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<document id="6BBC81A6A08F1276D7A528F47BD94165" ID-DOI="10.11646/zootaxa.5418.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10726419" ID-ZooBank="928EFF22-3EE9-4C29-92E0-4490C3FF0490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709201492302" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2024" docId="AB0F87F37932191BFF6EFCCEFADFF89C" docLanguage="en" docName="zootaxa.5418.3.5.pdf" docOrigin="Zootaxa 5418 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cirrothaumatia moyobamba Heppner &amp; Bae 2024, new species" docType="treatment" docVersion="2" lastPageNumber="273" masterDocId="5736FF8B7937191EFFF9FFD0FFD6FF99" masterDocTitle="Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)" masterLastPageNumber="278" masterPageNumber="268" pageNumber="273" updateTime="1709201945160" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="41BFB447BB3D67EA603D237954561153">Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)</mods:title>
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<mods:roleTerm id="532E2F14E3D3A9323710E01FEAE0D71C">Author</mods:roleTerm>
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<mods:namePart id="99203F9A048925CEE30E273899BAB007">Heppner, John B.</mods:namePart>
<mods:affiliation id="5C556796D8D3AA433AC11E3204115C62">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA.</mods:affiliation>
<mods:nameIdentifier id="722BFB171628F69AE0A594037D2CBC84" type="email">jheppner@flmnh.ufl.edu</mods:nameIdentifier>
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<mods:namePart id="0C2A6D30934B7FF3EF65819AA914EAAA">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="04B89B5CF1B7EDB20CC6EE6C53EFE7B6" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="3CE9EF9C2EABB5905DDAD702BADB457C">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="5863259B99F7666785EB9C33BC949E50" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="4889E25C929610FC7F8BB53DA1F87B6D">Zootaxa</mods:title>
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<mods:date id="377FE269DEC29C15890CDED570463B51">2024</mods:date>
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<mods:number id="6620C5313DADB04ED10687DA6C630591">2024-02-29</mods:number>
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<mods:number id="A7230B8F48F0BF2DBF9C78B6A707B3EB">5418</mods:number>
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<mods:number id="2F3EC35709671950FDAC2226039762F6">3</mods:number>
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<mods:classification id="A292AC3114C0FDACBE323CC432327871">journal article</mods:classification>
<mods:identifier id="3B54C24562F98DF478B4B883B4215378" type="DOI">10.11646/zootaxa.5418.3.5</mods:identifier>
<mods:identifier id="827F8D7B466AD97AA41144194CF5ED8A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="AB0F87F37932191BFF6EFCCEFADFF89C" ID-DOI="http://doi.org/10.5281/zenodo.10726439" ID-Zenodo-Dep="10726439" LSID="urn:lsid:plazi:treatment:AB0F87F37932191BFF6EFCCEFADFF89C" httpUri="http://treatment.plazi.org/id/AB0F87F37932191BFF6EFCCEFADFF89C" lastPageNumber="273" pageId="5" pageNumber="273">
<subSubSection id="6BBC656E7932191BFF6EFCCEFC8FFCA1" box="[151,857,798,825]" pageId="5" pageNumber="273" type="nomenclature">
<paragraph id="231936E57932191BFF6EFCCEFC8FFCA1" blockId="5.[151,857,798,861]" box="[151,857,798,825]" pageId="5" pageNumber="273">
<heading id="785181897932191BFF6EFCCEFC8FFCA1" bold="true" box="[151,857,798,825]" fontSize="11" level="1" pageId="5" pageNumber="273" reason="1">
<emphasis id="11D2EAF77932191BFF6EFCCEFC8FFCA1" bold="true" box="[151,857,798,825]" pageId="5" pageNumber="273">
<taxonomicName id="E4A64D667932191BFF6EFCCEFD6AFCA0" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[151,700,798,825]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="moyobamba" status="sp. nov.">
