840EE2DC19" box="[1187,1251,781,807]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="38" unit="km" value="1.0">1 km</quantity>
east of Daran, Pelasegan River tributary in the northern Zayandeh River basin,
<geoCoordinate id="E1B6B893E458B4391AAEFCBB0E48DC72" box="[985,1097,818,844]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="925" value="32.983334">32°59N</geoCoordinate>
,
<geoCoordinate id="E1B6B893E458B4391D2FFCBB0EC4DC72" box="[1112,1221,818,844]" direction="east" orientation="longitude" pageId="1" pageNumber="38" precision="925" value="50.433334">50°26E</geoCoordinate>
, B. W. Coad and Sh. Mansoorabadi,
<date id="F03CF894E458B439180CFCD1081BDC4C" box="[379,538,856,882]" pageId="1" pageNumber="38" value="1977-06-10">10 June 1977</date>
.
</paragraph>
</subSubSection>
<caption id="D0FD8EDCE458B43919E0FA400E9ADADE" box="[151,1179,1481,1505]" httpUri="https://zenodo.org/record/293381/files/figure.png" pageId="1" pageNumber="38" targetBox="[171,1414,938,1457]" targetPageId="1">
<paragraph id="843DDE54E458B43919E0FA400E9ADADE" blockId="1.[151,1179,1481,1505]" box="[151,1179,1481,1505]" pageId="1" pageNumber="38">
<emphasis id="B6F60246E458B43919E0FA400B18DADF" bold="true" box="[151,281,1481,1505]" pageId="1" pageNumber="38">FIGURE 1.</emphasis>
<taxonomicName id="4382A5D7E458B4391857FA43081DDADF" box="[288,540,1482,1505]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E458B4391857FA43081DDADF" box="[288,540,1482,1505]" italics="true" pageId="1" pageNumber="38">Petroleuciscus esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE458B4391B54FA40085EDADF" box="[547,607,1481,1505]" pageId="1" pageNumber="38" rank="species">
<emphasis id="B6F60246E458B4391B54FA40085EDADF" bold="true" box="[547,607,1481,1505]" pageId="1" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
, female holotype, CMNFI 1979-0249, 106.7 mm SL.
</paragraph>
</caption>
<caption id="D0FD8EDCE458B43919E0F81F0E79D890" box="[151,1144,1942,1966]" httpUri="https://zenodo.org/record/294009/files/figure.png" pageId="1" pageNumber="38" targetBox="[188,1398,1556,1921]" targetPageId="1">
<paragraph id="843DDE54E458B43919E0F81F0E79D890" blockId="1.[151,1144,1942,1966]" box="[151,1144,1942,1966]" pageId="1" pageNumber="38">
<emphasis id="B6F60246E458B43919E0F81F0B18D890" bold="true" box="[151,281,1942,1966]" pageId="1" pageNumber="38">FIGURE 2.</emphasis>
<taxonomicName id="4382A5D7E458B4391857F81E081DD890" box="[288,540,1943,1966]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E458B4391857F81E081DD890" box="[288,540,1943,1966]" italics="true" pageId="1" pageNumber="38">Petroleuciscus esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE458B4391B54F81F085ED890" box="[547,607,1942,1966]" pageId="1" pageNumber="38" rank="species">
<emphasis id="B6F60246E458B4391B54F81F085ED890" bold="true" box="[547,607,1942,1966]" pageId="1" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
, male paratype, CMNFI 1979-0251, 83.7 mm SL.
</paragraph>
</caption>
<caption id="D0FD8EDCE45BB43A19E0F8CF0AF7D8D4" httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="2" pageNumber="39" targetBox="[305,1279,204,1823]" targetPageId="2">
<paragraph id="843DDE54E45BB43A19E0F8CF0AF7D8D4" blockId="2.[151,1436,1862,2026]" pageId="2" pageNumber="39">
<emphasis id="B6F60246E45BB43A19E0F8CF0B18D860" bold="true" box="[151,281,1862,1886]" pageId="2" pageNumber="39">FIGURE 3.</emphasis>
Radiograph of
<taxonomicName id="4382A5D7E45BB43A18B6F8CE08BFD860" box="[449,702,1863,1886]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E45BB43A18B6F8CE08BFD860" box="[449,702,1863,1886]" italics="true" pageId="2" pageNumber="39">Petroleuciscus esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE45BB43A1BB2F8CF08FAD860" box="[709,763,1862,1886]" pageId="2" pageNumber="39" rank="species">
<emphasis id="B6F60246E45BB43A1BB2F8CF08FAD860" bold="true" box="[709,763,1862,1886]" pageId="2" pageNumber="39">sp. n</emphasis>
</taxonomicNameLabel>
, holotype (a);
<taxonomicName id="4382A5D7E45BB43A1AEFF8CE0E46D860" box="[920,1095,1863,1886]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="borysthenicus">
<emphasis id="B6F60246E45BB43A1AEFF8CE0E46D860" box="[920,1095,1863,1886]" italics="true" pageId="2" pageNumber="39">P. borysthenicus</emphasis>
</taxonomicName>
, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b);
<taxonomicName id="4382A5D7E45BB43A18B5F8E00854D8BE" box="[450,597,1897,1920]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="smyrnaeus">
<emphasis id="B6F60246E45BB43A18B5F8E00854D8BE" box="[450,597,1897,1920]" italics="true" pageId="2" pageNumber="39">P. smyrnaeus</emphasis>
</taxonomicName>
, syntype, BMNH 1895.12.28.19-28, Smyrna, Turkey, 136.0 mm SL (c);
<taxonomicName id="4382A5D7E45BB43A1CFFF8E00B2BD89D" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E45BB43A1CFFF8E00B2BD89D" italics="true" pageId="2" pageNumber="39">P. squaliusculus</emphasis>
</taxonomicName>
, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and
<taxonomicName id="4382A5D7E45BB43A1D63F8050E84D89D" box="[1044,1157,1932,1955]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E45BB43A1D63F8050E84D89D" box="[1044,1157,1932,1955]" italics="true" pageId="2" pageNumber="39">P. persidis</emphasis>
</taxonomicName>
, holotype, CMNFI 1979- 0154A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra.
</paragraph>
</caption>
<subSubSection id="CC988DDFE45AB43B19B1FF1E094DDE79" pageId="3" pageNumber="40" type="diagnosis">
<paragraph id="843DDE54E45AB43B19B1FF1E094DDE79" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">
<emphasis id="B6F60246E45AB43B19B1FF1E0B47DF8F" bold="true" box="[198,326,151,177]" pageId="3" pageNumber="40">Diagnosis.</emphasis>
The species is distinguished by a combination of characters from other members of the genus, including a mode of 8½ dorsal-fin branched rays, modally 1011½ anal-fin branched rays, small scales on the flank numbering 4454 pored scales to the hypural fold while total lateral-line scales number 4556, a pharyngeal-tooth formula commonly 2.54.2, and total vertebrae commonly 4042, the abdominal + caudal vertebral formulae being usually 22+19, 22+20 and 21+20.
</paragraph>
</subSubSection>
<subSubSection id="CC988DDFE45AB43C19B1FEDB0E9AD822" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="description">
<paragraph id="843DDE54E45AB43B19B1FEDB0E8DDD19" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">
<emphasis id="B6F60246E45AB43B19B1FEDB0B5EDE52" bold="true" box="[198,351,338,364]" pageId="3" pageNumber="40">Description.</emphasis>
<typeStatus id="5B3960F6E45AB43B1810FEDB0BD9DE52" box="[359,472,338,364]" pageId="3" pageNumber="40" type="holotype">Holotype</typeStatus>
. Dorsal-fin with 3 unbranched and 8½ branched rays, anal fin with 3 unbranched and 10½ branched rays, pectoral fin with 14 branched rays, and pelvic fin with 8 branched rays. Scales in lateral line to hypural fold 45, total lateral-line scales 47, scales around caudal peduncle 15, predorsal scales 23, scales between lateral line and dorsal-fin origin 9, scales between lateral line and anal-fin origin 6, and scales between lateral line and pelvic-fin origin 5. Total gill rakers in outer row of first gill arch 13, d total vertebrae 41: 22 abdominal (including 14 predorsal vertebrae) and 19 caudal (
<figureCitation id="1CB9C2D1E45AB43B1D5EFD840E6EDD19" box="[1065,1135,525,551]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="3" pageNumber="40">Fig. 3</figureCitation>
a).
</paragraph>
<paragraph id="843DDE54E45AB43B19B2FDBB0F22DC12" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">
The body is rounded in cross section with the upper body profile almost straight and the lower body profile gently convex. The snout is triangular and pointed. The lower jaw projects slightly compared to the upper jaw, forming a distinct chin such that the mouth is slightly superior. The tip of the mouth cleft is on a level with the upper margin of the pupil. The mouth cleft extends back to a level with the rear of the nostrils, and the lower jaw-quadrate junction is at the vertical through the anterior margin of the eye (
<figureCitation id="1CB9C2D1E45AB43B1D89FD410F47DDDC" box="[1278,1350,712,738]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="3" pageNumber="40">Fig. 3</figureCitation>
a). The anal-fin origin is behind the dorsal-fin insertion. The dorsal-fin is truncate and the anal-fin margin is rounded anteriorly, becoming truncate posteriorly. The caudal fin has a shallow fork with the lobes rounded.
</paragraph>
<paragraph id="843DDE54E45AB43B19B2FCB10F97DCFC" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">The lateral line is moderately decurved and is centrally located on the caudal peduncle. There is a pelvic axillary scale. Scales below the dorsal-fin have a subcentral anterior focus, numerous fine circuli, relatively few anterior and posterior radii with no radii in the lateral fields, a rounded posterior margin and a wavy anterior margin. Gill rakers are moderately elongate and extend to the second adjacent raker when appressed.</paragraph>
<paragraph id="843DDE54E45AB43B19B2FC440B12DBD2" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">
General topography of cephalic sensory canals and numbers of pores are typical of most
<taxonomicName id="4382A5D7E45AB43B1D9FFC440F95DCD8" box="[1256,1428,973,998]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45AB43B1D9FFC440F95DCD8" box="[1256,1428,973,998]" italics="true" pageId="3" pageNumber="40">Petroleuciscus</emphasis>
</taxonomicName>
, as described by
<bibRefCitation id="E013A3A5E45AB43B1824FC7B082ADB32" author="Bogutskaya" box="[339,555,1010,1036]" pageId="3" pageNumber="47" refString="Bogutskaya, N. G. (1995) Leuciscus kurui, a new cyprinid fish from the upper Tigris (Dicle) system. Mitteilungen aus dem hamburgischen Zoologischen Museum und Institut, 92, 149 - 154." type="journal article" year="1995">Bogutskaya (1995</bibRefCitation>
,
<bibRefCitation id="E013A3A5E45AB43B1B4FFC7B0874DB32" author="Bogutskaya" box="[568,629,1010,1036]" pageId="3" pageNumber="47" refString="Bogutskaya, N. G. (1996) Contribution to the knowledge of leuciscine fishes of Asia Minor. Part 1. Morphology and taxonomic relationships of Leuciscus borysthenicus (Kessler), Leuciscus smyrnaeus Boulenger and Ladigoesocypris ghigii (Gianferrari) (Cyprinidae, Pisces). Publicaciones Especiales Instituto Espanol de Oceanografia, 21, 25 - 44." type="journal article" year="1996">1996</bibRefCitation>
,
<bibRefCitation id="E013A3A5E45AB43B1BF6FC7B08C0DB32" author="Bogutskaya" box="[641,705,1010,1036]" pageId="3" pageNumber="47" refString="Bogutskaya, N. G. (2002) Petroleuciscus, a new genus for the Leuciscus borysthenicus species group (Teleostei: Cyprinidae). Zoosystematica Rossica, 11 (1), 235 - 237." type="journal article" year="2002">2002</bibRefCitation>
). The supraorbital canal is short and terminates at the parietalfrontal border (thus not entering the parietal), and has 9 pores, with 3 and 7 canal openings on the nasal and frontal bones, respectively. The infraorbital canal has 18 (left side) and 16 (right side) pores with 5 (left side) and 4 (right side) canal openings on the first infraorbital. The preopercular-mandibular canal is complete, with 17 (left side) and 16 (right side) pores with 3 (left side) and 2 canal openings on the dentary, 10 on the preoperculum and 2 on the operculum. The supratemporal canal is incomplete (mesially interrupted), with 3+3 pores.
</paragraph>
<paragraph id="843DDE54E45AB43B19B2FB710BBBDACC" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">There is no distinctive colour pattern in preserved fish. The flank above the lateral line bears a fine dark speckling of melanophores and is mostly unpigmented below the lateral line. Melanophores are present on the upper part of the operculum and extend down along the posterior edge. Melanophores on the cheek ring the lower orbit. The back bears a predorsal and postdorsal stripe. The dorsal and caudal fins have very fine melanophores on the rays only. The pectoral fin has similar fine melanophores on its anterior rays only. The anal and pelvic fins are almost unpigmented. The peritoneum is silvery to cream with very few, widely scattered, melanophores.</paragraph>
<paragraph id="843DDE54E45AB43B19B2FA7409F3D9B9" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">
<typeStatus id="5B3960F6E45AB43B19B2FA740B3AD929" box="[197,315,1533,1559]" pageId="3" pageNumber="40" type="paratype">Paratypes</typeStatus>
. General characters of the
<typeStatus id="5B3960F6E45AB43B1B04FA7408D8D929" box="[627,729,1533,1559]" pageId="3" pageNumber="40" type="holotype">holotype</typeStatus>
are also found in the
<typeStatus id="5B3960F6E45AB43B1AAFFA740E4DD929" box="[984,1100,1533,1559]" pageId="3" pageNumber="40" type="paratype">paratypes</typeStatus>
. Measurements are based on 15 fish of each sex, males
<quantity id="437A73B1E45AB43B1898F9AB08A6D902" box="[495,679,1570,1596]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.37" metricValueMin="3.63" pageId="3" pageNumber="40" unit="mm" value="60.0" valueMax="83.7" valueMin="36.3">36.383.7 mm</quantity>
SL and females 36.566.0 mm SL. Morphometric data are summarised in
<tableCitation id="C900EBEFE45AB43B183BF9C10BA4D95C" box="[332,421,1608,1634]" captionStart="TABLE 1" captionStartId="5.[151,240,300,324]" captionTargetBox="[151,1436,384,1870]" captionTargetPageId="5" captionText="TABLE 1. Morphometric data for the holotype, 15 male paratypes and 15 female paratypes (values are mean, standard deviation and range in parentheses)." httpUri="http://table.plazi.org/id/D0FD8EDCE45CB43D19E0FEA50819DE5A" pageId="3" pageNumber="40" tableUuid="D0FD8EDCE45CB43D19E0FEA50819DE5A">Table 1</tableCitation>
. Males had a significantly larger dorsal-fin base than females while this was reversed for the sexes in the anal-fin base, possibly associated with pair spawning.
