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<mods:title id="10551720A0CF940E1A43DA66705B61E5">An updated classification of the jumping plant-lice (Hemiptera: Psylloidea) integrating molecular and morphological evidence</mods:title>
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<mods:affiliation id="B2A5D2D0772158AB21DE159BA67A9509">Naturhistorisches Museum, Augustinergasse 2, 4001 Basel, Switzerland. ANSES, Plant Health Laboratory, Entomology and invasive plants unit, 755 avenue du campus Agropolis, CS 30016, 34988 Montferrier-sur-Lez Cedex, France. Department of Botany, University of British Columbia, 6270 University Boulevard, Vancouver V 6 T 1 Z 4, Canada.</mods:affiliation>
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Subfamily *
<taxonomicName id="438A4D63FFBBFF90FD2F81EAFC96FC38" authority="Burckhardt &amp; Ouvrard &amp; Percy, 2021" authorityName="Burckhardt &amp; Ouvrard &amp; Percy" authorityYear="2021" box="[684,889,898,924]" class="Insecta" family="Calophyidae" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="164" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Platycoryphinae">
<emphasis id="B6FEEAF2FFBBFF90FD2F81EAFC96FC38" bold="true" box="[684,889,898,924]" pageId="27" pageNumber="164">Platycoryphinae</emphasis>
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<figureCitation id="1CB12A65FFBBFF90FD7481AFFCD3FC46" box="[759,828,967,994]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8</figureCitation>
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<emphasis id="B6FEEAF2FFBBFF90FF3E8661FEA5FB87" bold="true" box="[189,330,1033,1059]" pageId="27" pageNumber="164">
<typeStatus id="5B318842FFBBFF90FF3E8661FF15FB87" box="[189,250,1033,1059]" pageId="27" pageNumber="164">Type</typeStatus>
genus
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<taxonomicName id="438A4D63FFBBFF90FF3E865FFE11FBF6" ID-CoL="6RRS" authority="Tuthill, 1945" authorityName="Tuthill" authorityYear="1945" box="[189,510,1079,1106]" class="Insecta" family="Psyllidae" genus="Platycorypha" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBBFF90FF3E865FFEB4FBF5" box="[189,347,1079,1105]" italics="true" pageId="27" pageNumber="164">Platycorypha</emphasis>
Tuthill, 1945
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.
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<subSubSection id="CC90656BFFBBFF90FF3E8610FD1EF9FB" pageId="27" pageNumber="164" type="diagnosis">
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<heading id="DF7D818CFFBBFF90FF3E8610FED9FB36" bold="true" box="[189,310,1144,1170]" fontSize="11" level="3" pageId="27" pageNumber="164" reason="3">
<emphasis id="B6FEEAF2FFBBFF90FF3E8610FED9FB36" bold="true" box="[189,310,1144,1170]" pageId="27" pageNumber="164">Diagnosis</emphasis>
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<emphasis id="B6FEEAF2FFBBFF90FF6986CEFEDDFB64" bold="true" box="[234,306,1190,1216]" pageId="27" pageNumber="164">Adult</emphasis>
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Head with oval frons, which is almost completely covered by median ocellus. Clypeus pear-shaped, in profile hidden by genae and not visible. Propleurites with episternum subequal to or smaller than epimeron. Metacoxa with large, pointed, horn-shaped meracanthus; metatibia with large genual spine, bearing 45 irregularly spaced, sclerotised, apical spurs; metabasitarsus with 2 spurs. Forewing rhomboidal; costal break and pterostigma developed; cell cu 1 large; anal break close to apex of vein Cu
<subScript id="180E34A5FFBBFF90FF5C87E2FF1EFA3C" attach="left" box="[223,241,1418,1432]" fontSize="6" pageId="27" pageNumber="164">1b</subScript>
. Male proctiger one-segmented.