<emphasis id="11D2EAF77932191BFF6EFCCEFE39FCA0" bold="true" box="[151,495,798,825]" italics="true" pageId="5" pageNumber="273">Cirrothaumatia moyobamba</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="0AE1578C7932191BFD30FCCFFC8FFCA1" box="[713,857,798,825]" pageId="5" pageNumber="273" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF6EFC93FC64FC3C" pageId="5" pageNumber="273" type="description">
<paragraph id="231936E57932191BFF6EFC93FEB0FCC4" blockId="5.[151,857,798,861]" box="[151,358,835,861]" pageId="5" pageNumber="273">
(
<figureCitation id="BB9D2A607932191BFF66FC93FF38FCC4" box="[159,238,835,861]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="5" pageNumber="273">Figs. 3</figureCitation>
,
<figureCitation id="BB9D2A607932191BFF02FC93FEDDFCC4" box="[251,267,835,861]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">6</figureCitation>
,
<figureCitation id="BB9D2A607932191BFEEEFC93FEE3FCC4" box="[279,309,835,861]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="5" pageNumber="273">12</figureCitation>
,
<figureCitation id="BB9D2A607932191BFEB8FC93FE88FCC4" box="[321,350,835,861]" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="5" pageNumber="273">18</figureCitation>
)
</paragraph>
<paragraph id="231936E57932191BFF6EFC5AFC64FC3C" blockId="5.[151,1437,906,1797]" box="[151,946,906,933]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF6EFC5AFEEAFC3D" bold="true" box="[151,316,906,932]" pageId="5" pageNumber="273">Type-locality:</emphasis>
Rumipata Lodge, Moyobamba, Dept.
<collectingRegion id="E162F8077932191BFD15FC5BFCBAFC3C" box="[748,876,907,933]" country="Peru" name="San Martin" pageId="5" pageNumber="273">San Martín</collectingRegion>
,
<collectingCountry id="5BB176757932191BFC8EFC5BFC78FC3C" box="[887,942,907,933]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFC7EFB32FC74" pageId="5" pageNumber="273" type="diagnosis">
<paragraph id="231936E57932191BFF3EFC7EFB32FC74" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFC7EFEECFC51" bold="true" box="[199,314,942,968]" pageId="5" pageNumber="273">Diagnosis</emphasis>
. This species can be distinguished by the very short ductus bursae (
<figureCitation id="BB9D2A607932191BFBD0FC7FFBBAFC50" box="[1065,1132,943,969]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">Fig. 6</figureCitation>
) (one-half the length of the previous new species) and a small ovate bursa with a large accessory bursa (larger than the bursa).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFC26FDD1FA04" pageId="5" pageNumber="273" type="description">
<paragraph id="231936E57932191BFF3EFC26FD1DFA94" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFC26FE87FB89" bold="true" box="[199,337,1014,1040]" pageId="5" pageNumber="273">Description</emphasis>
. Wing expanse: 12.4 mm female (n = 1). Female (
<figureCitation id="BB9D2A607932191BFC59FC27FC30FB88" box="[928,998,1015,1041]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="5" pageNumber="273">Fig. 3</figureCitation>
). As in
<taxonomicName id="E4A64D667932191BFBBCFC27FB60FB88" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1093,1206,1015,1041]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77932191BFBBCFC27FB60FB88" box="[1093,1206,1015,1041]" italics="true" pageId="5" pageNumber="273">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFB46FC27FAD6FB88" bold="true" box="[1215,1280,1015,1041]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFB46FC27FAD6FB88" box="[1215,1280,1015,1041]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but with the following particulars:
<emphasis id="11D2EAF77932191BFE6FFBCBFE05FBAC" box="[406,467,1051,1077]" italics="true" pageId="5" pageNumber="273">Head</emphasis>
(
<figureCitation id="BB9D2A607932191BFE1FFBCBFD8FFBAC" box="[486,601,1051,1077]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="5" pageNumber="273">Fig. 3a–b</figureCitation>
): caudo-lateral scale tufts white-tan, to dark brown-gray caudally; labial palpus mostly tan, with many tan to brown-gray scales laterally on all segments, with black-brown scales dorsally on apical segment; antenna scape brown-tan.