</paragraph>
<paragraph id="843DDE54E45AB43B19B2F91B088FD87C" blockId="3.[151,1437,151,2007]" pageId="3" pageNumber="40">The head length is about equal to body depth. The snout is short, its length equals to or slightly smaller than the orbit width. The tip of the mouth cleft is on a level with the upper margin of the pupil in large-sized individuals such that the mouth is slightly superior or somewhat below, at about the level of the middle of the eye, in small-sized ones. The lower jaw projects slightly compared to the upper jaw, often forming a distinct chin especially visible in larger specimens.</paragraph>
<paragraph id="843DDE54E45AB43C19B2F8C40F08DEE2" blockId="3.[151,1437,151,2007]" lastBlockId="4.[151,1437,152,2007]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
There were no significant differences (p&gt;0.05) between males and females for all meristic characters and data for the two sexes were combined. Dorsal-fin with 3 unbranched rays and 7½ (22), 8½ (107) or 9½ (5) branched rays (mean 7.9, standard deviation 0.43); anal fin with 3 unbranched rays and 9½ (14), 10½ (90), 11½ (25) or 12½ (5) branched rays (10.2, 0.65); pectoral fin with 14(4), 15(17) or 16(9) branched rays (15.2, 0.65); pelvic fin with 7(1), 8(27) or 9(2) branched rays (8.0, 0.32); lateral-line scales to hypural fold (in
<specimenCount id="928415DDE45DB43C1C09FF110B12DFE9" pageId="4" pageNumber="41" type="generic">30 specimens</specimenCount>
) 45(2), 46(1), 47(3), 48(8), 49(3), 50(1), 51(4), 52(4), 53(2), 54(1), 55(-) or 56(1) (49.5, 2.70); total lateral line scales (in
<specimenCount id="928415DDE45DB43C18EAFF6B0842DFC2" box="[413,579,226,252]" pageId="4" pageNumber="41" type="generic">60 specimens</specimenCount>
) 45(1), 46(2), 47(3), 48(2), 49(6), 50(14), 51(16), 52(7), 53(5), 54(2), 55(1) or 56(1) (50.5, 2.11); among 134
<typeStatus id="5B3960F6E45DB43C1BF5FE8108F6DE1C" box="[642,759,264,290]" pageId="4" pageNumber="41" type="paratype">paratypes</typeStatus>
there are also
<specimenCount id="928415DDE45DB43C1AD8FE810E47DE1C" box="[943,1094,264,290]" pageId="4" pageNumber="41" type="generic">8 specimens</specimenCount>
with the lateral line widely interrupted (2840 pored lateral-line scales from 5053 scales in lateral series); scales around caudal peduncle 14(3), 15(11), 16(13), 17(2) or 18(1) (15.6, 0.90); predorsal scales 23(11), 24(3), 25(5), 26(10) or 27(1) (24.6, 1.38); scales between lateral line and dorsal-fin origin 9(8), 10(17) or 11(5) (9.9, 0.66); scales between lateral line and anal-fin origin 4(6), 5(22) or 6(2) (4.9, 0.51); scales between lateral line and pelvic-fin origin 3(2), 4(17) or 5(11) (4.3, 0.60); total gill rakers 12(2), 13(3), 14(8), 15(10), 16(6) or 17(1) (14.6, 1.22).
</paragraph>
<paragraph id="843DDE54E45DB43C19B2FE610E1ADD72" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">
Pharyngeal teeth 2.54.2 (8) or 2.55.2(2), hooked at the tip and strongly serrated below it. The gut is an elongate s-shape. Pigmentation is generally the same as in the
<typeStatus id="5B3960F6E45DB43C1AFFFD8409F1DD19" box="[904,1008,525,551]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
except melanophores on the lateral sides of the head are larger and more evident, particularly in the smaller fish.
</paragraph>
<paragraph id="843DDE54E45DB43C19B2FDD10BFCDCFC" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">
The sensory canal system was examined in the
<typeStatus id="5B3960F6E45DB43C1B89FDD10967DD4C" box="[766,870,600,626]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
and 19
<typeStatus id="5B3960F6E45DB43C1ABFFDD10E3CDD4C" box="[968,1085,600,626]" pageId="4" pageNumber="41" type="paratype">paratypes</typeStatus>
, so the numbers of examined paired canals is 40. The supraorbital canal is not lengthened in its posterior section and has 812, commonly 9 or 10 pores (mean 9.6, standard deviation 1.33), with 3 or 4 (
<quantity id="437A73B1E45DB43C1A16FD2B098FDD82" box="[865,910,674,700]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="41" unit="in" value="3.0">3 in</quantity>
66%) and 58 (counts 6 and 7 found each in 30%) canal openings on the nasal and frontal bones, respectively. The infraorbital canal has (14, 15)1618 pores (a mode of 16 found in 38% of canals; mean 16.6, standard deviation 1.01) with 4 or 5 (in 75%) canal openings on the first infraorbital. The preoperculo-mandibular canal is complete, with 1517 pores (
<quantity id="437A73B1E45DB43C1C29FC9B0F9DDC12" box="[1374,1436,786,812]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="41" unit="in" value="15.0">15 in</quantity>
58%; 15.5, 0.74) with 4 or 5 (in 83%) and (8)910 (
<quantity id="437A73B1E45DB43C1A3CFCB10981DC6C" box="[843,896,824,850]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="41" unit="in" value="9.0">9 in</quantity>
50%) canal openings on the dentary and preoperculum, respectively. It always communicates with the infraorbital canal in the pterotic, passing through the antero-dorsal process of the operculum. The supratemporal canal is complete (60% of specimens) or incomplete, with 46 pores.
</paragraph>
<paragraph id="843DDE54E45DB43C19B2FC44097DDBD2" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">The total number of vertebrae is 39(2), 40(22), 41(70), 42(35) or 43(5) (41.1, 0.79); the number of abdominal (precaudal) vertebrae includes the Weberian and intermediate vertebrae (those vertebrae that lost articulation with ribs and differ in the degree of transformation of the parapophyses into the haemal arch with the haemal spine); it is 21(39), 22(86) or 23(9) (21.8, 0.56); the number of caudal vertebrae is 18(12), 19(65), 20(53) or 21(4) (19.4, 0.69); and the vertebral formula is 22+19(46), 22+20(28), 21+20(23), 21+19(13), 22+18(9), 23+19(6), 22+21(3), 23+20(2), 21+18(2), 23+18(1) or 21+21(1). The number of predorsal vertebrae (in front of the first dorsal pterygiophore) is 14(51), 15(76) or 16(7) (14.7, 0.57) and the number of intermediate vertebrae is 3(17), 4(69), 5(46) or 6(2) (4.2, 0.65).</paragraph>
<paragraph id="843DDE54E45DB43C19B2FB7108D5D9B9" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">
Other osteological characters available from the undissected specimens (n=10), the radiographs and three cleared-and-stained specimens are as follows. Maximum cranial width between the lateral margins of the pterotics (Lt pto) is 6874% of cranial roof length. The supraethmoid is relatively small, its width being 25 30% Lt pto. The neurocranium is moderately deep, with an angled parasphenoid and a well-defined interorbital septum of the orbitosphenoid. The pharyngeal process has a laterally compressed deep posterior portion. The length of the lower jaw, 3238% of HL, is smaller than the operculum depth (3841% HL). The 2nd4th infraorbitals are narrow. The 5th infraorbital is sometimes absent (on both sides of the head in 2 from 20 examined specimens, and on one side, left or right, in
<specimenCount id="928415DDE45DB43C1A4DFA7409D1D929" box="[826,976,1533,1559]" pageId="4" pageNumber="41" type="generic">5 specimens</specimenCount>
); if present, it is tube-like or has a very narrow lamellate portion. The postcleithrum is reduced to a very small, laterally compressed oval bone with a slightly pointed lower end; it almost entirely adjoins the posterior corner of the cleithrum along its ascending margin. The 2nd and 3rd vertebrae are not fused.
</paragraph>
<paragraph id="843DDE54E45DB43C19B2F91B0E9AD822" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">
Tubercles on two male fish,
<quantity id="437A73B1E45DB43C1B60F91B08C2D992" box="[535,707,1682,1708]" metricMagnitude="-2" metricUnit="m" metricValue="8.309999999999999" metricValueMax="8.37" metricValueMin="8.25" pageId="4" pageNumber="41" unit="mm" value="83.1" valueMax="83.7" valueMin="82.5">82.583.7 mm</quantity>
SL, are fine and scattered on the upper head extending down onto the upper half of the operculum but weak to absent on the snout (
<figureCitation id="1CB9C2D1E45DB43C1AA4F9310E1AD9EC" box="[979,1051,1720,1746]" captionStart="FIGURE 2" captionStartId="1.[151,255,1942,1966]" captionTargetBox="[188,1398,1556,1921]" captionTargetId="figure@1.[188,1399,1556,1921]" captionTargetPageId="1" captionText="FIGURE 2. Petroleuciscus esfahani sp. n., male paratype, CMNFI 1979 - 0251, 83.7 mm SL." httpUri="https://zenodo.org/record/294009/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
). Fine tubercles lining scales are prominent behind the head dorsally but become less evident posteriorly and ventrally. The pectoral fin bears fine tubercles on the anterior rays but numbers decrease to none on more posterior rays.
</paragraph>
</subSubSection>
<subSubSection id="CC988DDFE45DB43C19B2F8AE090CD859" pageId="4" pageNumber="41" type="etymology">
<paragraph id="843DDE54E45DB43C19B2F8AE090CD859" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="41">
<emphasis id="B6F60246E45DB43C19B2F8AE0B57D87F" bold="true" box="[197,342,1831,1857]" pageId="4" pageNumber="41">Etymology.</emphasis>
The new species is named after the central Iranian drainage basin which is named for its principal city of Esfahan, the third largest city in
<collectingCountry id="FC959EC4E45DB43C1BA1F8C40908D859" box="[726,777,1869,1895]" name="Iran" pageId="4" pageNumber="41">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CC988DDFE45DB43F19B2F8FB0940DD3C" lastPageId="7" lastPageNumber="44" pageId="4" pageNumber="41" type="distribution">
<paragraph id="843DDE54E45DB43D19B2F8FB0B8BDFC2" blockId="4.[151,1437,152,2007]" lastBlockId="5.[151,1436,152,252]" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
<emphasis id="B6F60246E45DB43C19B2F8FB0802D8B2" bold="true" box="[197,515,1906,1932]" pageId="4" pageNumber="41">Distribution and Habitat.</emphasis>
The new species is known only from two streams in the Esfahan basin, which has the Zayandeh as its major river (
<figureCitation id="1CB9C2D1E45DB43C1B33F811088FD88C" box="[580,654,1944,1970]" captionStart="FIGURE 4" captionStartId="6.[151,255,1150,1174]" captionTargetBox="[160,1426,185,1117]" captionTargetId="figure@6.[151,1436,175,1125]" captionTargetPageId="6" captionText="FIGURE 4. Distribution map (solid square = holotype of Petroleuciscus esfahani sp. n., open square = paratypes; solid circles = P. persidis holotype and paratypes (southern circle) and paratypes (northern circle), open circles = other material; solid triangles = P. ulanus syntypes (northern triangle), solid triangles = P. gaderanus syntypes (other triangles), open triangles = other material; star = P. kurui holotype and paratypes)." httpUri="https://zenodo.org/record/294010/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
). The Zayandeh and other, smaller rivers and streams of this basin flow from the eastern slopes of the Zagros Mountains to terminate in the salt swamp Gav Khuni. The stream at Dizaj is a southern or right bank tributary of the Zayandeh. The second locality is a stream tributary to the Pelasagan River which flowed into the northern or left bank of the Zayandeh. The Pelasegan now enters the Zayandeh Reservoir.
</paragraph>
<caption id="D0FD8EDCE45CB43D19E0FEA50819DE5A" ID-Table-UUID="D0FD8EDCE45CB43D19E0FEA50819DE5A" httpUri="http://table.plazi.org/id/D0FD8EDCE45CB43D19E0FEA50819DE5A" pageId="5" pageNumber="42" targetBox="[151,1436,384,1870]" targetIsTable="true" targetPageId="5">
<paragraph id="843DDE54E45CB43D19E0FEA50819DE5A" blockId="5.[151,1436,300,356]" pageId="5" pageNumber="42">
<emphasis id="B6F60246E45CB43D19E0FEA50B0ADE7A" bold="true" box="[151,267,300,324]" pageId="5" pageNumber="42">TABLE 1.</emphasis>
Morphometric data for the holotype, 15 male paratypes and 15 female paratypes (values are mean, standard deviation and range in parentheses).
