treatments-xml/data/AB/0E/18/AB0E185AC46FFFAA4FE91F0CAF372024.xml
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<mods:title id="E4643E7AA6CA9E228D77FA4F67CFB38B">New species of Cirratulidae (Annelida, Polychaeta) from abyssal depths of the Clarion-Clipperton Fracture Zone, North Equatorial Pacific Ocean</mods:title>
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<taxonomicName id="E4A7D2CFC46FFFA74FE91F0CADAD23B3" ID-CoL="85NQ2" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[151,393,621,648]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="174" phylum="Annelida" rank="species" species="grasslei" status="sp. nov.">
<emphasis id="11D3755EC46FFFA74FE91F0CADAD23B3" bold="true" box="[151,393,621,648]" italics="true" pageId="23" pageNumber="174">Chaetozone grasslei</emphasis>
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<figureCitation id="BB9CB5C9C46FFFA74FE91FF0AD1C2390" box="[151,312,656,683]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStartId-0="24.[151,250,1983,2008]" captionStartId-1="25.[151,250,1667,1692]" captionTargetBox-0="[171,1423,639,1948]" captionTargetBox-1="[163,1402,190,1641]" captionTargetId-0="figure@24.[151,1436,633,1964]" captionTargetId-1="figure@25.[151,1410,181,1647]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." captionText-1="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri-0="https://zenodo.org/record/3269019/files/figure.png" httpUri-1="https://zenodo.org/record/3269021/files/figure.png" pageId="23" pageNumber="174">Figures 1516</figureCitation>
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<taxonomicName id="E4A7D2CFC46FFFA74FE91E6CAD2B221E" ID-CoL="84M8M" authorityName="Malmgren" authorityYear="1867" box="[151,271,781,805]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="174" phylum="Annelida" rank="genus">
<emphasis id="11D3755EC46FFFA74FE91E6CAD2B221E" box="[151,271,781,805]" italics="true" pageId="23" pageNumber="174">Chaetozone</emphasis>
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sp. A:
<treatmentCitation id="A2068F5DC46FFFA74E281E6CAE4E221D" author="Wilson, G. D. F. &amp; Hessler, R. R." box="[342,618,781,806]" page="66" pageId="23" pageNumber="174" year="1987">
<bibRefCitation id="4736D4BDC46FFFA74E281E6CAE4E221D" author="Wilson, G. D. F. &amp; Hessler, R. R." box="[342,618,781,806]" pageId="23" pageNumber="174" pagination="24 - 86" refId="ref21781" refString="Wilson, G. D. F. &amp; Hessler, R. R. (1987) The effects of manganese nodule test mining on the benthic fauna in the North Equatorial Pacific. In: Spiess, F. N., Hessler, R., Wilson, G. &amp; Weydert, M. (Eds.), Environmental effects of deep sea dredging. Final Report prepared for the National Oceanographic and Atmospheric Administration under Contract NO. 83 - SAC- 00659. Scripps Institution of Oceanography, La Jolla, SIO Reference 87 - 5, pp. 24 - 86, appendices A-H. https: // doi. org / 10.13140 / RG. 2.1.1024.2080" type="journal article" year="1987">Wilson &amp; Hessler 1987: 66</bibRefCitation>
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Appendix E (in part).