</paragraph>
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<emphasis id="B6FEEAF2FFBBFF90FF6987D0FE14FA76" bold="true" box="[234,507,1464,1490]" pageId="27" pageNumber="164">Fifth instar immature</emphasis>
</paragraph>
<paragraph id="843536E0FFBBFF90FF3E87B3FD1EF9FB" blockId="27.[189,1399,1464,1631]" pageId="27" pageNumber="164">Body broadly oval, lacking capitate setae. Antenna with 4 rhinaria. Meso and metathoracic sclerites small. Forewing pad lacking humeral lobes. Margin of hindwing pad usually with one sectaseta. Legs lacking capitate setae; tarsal arolium fan-shaped with unguitractor and pedicel. Caudal plate developed, semi-circular; margin with up to 3+3 sectasetae.</paragraph>
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<paragraph id="843536E0FFBBFF90FF3E84EDFEA1F93B" blockId="27.[189,334,1669,1695]" box="[189,334,1669,1695]" pageId="27" pageNumber="164">
<heading id="DF7D818CFFBBFF90FF3E84EDFEA1F93B" bold="true" box="[189,334,1669,1695]" fontSize="11" level="3" pageId="27" pageNumber="164" reason="3">
<emphasis id="B6FEEAF2FFBBFF90FF3E84EDFEA1F93B" bold="true" box="[189,334,1669,1695]" pageId="27" pageNumber="164">Description</emphasis>
</heading>
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<paragraph id="843536E0FFBBFF90FF6984DBFEDDF969" blockId="27.[188,1400,1715,2025]" box="[234,306,1715,1741]" pageId="27" pageNumber="164">
<emphasis id="B6FEEAF2FFBBFF90FF6984DBFEDDF969" bold="true" box="[234,306,1715,1741]" pageId="27" pageNumber="164">Adult</emphasis>
</paragraph>
<paragraph id="843536E0FFBBFF97FF3E84BEFED8FDC4" blockId="27.[188,1400,1715,2025]" lastBlockId="28.[186,1400,265,608]" lastPageId="28" lastPageNumber="165" pageId="27" pageNumber="164">
Head, in profile, weakly to strongly inclined at 3090° from longitudinal body axis (
<figureCitation id="1CB12A65FFBBFF90FB4484BEFF39F8B7" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8A, C, E, G</figureCitation>
). Vertex subrectangular to transversely subtrapezoidal (
<figureCitation id="1CB12A65FFBBFF90FC038491FBB9F8B0" box="[896,1110,1785,1812]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8B, D, F, H</figureCitation>
); separated from genae sometimes by transverse or oblique suture, sometimes passing smoothly into genae; genae smoothly rounded (
<figureCitation id="1CB12A65FFBBFF90FEAF8557FE44F8FE" box="[300,427,1855,1882]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8D, H</figureCitation>
) or forming short (
<figureCitation id="1CB12A65FFBBFF90FD0B8557FD31F8FE" box="[648,734,1855,1882]" captionStart="Fig" captionStartId="25.[189,232,1839,1865]" captionTargetBox="[189,1399,265,1772]" captionTargetId="figure-55@25.[189,1399,265,1772]" captionTargetPageId="25" captionText="Fig. 7. Katacephalinae subfam. nov. A, G. Katacephala longiramis (Burckhardt, 1987). B. Katacephala tenuipennis Tuthill, 1944. C, I. Lautereropsis sp. D, J. Notophorina brevicornis Burckhardt, 1987. E, K. Notophorina sp. F, L. Tuthillia sp. H. Katacephala sp. AF. Habitus, in profile view. GL. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594346" httpUri="https://zenodo.org/record/4594346/files/figure.png" pageId="27" pageNumber="164">Fig. 7F</figureCitation>
) or long conical processes (
<figureCitation id="1CB12A65FFBBFF90FBA28557FB93F8FD" box="[1057,1148,1855,1881]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8B</figureCitation>
); coronal suture fully developed or completely reduced; frons oval, almost completely covered by median ocellus; anteorbital tubercle sometimes developed (
<figureCitation id="1CB12A65FFBBFF90FDB385EFFD60F805" box="[560,655,1927,1953]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="27" pageNumber="164">Fig. 8D</figureCitation>
: arrow). Antenna 10-segmented, filiform, ranging from slightly longer than head width to distinctly longer than forewing, segment 3 shorter or longer than segments 7 or 8. Clypeus pear-shaped, in profile hidden by genae and not visible. Rostrum usually short, only tip exceeding procoxae, sometimes longer (in some
<taxonomicName id="438A4D63FFBCFF97FCB78361FC3DFE87" authorityName="Tuthill" authorityYear="1945" box="[820,978,265,291]" class="Insecta" family="Psyllidae" genus="Platycorypha" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBCFF97FCB78361FC3DFE87" box="[820,978,265,291]" italics="true" pageId="28" pageNumber="165">Platycorypha</emphasis>