</paragraph>
</caption>
<paragraph id="843DDE54E45CB43D19E8FE090F51D877" pageId="5" pageNumber="42">
<table id="F6822CF4E45C4BC719E0FE090F9DD870" box="[151,1436,384,1870]" gridcols="5" gridrows="22" pageId="5" pageNumber="42">
<tr id="3AB2DC16E45C4BC719E0FE090F9DDEFF" box="[151,1436,384,449]" gridrow="0" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FE090BECDEFF" box="[151,493,384,449]" gridcol="0" gridrow="0" pageId="5" pageNumber="42">Character Size (mm SL)</th>
<th id="7963B56AE45C4BC71B34FE0908A7DEFF" box="[579,678,384,449]" gridcol="1" gridrow="0" pageId="5" pageNumber="42">Holotype 106.7</th>
<th id="7963B56AE45C4BC71B9FFE0909C5DEFF" box="[744,964,384,449]" gridcol="2" gridrow="0" pageId="5" pageNumber="42">Male paratypes</th>
<th id="7963B56AE45C4BC71A8EFE090EC6DEFF" box="[1017,1223,384,449]" gridcol="3" gridrow="0" pageId="5" pageNumber="42">Female paratypes</th>
<th id="7963B56AE45C4BC71C63FE090F9DDEFF" box="[1300,1436,384,449]" gridcol="4" gridrow="0" pageId="5" pageNumber="42">p</th>
</tr>
<tr id="3AB2DC16E45C4BC719E0FE5C0F9DDED2" box="[151,1436,469,492]" gridrow="1" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FE5C0BECDED2" box="[151,493,469,492]" gridcol="0" gridrow="1" pageId="5" pageNumber="42">SL/HL</th>
<td id="7963B56AE45C4BC71B34FE5C08A7DED2" box="[579,678,469,492]" gridcol="1" gridrow="1" pageId="5" pageNumber="42">4.4</td>
<td id="7963B56AE45C4BC71B9FFE5C09C5DED2" box="[744,964,469,492]" gridcol="2" gridrow="1" pageId="5" pageNumber="42">4.0, 0.17 (3.74.4)</td>
<td id="7963B56AE45C4BC71A8EFE5C0EC6DED2" box="[1017,1223,469,492]" gridcol="3" gridrow="1" pageId="5" pageNumber="42">4.0, 0.14 (3.74.3)</td>
<td id="7963B56AE45C4BC71C63FE5C0F9DDED2" box="[1300,1436,469,492]" gridcol="4" gridrow="1" pageId="5" pageNumber="42">0.807</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FD890F9DDD7F" box="[151,1436,512,577]" gridrow="2" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FD890BECDD7F" box="[151,493,512,577]" gridcol="0" gridrow="2" pageId="5" pageNumber="42">SL/Head depth SL/Head width</th>
<td id="7963B56AE45C4BC71B34FD8908A7DD7F" box="[579,678,512,577]" gridcol="1" gridrow="2" pageId="5" pageNumber="42">5.6 7.3</td>
<td id="7963B56AE45C4BC71B9FFD8909C5DD7F" box="[744,964,512,577]" gridcol="2" gridrow="2" pageId="5" pageNumber="42">5.3, 0.29 (5.05.9) 7.3, 0.40 (6.77.9)</td>
<td id="7963B56AE45C4BC71A8EFD890EC6DD7F" box="[1017,1223,512,577]" gridcol="3" gridrow="2" pageId="5" pageNumber="42">5.2, 0.36 (4.25.6) 7.3, 0.32 (6.77.9)</td>
<td id="7963B56AE45C4BC71C63FD890F9DDD7F" box="[1300,1436,512,577]" gridcol="4" gridrow="2" pageId="5" pageNumber="42">0.681 0.650</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FDDC0F9DDD52" box="[151,1436,597,620]" gridrow="3" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FDDC0BECDD52" box="[151,493,597,620]" gridcol="0" gridrow="3" pageId="5" pageNumber="42">SL/Body depth</th>
<td id="7963B56AE45C4BC71B34FDDC08A7DD52" box="[579,678,597,620]" gridcol="1" gridrow="3" pageId="5" pageNumber="42">4.0</td>
<td id="7963B56AE45C4BC71B9FFDDC09C5DD52" box="[744,964,597,620]" gridcol="2" gridrow="3" pageId="5" pageNumber="42">4.1, 0.18 (3.84.5)</td>
<td id="7963B56AE45C4BC71A8EFDDC0EC6DD52" box="[1017,1223,597,620]" gridcol="3" gridrow="3" pageId="5" pageNumber="42">4.2, 0.34 (3.75.1)</td>
<td id="7963B56AE45C4BC71C63FDDC0F9DDD52" box="[1300,1436,597,620]" gridcol="4" gridrow="3" pageId="5" pageNumber="42">0.256</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FD090F9DDDCF" box="[151,1436,640,753]" gridrow="4" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FD090BECDDCF" box="[151,493,640,753]" gridcol="0" gridrow="4" pageId="5" pageNumber="42">SL/Pectoral fin length (P1) SL/Pelvic fin length (P2) SL/Longest dorsal-fin ray</th>
<td id="7963B56AE45C4BC71B34FD0908A7DDCF" box="[579,678,640,753]" gridcol="1" gridrow="4" pageId="5" pageNumber="42">5.6 7.0 5.7</td>
<td id="7963B56AE45C4BC71B9FFD0909C5DDCF" box="[744,964,640,753]" gridcol="2" gridrow="4" pageId="5" pageNumber="42">5.3, 0.26 (5.05.7) 6.7, 0.33 (6.37.6) 6.4, 1.02 (5.38.8)</td>
<td id="7963B56AE45C4BC71A8EFD090EC6DDCF" box="[1017,1223,640,753]" gridcol="3" gridrow="4" pageId="5" pageNumber="42">5.3, 0.20 (5.05.7) 6.7, 0.25 (6.37.0) 6.1, 0.42 (5.56.8)</td>
<td id="7963B56AE45C4BC71C63FD090F9DDDCF" box="[1300,1436,640,753]" gridcol="4" gridrow="4" pageId="5" pageNumber="42">0.429 0.803 0.267</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FC8C0F9DDC22" box="[151,1436,773,796]" gridrow="5" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FC8C0BECDC22" box="[151,493,773,796]" gridcol="0" gridrow="5" pageId="5" pageNumber="42">SL/Longest anal fin ray</th>
<td id="7963B56AE45C4BC71B34FC8C08A7DC22" box="[579,678,773,796]" gridcol="1" gridrow="5" pageId="5" pageNumber="42">7.9</td>
<td id="7963B56AE45C4BC71B9FFC8C09C5DC22" box="[744,964,773,796]" gridcol="2" gridrow="5" pageId="5" pageNumber="42">7.8, 0.56 (6.78.6)</td>
<td id="7963B56AE45C4BC71A8EFC8C0EC6DC22" box="[1017,1223,773,796]" gridcol="3" gridrow="5" pageId="5" pageNumber="42">7.6, 0.53 (6.78.5)</td>
<td id="7963B56AE45C4BC71C63FC8C0F9DDC22" box="[1300,1436,773,796]" gridcol="4" gridrow="5" pageId="5" pageNumber="42">0.523</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FCB90F9DDC4F" box="[151,1436,816,881]" gridrow="6" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FCB90BECDC4F" box="[151,493,816,881]" gridcol="0" gridrow="6" pageId="5" pageNumber="42">SL/Dorsal-fin base SL/Anal fin base</th>
<td id="7963B56AE45C4BC71B34FCB908A7DC4F" box="[579,678,816,881]" gridcol="1" gridrow="6" pageId="5" pageNumber="42">8.5 7.6</td>
<td id="7963B56AE45C4BC71B9FFCB909C5DC4F" box="[744,964,816,881]" gridcol="2" gridrow="6" pageId="5" pageNumber="42">8.1, 1.11 (6.09.8) 7.6, 0.51 (6.98.4)</td>
<td id="7963B56AE45C4BC71A8EFCB90EC6DC4F" box="[1017,1223,816,881]" gridcol="3" gridrow="6" pageId="5" pageNumber="42">8.7, 0.52 (8.09.9) 8.0, 0.49 (7.29.1)</td>
<td id="7963B56AE45C4BC71C63FCB90F9DDC4F" box="[1300,1436,816,881]" gridcol="4" gridrow="6" pageId="5" pageNumber="42">0.035 0.015</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FC0C0F9DDCA2" box="[151,1436,901,924]" gridrow="7" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FC0C0BECDCA2" box="[151,493,901,924]" gridcol="0" gridrow="7" pageId="5" pageNumber="42">SL/Predorsal length</th>
<td id="7963B56AE45C4BC71B34FC0C08A7DCA2" box="[579,678,901,924]" gridcol="1" gridrow="7" pageId="5" pageNumber="42">1.8</td>
<td id="7963B56AE45C4BC71B9FFC0C09C5DCA2" box="[744,964,901,924]" gridcol="2" gridrow="7" pageId="5" pageNumber="42">1.8, 0.04 (1.81.9)</td>
<td id="7963B56AE45C4BC71A8EFC0C0EC6DCA2" box="[1017,1223,901,924]" gridcol="3" gridrow="7" pageId="5" pageNumber="42">1.8, 0.03 (1.81.9)</td>
<td id="7963B56AE45C4BC71C63FC0C0F9DDCA2" box="[1300,1436,901,924]" gridcol="4" gridrow="7" pageId="5" pageNumber="42">0.540</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FC390F9DDCCF" box="[151,1436,944,1009]" gridrow="8" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FC390BECDCCF" box="[151,493,944,1009]" gridcol="0" gridrow="8" pageId="5" pageNumber="42">SL/Prepelvic length SL/Preanal length</th>
<td id="7963B56AE45C4BC71B34FC3908A7DCCF" box="[579,678,944,1009]" gridcol="1" gridrow="8" pageId="5" pageNumber="42">2.0 1.8</td>
<td id="7963B56AE45C4BC71B9FFC3909C5DCCF" box="[744,964,944,1009]" gridcol="2" gridrow="8" pageId="5" pageNumber="42">2.1, 0.06 (2.02.2) 1.6, 0.15 (1.51.9)</td>
<td id="7963B56AE45C4BC71A8EFC390EC6DCCF" box="[1017,1223,944,1009]" gridcol="3" gridrow="8" pageId="5" pageNumber="42">2.1, 0.04 (2.02.2) 1.6, 0.14 (1.51.9)</td>
<td id="7963B56AE45C4BC71C63FC390F9DDCCF" box="[1300,1436,944,1009]" gridcol="4" gridrow="8" pageId="5" pageNumber="42">0.816 0.779</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FB8C0F9DDB22" box="[151,1436,1029,1052]" gridrow="9" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FB8C0BECDB22" box="[151,493,1029,1052]" gridcol="0" gridrow="9" pageId="5" pageNumber="42">SL/Caudal peduncle length</th>
<td id="7963B56AE45C4BC71B34FB8C08A7DB22" box="[579,678,1029,1052]" gridcol="1" gridrow="9" pageId="5" pageNumber="42">4.4</td>
<td id="7963B56AE45C4BC71B9FFB8C09C5DB22" box="[744,964,1029,1052]" gridcol="2" gridrow="9" pageId="5" pageNumber="42">4.6, 0.16 (4.34.9)</td>
<td id="7963B56AE45C4BC71A8EFB8C0EC6DB22" box="[1017,1223,1029,1052]" gridcol="3" gridrow="9" pageId="5" pageNumber="42">4.5, 0.24 (4.04.8)</td>
<td id="7963B56AE45C4BC71C63FB8C0F9DDB22" box="[1300,1436,1029,1052]" gridcol="4" gridrow="9" pageId="5" pageNumber="42">0.419</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FBB90F9DDB4F" box="[151,1436,1072,1137]" gridrow="10" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FBB90BECDB4F" box="[151,493,1072,1137]" gridcol="0" gridrow="10" pageId="5" pageNumber="42">SL/Caudal peduncle depth SL/Pectoral to pelvic fin length</th>
<td id="7963B56AE45C4BC71B34FBB908A7DB4F" box="[579,678,1072,1137]" gridcol="1" gridrow="10" pageId="5" pageNumber="42">9.7 3.6</td>
<td id="7963B56AE45C4BC71B9FFBB909C5DB4F" box="[744,964,1072,1137]" gridcol="2" gridrow="10" pageId="5" pageNumber="42">9.5, 0.40 (8.810.3) 4.2, 0.18 (3.94.5)</td>
<td id="7963B56AE45C4BC71A8EFBB90EC6DB4F" box="[1017,1223,1072,1137]" gridcol="3" gridrow="10" pageId="5" pageNumber="42">9.3, 0.45 (8.39.9) 4.2, 0.25 (3.84.6)</td>
<td id="7963B56AE45C4BC71C63FBB90F9DDB4F" box="[1300,1436,1072,1137]" gridcol="4" gridrow="10" pageId="5" pageNumber="42">0.597 0.633</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FB0C0F9DDBA2" box="[151,1436,1157,1180]" gridrow="11" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FB0C0BECDBA2" box="[151,493,1157,1180]" gridcol="0" gridrow="11" pageId="5" pageNumber="42">SL/Pelvic to anal fin length</th>
<td id="7963B56AE45C4BC71B34FB0C08A7DBA2" box="[579,678,1157,1180]" gridcol="1" gridrow="11" pageId="5" pageNumber="42">4.7</td>
<td id="7963B56AE45C4BC71B9FFB0C09C5DBA2" box="[744,964,1157,1180]" gridcol="2" gridrow="11" pageId="5" pageNumber="42">4.2, 0.18 (3.94.5)</td>
<td id="7963B56AE45C4BC71A8EFB0C0EC6DBA2" box="[1017,1223,1157,1180]" gridcol="3" gridrow="11" pageId="5" pageNumber="42">4.2, 0.22 (3.84.5)</td>
<td id="7963B56AE45C4BC71C63FB0C0F9DDBA2" box="[1300,1436,1157,1180]" gridcol="4" gridrow="11" pageId="5" pageNumber="42">0.641</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FB390F9DDBCF" box="[151,1436,1200,1265]" gridrow="12" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FB390BECDBCF" box="[151,493,1200,1265]" gridcol="0" gridrow="12" pageId="5" pageNumber="42">HL/Mouth width HL/Head depth</th>