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<emphasis id="11D3755EC46FFFA74FE91E32A9B82255" bold="true" box="[151,1436,851,878]" pageId="23" pageNumber="174">Material examined. North Equatorial Pacific Ocean, abyssal plain, Clarion-Clipperton Fracture Zone.</emphasis>
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Project site
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<collectingDate id="475D7664C46FFFA74B1A1969A8DD2519" box="[1124,1273,1032,1058]" pageId="23" pageNumber="174" value="1993-09-04">04 Sep 1993</collectingDate>
</date>
,
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,
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,
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</quantity>
(1,
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)
</materialsCitation>
;
<materialsCitation id="93CFA311C46FFFA74DEE194DAE732551" ID-GBIF-Occurrence="2412500508" collectingDate="1994-07-29" collectionCode="USNN" elevation="4880" latitude="12.917" longLatPrecision="1" longitude="-128.59" pageId="23" pageNumber="174" specimenCode="USNN 1557555" specimenCount="1" typeStatus="paratype">
Sta. DDT-8-94, on a nodule,
<date id="57198F8CC46FFFA74CB3194DA874257D" box="[973,1104,1068,1094]" pageId="23" pageNumber="174" value="1994-07-29">
<collectingDate id="475D7664C46FFFA74CB3194DA874257D" box="[973,1104,1068,1094]" pageId="23" pageNumber="174" value="1994-07-29">29 Jul 1994</collectingDate>
</date>
,
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,
<geoCoordinate id="4693CF8BC46FFFA74B8B194AA9BC257D" box="[1269,1432,1067,1094]" degrees="128" direction="west" minutes="35.400" orientation="longitude" pageId="23" pageNumber="174" precision="1" value="-128.59">128°35.400W</geoCoordinate>
,
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<elevation id="A88A4E7FC46FFFA74FE91931ACD52551" box="[151,241,1104,1130]" metricMagnitude="3" metricUnit="m" metricValue="4.88" pageId="23" pageNumber="174" unit="m" value="4880.0">4880 m</elevation>
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, 1
<emphasis id="11D3755EC46FFFA74E681931ADA52551" bold="true" box="[278,385,1104,1130]" pageId="23" pageNumber="174">
<typeStatus id="FC1C17EEC46FFFA74E681931ADA52551" box="[278,385,1104,1130]" pageId="23" pageNumber="174" type="paratype">paratype</typeStatus>
</emphasis>
(
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)
</materialsCitation>
.
<materialsCitation id="93CFA311C46FFFA74D24192EADE72589" ID-GBIF-Occurrence="2412500497" collectingDate="1983-06-14" collectionCode="DOMES, LACM-AHF" collectorName="R. Hessler" elevation="3" latitude="14.635377" location="Melville" longLatPrecision="1" longitude="-125.447014" municipality="Site" pageId="23" pageNumber="174" specimenCount="2" specimenCount-juvenile="2">
<emphasis id="11D3755EC46FFFA74D06192EAF8E2551" bold="true" box="[632,938,1103,1130]" pageId="23" pageNumber="174">
<taxonomicName id="E4A7D2CFC46FFFA74D06192EAEEE2551" box="[632,714,1103,1130]" class="Insecta" family="Calopterygidae" genus="Echo" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="186" phylum="Arthropoda" rank="genus">ECHO</taxonomicName>
I,
<collectionCode id="45B63189C46FFFA74D91192EAF722551" box="[751,854,1103,1130]" pageId="23" pageNumber="174">DOMES</collectionCode>
<collectingMunicipality id="C37C3336C46FFFA74C21192EAFA82551" box="[863,908,1103,1130]" pageId="23" pageNumber="174">Site</collectingMunicipality>
C
</emphasis>
, R/V
<emphasis id="11D3755EC46FFFA74C8C1931A86A2551" box="[1010,1102,1104,1130]" italics="true" pageId="23" pageNumber="174">
<location id="2678FF97C46FFFA74C8C1931A86A2551" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74C8C1931A86A2551" box="[1010,1102,1104,1130]" latitude="14.635377" longLatPrecision="1" longitude="-125.447014" municipality="Site" name="Melville" pageId="23" pageNumber="174">Melville</location>
</emphasis>
cruise, coll.