</taxonomicName>
spp.). Thorax weakly (
<figureCitation id="1CB12A65FFBCFF97FB6D8361FAA2FE87" box="[1262,1357,265,291]" captionStart="Fig" captionStartId="25.[189,232,1839,1865]" captionTargetBox="[189,1399,265,1772]" captionTargetId="figure-55@25.[189,1399,265,1772]" captionTargetPageId="25" captionText="Fig. 7. Katacephalinae subfam. nov. A, G. Katacephala longiramis (Burckhardt, 1987). B. Katacephala tenuipennis Tuthill, 1944. C, I. Lautereropsis sp. D, J. Notophorina brevicornis Burckhardt, 1987. E, K. Notophorina sp. F, L. Tuthillia sp. H. Katacephala sp. AF. Habitus, in profile view. GL. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594346" httpUri="https://zenodo.org/record/4594346/files/figure.png" pageId="28" pageNumber="165">Fig. 7G</figureCitation>
) to strongly (
<figureCitation id="1CB12A65FFBCFF97FEAC8344FE63FEE2" box="[303,396,300,326]" captionStart="Fig" captionStartId="28.[188,231,1925,1951]" captionTargetBox="[189,1399,659,1883]" captionTargetId="figure-197@28.[189,1399,659,1883]" captionTargetPageId="28" captionText="Fig. 8. Platycoryphinae subfam. nov.AB. Allophorina sp.CD. Limbopsylla nata Brown &amp; Hodkinson, 1988. EF. Padaukia macrolobii Burckhardt &amp; Queiroz, 2018. GH. Platycorypha nigrivirga Burckhardt, 1987. A, C, E, G. Habitus, in profile view. B, D, F, H. Head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="28" pageNumber="165">Fig. 8A</figureCitation>
) arched dorsally, about as wide as head; pronotum weakly or very strongly inclined from longitudinal body axis; propleurites narrow to broad, with episternum subequal to or smaller than epimeron. Legs moderately long, tibiae often shorter than femora, sometimes subequal or longer; basitarsi not much longer than broad; metacoxa with large, pointed, horn-shaped meracanthus; metatibia with large genual spine, bearing 45 irregularly spaced, sclerotised, apical spurs; metabasitarsus with 2 spurs. Forewing rhomboidal, broadest in apical third or in the middle, narrowly rounded or angular apically; membrane semitransparent, covered in surface spinules; costal break and pterostigma developed; vein C+Sc weakly or strongly widened; vein R longer than M+Cu; cell cu
<subScript id="180E34A5FFBCFF97FC64805DFC1FFDE7" attach="left" box="[999,1008,565,579]" fontSize="6" pageId="28" pageNumber="165">1</subScript>
large; caudal break close to apex of vein Cu
</paragraph>
<paragraph id="843536E0FFBCFF97FEB48031FEA6FDC3" box="[311,329,601,615]" pageId="28" pageNumber="165">1b</paragraph>
<paragraph id="843536E0FFBCFF96FECB802EFB3DFECD" blockId="28.[186,1400,265,608]" lastBlockId="29.[189,1399,265,362]" lastPageId="29" lastPageNumber="166" pageId="28" pageNumber="165">. Hindwing slightly shorter than forewing, membranous; costal setae ungrouped or grouped; vein R+M+Cu indistinctly trifurcating or splitting into R and M+Cu. Male proctiger one-segmented; in profile, tubular or with posterior lobe. Male subgenital plate subglobular or elongate. Paramere lamellar or complex. Female terminalia short or moderaly long; proctiger often with dorsal hump.</paragraph>
<caption id="D0F56668FFBCFF97FF3F85EDFB41F842" ID-DOI="http://doi.org/10.5281/zenodo.4594349" ID-Zenodo-Dep="4594349" httpUri="https://zenodo.org/record/4594349/files/figure.png" pageId="28" pageNumber="165" startId="28.[188,231,1925,1951]" targetBox="[189,1399,659,1883]" targetPageId="28">
<paragraph id="843536E0FFBCFF97FF3F85EDFB41F842" blockId="28.[187,1399,1925,2022]" pageId="28" pageNumber="165">
<emphasis id="B6FEEAF2FFBCFF97FF3F85EDFEE7F83B" bold="true" box="[188,264,1925,1951]" pageId="28" pageNumber="165">Fig. 8.</emphasis>
<taxonomicName id="438A4D63FFBCFF97FE8E85EDFE23F83B" authority="Burckhardt &amp; Ouvrard &amp; Percy, 2021" authorityName="Burckhardt &amp; Ouvrard &amp; Percy" authorityYear="2021" box="[269,460,1925,1951]" class="Insecta" family="Calophyidae" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Platycoryphinae">Platycoryphinae</taxonomicName>
<taxonomicNameLabel id="ADCD5789FFBCFF97FE5285EDFD89F804" box="[465,614,1925,1952]" pageId="28" pageNumber="165" rank="subFamily">subfam. nov.</taxonomicNameLabel>
<emphasis id="B6FEEAF2FFBCFF97FDEA85EDFD4FF83B" bold="true" box="[617,672,1925,1951]" pageId="28" pageNumber="165">AB</emphasis>
.