<td id="7963B56AE45C4BC71B34FB3908A7DBCF" box="[579,678,1200,1265]" gridcol="1" gridrow="12" pageId="5" pageNumber="42">3.8 1.3</td>
<td id="7963B56AE45C4BC71B9FFB3909C5DBCF" box="[744,964,1200,1265]" gridcol="2" gridrow="12" pageId="5" pageNumber="42">3.6, 0.23 (3.24.0) 1.3, 0.04 (1.31.4)</td>
<td id="7963B56AE45C4BC71A8EFB390EC6DBCF" box="[1017,1223,1200,1265]" gridcol="3" gridrow="12" pageId="5" pageNumber="42">3.5, 0.14 (3.33.7) 1.3, 0.08 (1.11.4)</td>
<td id="7963B56AE45C4BC71C63FB390F9DDBCF" box="[1300,1436,1200,1265]" gridcol="4" gridrow="12" pageId="5" pageNumber="42">0.567 0.410</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FA8C0F9DDA22" box="[151,1436,1285,1308]" gridrow="13" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FA8C0BECDA22" box="[151,493,1285,1308]" gridcol="0" gridrow="13" pageId="5" pageNumber="42">HL/Head width</th>
<td id="7963B56AE45C4BC71B34FA8C08A7DA22" box="[579,678,1285,1308]" gridcol="1" gridrow="13" pageId="5" pageNumber="42">1.7</td>
<td id="7963B56AE45C4BC71B9FFA8C09C5DA22" box="[744,964,1285,1308]" gridcol="2" gridrow="13" pageId="5" pageNumber="42">1.8, 0.07 (1.71.9)</td>
<td id="7963B56AE45C4BC71A8EFA8C0EC6DA22" box="[1017,1223,1285,1308]" gridcol="3" gridrow="13" pageId="5" pageNumber="42">1.8, 0.06 (1.71.9)</td>
<td id="7963B56AE45C4BC71C63FA8C0F9DDA22" box="[1300,1436,1285,1308]" gridcol="4" gridrow="13" pageId="5" pageNumber="42">0.683</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FAB90F9DDA4F" box="[151,1436,1328,1393]" gridrow="14" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FAB90BECDA4F" box="[151,493,1328,1393]" gridcol="0" gridrow="14" pageId="5" pageNumber="42">HL/Snout length HL/Orbit width</th>
<td id="7963B56AE45C4BC71B34FAB908A7DA4F" box="[579,678,1328,1393]" gridcol="1" gridrow="14" pageId="5" pageNumber="42">3.8 3.7</td>
<td id="7963B56AE45C4BC71B9FFAB909C5DA4F" box="[744,964,1328,1393]" gridcol="2" gridrow="14" pageId="5" pageNumber="42">3.7, 0.22 (3.24.0) 3.4, 0.20 (3.23.8)</td>
<td id="7963B56AE45C4BC71A8EFAB90EC6DA4F" box="[1017,1223,1328,1393]" gridcol="3" gridrow="14" pageId="5" pageNumber="42">3.6, 0.23 (3.34.0) 3.3, 0.12 (3.13.5)</td>
<td id="7963B56AE45C4BC71C63FAB90F9DDA4F" box="[1300,1436,1328,1393]" gridcol="4" gridrow="14" pageId="5" pageNumber="42">0.817 0.261</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FA0C0F9DDAA2" box="[151,1436,1413,1436]" gridrow="15" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FA0C0BECDAA2" box="[151,493,1413,1436]" gridcol="0" gridrow="15" pageId="5" pageNumber="42">HL/Interorbital width</th>
<td id="7963B56AE45C4BC71B34FA0C08A7DAA2" box="[579,678,1413,1436]" gridcol="1" gridrow="15" pageId="5" pageNumber="42">3.1</td>
<td id="7963B56AE45C4BC71B9FFA0C09C5DAA2" box="[744,964,1413,1436]" gridcol="2" gridrow="15" pageId="5" pageNumber="42">3.1, 0.10 (3.03.3)</td>
<td id="7963B56AE45C4BC71A8EFA0C0EC6DAA2" box="[1017,1223,1413,1436]" gridcol="3" gridrow="15" pageId="5" pageNumber="42">3.1, 0.14 (2.83.3)</td>
<td id="7963B56AE45C4BC71C63FA0C0F9DDAA2" box="[1300,1436,1413,1436]" gridcol="4" gridrow="15" pageId="5" pageNumber="42">0.551</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0FA390F9DDACF" box="[151,1436,1456,1521]" gridrow="16" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0FA390BECDACF" box="[151,493,1456,1521]" gridcol="0" gridrow="16" pageId="5" pageNumber="42">HL/Postorbital length HL/Longest dorsal-fin ray</th>
<td id="7963B56AE45C4BC71B34FA3908A7DACF" box="[579,678,1456,1521]" gridcol="1" gridrow="16" pageId="5" pageNumber="42">2.0 1.3</td>
<td id="7963B56AE45C4BC71B9FFA3909C5DACF" box="[744,964,1456,1521]" gridcol="2" gridrow="16" pageId="5" pageNumber="42">2.1, 0.10 (2.02.4) 1.6, 0.27 (1.42.2)</td>
<td id="7963B56AE45C4BC71A8EFA390EC6DACF" box="[1017,1223,1456,1521]" gridcol="3" gridrow="16" pageId="5" pageNumber="42">2.2, 0.12 (2.02.5) 1.5, 0.13 (1.41.8)</td>
<td id="7963B56AE45C4BC71C63FA390F9DDACF" box="[1300,1436,1456,1521]" gridcol="4" gridrow="16" pageId="5" pageNumber="42">0.084 0.281</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0F98C0F9DD922" box="[151,1436,1541,1564]" gridrow="17" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0F98C0BECD922" box="[151,493,1541,1564]" gridcol="0" gridrow="17" pageId="5" pageNumber="42">HL/Longest anal fin ray</th>
<td id="7963B56AE45C4BC71B34F98C08A7D922" box="[579,678,1541,1564]" gridcol="1" gridrow="17" pageId="5" pageNumber="42">1.8</td>
<td id="7963B56AE45C4BC71B9FF98C09C5D922" box="[744,964,1541,1564]" gridcol="2" gridrow="17" pageId="5" pageNumber="42">1.9, 0.14 (1.72.2)</td>
<td id="7963B56AE45C4BC71A8EF98C0EC6D922" box="[1017,1223,1541,1564]" gridcol="3" gridrow="17" pageId="5" pageNumber="42">1.9, 0.15 (1.72.2)</td>
<td id="7963B56AE45C4BC71C63F98C0F9DD922" box="[1300,1436,1541,1564]" gridcol="4" gridrow="17" pageId="5" pageNumber="42">0.469</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0F9B90F9DD94F" box="[151,1436,1584,1649]" gridrow="18" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0F9B90BECD94F" box="[151,493,1584,1649]" gridcol="0" gridrow="18" pageId="5" pageNumber="42">HL/Pectoral fin length HL/Pelvic fin length</th>
<td id="7963B56AE45C4BC71B34F9B908A7D94F" box="[579,678,1584,1649]" gridcol="1" gridrow="18" pageId="5" pageNumber="42">1.3 1.6</td>
<td id="7963B56AE45C4BC71B9FF9B909C5D94F" box="[744,964,1584,1649]" gridcol="2" gridrow="18" pageId="5" pageNumber="42">1.3, 0.06 (1.21.4) 1.7, 0.10 (1.51.9)</td>
<td id="7963B56AE45C4BC71A8EF9B90EC6D94F" box="[1017,1223,1584,1649]" gridcol="3" gridrow="18" pageId="5" pageNumber="42">1.3, 0.08 (1.21.5) 1.7, 0.08 (1.51.8)</td>
<td id="7963B56AE45C4BC71C63F9B90F9DD94F" box="[1300,1436,1584,1649]" gridcol="4" gridrow="18" pageId="5" pageNumber="42">0.592 0.992</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0F90C0F9DD9A2" box="[151,1436,1669,1692]" gridrow="19" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0F90C0BECD9A2" box="[151,493,1669,1692]" gridcol="0" gridrow="19" pageId="5" pageNumber="42">HL/Dorsal-fin base</th>
<td id="7963B56AE45C4BC71B34F90C08A7D9A2" box="[579,678,1669,1692]" gridcol="1" gridrow="19" pageId="5" pageNumber="42">2.0</td>
<td id="7963B56AE45C4BC71B9FF90C09C5D9A2" box="[744,964,1669,1692]" gridcol="2" gridrow="19" pageId="5" pageNumber="42">2.0, 0.28 (1.52.5)</td>
<td id="7963B56AE45C4BC71A8EF90C0EC6D9A2" box="[1017,1223,1669,1692]" gridcol="3" gridrow="19" pageId="5" pageNumber="42">2.2, 0.15 (2.02.5)</td>
<td id="7963B56AE45C4BC71C63F90C0F9DD9A2" box="[1300,1436,1669,1692]" gridcol="4" gridrow="19" pageId="5" pageNumber="42">0.055</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0F9390F9DD9CF" box="[151,1436,1712,1777]" gridrow="20" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0F9390BECD9CF" box="[151,493,1712,1777]" gridcol="0" gridrow="20" pageId="5" pageNumber="42">HL/Anal fin base Caudal peduncle length/depth</th>
<td id="7963B56AE45C4BC71B34F93908A7D9CF" box="[579,678,1712,1777]" gridcol="1" gridrow="20" pageId="5" pageNumber="42">1.8 2.2</td>
<td id="7963B56AE45C4BC71B9FF93909C5D9CF" box="[744,964,1712,1777]" gridcol="2" gridrow="20" pageId="5" pageNumber="42">1.9, 0.14 (1.72.2) 2.1, 0.11 (1.92.3)</td>
<td id="7963B56AE45C4BC71A8EF9390EC6D9CF" box="[1017,1223,1712,1777]" gridcol="3" gridrow="20" pageId="5" pageNumber="42">1.9, 0.15 (1.72.2) 2.1, 0.17 (1.82.4)</td>
<td id="7963B56AE45C4BC71C63F9390F9DD9CF" box="[1300,1436,1712,1777]" gridcol="4" gridrow="20" pageId="5" pageNumber="42">0.469 0.815</td>
</tr>
<tr id="3AB2DC16E45C4BC719E0F88C0F9DD870" box="[151,1436,1797,1870]" gridrow="21" pageId="5" pageNumber="42">
<th id="7963B56AE45C4BC719E0F88C0BECD870" box="[151,493,1797,1870]" gridcol="0" gridrow="21" pageId="5" pageNumber="42">P1 origin to P2 origin/P1 P2 origin to anal fin origin/P2</th>
<td id="7963B56AE45C4BC71B34F88C08A7D870" box="[579,678,1797,1870]" gridcol="1" gridrow="21" pageId="5" pageNumber="42">1.7 1.5</td>
<td id="7963B56AE45C4BC71B9FF88C09C5D870" box="[744,964,1797,1870]" gridcol="2" gridrow="21" pageId="5" pageNumber="42">1.3, 0.09 (1.11.5) 1.3, 0.12 (1.11.5)</td>
<td id="7963B56AE45C4BC71A8EF88C0EC6D870" box="[1017,1223,1797,1870]" gridcol="3" gridrow="21" pageId="5" pageNumber="42">1.3, 0.10 (1.21.5) 1.2, 0.07 (1.11.3)</td>
<td id="7963B56AE45C4BC71C63F88C0F9DD870" box="[1300,1436,1797,1870]" gridcol="4" gridrow="21" pageId="5" pageNumber="42">0.395 0.530</td>
</tr>
</table>
</paragraph>
<caption id="D0FD8EDCE45FB43E19E0FBF70E04DBCB" httpUri="https://zenodo.org/record/294010/files/figure.png" pageId="6" pageNumber="43" targetBox="[160,1426,185,1117]" targetPageId="6">
<paragraph id="843DDE54E45FB43E19E0FBF70E04DBCB" blockId="6.[151,1437,1150,1269]" pageId="6" pageNumber="43">
<emphasis id="B6F60246E45FB43E19E0FBF70B12DBA8" bold="true" box="[151,275,1150,1174]" pageId="6" pageNumber="43">FIGURE 4</emphasis>
. Distribution map (solid square = holotype of
<taxonomicName id="4382A5D7E45FB43E1A70FBF70E07DBAB" box="[775,1030,1150,1173]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E45FB43E1A70FBF70E07DBAB" box="[775,1030,1150,1173]" italics="true" pageId="6" pageNumber="43">Petroleuciscus esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE45FB43E1D7AFBF70E49DBAB" box="[1037,1096,1150,1173]" pageId="6" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
, open square = paratypes; solid circles =
<taxonomicName id="4382A5D7E45FB43E1877FB170B78DB8B" box="[256,377,1182,1205]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E45FB43E1877FB170B78DB8B" box="[256,377,1182,1205]" italics="true" pageId="6" pageNumber="43">P. persidis</emphasis>
</taxonomicName>
holotype and paratypes (southern circle) and paratypes (northern circle), open circles = other material; solid triangles =
<taxonomicName id="4382A5D7E45FB43E18B0FB370835DBEB" box="[455,564,1214,1237]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="ulanus">
<emphasis id="B6F60246E45FB43E18B0FB370835DBEB" box="[455,564,1214,1237]" italics="true" pageId="6" pageNumber="43">P. ulanus</emphasis>
</taxonomicName>
syntypes (northern triangle), solid triangles =
<taxonomicName id="4382A5D7E45FB43E1D3BFB370EE4DBEB" box="[1100,1253,1214,1237]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="gaderanus">
<emphasis id="B6F60246E45FB43E1D3BFB370EE4DBEB" box="[1100,1253,1214,1237]" italics="true" pageId="6" pageNumber="43">P. gaderanus</emphasis>
</taxonomicName>
syntypes (other triangles), open triangles = other material; star =
<taxonomicName id="4382A5D7E45FB43E1BD6FB5708F6DBCB" box="[673,759,1246,1269]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="kurui">
<emphasis id="B6F60246E45FB43E1BD6FB5708F6DBCB" box="[673,759,1246,1269]" italics="true" pageId="6" pageNumber="43">P. kurui</emphasis>
</taxonomicName>
holotype and paratypes).