<collectorName id="8E52CC9AC46FFFA74B9B1931A97A2551" box="[1253,1374,1104,1130]" pageId="23" pageNumber="174">R. Hessler</collectorName>
,
<quantity id="E45F04A9C46FFFA74A15192EAC8925B5" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="174" unit="m" value="0.25">
<elevation id="A88A4E7FC46FFFA74A15192EAC8925B5" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="174" unit="m" value="0.25">0.25 m</elevation>
</quantity>
<superScript id="D4D20404C46FFFA74FD31913AC9125BB" attach="left" box="[173,181,1138,1152]" fontSize="6" pageId="23" pageNumber="174">2</superScript>
<location id="2678FF97C46FFFA74FC21915AD2C25B5" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74FC21915AD2C25B5" box="[188,264,1140,1166]" latitude="14.635377" longLatPrecision="1" longitude="-125.447014" municipality="Site" name="Sandia" pageId="23" pageNumber="174">Sandia</location>
box core, Sta. H350,
<quantity id="E45F04A9C46FFFA74E811915AE7625B5" box="[511,594,1140,1166]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="23" pageNumber="174" unit="cm" value="0.5" valueMax="1.0" valueMin="0.0">01 cm</quantity>
fraction,
<date id="57198F8CC46FFFA74DC11915AF6F25B5" box="[703,843,1140,1166]" pageId="23" pageNumber="174" value="1983-06-14">
<collectingDate id="475D7664C46FFFA74DC11915AF6F25B5" box="[703,843,1140,1166]" pageId="23" pageNumber="174" value="1983-06-14">14 Jun 1983</collectingDate>
</date>
,
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,
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,
<quantity id="E45F04A9C46FFFA74BC01912A93225B5" box="[1214,1302,1139,1166]" metricMagnitude="3" metricUnit="m" metricValue="4.506" pageId="23" pageNumber="174" unit="m" value="4506.0">
<elevation id="A88A4E7FC46FFFA74BC01912A93225B5" box="[1214,1302,1139,1166]" metricMagnitude="3" metricUnit="m" metricValue="4.506" pageId="23" pageNumber="174" unit="m" value="4506.0">4506 m</elevation>
</quantity>
,
<specimenCount id="35A162C5C46FFFA74A5C1915A9B825B5" box="[1314,1436,1140,1166]" pageId="23" pageNumber="174" type="juvenile">2 juveniles</specimenCount>
(
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<location id="2678FF97C46FFFA74E4619F9AD4F2589" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74E4619F9AD4F2589" box="[312,363,1176,1202]" latitude="14.635377" longLatPrecision="1" longitude="-125.447014" municipality="Site" name="Poly" pageId="23" pageNumber="174">Poly</location>
11282)
</materialsCitation>
;
<materialsCitation id="93CFA311C46FFFA74EB319F9AE2C25ED" ID-GBIF-Occurrence="2412500506" collectingDate="1983-06-21" collectingDateMax="43-11-28" collectingDateMin="1983-06-21" collectionCode="LACM-AHF" elevation="4518" latitude="14.707568" location="Poly" longLatPrecision="1" longitude="-125.40444" pageId="23" pageNumber="174" specimenCount="1">
Sta. 356,
<quantity id="E45F04A9C46FFFA74D4A19F9AEA22589" box="[564,646,1176,1202]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="23" pageNumber="174" unit="cm" value="0.5" valueMax="1.0" valueMin="0.0">01 cm</quantity>
fraction,
<date id="57198F8CC46FFFA74D9119F9AF5C2589" box="[751,888,1176,1202]" pageId="23" pageNumber="174" value="1983-06-21">
<collectingDate id="475D7664C46FFFA74D9119F9AF5C2589" box="[751,888,1176,1202]" pageId="23" pageNumber="174" value="1983-06-21">21 Jun 1983</collectingDate>
</date>
,
<geoCoordinate id="4693CF8BC46FFFA74CFC19F6A8052589" box="[898,1057,1175,1202]" degrees="14" direction="north" minutes="42.4541" orientation="latitude" pageId="23" pageNumber="174" precision="1" value="14.707568">14°42.4541N</geoCoordinate>
,
<geoCoordinate id="4693CF8BC46FFFA74B5519F6A8F82589" box="[1067,1244,1175,1202]" degrees="125" direction="west" minutes="24.2664" orientation="longitude" pageId="23" pageNumber="174" precision="1" value="-125.40444">125°24.2664W</geoCoordinate>
,
<quantity id="E45F04A9C46FFFA74B9819F6A91F2589" box="[1254,1339,1175,1202]" metricMagnitude="3" metricUnit="m" metricValue="4.518" pageId="23" pageNumber="174" unit="m" value="4518.0">
<elevation id="A88A4E7FC46FFFA74B9819F6A91F2589" box="[1254,1339,1175,1202]" metricMagnitude="3" metricUnit="m" metricValue="4.518" pageId="23" pageNumber="174" unit="m" value="4518.0">4518 m</elevation>
</quantity>
, 1 speci- men (
<collectionCode id="45B63189C46FFFA74FAB19DDAD4D25ED" box="[213,361,1212,1238]" pageId="23" pageNumber="174">LACM-AHF</collectionCode>