<taxonomicName id="438A4D63FFBCFF97FD2C85EDFCD2F83B" authorityName="Hodkinson" authorityYear="1991" box="[687,829,1925,1951]" class="Insecta" family="Calophyidae" genus="Allophorina" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBCFF97FD2C85EDFCD2F83B" box="[687,829,1925,1951]" italics="true" pageId="28" pageNumber="165">Allophorina</emphasis>
</taxonomicName>
sp.
<emphasis id="B6FEEAF2FFBCFF97FCE585EDFC70F83B" bold="true" box="[870,927,1925,1951]" pageId="28" pageNumber="165">CD</emphasis>
.
<taxonomicName id="438A4D63FFBCFF97FC2D85EDFF16F867" authority="Brown &amp; Hodkinson, 1988" authorityName="Brown &amp; Hodkinson" authorityYear="1988" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="species" species="nata">
<emphasis id="B6FEEAF2FFBCFF97FC2D85EDFB97F83B" box="[942,1144,1925,1951]" italics="true" pageId="28" pageNumber="165">Limbopsylla nata</emphasis>
<bibRefCitation id="E01B4B11FFBCFF97FBFE85EEFF16F867" author="Brown R. G. &amp; Hodkinson I. D." pageId="28" pageNumber="165" refId="ref20845" refString="Brown R. G. &amp; Hodkinson I. D. 1988. Taxonomy and Ecology of the jumping Plant-Lice of Panama (Homoptera: Psylloidea). E. J. Brill / Scandinavian Science Press Ltd, Leiden." type="book" year="1988">Brown &amp; Hodkinson, 1988</bibRefCitation>
</taxonomicName>
.
<emphasis id="B6FEEAF2FFBCFF97FE9285C1FEAAF867" bold="true" box="[273,325,1961,1987]" pageId="28" pageNumber="165">EF</emphasis>
.
<taxonomicName id="438A4D63FFBCFF97FEDD85C0FC09F867" authority="Burckhardt &amp; Queiroz, 2018" authorityName="Burckhardt &amp; Queiroz" authorityYear="2018" box="[350,998,1960,1987]" class="Insecta" family="Psyllidae" genus="Padaukia" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="species" species="macrolobii">
<emphasis id="B6FEEAF2FFBCFF97FEDD85C0FD8FF866" box="[350,608,1960,1986]" italics="true" pageId="28" pageNumber="165">Padaukia macrolobii</emphasis>
Burckhardt &amp; Queiroz, 2018
</taxonomicName>
.
<emphasis id="B6FEEAF2FFBCFF97FC7885C0FBD8F866" bold="true" box="[1019,1079,1960,1986]" pageId="28" pageNumber="165">GH</emphasis>
.
<taxonomicName id="438A4D63FFBCFF97FBD285C0FE61F842" authority="Burckhardt, 1987" authorityName="Burckhardt" authorityYear="1987" class="Insecta" family="Psyllidae" genus="Platycorypha" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="165" phylum="Arthropoda" rank="species" species="nigrivirga">
<emphasis id="B6FEEAF2FFBCFF97FBD285C0FA98F866" box="[1105,1399,1960,1986]" italics="true" pageId="28" pageNumber="165">Platycorypha nigrivirga</emphasis>
<bibRefCitation id="E01B4B11FFBCFF97FF3E85A4FE61F842" author="Burckhardt D." box="[189,398,1996,2022]" pageId="28" pageNumber="165" pagination="299 - 392" refId="ref20886" refString="Burckhardt D. 1987. Jumping plant lice (Homoptera: Psylloidea) of the temperate Neotropical region. Part 1: Psyllidae (subfamilies Aphalarinae, Rhinocolinae and Aphalaroidinae). Zoological Journal of the Linnean Society 89: 299 - 392. https: // doi. org / 10.1111 / j. 1096 - 3642.1987. tb 01568. x" type="journal article" year="1987">Burckhardt, 1987</bibRefCitation>
</taxonomicName>
.