<emphasis id="11D2EAF77932191BFD5FFBB3FD21FBE4" box="[678,759,1123,1149]" italics="true" pageId="5" pageNumber="273">Thorax</emphasis>
: mostly dark brown mottled with orange scales, and more orange on metathorax; patagia and tegula similar.
<emphasis id="11D2EAF77932191BFD21FB57FC92FB39" box="[728,836,1159,1184]" italics="true" pageId="5" pageNumber="273">Forewing</emphasis>
(
<figureCitation id="BB9D2A607932191BFCACFB57FC4CFB38" box="[853,922,1159,1185]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="5" pageNumber="273">Fig. 3</figureCitation>
) darker than in
<taxonomicName id="E4A64D667932191BFBAEFB57FB1EFB38" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1111,1224,1159,1185]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77932191BFBAEFB57FB1EFB38" box="[1111,1224,1159,1185]" italics="true" pageId="5" pageNumber="273">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFB28FB57FAC4FB38" bold="true" box="[1233,1298,1159,1185]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFB28FB57FAC4FB38" box="[1233,1298,1159,1185]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with much smaller areas of white scaling; black-brown bar adjacent to orange tornal spot mesally larger, and subterminal brown-gray fascia wider, more curved and with distal knob-like extension obsolete.
<emphasis id="11D2EAF77932191BFBA8FB1FFB69FB70" box="[1105,1215,1231,1257]" italics="true" pageId="5" pageNumber="273">Hindwing</emphasis>
(
<figureCitation id="BB9D2A607932191BFB36FB1FFAC5FB70" box="[1231,1299,1231,1257]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="5" pageNumber="273">Fig. 3</figureCitation>
) brown and somewhat less translucent than in
<taxonomicName id="E4A64D667932191BFDEEFB23FD53FA94" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[535,645,1267,1293]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77932191BFDEEFB23FD53FA94" box="[535,645,1267,1293]" italics="true" pageId="5" pageNumber="273">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFD75FB23FD1DFA94" bold="true" box="[652,715,1267,1293]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFD75FB23FD1DFA94" box="[652,715,1267,1293]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="231936E57932191BFF3EFAC7FDD1FA04" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFAC7FEE7FAA8" box="[199,305,1303,1329]" italics="true" pageId="5" pageNumber="273">Abdomen</emphasis>
.
<emphasis id="11D2EAF77932191BFEC6FAC7FE28FAA8" box="[319,510,1303,1329]" italics="true" pageId="5" pageNumber="273">Female genitalia</emphasis>
(
<figureCitation id="BB9D2A607932191BFDF4FAC7FD86FAA8" box="[525,592,1303,1329]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">Fig. 6</figureCitation>
) as in
<taxonomicName id="E4A64D667932191BFD62FAC7FCE8FAA8" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[667,830,1303,1329]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="huemboana" status="n. sp.">
<emphasis id="11D2EAF77932191BFD62FAC7FCE8FAA8" box="[667,830,1303,1329]" italics="true" pageId="5" pageNumber="273">C. huemboana</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFCBCFAC7FC52FAA8" bold="true" box="[837,900,1303,1329]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFCBCFAC7FC52FAA8" box="[837,900,1303,1329]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but caudal end of ductus bursae short and more funnel-like (
<figureCitation id="BB9D2A607932191BFEDDFAEBFEA1FACC" box="[292,375,1339,1365]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">Fig. 6a</figureCitation>
), with slight constriction prior to remainder of short ductus bursae; corpus bursae ovate and small (less than one-half size of bursa of other species); cestum reduced; accessory bursae very large (larger than corpus bursae).