<location id="2678FF97C46FFFA74E1119DDAD8625ED" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74E1119DDAD8625ED" box="[367,418,1212,1238]" latitude="14.707568" longLatPrecision="1" longitude="-125.40444" name="Poly" pageId="23" pageNumber="174">Poly</location>
<date id="57198F8CC46FFFA74ED619DDAE2425ED" box="[424,512,1212,1238]" pageId="23" pageNumber="174" value="43-11-28">
<collectingDate id="475D7664C46FFFA74ED619DDAE2425ED" box="[424,512,1212,1238]" pageId="23" pageNumber="174" value="43-11-28">112843</collectingDate>
</date>
)
</materialsCitation>
;
<materialsCitation id="93CFA311C46FFFA74D6C19DDAE6825C1" ID-GBIF-Occurrence="2412500495" collectingDate="1983-06-23" collectionCode="LACM-AHF" elevation="4510" latitude="14.70234" location="Poly" longLatPrecision="1" longitude="-125.40465" pageId="23" pageNumber="174" specimenCount="1">
Sta. H357,
<quantity id="E45F04A9C46FFFA74DF019DDAEC425ED" box="[654,736,1212,1238]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="23" pageNumber="174" unit="cm" value="0.5" valueMax="1.0" valueMin="0.0">01 cm</quantity>
fraction,
<date id="57198F8CC46FFFA74C3419DDAFF725ED" box="[842,979,1212,1238]" pageId="23" pageNumber="174" value="1983-06-23">
<collectingDate id="475D7664C46FFFA74C3419DDAFF725ED" box="[842,979,1212,1238]" pageId="23" pageNumber="174" value="1983-06-23">23 Jun 1983</collectingDate>
</date>
,
<geoCoordinate id="4693CF8BC46FFFA74CA319DAA85825ED" box="[989,1148,1211,1238]" degrees="14" direction="north" minutes="42.1404" orientation="latitude" pageId="23" pageNumber="174" precision="1" value="14.70234">14°42.1404N</geoCoordinate>
,
<geoCoordinate id="4693CF8BC46FFFA74BF919DAA91C25ED" box="[1159,1336,1211,1238]" degrees="125" direction="west" minutes="24.2787" orientation="longitude" pageId="23" pageNumber="174" precision="1" value="-125.40465">125°24.2787W</geoCoordinate>
,
<quantity id="E45F04A9C46FFFA74A3C19DAA9B325ED" box="[1346,1431,1211,1238]" metricMagnitude="3" metricUnit="m" metricValue="4.51" pageId="23" pageNumber="174" unit="m" value="4510.0">
<elevation id="A88A4E7FC46FFFA74A3C19DAA9B325ED" box="[1346,1431,1211,1238]" metricMagnitude="3" metricUnit="m" metricValue="4.51" pageId="23" pageNumber="174" unit="m" value="4510.0">4510 m</elevation>
</quantity>
,
<specimenCount id="35A162C5C46FFFA74FE91981AD3225C1" box="[151,278,1248,1274]" pageId="23" pageNumber="174" type="generic">1 specimen</specimenCount>
(
<collectionCode id="45B63189C46FFFA74E5A1981AD9C25C1" box="[292,440,1248,1274]" pageId="23" pageNumber="174">LACM-AHF</collectionCode>
<location id="2678FF97C46FFFA74EC11981ADD625C1" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74EC11981ADD625C1" box="[447,498,1248,1274]" latitude="14.70234" longLatPrecision="1" longitude="-125.40465" name="Poly" pageId="23" pageNumber="174">Poly</location>
11284)
</materialsCitation>
;
<materialsCitation id="93CFA311C46FFFA74D291981A9BE25C1" ID-GBIF-Occurrence="2412500496" box="[599,1434,1247,1274]" collectionCode="LACM-AHF" location="Poly" pageId="23" pageNumber="174" specimenCount="1" specimenCount-juvenile="1">
Sta. H357, top water from box core,
<specimenCount id="35A162C5C46FFFA74C8F1981A84425C1" box="[1009,1120,1248,1274]" pageId="23" pageNumber="174" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="45B63189C46FFFA74B111981A92725C1" box="[1135,1283,1248,1274]" pageId="23" pageNumber="174">LACM-AHF</collectionCode>
<location id="2678FF97C46FFFA74A741981A91925C1" LSID="urn:lsid:plazi:treatment:AB0E185AC46FFFAA4FE91F0CAF372024:2678FF97C46FFFA74A741981A91925C1" box="[1290,1341,1248,1274]" name="Poly" pageId="23" pageNumber="174">Poly</location>
11285).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6BBDFAC7C46FFFA84FB91862ACC720B0" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="description">
<paragraph id="2318A94CC46FFFA74FB91862A8782721" blockId="23.[151,1437,851,2030]" pageId="23" pageNumber="174">
<emphasis id="11D3755EC46FFFA74FB91862AD752425" bold="true" box="[199,337,1283,1310]" pageId="23" pageNumber="174">Description</emphasis>
. An elongate, slender species (
<figureCitation id="BB9CB5C9C46FFFA74DC51865AF032425" box="[699,807,1283,1310]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="23" pageNumber="174">Fig. 16A</figureCitation>
).