<emphasis id="B6FEEAF2FFBCFF97FE1A85A3FDF5F841" bold="true" box="[409,538,1995,2022]" pageId="28" pageNumber="165">A, C, E, G</emphasis>
. Habitus, in profile view.
<emphasis id="B6FEEAF2FFBCFF97FCC985A4FC2AF842" bold="true" box="[842,965,1996,2022]" pageId="28" pageNumber="165">B, D, F, H</emphasis>
. Head, dorsal view.
</paragraph>
</caption>
<paragraph id="843536E0FFBDFF96FF6983F9FE14FE0F" blockId="29.[188,1399,401,675]" box="[234,507,401,427]" pageId="29" pageNumber="166">
<emphasis id="B6FEEAF2FFBDFF96FF6983F9FE14FE0F" bold="true" box="[234,507,401,427]" pageId="29" pageNumber="166">Fifth instar immature</emphasis>
</paragraph>
<paragraph id="843536E0FFBDFF96FF3E83DCFEAFFD06" blockId="29.[188,1399,401,675]" pageId="29" pageNumber="166">Body broadly oval, lacking capitate setae. Antenna 7, 9 or 10-segmented, with 4 rhinaria. Meso and metathoracic sclerites small. Forewing pad lacking humeral lobes. Margin of hindwing pad usually with one sectaseta. Legs lacking capitate setae; tarsal arolium shorter or longer than claws, fan-shaped with unguitractor and pedicel. Caudal plate developed, semi-circular; margin with up to 3+3 sectasetae. Anus in ventral or terminal position; circumanal ring small heart-shaped to large undulate, restricted to ventral side or extended to dorsal side; consisting of a single row or multiple rows of pores, without additional pore fields.</paragraph>
</subSubSection>
<subSubSection id="CC90656BFFBDFF96FF3E80A1FD76FC42" pageId="29" pageNumber="166" type="discussion">
<paragraph id="843536E0FFBDFF96FF3E80A1FEABFD47" blockId="29.[189,324,713,739]" box="[189,324,713,739]" pageId="29" pageNumber="166">
<heading id="DF7D818CFFBDFF96FF3E80A1FEABFD47" bold="true" box="[189,324,713,739]" fontSize="11" level="3" pageId="29" pageNumber="166" reason="3">
<emphasis id="B6FEEAF2FFBDFF96FF3E80A1FEABFD47" bold="true" box="[189,324,713,739]" pageId="29" pageNumber="166">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="843536E0FFBDFF96FF3E8091FD76FC42" blockId="29.[187,1399,760,999]" pageId="29" pageNumber="166">
<bibRefCitation id="E01B4B11FFBDFF96FF3E8091FDE7FCB6" author="Brown R. G. &amp; Hodkinson I. D." box="[189,520,760,787]" pageId="29" pageNumber="166" refId="ref20845" refString="Brown R. G. &amp; Hodkinson I. D. 1988. Taxonomy and Ecology of the jumping Plant-Lice of Panama (Homoptera: Psylloidea). E. J. Brill / Scandinavian Science Press Ltd, Leiden." type="book" year="1988">Brown &amp; Hodkinson (1988)</bibRefCitation>
created
<taxonomicName id="438A4D63FFBDFF96FDE88090FD13FCB6" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[619,764,760,786]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FDE88090FD13FCB6" box="[619,764,760,786]" italics="true" pageId="29" pageNumber="166">Limbopsylla</emphasis>
</taxonomicName>
as a polyphyletic holding place for ten “species of the subfamilies
<taxonomicName id="438A4D63FFBDFF96FEC48174FE2BFC92" authorityName="White &amp; Hodkinson" authorityYear="1985" box="[327,452,796,822]" class="Insecta" family="Psyllidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="subFamily" subFamily="Acizziinae">Acizziinae</taxonomicName>
and
<taxonomicName id="438A4D63FFBDFF96FE7A8174FD70FC92" authorityName="Enderlein" authorityYear="1910" box="[505,671,796,822]" class="Insecta" family="Psyllidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="subFamily" subFamily="Ciriacreminae">Ciriacreminae</taxonomicName>
which cannot be placed in existing genera”. Three species have been removed previously and three species are transferred here (