</paragraph>
</caption>
<caption id="D0FD8EDCE45FB43E19E0F8900EE2D80E" box="[151,1251,1817,1841]" httpUri="https://zenodo.org/record/293385/files/figure.png" pageId="6" pageNumber="43" targetBox="[173,1412,1323,1793]" targetPageId="6">
<paragraph id="843DDE54E45FB43E19E0F8900EE2D80E" blockId="6.[151,1251,1817,1841]" box="[151,1251,1817,1841]" pageId="6" pageNumber="43">
<emphasis id="B6F60246E45FB43E19E0F8900B13D80F" bold="true" box="[151,274,1817,1841]" pageId="6" pageNumber="43">FIGURE 5</emphasis>
. Syntype of
<taxonomicName id="4382A5D7E45FB43E18ECF890091FD80E" authority="Gunther, 1899" authorityName="Gunther" authorityYear="1899" box="[411,798,1817,1840]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="gaderanus">
<emphasis id="B6F60246E45FB43E18ECF8900805D80E" box="[411,516,1817,1840]" italics="true" pageId="6" pageNumber="43">Leuciscus</emphasis>
<emphasis id="B6F60246E45FB43E1B7CF890087AD80E" box="[523,635,1817,1840]" italics="true" pageId="6" pageNumber="43">gaderanus</emphasis>
Günther, 1899
</taxonomicName>
, BMNH 1899.9.30.113-115, 73.0 mm SL.
</paragraph>
</caption>
<paragraph id="843DDE54E45FB43F19B2F8E80F95DFC2" blockId="6.[151,1436,1889,2027]" lastBlockId="7.[151,1437,152,1188]" lastPageId="7" lastPageNumber="44" pageId="6" pageNumber="43">
The
<typeStatus id="5B3960F6E45FB43E198DF8E80B2DD845" box="[250,300,1889,1915]" pageId="6" pageNumber="43">type</typeStatus>
locality was a fresh, clear stream at an altitude of
<quantity id="437A73B1E45FB43E1A0EF8E809D7D845" box="[889,982,1889,1915]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="43" unit="m" value="2300.0">2300 m</quantity>
. Water temperature at 1705 hours on
<date id="F03CF894E45FB43E1CFBF8E80ACCD89F" pageId="6" pageNumber="43">9 June</date>
was 22°C,
<collectingCountry id="FC959EC4E45FB43E1822F80E0B78D89F" box="[341,377,1927,1953]" name="Philippines" pageId="6" pageNumber="43">pH</collectingCountry>
was 6.2, conductivity was 0.455 mS, stream width was
<quantity id="437A73B1E45FB43E1D67F80E0E5FD89F" box="[1040,1118,1927,1953]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="6" pageNumber="43" unit="m" value="5.0" valueMax="8.0" valueMin="2.0">28 m</quantity>
, maximum depth was
<quantity id="437A73B1E45FB43E1C10F80E0F99D89F" box="[1383,1432,1927,1953]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="43" unit="m" value="1.0">1 m</quantity>
, current was slow to moderate, aquatic plant material was of the submergent
<typeStatus id="5B3960F6E45FB43E1D7CF8250E41D8F8" box="[1035,1088,1964,1990]" pageId="6" pageNumber="43">type</typeStatus>
, the shore was grassy, and the stream bottom was a mix of pebbles and mud. The second locality was a fresh stream with clear water at an altitude of
<quantity id="437A73B1E45EB43F1860FF110B75DF8C" box="[279,372,152,178]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="7" pageNumber="44" unit="m" value="2410.0">2410 m</quantity>
. Water temperature at 1225 hours on
<date id="F03CF894E45EB43F1A59FF110988DF8C" box="[814,905,152,178]" pageId="7" pageNumber="44" value="1910-06">10 June</date>
was 17°C,
<collectingCountry id="FC959EC4E45EB43F1D65FF110E37DF8C" box="[1042,1078,152,178]" name="Philippines" pageId="7" pageNumber="44">pH</collectingCountry>
was 6.2, conductivity was 0.4 mS, stream width was
<quantity id="437A73B1E45EB43F18D4FF340BD2DFE9" box="[419,467,189,215]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="44" unit="m" value="4.0">4 m</quantity>
, maximum depth was
<quantity id="437A73B1E45EB43F1BABFF340929DFE9" box="[732,808,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="44" unit="cm" value="70.0">70 cm</quantity>
, current was slow to moderate, aquatic plant material was of the encrusting
<typeStatus id="5B3960F6E45EB43F18EFFF6B0BCCDFC2" box="[408,461,226,252]" pageId="7" pageNumber="44">type</typeStatus>
, the shore was grassy, and the stream bottom was a mix of pebbles, sand and mud.
</paragraph>
<paragraph id="843DDE54E45EB43F19B1FE8E0940DD3C" blockId="7.[151,1437,152,1188]" pageId="7" pageNumber="44">
<emphasis id="B6F60246E45EB43F19B1FE8E0B74DE1F" bold="true" box="[198,373,263,289]" pageId="7" pageNumber="44">Conservation.</emphasis>
Construction of the dam on the Zayandeh River has altered the ecology of the river system but its effects, positive or negative, on the distribution and numbers of the new species is unknown. It may be relatively secure in higher tributary streams. However, demands on water resources from this and other rivers on the central desert plateau of
<collectingCountry id="FC959EC4E45EB43F1B6DFEF1084ADEAC" box="[538,587,376,402]" name="Iran" pageId="7" pageNumber="44">Iran</collectingCountry>
have increased markedly as the human population of the country has doubled in 30 years (from ca. 35 million in mid-1970s to 72 million in 2008, http://en.wikipedia.org/wiki/ Demographics_of_
<collectingCountry id="FC959EC4E45EB43F18F1FE4B0BB9DEE2" box="[390,440,450,476]" name="Iran" pageId="7" pageNumber="44">Iran</collectingCountry>
). Freshwater resources are under threat, exacerbated by droughts, and all species found in these desert or semi-desert areas are under stress.
</paragraph>
</subSubSection>
<subSubSection id="CC988DDFE45EB43019B1FD840E32D934" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="description">
<paragraph id="843DDE54E45EB43F19B1FD840830DB9A" blockId="7.[151,1437,152,1188]" pageId="7" pageNumber="44">
<emphasis id="B6F60246E45EB43F19B1FD840BE2DD19" bold="true" box="[198,483,525,551]" pageId="7" pageNumber="44">Comparative remarks.</emphasis>
The species is assigned to the genus
<taxonomicName id="4382A5D7E45EB43F1AECFD840E46DD18" box="[923,1095,525,550]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1AECFD840E46DD18" box="[923,1095,525,550]" italics="true" pageId="7" pageNumber="44">Petroleuciscus</emphasis>
</taxonomicName>
and not to either
<taxonomicName id="4382A5D7E45EB43F1C60FD840F7DDD18" box="[1303,1404,525,550]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1C60FD840F7DDD18" box="[1303,1404,525,550]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
or
<taxonomicName id="4382A5D7E45EB43F19E0FDBB0B0ADD75" box="[151,267,562,587]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F19E0FDBB0B0ADD75" box="[151,267,562,587]" italics="true" pageId="7" pageNumber="44">Leuciscus</emphasis>
</taxonomicName>
for it shares most of the diagnostic characters of the former (
<bibRefCitation id="E013A3A5E45EB43F1A97FDBB0EB2DD72" author="Bogutskaya" box="[992,1203,562,588]" pageId="7" pageNumber="47" refString="Bogutskaya, N. G. (2002) Petroleuciscus, a new genus for the Leuciscus borysthenicus species group (Teleostei: Cyprinidae). Zoosystematica Rossica, 11 (1), 235 - 237." type="journal article" year="2002">Bogutskaya 2002</bibRefCitation>
), namely small size in adults, commonly around
<quantity id="437A73B1E45EB43F189DFDD10861DD4C" box="[490,608,600,626]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="7" pageNumber="44" unit="cm" value="11.0" valueMax="12.0" valueMin="10.0">
10
<emphasis id="B6F60246E45EB43F1B7FFDD10816DD4F" box="[520,535,600,625]" italics="true" pageId="7" pageNumber="44"></emphasis>
12 cm
</quantity>
(vs. about
<quantity id="437A73B1E45EB43F1B92FDD1092FDD4C" box="[741,814,600,626]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="44" unit="cm" value="20.0">20 cm</quantity>
and over in
<taxonomicName id="4382A5D7E45EB43F1AB3FDD10E28DD4F" box="[964,1065,600,625]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1AB3FDD10E28DD4F" box="[964,1065,600,625]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
and
<taxonomicName id="4382A5D7E45EB43F1D13FDD10ED9DD4F" box="[1124,1240,600,625]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1D13FDD10ED9DD4F" box="[1124,1240,600,625]" italics="true" pageId="7" pageNumber="44">Leuciscus</emphasis>
</taxonomicName>
); commonly 8½ dorsal-fin branched rays (vs. commonly 7½ in
<taxonomicName id="4382A5D7E45EB43F1BC0FDF4092ADDA8" box="[695,811,637,662]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1BC0FDF4092ADDA8" box="[695,811,637,662]" italics="true" pageId="7" pageNumber="44">Leuciscus</emphasis>
</taxonomicName>
); anal-fin margin convex in its anterior part (vs. analfin margin clearly concave in
<taxonomicName id="4382A5D7E45EB43F1B7FFD2B0881DD85" box="[520,640,674,699]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1B7FFD2B0881DD85" box="[520,640,674,699]" italics="true" pageId="7" pageNumber="44">Leuciscus</emphasis>
</taxonomicName>
), and two teeth in the short row (5.2 or 4.2 vs. commonly
<quantity id="437A73B1E45EB43F1C21FD2B0F9DDD82" box="[1366,1436,674,700]" metricMagnitude="-1" metricUnit="m" metricValue="1.3462" pageId="7" pageNumber="44" unit="in" value="5.3">5.3 in</quantity>
<taxonomicName id="4382A5D7E45EB43F19E0FD410B0ADDDF" box="[151,267,712,737]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F19E0FD410B0ADDDF" box="[151,267,712,737]" italics="true" pageId="7" pageNumber="44">Leuciscus</emphasis>
</taxonomicName>
); few sensory cephalic pores,
<quantity id="437A73B1E45EB43F1B00FD4108D1DDDC" box="[631,720,712,738]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="7" pageNumber="44" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">
8
<emphasis id="B6F60246E45EB43F1BF1FD410894DDDF" box="[646,661,712,737]" italics="true" pageId="7" pageNumber="44"></emphasis>
11 in
</quantity>
the supraorbital,
<quantity id="437A73B1E45EB43F1AD5FD410E0ADDDC" box="[930,1035,712,738]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.572" metricValueMin="3.556" pageId="7" pageNumber="44" unit="in" value="16.0" valueMax="18.0" valueMin="14.0">
14
<emphasis id="B6F60246E45EB43F1AC8FD4109CFDDDF" box="[959,974,712,737]" italics="true" pageId="7" pageNumber="44"></emphasis>
18 in
</quantity>
the infraorbital, and
<quantity id="437A73B1E45EB43F1C70FD410F71DDDC" box="[1287,1392,712,738]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" pageId="7" pageNumber="44" unit="in" value="16.0" valueMax="17.0" valueMin="15.0">
15
<emphasis id="B6F60246E45EB43F1C53FD410F32DDDF" box="[1316,1331,712,737]" italics="true" pageId="7" pageNumber="44"></emphasis>
17 in
</quantity>
the preoperculo-mandibular canal (vs. 9
<emphasis id="B6F60246E45EB43F1B30FD640857DC38" box="[583,598,749,774]" italics="true" pageId="7" pageNumber="44"></emphasis>
14 supraorbital, 16
<emphasis id="B6F60246E45EB43F1A41FD640944DC38" box="[822,837,749,774]" italics="true" pageId="7" pageNumber="44"></emphasis>
27 infraorbital and 15
<emphasis id="B6F60246E45EB43F1D3EFD640E59DC38" box="[1097,1112,749,774]" italics="true" pageId="7" pageNumber="44"></emphasis>
22 preoperculo-mandibular pores in
<taxonomicName id="4382A5D7E45EB43F1988FC9B0B67DC15" box="[255,358,786,811]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1988FC9B0B67DC15" box="[255,358,786,811]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
); relatively small supraethmoid-mesethmoid block, supraethmoid width less than 30% of cranial roof length (vs. over 36% in
<taxonomicName id="4382A5D7E45EB43F1B35FCB108A6DC6F" box="[578,679,824,849]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1B35FCB108A6DC6F" box="[578,679,824,849]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
); infraorbitals narrow, posterior margin of fourth infraorbital far from reaching the posterior margin of the preoperculum (vs. broad, posterior margin of 4th infraorbital reaching almost to posterior margin of preoperculum in
<taxonomicName id="4382A5D7E45EB43F1A4CFC0F09A3DCA1" box="[827,930,902,927]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1A4CFC0F09A3DCA1" box="[827,930,902,927]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
); and a deep neurocranium with normally developed interorbital septum (vs. a low neurocranium with low or completely reduced septum in
<taxonomicName id="4382A5D7E45EB43F1C52FC220F8BDCFA" box="[1317,1418,939,964]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1C52FC220F8BDCFA" box="[1317,1418,939,964]" italics="true" pageId="7" pageNumber="44">Squalius</emphasis>
</taxonomicName>
). However, the new species does not possess a low number of vertebrae typical for most
<taxonomicName id="4382A5D7E45EB43F1DE3FC590F41DCD7" box="[1172,1344,976,1001]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1DE3FC590F41DCD7" box="[1172,1344,976,1001]" italics="true" pageId="7" pageNumber="44">Petroleuciscus</emphasis>
</taxonomicName>
species (
<tableCitation id="C900EBEFE45EB43F19E9FC7C0AF8DB31" box="[158,249,1013,1039]" captionStart="TABLE 2" captionStartId="7.[151,242,1237,1261]" captionTargetBox="[151,1436,1320,1855]" captionTargetPageId="7" captionText="TABLE 2. Vertebral counts (range, mode and mean) and most frequent vertebral formulae in species of the genus Petroleuciscus." httpUri="http://table.plazi.org/id/D0FD8EDCE45EB43F19E0FB5C0B3BDA32" pageId="7" pageNumber="44" tableUuid="D0FD8EDCE45EB43F19E0FB5C0B3BDA32">Table 2</tableCitation>
;
<figureCitation id="1CB9C2D1E45EB43F1871FC7C0B48DB31" box="[262,329,1013,1039]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="7" pageNumber="44">Fig. 3</figureCitation>
b e). The new species is not assigned to the genus
<taxonomicName id="4382A5D7E45EB43F1AE8FC7F09FADB31" box="[927,1019,1014,1039]" class="Actinopterygii" family="Cyprinidae" genus="Telestes" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1AE8FC7F09FADB31" box="[927,1019,1014,1039]" italics="true" pageId="7" pageNumber="44">Telestes</emphasis>
</taxonomicName>
because it differs from the latter by having an indistinct colour pattern with no conspicuous black mid-lateral stripe and/or a conspicuous black dotted line along the lateral line (when preserved) and a broad gold-orange stripe with the lateral line yellow, margined on each side by a narrow black line (in life) characteristic for
<taxonomicName id="4382A5D7E45EB43F1A8FFBEF0E52DB41" box="[1016,1107,1126,1151]" class="Actinopterygii" family="Cyprinidae" genus="Telestes" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1A8FFBEF0E52DB41" box="[1016,1107,1126,1151]" italics="true" pageId="7" pageNumber="44">Telestes</emphasis>
</taxonomicName>
(our observations and data from
<bibRefCitation id="E013A3A5E45EB43F19A0FB03082ADB9A" author="Kottelat" box="[215,555,1162,1188]" pageId="7" pageNumber="47" refString="Kottelat, M. &amp; Freyhof, J. (2007) Handbook of European freshwater fishes. Published by the authors. Printed by Imprimerie du Democrate SA, Delemont, Switzerland, 646 pp." type="book" year="2007">Kottelat and Freyhof (2007))</bibRefCitation>
.