<typeStatus id="FC1C17EEC46FFFA74C431865AF812425" box="[829,933,1284,1310]" pageId="23" pageNumber="174" type="holotype">Holotype</typeStatus>
(USNM 1557550) complete,
<quantity id="E45F04A9C46FFFA74A7F1862A97E2425" box="[1281,1370,1283,1310]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="174" unit="mm" value="3.5">3.5 mm</quantity>
long,
<quantity id="E45F04A9C46FFFA74FE91846AD2D2479" box="[151,265,1319,1346]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="174" unit="mm" value="0.15">0.150 mm</quantity>
wide across anterior setigers, with 36 setigerous segments;
<typeStatus id="FC1C17EEC46FFFA74CD41849A82E2479" box="[938,1034,1320,1346]" pageId="23" pageNumber="174" type="paratype">paratype</typeStatus>
(USNM 1557553) in two parts,
<quantity id="E45F04A9C46FFFA74A071849ACE7245D" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="23" pageNumber="174" unit="mm" value="4.9">4.9 mm</quantity>
long,
<quantity id="E45F04A9C46FFFA74E71182DAD51245D" box="[271,373,1356,1382]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="174" unit="mm" value="0.22">0.22 mm</quantity>
wide across anterior segments, with 46 setigerous segments. Anterior 56 segments narrow, about three times wider than long (
<figureCitation id="BB9CB5C9C46FFFA74D551811AEB124B1" box="[555,661,1391,1418]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
), middle and posterior segments becoming longer, eventually about 1.5 times as long as wide, sometimes appearing oval or moniliform (
<figureCitation id="BB9CB5C9C46FFFA74CEB18F5A8212495" box="[917,1029,1427,1454]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Figs. 15B</figureCitation>
,
<figureCitation id="BB9CB5C9C46FFFA74B6F18F2A8092495" box="[1041,1069,1427,1454]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="23" pageNumber="174">16</figureCitation>
AB).
<typeStatus id="FC1C17EEC46FFFA74B0718F5A8C52495" box="[1145,1249,1428,1454]" pageId="23" pageNumber="174" type="holotype">Holotype</typeStatus>
and a few larger
<typeStatus id="FC1C17EEC46FFFA74FE918D9AD2624E9" box="[151,258,1464,1490]" pageId="23" pageNumber="174" type="paratype">paratypes</typeStatus>
with some individual segments along body enlarged, appearing distended due to concentration of ingested sediment particles in intestine (
<figureCitation id="BB9CB5C9C46FFFA74E8D18BDAE6024CD" box="[499,580,1499,1526]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="23" pageNumber="174">Fig. 16</figureCitation>
AB). Dorsum rounded, venter flattened or with shallow groove along much of body. Color in alcohol opaque white, with diffuse reddish pigment on peristomium.
</paragraph>
<paragraph id="2318A94CC46FFFA74FB91B45ADD2262D" blockId="23.[151,1437,851,2030]" pageId="23" pageNumber="174">
Pre-setiger region relatively narrow, about twice as long as wide (
<figureCitation id="BB9CB5C9C46FFFA74CC71B45A8002705" box="[953,1060,1571,1598]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
). Prostomium broadly triangular, narrowing to rounded tip (
<figureCitation id="BB9CB5C9C46FFFA74EBA1B29AE0A2759" box="[452,558,1607,1634]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
); eyespots absent; nuchal organs not observed. Peristomium divided into two rings, first ring broadly rounded ending at narrower second ring bearing dorsal tentacles and first pair of branchiae (
<figureCitation id="BB9CB5C9C46FFFA74FDE1BF1AD2D2791" box="[160,265,1679,1706]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
); dorsal crest absent. Dorsal tentacles arising from anterior half of second peristomial ring, immediately followed by first pair of branchiae (
<figureCitation id="BB9CB5C9C46FFFA74D561BD5AEB527F5" box="[552,657,1715,1742]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
). Second pair of branchiae arising in a line posterior to the first branchiae dorsal to notosetae (
<figureCitation id="BB9CB5C9C46FFFA74EC71BB9AE0627C9" box="[441,546,1751,1778]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15A</figureCitation>
). Subsequent branchiae arising from similar location; branchiae not observed in middle and posterior segments.