<emphasis id="11D2EAF77932191BFEB0FA53FE55FA04" box="[329,387,1411,1437]" italics="true" pageId="5" pageNumber="273">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFA77FE52F990" pageId="5" pageNumber="273" type="materials_examined">
<paragraph id="231936E57932191BFF3EFA77FE52F990" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFA77FE54FA59" bold="true" box="[199,386,1446,1473]" pageId="5" pageNumber="273">Type materials.</emphasis>
<materialsCitation id="93CE3CB87932191BFE72FA77FE52F990" collectingDate="2012-10-13" collectingDateMax="2012-10-17" collectingDateMin="2012-10-13" collectionCode="MGCL, UNALM" collectorName="J. B. Heppner &amp; C. Carrera" country="Peru" county="Moyobamba" elevation="970" location="Female" municipality="Rumipata Lodge" pageId="5" pageNumber="273" specimenCount="1" stateProvince="San Martin" typeStatus="holotype">
<emphasis id="11D2EAF77932191BFE72FA77FE2EFA58" bold="true" box="[395,504,1447,1473]" pageId="5" pageNumber="273">
<typeStatus id="FC1D88477932191BFE72FA77FE2EFA58" box="[395,504,1447,1473]" pageId="5" pageNumber="273" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="2679603E7932191BFDF0FA77FD88FA58" LSID="urn:lsid:plazi:treatment:AB0F87F37932191BFF6EFCCEFADFF89C:2679603E7932191BFDF0FA77FD88FA58" box="[521,606,1447,1473]" country="Peru" county="Moyobamba" municipality="Rumipata Lodge" name="Female" pageId="5" pageNumber="273" stateProvince="San Martin">Female</location>
,
<collectingMunicipality id="C37DAC9F7932191BFD92FA77FCFFFA58" box="[619,809,1447,1473]" pageId="5" pageNumber="273">Rumipata Lodge</collectingMunicipality>
(
<quantity id="E45E9B007932191BFCC3FA77FC5EFA58" box="[826,904,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="273" unit="m" value="970.0">
<elevation id="A88BD1D67932191BFCC3FA77FC5EFA58" box="[826,904,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="273" unit="m" value="970.0">970 m</elevation>
</quantity>
),
<collectingCounty id="CA784E697932191BFC67FA77FBF8FA58" box="[926,1070,1447,1473]" pageId="5" pageNumber="273">Moyobamba</collectingCounty>
, Dept.
<collectingRegion id="E162F8077932191BFB7AFA77FAD3FA58" box="[1155,1285,1447,1473]" country="Peru" name="San Martin" pageId="5" pageNumber="273">San Martín</collectingRegion>
,
<collectingCountry id="5BB176757932191BFAEAFA77FA9FFA58" box="[1299,1353,1447,1473]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
,
<date id="571810257932191BFAAFFA77FED7FA7C" pageId="5" pageNumber="273" value="2012-10-13" valueMax="2012-10-17" valueMin="2012-10-13">
<collectingDate id="475CE9CD7932191BFAAFFA77FED7FA7C" pageId="5" pageNumber="273" value="2012-10-13" valueMax="2012-10-17" valueMin="2012-10-13">13–17 Oct 2012</collectingDate>
</date>
,
<collectorName id="8E5353337932191BFEF5FA1BFE73FA7C" box="[268,421,1483,1509]" pageId="5" pageNumber="273">J. B. Heppner</collectorName>
&amp;
<collectorName id="8E5353337932191BFE31FA1BFDEAFA7C" box="[456,572,1483,1509]" pageId="5" pageNumber="273">C. Carrera</collectorName>
(gen. slide JBH-4470; adult photo 15858) (deposited at
<collectionCode id="45B7AE207932191BFB49FA1BFAD2FA7C" box="[1200,1284,1483,1509]" pageId="5" pageNumber="273">MGCL</collectionCode>
, on indefinite loan from
<collectionCode id="45B7AE207932191BFEF5FA3FFEAEF990" box="[268,376,1519,1545]" pageId="5" pageNumber="273">UNALM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EF9C3FAB1F9B4" box="[199,1383,1555,1581]" pageId="5" pageNumber="273" type="etymology">
<paragraph id="231936E57932191BFF3EF9C3FAB1F9B4" blockId="5.[151,1437,906,1797]" box="[199,1383,1555,1581]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EF9C3FE9EF9B4" bold="true" box="[199,328,1555,1581]" pageId="5" pageNumber="273">Etymology</emphasis>
. The species is named after Moyobamba, a town in northern Dept.