<tableCitation id="C908035BFFBDFF96FC2F8157FBEBFCFD" box="[940,1028,831,857]" captionStart="Table 1" captionStartId="21.[189,258,265,291]" captionTargetPageId="21" captionText="Table 1. Species in Psyllidae originally assigned to Limbopsylla Brown &amp; Hodkinson, 1988 and their current generic placement." httpUri="http://table.plazi.org/id/D0F56668FFB5FF9EFF3E8361FE18FEE2" pageId="29" pageNumber="166" tableUuid="D0F56668FFB5FF9EFF3E8361FE18FEE2">Table 1</tableCitation>
). Based on adult and immature material of an undescribed species from
<collectingCountry id="FC9D7670FFBDFF96FD21810AFD05FCD8" box="[674,746,866,892]" name="Brazil" pageId="29" pageNumber="166">Brazil</collectingCountry>
associated with
<taxonomicName id="438A4D63FFBDFF96FC31810AFAEDFCD9" box="[946,1282,866,893]" class="Magnoliopsida" family="Fabaceae" genus="Tachigali" kingdom="Plantae" order="Fabales" pageId="29" pageNumber="166" phylum="Tracheophyta" rank="species" species="rugosa">
<emphasis id="B6FEEAF2FFBDFF96FC31810AFB95FCD8" box="[946,1146,866,892]" italics="true" pageId="29" pageNumber="166">Tachigali rugosa</emphasis>
(Fabaceae)
</taxonomicName>
(NHMB) which is congeneric with
<taxonomicName id="438A4D63FFBDFF96FE7081EEFDA9FC3B" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[499,582,902,927]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="species" species="nata">
<emphasis id="B6FEEAF2FFBDFF96FE7081EEFDA9FC3B" box="[499,582,902,927]" italics="true" pageId="29" pageNumber="166">L. nata</emphasis>
</taxonomicName>
, the
<typeStatus id="5B318842FFBDFF96FD0081EEFD59FC04" box="[643,694,902,928]" pageId="29" pageNumber="166">type</typeStatus>
species of
<taxonomicName id="438A4D63FFBDFF96FCC381EDFC20FC3B" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[832,975,901,927]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FCC381EDFC20FC3B" box="[832,975,901,927]" italics="true" pageId="29" pageNumber="166">Limbopsylla</emphasis>
</taxonomicName>
, we conclude that
<taxonomicName id="438A4D63FFBDFF96FB3781EDFAAAFC3B" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[1204,1349,901,927]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FB3781EDFAAAFC3B" box="[1204,1349,901,927]" italics="true" pageId="29" pageNumber="166">Limbopsylla</emphasis>
</taxonomicName>
is a valid genus closely related to
<taxonomicName id="438A4D63FFBDFF96FD9681C1FD5CFC67" authorityName="Tuthill" authorityYear="1945" box="[533,691,937,963]" class="Insecta" family="Psyllidae" genus="Platycorypha" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FD9681C1FD5CFC67" box="[533,691,937,963]" italics="true" pageId="29" pageNumber="166">Platycorypha</emphasis>
</taxonomicName>
, and that the other species included in
<taxonomicName id="438A4D63FFBDFF96FBF681C1FAE9FC67" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[1141,1286,937,963]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FBF681C1FAE9FC67" box="[1141,1286,937,963]" italics="true" pageId="29" pageNumber="166">Limbopsylla</emphasis>
</taxonomicName>
(
<tableCitation id="C908035BFFBDFF96FA9681C1FA84FC67" box="[1301,1387,937,963]" captionStart="Table 1" captionStartId="21.[189,258,265,291]" captionTargetPageId="21" captionText="Table 1. Species in Psyllidae originally assigned to Limbopsylla Brown &amp; Hodkinson, 1988 and their current generic placement." httpUri="http://table.plazi.org/id/D0F56668FFB5FF9EFF3E8361FE18FEE2" pageId="29" pageNumber="166" tableUuid="D0F56668FFB5FF9EFF3E8361FE18FEE2">Table 1</tableCitation>
) are not congeneric with the