</paragraph>
<paragraph id="2318A94CC46FFFA84FB91A41ACD52024" blockId="23.[151,1437,851,2030]" lastBlockId="24.[151,1437,152,611]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
Parapodia reduced, lateral shoulders not developed, distinct podial lobes absent, setae appearing to arise directly from body wall, some anterior setigers with low mound posterior to origin of setae. Posterior segments with spines and capillaries forming cinctures, but these arising from low ridges, not elevated into high membranes (
<figureCitation id="BB9CB5C9C46FFFA74A111A09ACEE269D" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="23" pageNumber="174">Fig. 15B</figureCitation>
). Anterior setae all capillaries, 34 per noto- and neuropodia; additional long natatory capillaries on variable number of anterior and middle segments. Noto- and neuropodial acicular spines from setiger 16 on all larger specimens examined including
<typeStatus id="FC1C17EEC46FFFA74EBB1AB5AE0326D5" box="[453,551,2004,2030]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
and a
<typeStatus id="FC1C17EEC46FFFA74D091AB5AEF326D5" box="[631,727,2004,2030]" pageId="23" pageNumber="174" type="paratype">paratype</typeStatus>
(USNM 1557553). Spines initially numbering 12 per podia, increasing to 56 spines in notopodia and
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neuropodia in posterior 1012 setigers; with up to 1114 spines on a side; spines alternating with slender capillaries. Individual spines curved, tapering to pointed tip; notoacicular spines longer and thinner (
<figureCitation id="BB9CB5C9C460FFA84EBA1D80AE1121C0" box="[452,565,224,251]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="24" pageNumber="175">Figs. 15C</figureCitation>
,
<figureCitation id="BB9CB5C9C460FFA84D3F1D80AE5721C7" box="[577,627,225,252]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="24" pageNumber="175">16D</figureCitation>
) than neuropodial spines (
<figureCitation id="BB9CB5C9C460FFA84CE01D80A83521C0" box="[926,1041,224,251]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="24" pageNumber="175">Figs. 15D</figureCitation>
,
<figureCitation id="BB9CB5C9C460FFA84B631D80A86A21C7" box="[1053,1102,225,252]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="24" pageNumber="175">16E</figureCitation>
); core of spines clear, weakly striated.
</paragraph>
<paragraph id="2318A94CC460FFA84FB91C48A92B207F" blockId="24.[151,1437,152,611]" box="[199,1295,296,324]" pageId="24" pageNumber="175">
Posterior segments narrow, pygidium with rounded lobe ventral to anal opening (
<figureCitation id="BB9CB5C9C460FFA84B2A1C48A8E02078" box="[1108,1220,296,323]" captionStart="FIGURE 15" captionStartId="24.[151,250,1983,2008]" captionTargetBox="[171,1423,639,1948]" captionTargetId="figure@24.[151,1436,633,1964]" captionTargetPageId="24" captionText="FIGURE 15. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine." httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="24" pageNumber="175">Figs. 15B</figureCitation>
,
<figureCitation id="BB9CB5C9C460FFA84BAE1C48A926207F" box="[1232,1282,297,324]" captionStart="FIGURE 16" captionStartId="25.[151,250,1667,1692]" captionTargetBox="[163,1402,190,1641]" captionTargetId="figure@25.[151,1410,181,1647]" captionTargetPageId="25" captionText="FIGURE 16. Chaetozone grasslei n. sp. Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="24" pageNumber="175">16C</figureCitation>
).
</paragraph>
<paragraph id="2318A94CC460FFA84FB91C2CACC720B0" blockId="24.[151,1437,152,611]" pageId="24" pageNumber="175">
<emphasis id="11D3755EC460FFA84FB91C2CAD91205C" bold="true" box="[199,437,332,359]" pageId="24" pageNumber="175">Methyl Green stain</emphasis>
. A weak light-green peristomial band posterior to prostomium, otherwise stain not retained.