</paragraph>
<caption id="D0FD8EDCE45EB43F19E0FB5C0B3BDA32" ID-Table-UUID="D0FD8EDCE45EB43F19E0FB5C0B3BDA32" httpUri="http://table.plazi.org/id/D0FD8EDCE45EB43F19E0FB5C0B3BDA32" pageId="7" pageNumber="44" targetBox="[151,1436,1320,1855]" targetIsTable="true" targetPageId="7">
<paragraph id="843DDE54E45EB43F19E0FB5C0B3BDA32" blockId="7.[151,1436,1237,1292]" pageId="7" pageNumber="44">
<emphasis id="B6F60246E45EB43F19E0FB5C0B0EDBD3" bold="true" box="[151,271,1237,1261]" pageId="7" pageNumber="44">TABLE 2.</emphasis>
Vertebral counts (range, mode and mean) and most frequent vertebral formulae in species of the genus
<taxonomicName id="4382A5D7E45EB43F19E0FB7C0B32DA32" box="[151,307,1269,1292]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F19E0FB7C0B32DA32" box="[151,307,1269,1292]" italics="true" pageId="7" pageNumber="44">Petroleuciscus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="843DDE54E45EB43F18EAFAA10F12D801" pageId="7" pageNumber="44">
<table id="F6822CF4E45E4BC719E0FAA10F9DD801" box="[151,1436,1320,1855]" gridcols="6" gridrows="8" pageId="7" pageNumber="44">
<tr id="3AB2DC16E45E4BC719E0FAA10F9DDA64" box="[151,1436,1320,1370]" gridrow="0" pageId="7" pageNumber="44" rowspan-0="1">
<th id="7963B56AE45E4BC718EAFAA10837DA64" box="[413,566,1320,1370]" gridcol="1" gridrow="0" pageId="7" pageNumber="44">Total vertebrae</th>
<th id="7963B56AE45E4BC71B04FAA1091FDA64" box="[627,798,1320,1370]" gridcol="2" gridrow="0" pageId="7" pageNumber="44">Abdominal vertebrae</th>
<th id="7963B56AE45E4BC71A43FAA109D6DA64" box="[820,983,1320,1370]" gridcol="3" gridrow="0" pageId="7" pageNumber="44">Caudal vertebrae</th>
<th id="7963B56AE45E4BC71D77FAA10EC3DA64" box="[1024,1218,1320,1370]" gridcol="4" gridrow="0" pageId="7" pageNumber="44">Predorsalabdominal vertebrae</th>
<th id="7963B56AE45E4BC71DA1FAA10F9DDA64" box="[1238,1436,1320,1370]" gridcol="5" gridrow="0" pageId="7" pageNumber="44">Most frequent vertebral formulae</th>
</tr>
<tr id="3AB2DC16E45E4BC719E0FAE40F9DDAA1" box="[151,1436,1389,1439]" gridrow="1" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0FAE40B81DAA1" box="[151,384,1389,1439]" gridcol="0" gridrow="1" pageId="7" pageNumber="44">
<emphasis id="B6F60246E45EB43F19E8FAE40B1BDABC" box="[159,282,1389,1410]" italics="true" pageId="7" pageNumber="44">P. s m y r n a e u s</emphasis>
(n=35)
</th>
<td id="7963B56AE45E4BC718EAFAE40837DAA1" box="[413,566,1389,1439]" gridcol="1" gridrow="1" pageId="7" pageNumber="44">3436; 35; 35.3</td>
<td id="7963B56AE45E4BC71B04FAE4091FDAA1" box="[627,798,1389,1439]" gridcol="2" gridrow="1" pageId="7" pageNumber="44">2021; 20; 20.2</td>
<td id="7963B56AE45E4BC71A43FAE409D6DAA1" box="[820,983,1389,1439]" gridcol="3" gridrow="1" pageId="7" pageNumber="44">1416; 15; 15.0</td>
<td id="7963B56AE45E4BC71D77FAE40EC3DAA1" box="[1024,1218,1389,1439]" gridcol="4" gridrow="1" pageId="7" pageNumber="44">1213; 13; 12.6</td>
<td id="7963B56AE45E4BC71DA1FAE40F9DDAA1" box="[1238,1436,1389,1439]" gridcol="5" gridrow="1" pageId="7" pageNumber="44">20+15, 20+16, 21+15</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0FA3B0F9DDADB" box="[151,1436,1458,1509]" gridrow="2" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0FA3B0B81DADB" box="[151,384,1458,1509]" gridcol="0" gridrow="2" pageId="7" pageNumber="44">
<taxonomicName id="4382A5D7E45EB43F19E8FA3A0B03DAF6" box="[159,258,1459,1480]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E45EB43F19E8FA3A0B03DAF6" box="[159,258,1459,1480]" italics="true" pageId="7" pageNumber="44">P. persidis</emphasis>
</taxonomicName>
(n=126)
</th>
<td id="7963B56AE45E4BC718EAFA3B0837DADB" box="[413,566,1458,1509]" gridcol="1" gridrow="2" pageId="7" pageNumber="44">3437; 36; 35.6</td>
<td id="7963B56AE45E4BC71B04FA3B091FDADB" box="[627,798,1458,1509]" gridcol="2" gridrow="2" pageId="7" pageNumber="44">1820; 19; 18.9</td>
<td id="7963B56AE45E4BC71A43FA3B09D6DADB" box="[820,983,1458,1509]" gridcol="3" gridrow="2" pageId="7" pageNumber="44">1518; 17; 16.7</td>
<td id="7963B56AE45E4BC71D77FA3B0EC3DADB" box="[1024,1218,1458,1509]" gridcol="4" gridrow="2" pageId="7" pageNumber="44">1012; 12; 11.5</td>
<td id="7963B56AE45E4BC71DA1FA3B0F9DDADB" box="[1238,1436,1458,1509]" gridcol="5" gridrow="2" pageId="7" pageNumber="44">19+17, 19+16, 18+17</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0FA710F9DD914" box="[151,1436,1528,1578]" gridrow="3" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0FA710B81D914" box="[151,384,1528,1578]" gridcol="0" gridrow="3" pageId="7" pageNumber="44">
<taxonomicName id="4382A5D7E45EB43F19E8FA710B3AD933" box="[159,315,1528,1549]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="borysthenicus">
<emphasis id="B6F60246E45EB43F19E8FA710B3AD933" box="[159,315,1528,1549]" italics="true" pageId="7" pageNumber="44">P. borysthenicus</emphasis>
</taxonomicName>
(n=277)
</th>
<td id="7963B56AE45E4BC718EAFA710837D914" box="[413,566,1528,1578]" gridcol="1" gridrow="3" pageId="7" pageNumber="44">3639; 37; 37.3</td>
<td id="7963B56AE45E4BC71B04FA71091FD914" box="[627,798,1528,1578]" gridcol="2" gridrow="3" pageId="7" pageNumber="44">1921; 20; 20.1</td>
<td id="7963B56AE45E4BC71A43FA7109D6D914" box="[820,983,1528,1578]" gridcol="3" gridrow="3" pageId="7" pageNumber="44">1619; 17; 17.3</td>
<td id="7963B56AE45E4BC71D77FA710EC3D914" box="[1024,1218,1528,1578]" gridcol="4" gridrow="3" pageId="7" pageNumber="44">1214; 13; 13.1</td>
<td id="7963B56AE45E4BC71DA1FA710F9DD914" box="[1238,1436,1528,1578]" gridcol="5" gridrow="3" pageId="7" pageNumber="44">20+17, 20+18, 19+18</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0F9B40F9DD951" box="[151,1436,1597,1647]" gridrow="4" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0F9B40B81D951" box="[151,384,1597,1647]" gridcol="0" gridrow="4" pageId="7" pageNumber="44">
<taxonomicName id="4382A5D7E45EB43F19E8F9B40AF7D96C" box="[159,246,1597,1618]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="ulanus">
<emphasis id="B6F60246E45EB43F19E8F9B40AF7D96C" box="[159,246,1597,1618]" italics="true" pageId="7" pageNumber="44">P. ulanus</emphasis>
</taxonomicName>
(n=14)
</th>
<td id="7963B56AE45E4BC718EAF9B40837D951" box="[413,566,1597,1647]" gridcol="1" gridrow="4" pageId="7" pageNumber="44">3739; 37; 37.4</td>
<td id="7963B56AE45E4BC71B04F9B4091FD951" box="[627,798,1597,1647]" gridcol="2" gridrow="4" pageId="7" pageNumber="44">2021; 20 and 21; 20.5</td>
<td id="7963B56AE45E4BC71A43F9B409D6D951" box="[820,983,1597,1647]" gridcol="3" gridrow="4" pageId="7" pageNumber="44">1618; 17; 16.9</td>
<td id="7963B56AE45E4BC71D77F9B40EC3D951" box="[1024,1218,1597,1647]" gridcol="4" gridrow="4" pageId="7" pageNumber="44">1315; 14; 14.0</td>
<td id="7963B56AE45E4BC71DA1F9B40F9DD951" box="[1238,1436,1597,1647]" gridcol="5" gridrow="4" pageId="7" pageNumber="44">20+17, 21+17, 21+16</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0F90B0F9DD98B" box="[151,1436,1666,1717]" gridrow="5" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0F90B0B81D98B" box="[151,384,1666,1717]" gridcol="0" gridrow="5" pageId="7" pageNumber="44">
<emphasis id="B6F60246E45EB43F19E8F90A0AE7D9A6" box="[159,230,1667,1688]" italics="true" pageId="7" pageNumber="44">P. k u r u i</emphasis>
(n=4)
</th>
<td id="7963B56AE45E4BC718EAF90B0837D98B" box="[413,566,1666,1717]" gridcol="1" gridrow="5" pageId="7" pageNumber="44">3940; 39; 39.3</td>
<td id="7963B56AE45E4BC71B04F90B091FD98B" box="[627,798,1666,1717]" gridcol="2" gridrow="5" pageId="7" pageNumber="44">2223; 22; 22.3</td>
<td id="7963B56AE45E4BC71A43F90B09D6D98B" box="[820,983,1666,1717]" gridcol="3" gridrow="5" pageId="7" pageNumber="44">17</td>
<td id="7963B56AE45E4BC71D77F90B0EC3D98B" box="[1024,1218,1666,1717]" gridcol="4" gridrow="5" pageId="7" pageNumber="44">1415; 14 and 15; 14.5</td>
<td id="7963B56AE45E4BC71DA1F90B0F9DD98B" box="[1238,1436,1666,1717]" gridcol="5" gridrow="5" pageId="7" pageNumber="44">22+17, 22+18</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0F9410F9DD9C4" box="[151,1436,1736,1786]" gridrow="6" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0F9410B81D9C4" box="[151,384,1736,1786]" gridcol="0" gridrow="6" pageId="7" pageNumber="44">
<taxonomicName id="4382A5D7E45EB43F19E8F9410B39D9E3" box="[159,312,1736,1757]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E45EB43F19E8F9410B39D9E3" box="[159,312,1736,1757]" italics="true" pageId="7" pageNumber="44">P. squaliusculus</emphasis>
</taxonomicName>
(n=38)
</th>
<td id="7963B56AE45E4BC718EAF9410837D9C4" box="[413,566,1736,1786]" gridcol="1" gridrow="6" pageId="7" pageNumber="44">3840; 39; 39.6</td>
<td id="7963B56AE45E4BC71B04F941091FD9C4" box="[627,798,1736,1786]" gridcol="2" gridrow="6" pageId="7" pageNumber="44">2023; 22; 22.1</td>
<td id="7963B56AE45E4BC71A43F94109D6D9C4" box="[820,983,1736,1786]" gridcol="3" gridrow="6" pageId="7" pageNumber="44">1619; 17; 17.5</td>
<td id="7963B56AE45E4BC71D77F9410EC3D9C4" box="[1024,1218,1736,1786]" gridcol="4" gridrow="6" pageId="7" pageNumber="44">1215; 14; 13.8</td>