<typeStatus id="5B318842FFBDFF96FD8081A5FDD9FC43" box="[515,566,973,999]" pageId="29" pageNumber="166">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="CC90656BFFBDFF96FF3E8666FE20FB39" pageId="29" pageNumber="166" type="multiple">
<paragraph id="843536E0FFBDFF96FF3E8666FE67FB8C" blockId="29.[189,392,1038,1064]" box="[189,392,1038,1064]" pageId="29" pageNumber="166">
<heading id="DF7D818CFFBDFF96FF3E8666FE67FB8C" bold="true" box="[189,392,1038,1064]" fontSize="11" level="3" pageId="29" pageNumber="166" reason="3">
<emphasis id="B6FEEAF2FFBDFF96FF3E8666FE67FB8C" bold="true" box="[189,392,1038,1064]" pageId="29" pageNumber="166">Included genera</emphasis>
</heading>
</paragraph>
<paragraph id="843536E0FFBDFF96FF3E8654FE20FB39" blockId="29.[189,1398,1084,1182]" pageId="29" pageNumber="166">
<taxonomicName id="438A4D63FFBDFF96FF3E8654FDC8FBF2" authority="Hodkinson, 1991" authorityName="Hodkinson" authorityYear="1991" box="[189,551,1084,1110]" class="Insecta" family="Calophyidae" genus="Allophorina" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FF3E8654FEA4FBF2" box="[189,331,1084,1110]" italics="true" pageId="29" pageNumber="166">Allophorina</emphasis>
Hodkinson, 1991
</taxonomicName>
;
<taxonomicName id="438A4D63FFBDFF96FDB58654FBCFFBF2" authority="Brown &amp; Hodkinson, 1988" authorityName="Brown &amp; Hodkinson" authorityYear="1988" box="[566,1056,1084,1111]" class="Insecta" family="Psyllidae" genus="Limbopsylla" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FDB58654FD28FBF2" box="[566,711,1084,1110]" italics="true" pageId="29" pageNumber="166">Limbopsylla</emphasis>
<bibRefCitation id="E01B4B11FFBDFF96FD518655FBCFFBF2" author="Brown R. G. &amp; Hodkinson I. D." box="[722,1056,1084,1111]" pageId="29" pageNumber="166" refId="ref20845" refString="Brown R. G. &amp; Hodkinson I. D. 1988. Taxonomy and Ecology of the jumping Plant-Lice of Panama (Homoptera: Psylloidea). E. J. Brill / Scandinavian Science Press Ltd, Leiden." type="book" year="1988">Brown &amp; Hodkinson, 1988</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="438A4D63FFBDFF96FBB38654FF17FBDE" authority="Hollis &amp; Martin, 1993" authorityName="Hollis &amp; Martin" authorityYear="1993" class="Insecta" family="Psyllidae" genus="Padaukia" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FBB38654FB4FFBF2" box="[1072,1184,1084,1110]" italics="true" pageId="29" pageNumber="166">Padaukia</emphasis>
Hollis &amp; Martin, 1993
</taxonomicName>
(syn.
<taxonomicName id="438A4D63FFBDFF96FEC28608FD03FBDE" authority="Heslop-Harrison" authorityName="Heslop-Harrison" box="[321,748,1120,1146]" class="Insecta" family="Psyllidae" genus="Peltapaurocephala" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FEC28608FDCEFBDE" box="[321,545,1120,1146]" italics="true" pageId="29" pageNumber="166">Peltapaurocephala</emphasis>
Heslop-Harrison
</taxonomicName>
nomen nudum, no description); *
<taxonomicName id="438A4D63FFBDFF96FBF88608FF17FB3A" authority="Tuthill, 1945" authorityName="Tuthill" authorityYear="1945" class="Insecta" family="Psyllidae" genus="Platycorypha" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FBF88608FAF6FBDE" box="[1147,1305,1120,1146]" italics="true" pageId="29" pageNumber="166">Platycorypha</emphasis>
Tuthill, 1945
</taxonomicName>
(syn.
<taxonomicName id="438A4D63FFBDFF96FEC386EBFE52FB39" authorityName="Caldwell" authorityYear="1948" box="[320,445,1155,1181]" class="Insecta" family="Psyllidae" genus="Neopsyllia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="B6FEEAF2FFBDFF96FEC386EBFE52FB39" box="[320,445,1155,1181]" italics="true" pageId="29" pageNumber="166">Neopsyllia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>