</paragraph>
</subSubSection>
<subSubSection id="6BBDFAC7C460FFA84FB91CF4AE602358" pageId="24" pageNumber="175" type="etymology">
<paragraph id="2318A94CC460FFA84FB91CF4AE602358" blockId="24.[151,1437,152,611]" pageId="24" pageNumber="175">
<emphasis id="11D3755EC460FFA84FB91CF4AD6C2094" bold="true" box="[199,328,405,431]" pageId="24" pageNumber="175">Etymology</emphasis>
. This species is named for the late Dr. J. Frederick Grassle, prominent deep-sea benthic ecologist of the Woods Hole Oceanographic Institution and the Rutgers University Institute of Marine and Coastal Sciences. Dr. Grassle introduced this author to offshore benthos, including how to collect specimens with care and analyze and interpret the results with the most relevant statistical methods. From his numerous deep-sea surveys, he also provided this author and other investigators with numerous polychaetes from vents and other habitats that have resulted in a rich harvest of species new to science.
</paragraph>
</subSubSection>
<caption id="77D8F9C4C460FFA84FE91ADEAF1426C7" httpUri="https://zenodo.org/record/3269019/files/figure.png" pageId="24" pageNumber="175" startId="24.[151,250,1983,2008]" targetBox="[171,1423,639,1948]" targetPageId="24">
<paragraph id="2318A94CC460FFA84FE91ADEAF1426C7" blockId="24.[151,1436,1983,2044]" pageId="24" pageNumber="175">
<emphasis id="11D3755EC460FFA84FE91ADEAE6426E3" bold="true" box="[151,576,1983,2008]" pageId="24" pageNumber="175">
FIGURE 15.
<taxonomicName id="E4A7D2CFC460FFA84E541ADEADD926E3" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[298,509,1983,2008]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="24" pageNumber="175" phylum="Annelida" rank="species" species="grasslei" status="sp. nov.">
<emphasis id="11D3755EC460FFA84E541ADEADD926E3" bold="true" box="[298,509,1983,2008]" italics="true" pageId="24" pageNumber="175">Chaetozone grasslei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AE0C825C460FFA84D7B1AA1AE6426E3" box="[517,576,1984,2008]" pageId="24" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype (USNM 1557550): A, anterior end, dorsal view; B, posterior end, dorsal view; C, notopodial acicular spine, D, neuropodial acicular spine.
</paragraph>
</caption>
<caption id="77D8F9C4C461FFA94FE91BE2ADF427DF" httpUri="https://zenodo.org/record/3269021/files/figure.png" pageId="25" pageNumber="176" startId="25.[151,250,1667,1692]" targetBox="[163,1402,190,1641]" targetPageId="25">
<paragraph id="2318A94CC461FFA94FE91BE2ADF427DF" blockId="25.[151,1436,1667,1764]" pageId="25" pageNumber="176">
<emphasis id="11D3755EC461FFA94FE91BE2AE6527A7" bold="true" box="[151,577,1667,1692]" pageId="25" pageNumber="176">
FIGURE 16.
<taxonomicName id="E4A7D2CFC461FFA94E541BE2ADD927A7" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[298,509,1667,1692]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="176" phylum="Annelida" rank="species" species="grasslei" status="sp. nov.">
<emphasis id="11D3755EC461FFA94E541BE2ADD927A7" bold="true" box="[298,509,1667,1692]" italics="true" pageId="25" pageNumber="176">Chaetozone grasslei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AE0C825C461FFA94D7B1BE5AE6527A7" box="[517,577,1668,1692]" pageId="25" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype (USNM 1557550): A, entire worm, mostly lateral view; B, anterior end, right lateral view; C, posterior end, dorsal view; D, notopodial acicular spine; E. neuropodial acicular spine (same scale as in D). Stained with Shirlastain A.
</paragraph>
</caption>
<subSubSection id="6BBDFAC7C461FFA94FB91A73A83026DA" pageId="25" pageNumber="176" type="distribution">
<paragraph id="2318A94CC461FFA94FB91A73A88926A2" blockId="25.[151,1437,1810,2017]" pageId="25" pageNumber="176">
<emphasis id="11D3755EC461FFA94FB91A73AD102617" bold="true" box="[199,308,1810,1836]" pageId="25" pageNumber="176">Remarks</emphasis>
.