<collectingRegion id="E162F8077932191BFBC2F9C3FB6DF9B4" box="[1083,1211,1555,1581]" country="Peru" name="San Martin" pageId="5" pageNumber="273">San Martín</collectingRegion>
, northern
<collectingCountry id="5BB176757932191BFAD5F9C3FAB5F9B4" box="[1324,1379,1555,1581]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EF9E6FC76F9C8" box="[199,928,1590,1617]" pageId="5" pageNumber="273" type="biology_ecology">
<paragraph id="231936E57932191BFF3EF9E6FC76F9C8" blockId="5.[151,1437,906,1797]" box="[199,928,1590,1617]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EF9E6FEF7F9C9" bold="true" box="[199,289,1590,1616]" pageId="5" pageNumber="273">Biology</emphasis>
. Unknown. Habitat submontane Andean forest (
<figureCitation id="BB9D2A607932191BFCC6F9E7FC45F9C8" box="[831,915,1591,1617]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="5" pageNumber="273">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EF98AFD8AF924" pageId="5" pageNumber="273" type="distribution">
<paragraph id="231936E57932191BFF3EF98AFD8AF924" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EF98AFE8FF9ED" bold="true" box="[199,345,1626,1652]" pageId="5" pageNumber="273">Distribution</emphasis>
. Known only from the eastern Andean submontane site at Rumipata Lodge (
<quantity id="E45E9B007932191BFB4EF98BFAD5F9EC" box="[1207,1283,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="273" unit="m" value="970.0">970 m</quantity>
), by the town of Moyobamba (
<figureCitation id="BB9D2A607932191BFEAFF9AFFE7CF900" box="[342,426,1663,1689]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="5" pageNumber="273">Fig. 12</figureCitation>
), a mountain upland area in Dept.
<collectingRegion id="E162F8077932191BFCC2F9AFFC6BF900" box="[827,957,1663,1689]" country="Peru" name="San Martin" pageId="5" pageNumber="273">San Martín</collectingRegion>
. This locality is on the eastern side of the Andes Range of northern
<collectingCountry id="5BB176757932191BFE41F973FE3DF924" box="[440,491,1699,1725]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
(
<figureCitation id="BB9D2A607932191BFE02F973FD98F924" box="[507,590,1699,1725]" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="5" pageNumber="273">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EF916FADFF89C" pageId="5" pageNumber="273" type="discussion">
<paragraph id="231936E57932191BFF3EF916FADFF89C" blockId="5.[151,1437,906,1797]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EF916FE93F979" bold="true" box="[199,325,1734,1760]" pageId="5" pageNumber="273">Discussion</emphasis>
. The female genitalia of
<taxonomicName id="E4A64D667932191BFDA5F917FCD3F978" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[604,773,1735,1761]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="moyobamba">
<emphasis id="11D2EAF77932191BFDA5F917FCD3F978" box="[604,773,1735,1761]" italics="true" pageId="5" pageNumber="273">C. moyobamba</emphasis>
</taxonomicName>
differ considerably from other species of the genus, but the wing maculation is much the same, so discovery of the male is needed to help elucidate relationships.
</paragraph>
</subSubSection>
</treatment>
</document>
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