<td id="7963B56AE45E4BC71DA1F9410F9DD9C4" box="[1238,1436,1736,1786]" gridcol="5" gridrow="6" pageId="7" pageNumber="44">22+17, 22+18, 21+18</td>
</tr>
<tr id="3AB2DC16E45E4BC719E0F8840F9DD801" box="[151,1436,1805,1855]" gridrow="7" pageId="7" pageNumber="44">
<th id="7963B56AE45E4BC719E0F8840B81D801" box="[151,384,1805,1855]" gridcol="0" gridrow="7" pageId="7" pageNumber="44">
<taxonomicName id="4382A5D7E45EB43F19E8F8840B06D81C" box="[159,263,1805,1826]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E45EB43F19E8F8840B06D81C" box="[159,263,1805,1826]" italics="true" pageId="7" pageNumber="44">P. esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE45EB43F187AF8840B3FD81C" box="[269,318,1805,1826]" pageId="7" pageNumber="44" rank="species">
<emphasis id="B6F60246E45EB43F187AF8840B3FD81C" bold="true" box="[269,318,1805,1826]" pageId="7" pageNumber="44">sp.n.</emphasis>
</taxonomicNameLabel>
(n=135)
</th>
<td id="7963B56AE45E4BC718EAF8840837D801" box="[413,566,1805,1855]" gridcol="1" gridrow="7" pageId="7" pageNumber="44">3943; 41; 41.1</td>
<td id="7963B56AE45E4BC71B04F884091FD801" box="[627,798,1805,1855]" gridcol="2" gridrow="7" pageId="7" pageNumber="44">2123; 22; 21.8</td>
<td id="7963B56AE45E4BC71A43F88409D6D801" box="[820,983,1805,1855]" gridcol="3" gridrow="7" pageId="7" pageNumber="44">1821; 20; 19.4</td>
<td id="7963B56AE45E4BC71D77F8840EC3D801" box="[1024,1218,1805,1855]" gridcol="4" gridrow="7" pageId="7" pageNumber="44">1416; 15; 14.7</td>
<td id="7963B56AE45E4BC71DA1F8840F9DD801" box="[1238,1436,1805,1855]" gridcol="5" gridrow="7" pageId="7" pageNumber="44">22+19, 22+20, 21+20</td>
</tr>
</table>
</paragraph>
<paragraph id="843DDE54E45EB43019B2F80C0B76DD3C" blockId="7.[151,1437,1925,2026]" lastBlockId="8.[151,1437,151,850]" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
The new species is distinguished from all other members of the genus
<taxonomicName id="4382A5D7E45EB43F1D6DF80F0EC8D8A1" box="[1050,1225,1926,1951]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E45EB43F1D6DF80F0EC8D8A1" box="[1050,1225,1926,1951]" italics="true" pageId="7" pageNumber="44">Petroleuciscus</emphasis>
</taxonomicName>
by having a high total vertebrae count, modally 41 (vs. 35
<emphasis id="B6F60246E45EB43F1BFBF822089AD8FA" box="[652,667,1963,1988]" italics="true" pageId="7" pageNumber="44"></emphasis>
39) and a high caudal vertebrae count, modally 20 (vs. 1517) (
<tableCitation id="C900EBEFE45EB43F19E9F8590AFBD8D4" box="[158,250,2000,2026]" captionStart="TABLE 2" captionStartId="7.[151,242,1237,1261]" captionTargetBox="[151,1436,1320,1855]" captionTargetPageId="7" captionText="TABLE 2. Vertebral counts (range, mode and mean) and most frequent vertebral formulae in species of the genus Petroleuciscus." httpUri="http://table.plazi.org/id/D0FD8EDCE45EB43F19E0FB5C0B3BDA32" pageId="7" pageNumber="44" tableUuid="D0FD8EDCE45EB43F19E0FB5C0B3BDA32">Table 2</tableCitation>
,
<figureCitation id="1CB9C2D1E45EB43F1871F8590B4DD8D4" box="[262,332,2000,2026]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="7" pageNumber="44">Fig. 3</figureCitation>
).
<collectingCountry id="FC959EC4E45EB43F1814F8590B85D8D4" box="[355,388,2000,2026]" name="American Samoa" pageId="7" pageNumber="44">As</collectingCountry>
can be seen from
<tableCitation id="C900EBEFE45EB43F1B2DF85908B2D8D4" box="[602,691,2000,2026]" captionStart="TABLE 2" captionStartId="7.[151,242,1237,1261]" captionTargetBox="[151,1436,1320,1855]" captionTargetPageId="7" captionText="TABLE 2. Vertebral counts (range, mode and mean) and most frequent vertebral formulae in species of the genus Petroleuciscus." httpUri="http://table.plazi.org/id/D0FD8EDCE45EB43F19E0FB5C0B3BDA32" pageId="7" pageNumber="44" tableUuid="D0FD8EDCE45EB43F19E0FB5C0B3BDA32">Table 2</tableCitation>
, the new species has considerably more vertebrae in all regions than
<taxonomicName id="4382A5D7E451B43019A6FF110B4DDF8F" box="[209,332,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E451B43019A6FF110B4DDF8F" box="[209,332,152,177]" italics="true" pageId="8" pageNumber="45">P. persidis</emphasis>
</taxonomicName>
(
<collectingCountry id="FC959EC4E451B430182BFF110B92DF8C" box="[348,403,152,178]" name="Iran" pageId="8" pageNumber="45">Fars</collectingCountry>
, southern
<collectingCountry id="FC959EC4E451B4301B7BFF11083FDF8C" box="[524,574,152,178]" name="Iran" pageId="8" pageNumber="45">Iran</collectingCountry>
,
<figureCitation id="1CB9C2D1E451B4301B3CFF110891DF8C" box="[587,656,152,178]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="8" pageNumber="45">Fig. 3</figureCitation>
e), and smaller lateral-line scales (4556) than those in
<taxonomicName id="4382A5D7E451B4301C55FF110F9ADF8F" box="[1314,1435,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E451B4301C55FF110F9ADF8F" box="[1314,1435,152,177]" italics="true" pageId="8" pageNumber="45">P. persidis</emphasis>
</taxonomicName>
(3543). Besides vertebral counts, a terminal or superior mouth with a projecting lower jaw forming a chin is a character that makes
<taxonomicName id="4382A5D7E451B43018D0FF6B0828DFC5" box="[423,553,226,251]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E451B43018D0FF6B0828DFC5" box="[423,553,226,251]" italics="true" pageId="8" pageNumber="45">P. esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE451B4301B47FF6B0872DFC2" box="[560,627,226,252]" pageId="8" pageNumber="45" rank="species">
<emphasis id="B6F60246E451B4301B47FF6B0872DFC2" bold="true" box="[560,627,226,252]" pageId="8" pageNumber="45">sp. n.</emphasis>
</taxonomicNameLabel>
different from known species of the genus. The only exception is
<taxonomicName id="4382A5D7E451B4301CF1FF6B0B12DE1F" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="gaderanus">
<emphasis id="B6F60246E451B4301CF1FF6B0B12DE1F" italics="true" pageId="8" pageNumber="45">P. gaderanus</emphasis>
</taxonomicName>
that we tentatively consider as a synonym of
<taxonomicName id="4382A5D7E451B4301A59FE81099BDE1F" box="[814,922,264,289]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="ulanus">
<emphasis id="B6F60246E451B4301A59FE81099BDE1F" box="[814,922,264,289]" italics="true" pageId="8" pageNumber="45">P. ulanus</emphasis>
</taxonomicName>
(both are from the Lake Orumiyeh basin in northwestern
<collectingCountry id="FC959EC4E451B4301840FEA40B6ADE79" box="[311,363,301,327]" name="Iran" pageId="8" pageNumber="45">Iran</collectingCountry>
). The three
<typeStatus id="5B3960F6E451B4301883FEA4085BDE79" box="[500,602,301,327]" pageId="8" pageNumber="45" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4382A5D7E451B4301BF6FEA40976DE78" box="[641,887,301,326]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="gaderanus">
<emphasis id="B6F60246E451B4301BF6FEA40976DE78" box="[641,887,301,326]" italics="true" pageId="8" pageNumber="45">Leuciscus gaderanus</emphasis>
</taxonomicName>
(BMNH 1899.9.30.113-115) possess a distinct chin and a straight mouth cleft with the tip of the mouth above the middle of the eye (
<figureCitation id="1CB9C2D1E451B4301DE7FEDB0ED8DE52" box="[1168,1241,338,364]" captionStart="FIGURE 5" captionStartId="6.[151,255,1817,1841]" captionTargetBox="[173,1412,1323,1793]" captionTargetId="figure@6.[173,1413,1323,1793]" captionTargetPageId="6" captionText="FIGURE 5. Syntype of Leuciscus gaderanus Günther, 1899, BMNH 1899.9.30.113 - 115, 73.0 mm SL." httpUri="https://zenodo.org/record/293385/files/figure.png" pageId="8" pageNumber="45">Fig. 5</figureCitation>
).
<taxonomicName id="4382A5D7E451B4301D87FEDB0AFBDEAF" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E451B4301D87FEDB0AFBDEAF" italics="true" pageId="8" pageNumber="45">Petroleuciscus esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE451B4301875FEFE0B45DEAF" box="[258,324,375,401]" pageId="8" pageNumber="45" rank="species">
<emphasis id="B6F60246E451B4301875FEFE0B45DEAF" bold="true" box="[258,324,375,401]" pageId="8" pageNumber="45">sp. n.</emphasis>
</taxonomicNameLabel>
, however, is easily distinguishable from the
<typeStatus id="5B3960F6E451B4301A22FEF109BADEAC" box="[853,955,376,402]" pageId="8" pageNumber="45" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4382A5D7E451B4301A93FEF10E7FDEAF" box="[996,1150,376,401]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="gaderanus">
<emphasis id="B6F60246E451B4301A93FEF10E7FDEAF" box="[996,1150,376,401]" italics="true" pageId="8" pageNumber="45">P. gaderanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4382A5D7E451B4301DCFFEF10F22DEAF" box="[1208,1315,376,401]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="ulanus">
<emphasis id="B6F60246E451B4301DCFFEF10F22DEAF" box="[1208,1315,376,401]" italics="true" pageId="8" pageNumber="45">P. ulanus</emphasis>
</taxonomicName>
(
<typeStatus id="5B3960F6E451B4301C43FEF10F9DDEAC" box="[1332,1436,376,402]" pageId="8" pageNumber="45" type="syntype">syntypes</typeStatus>
and additional material as listed below), besides the vertebral counts mentioned above, in having a higher range and modes for anal fin branched rays (912½, modes 1011½ vs. 710, mode 9½) and smaller scales (4556 vs. 3645).