<taxonomicName id="E4A7D2CFC461FFA94E3C1A73AE032617" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[322,551,1810,1836]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="176" phylum="Annelida" rank="species" species="grasslei" status="sp. nov.">
<emphasis id="11D3755EC461FFA94E3C1A73AE032617" box="[322,551,1810,1836]" italics="true" pageId="25" pageNumber="176">Chaetozone grasslei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AE0C825C461FFA94D501A73AE492616" box="[558,621,1810,1837]" pageId="25" pageNumber="176" rank="species">
<emphasis id="11D3755EC461FFA94D501A73AE422617" bold="true" box="[558,614,1810,1836]" pageId="25" pageNumber="176">n. sp</emphasis>
.
</taxonomicNameLabel>
is a long slender species with only a slight widening of the anterior segments. Middle and posterior segments are longer and somewhat rounded, sometimes weakly moniliform. The distension of some body segments is due to concentration of sediment in the intestine. These enlarged segments, when present, are conspicuous owing to the otherwise elongate slender appearance of these worms.
</paragraph>
<paragraph id="2318A94CC461FFA94FB91AC3A83026DA" blockId="25.[151,1437,1810,2017]" pageId="25" pageNumber="176">
There are no described species of
<taxonomicName id="E4A7D2CFC461FFA94D321AC3AEF42687" authorityName="Malmgren" authorityYear="1867" box="[588,720,1954,1980]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="176" phylum="Annelida" rank="genus">
<emphasis id="11D3755EC461FFA94D321AC3AEF42687" box="[588,720,1954,1980]" italics="true" pageId="25" pageNumber="176">Chaetozone</emphasis>
</taxonomicName>
that have the combination of a long, slender appearance to the body and two peristomial rings. See comments for
<taxonomicName id="E4A7D2CFC461FFA94DAD1AA6AF4C26DB" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[723,872,1990,2017]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="176" phylum="Annelida" rank="species" species="truebloodi" status="sp. nov.">
<emphasis id="11D3755EC461FFA94DAD1AA6AF4C26DB" box="[723,872,1990,2017]" italics="true" pageId="25" pageNumber="176">C. truebloodi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AE0C825C461FFA94C111AA7AF8A26DB" box="[879,942,1990,2016]" pageId="25" pageNumber="176" rank="species">
<emphasis id="11D3755EC461FFA94C111AA7AF8A26DB" bold="true" box="[879,942,1990,2016]" pageId="25" pageNumber="176">n. sp.</emphasis>
</taxonomicNameLabel>
(below).
</paragraph>
</subSubSection>
<subSubSection id="6BBDFAC7C462FFAA4FB91DF9AFA321C1" pageId="26" pageNumber="177" type="biology_ecology">
<paragraph id="2318A94CC462FFAA4FB91DF9AFA321C1" blockId="26.[151,1436,152,287]" pageId="26" pageNumber="177">
<emphasis id="11D3755EC462FFAA4FB91DF9AD852188" bold="true" box="[199,417,152,179]" pageId="26" pageNumber="177">Habitat &amp; biology</emphasis>
. The fine sediment in the gut of these worms suggests a silt-clay habitat. The worms were typically found in the upper
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of the sediment column: two specimens including the
<typeStatus id="FC1C17EEC462FFAA4B1C1DDDA8E021ED" box="[1122,1220,188,214]" pageId="26" pageNumber="177" type="holotype">holotype</typeStatus>
were in the
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level in the box core and three specimens were at the
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level.
</paragraph>
</subSubSection>
<subSubSection id="6BBDFAC7C462FFAA4FB91C65AF372024" box="[199,787,260,287]" pageId="26" pageNumber="177" type="distribution">
<paragraph id="2318A94CC462FFAA4FB91C65AF372024" blockId="26.[151,1436,152,287]" box="[199,787,260,287]" pageId="26" pageNumber="177">
<emphasis id="11D3755EC462FFAA4FB91C65AD7D2024" bold="true" box="[199,345,260,287]" pageId="26" pageNumber="177">Distribution</emphasis>
. Abyssal Pacific Ocean,
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.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>