</paragraph>
<paragraph id="843DDE54E451B43019B2FD840EA7DC6C" blockId="8.[151,1437,151,850]" pageId="8" pageNumber="45">
The new species differs from both
<taxonomicName id="4382A5D7E451B4301B0EFD840943DD18" box="[633,834,525,550]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E451B4301B0EFD840943DD18" box="[633,834,525,550]" italics="true" pageId="8" pageNumber="45">P. squaliusculus</emphasis>
</taxonomicName>
and
<emphasis id="B6F60246E451B4301A08FD8409DDDD18" box="[895,988,525,550]" italics="true" pageId="8" pageNumber="45">P. k u r u i</emphasis>
in the shape of the mouth cleft and position of the mouth (an upturned straight mouth cleft with the tip of the mouth on a level with the upper margin of the pupil or only slightly below vs. an oblique curved mouth cleft and the tip of the mouth on the level of the inferior margin of the pupil or below) (
<taxonomicName id="4382A5D7E451B4301B9CFDF409A6DDA8" box="[747,935,637,662]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E451B4301B9CFDF409A6DDA8" box="[747,935,637,662]" italics="true" pageId="8" pageNumber="45">P. squaliusculus</emphasis>
</taxonomicName>
is shown in
<figureCitation id="1CB9C2D1E451B4301D4CFDF40E81DDA9" box="[1083,1152,637,663]" captionStart="FIGURE 3" captionStartId="2.[151,255,1862,1886]" captionTargetBox="[305,1279,204,1823]" captionTargetId="figure@2.[300,1289,194,1828]" captionTargetPageId="2" captionText="FIGURE 3. Radiograph of Petroleuciscus esfahani sp. n, holotype (a); P. borysthenicus, ZISP 15324, Paleostomi Lake, Georgia, 96.8 mm SL (b); P. smyrnaeus, syntype, BMNH 1895.12.28.19 - 28, Smyrna, Turkey, 136.0 mm SL (c); P. squaliusculus, syntype, ZISP 2075, Khodzhent, Tadjikistan, 74.7 mm SL (d); and P. persidis, holotype, CMNFI 1979 - 0154 A, holotype, upper Shur River, Iran, 54.7 mm SL (e). Arrows show position of tip of mouth cleft and first caudal vertebra." httpUri="https://zenodo.org/record/293383/files/figure.png" pageId="8" pageNumber="45">Fig. 3</figureCitation>
d). It also has more total gill rakers (1217) in contrast to
<emphasis id="B6F60246E451B4301B50FD2B0884DD85" box="[551,645,674,699]" italics="true" pageId="8" pageNumber="45">P. k u r u i</emphasis>
(1112). The new species can be further distinguished from
<taxonomicName id="4382A5D7E451B4301CF1FD2B0B3CDDDF" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E451B4301CF1FD2B0B3CDDDF" italics="true" pageId="8" pageNumber="45">P. squaliusculus</emphasis>
</taxonomicName>
by having 1217 gill rakers (vs. 911), 4556 total lateral line scales (vs. 4047), 912½, commonly 1011½ branched anal fin rays (vs. 79½, commonly 8½). Data for some meristic characters other than vertebral counts are summarised in
<tableCitation id="C900EBEFE451B4301BE4FC9B08F0DC12" box="[659,753,786,812]" captionStart="TABLE 3" captionStartId="8.[151,239,898,922]" captionTargetBox="[151,1436,1050,1476]" captionTargetPageId="8" captionText="TABLE 3. Total lateral line, dorsal and anal fin branched rays, pharyngeal teeth and gill raker counts (range and mode)" httpUri="http://table.plazi.org/id/D0FD8EDCE451B43019E0FC0B0F9DDCA7" pageId="8" pageNumber="45" tableUuid="D0FD8EDCE451B43019E0FC0B0F9DDCA7">Table 3</tableCitation>
, and those distinguishing
<taxonomicName id="4382A5D7E451B4301D4FFC9B0EC0DC15" box="[1080,1217,786,811]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E451B4301D4FFC9B0EC0DC15" box="[1080,1217,786,811]" italics="true" pageId="8" pageNumber="45">P. esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE451B4301DBDFC9B0F0EDC12" box="[1226,1295,786,812]" pageId="8" pageNumber="45" rank="species">
<emphasis id="B6F60246E451B4301DBDFC9B0F0EDC12" bold="true" box="[1226,1295,786,812]" pageId="8" pageNumber="45">sp. n.</emphasis>
</taxonomicNameLabel>
from other members of the genus in
<collectingCountry id="FC959EC4E451B43018C9FCB10BECDC6C" box="[446,493,824,850]" name="Iran" pageId="8" pageNumber="45">Iran</collectingCountry>
and the Tigris-Euphrates basin are outlined below in a key.
</paragraph>
<caption id="D0FD8EDCE451B43019E0FC0B0F9DDCA7" ID-Table-UUID="D0FD8EDCE451B43019E0FC0B0F9DDCA7" box="[151,1436,898,922]" httpUri="http://table.plazi.org/id/D0FD8EDCE451B43019E0FC0B0F9DDCA7" pageId="8" pageNumber="45" targetBox="[151,1436,1050,1476]" targetIsTable="true" targetPageId="8">
<paragraph id="843DDE54E451B43019E0FC0B0F9DDCA7" blockId="8.[151,1436,898,1031]" box="[151,1436,898,922]" pageId="8" pageNumber="45">
<emphasis id="B6F60246E451B43019E0FC0B0B0BDCA4" bold="true" box="[151,266,898,922]" pageId="8" pageNumber="45">TABLE 3.</emphasis>
Total lateral line, dorsal and anal fin branched rays, pharyngeal teeth and gill raker counts (range and mode)
</paragraph>
</caption>
<paragraph id="843DDE54E451B43019E0FC2B083ADC87" blockId="8.[151,1436,898,1031]" box="[151,571,930,953]" pageId="8" pageNumber="45">
in species of the genus
<taxonomicName id="4382A5D7E451B43018F8FC2B082DDC87" box="[399,556,930,953]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B6F60246E451B43018F8FC2B082DDC87" box="[399,556,930,953]" italics="true" pageId="8" pageNumber="45">Petroleuciscus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="843DDE54E451B43018EAFC5C094ADB39" blockId="8.[151,1436,898,1031]" pageId="8" pageNumber="45">Total lateral line Dorsal-fin Anal fin branched Pharyngeal teeth Gill rakers scales branched rays rays</paragraph>
<paragraph id="843DDE54E451B43019E8FB930E89DAFA" pageId="8" pageNumber="45">
<table id="F6822CF4E4514BC719E0FB930F9DDAFA" box="[151,1436,1050,1476]" gridcols="4" gridrows="6" pageId="8" pageNumber="45">
<tr id="3AB2DC16E4514BC719E0FB930F9DDB72" box="[151,1436,1050,1100]" gridrow="0" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FB930B3ADB72" box="[151,315,1050,1100]" gridcol="0" gridrow="0" pageId="8" pageNumber="45">
<emphasis id="B6F60246E451B43019E8FB930B1BDB11" box="[159,282,1050,1071]" italics="true" pageId="8" pageNumber="45">P. s m y r n a e u s</emphasis>
</th>
<th id="7963B56AE4514BC718EAFB9308DCDB72" box="[413,733,1050,1100]" gridcol="1" gridrow="0" pageId="8" pageNumber="45">1233, incomplete; 67½; 7½ 2226</th>
<th id="7963B56AE4514BC71A54FB93099ADB72" box="[803,923,1050,1100]" gridcol="2" gridrow="0" pageId="8" pageNumber="45">79½; 8½</th>
<th id="7963B56AE4514BC71A9CFB930F9DDB72" box="[1003,1436,1050,1100]" gridcol="3" gridrow="0" pageId="8" pageNumber="45">2.55.2 710; 8</th>
</tr>
<tr id="3AB2DC16E4514BC719E0FBD60F9DDBAC" box="[151,1436,1119,1170]" gridrow="1" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FBD60B3ADBAC" box="[151,315,1119,1170]" gridcol="0" gridrow="1" pageId="8" pageNumber="45">
<taxonomicName id="4382A5D7E451B43019E8FBE90B03DB4B" box="[159,258,1120,1141]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="persidis">
<emphasis id="B6F60246E451B43019E8FBE90B03DB4B" box="[159,258,1120,1141]" italics="true" pageId="8" pageNumber="45">P. persidis</emphasis>
</taxonomicName>
</th>
<td id="7963B56AE4514BC718EAFBD608DCDBAC" box="[413,733,1119,1170]" gridcol="1" gridrow="1" pageId="8" pageNumber="45">3543; 3638 67½; 7½</td>
<td id="7963B56AE4514BC71A54FBD6099ADBAC" box="[803,923,1119,1170]" gridcol="2" gridrow="1" pageId="8" pageNumber="45">79½; 8½</td>
<td id="7963B56AE4514BC71A9CFBD60F9DDBAC" box="[1003,1436,1119,1170]" gridcol="3" gridrow="1" pageId="8" pageNumber="45">1.54.2, 1.54.1 or 1014; 12 1.54.0; 1.54.2</td>
</tr>
<tr id="3AB2DC16E4514BC719E0FB2C0F9DDBE9" box="[151,1436,1189,1239]" gridrow="2" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FB2C0B3ADBE9" box="[151,315,1189,1239]" gridcol="0" gridrow="2" pageId="8" pageNumber="45">
<taxonomicName id="4382A5D7E451B43019E8FB2C0B3ADB84" box="[159,315,1189,1210]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="borysthenicus">
<emphasis id="B6F60246E451B43019E8FB2C0B3ADB84" box="[159,315,1189,1210]" italics="true" pageId="8" pageNumber="45">P. borysthenicus</emphasis>
</taxonomicName>
</th>
<td id="7963B56AE4514BC718EAFB2C08DCDBE9" box="[413,733,1189,1239]" gridcol="1" gridrow="2" pageId="8" pageNumber="45">3541; 3739 78½; 8½</td>
<td id="7963B56AE4514BC71A54FB2C099ADBE9" box="[803,923,1189,1239]" gridcol="2" gridrow="2" pageId="8" pageNumber="45">911½; 10½</td>
<td id="7963B56AE4514BC71A9CFB2C0F9DDBE9" box="[1003,1436,1189,1239]" gridcol="3" gridrow="2" pageId="8" pageNumber="45">2.55.2, 1.55.2 or 810; 9 2.54.2; 2.55.2</td>
</tr>
<tr id="3AB2DC16E4514BC719E0FB630F9DDA04" box="[151,1436,1258,1338]" gridrow="3" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FB630B3ADA04" box="[151,315,1258,1338]" gridcol="0" gridrow="3" pageId="8" pageNumber="45">
<taxonomicName id="4382A5D7E451B43019E8FB630AF7DBC1" box="[159,246,1258,1279]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="ulanus">
<emphasis id="B6F60246E451B43019E8FB630AF7DBC1" box="[159,246,1258,1279]" italics="true" pageId="8" pageNumber="45">P. ulanus</emphasis>
</taxonomicName>
</th>
<td id="7963B56AE4514BC718EAFB6308DCDA04" box="[413,733,1258,1338]" gridcol="1" gridrow="3" pageId="8" pageNumber="45">3845; 3940 78½; 8½</td>
<td id="7963B56AE4514BC71A54FB63099ADA04" box="[803,923,1258,1338]" gridcol="2" gridrow="3" pageId="8" pageNumber="45">710; 9½</td>
<td id="7963B56AE4514BC71A9CFB630F9DDA04" box="[1003,1436,1258,1338]" gridcol="3" gridrow="3" pageId="8" pageNumber="45">2.5 4. 2, 2.4 5. 2, 1216; 1314 2.55.2 or 2.44.2; 2.54.2</td>
</tr>
<tr id="3AB2DC16E4514BC719E0FAC40F9DDA5C" box="[151,1436,1357,1378]" gridrow="4" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FAC40B3ADA5C" box="[151,315,1357,1378]" gridcol="0" gridrow="4" pageId="8" pageNumber="45">
<emphasis id="B6F60246E451B43019E8FAC40AE7DA5C" box="[159,230,1357,1378]" italics="true" pageId="8" pageNumber="45">P. k u r u i</emphasis>
</th>
<td id="7963B56AE4514BC718EAFAC408DCDA5C" box="[413,733,1357,1378]" gridcol="1" gridrow="4" pageId="8" pageNumber="45">5057 8½</td>
<td id="7963B56AE4514BC71A54FAC4099ADA5C" box="[803,923,1357,1378]" gridcol="2" gridrow="4" pageId="8" pageNumber="45">810½</td>
<td id="7963B56AE4514BC71A9CFAC40F9DDA5C" box="[1003,1436,1357,1378]" gridcol="3" gridrow="4" pageId="8" pageNumber="45">2.54.2 or 1.54.2 1112</td>
</tr>
<tr id="3AB2DC16E4514BC719E0FAFC0F9DDAFA" box="[151,1436,1397,1476]" gridrow="5" pageId="8" pageNumber="45">
<th id="7963B56AE4514BC719E0FAFC0B3ADAFA" box="[151,315,1397,1476]" gridcol="0" gridrow="5" pageId="8" pageNumber="45">
<taxonomicName id="4382A5D7E451B43019E8FAFC0B39DAB4" box="[159,312,1397,1418]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="squaliusculus">
<emphasis id="B6F60246E451B43019E8FAFC0B39DAB4" box="[159,312,1397,1418]" italics="true" pageId="8" pageNumber="45">P. squaliusculus</emphasis>
</taxonomicName>
</th>
<td id="7963B56AE4514BC718EAFAFC08DCDAFA" box="[413,733,1397,1476]" gridcol="1" gridrow="5" pageId="8" pageNumber="45">4047; 4345 68½; 7½</td>
<td id="7963B56AE4514BC71A54FAFC099ADAFA" box="[803,923,1397,1476]" gridcol="2" gridrow="5" pageId="8" pageNumber="45">79½; 8½</td>
<td id="7963B56AE4514BC71A9CFAFC0F9DDAFA" box="[1003,1436,1397,1476]" gridcol="3" gridrow="5" pageId="8" pageNumber="45">2. 5 5.2, 1. 5 5.2, 911; 15 2.55.3, 3.5-5.2 or 3.55.3; 2.55.2</td>
</tr>
</table>
</paragraph>
<paragraph id="843DDE54E451B43019E8FA510E32D934" blockId="8.[159,1322,1495,1546]" pageId="8" pageNumber="45">
<taxonomicName id="4382A5D7E451B43019E8FA510B06DAD3" box="[159,263,1496,1517]" class="Actinopterygii" family="Cyprinidae" genus="Petroleuciscus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="esfahani" status="sp. nov.">
<emphasis id="B6F60246E451B43019E8FA510B06DAD3" box="[159,263,1496,1517]" italics="true" pageId="8" pageNumber="45">P. esfahani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADC5BF3DE451B430187AFA5E0B3FDAD2" box="[269,318,1495,1516]" pageId="8" pageNumber="45" rank="species">
<emphasis id="B6F60246E451B430187AFA5E0B3FDAD2" bold="true" box="[269,318,1495,1516]" pageId="8" pageNumber="45">sp.n.</emphasis>
</taxonomicNameLabel>
4556; 50
<date id="F03CF894E451B4301B7AFA5E0822DAD2" box="[525,547,1495,1516]" pageId="8" pageNumber="45" value="1951-09-07">51</date>
79½; 8½ 912½; 1011½ 2.54.2 or 2.55.2; 1217; 15 2.54.2
</paragraph>
</subSubSection>
</treatment>
</document>