610 lines
55 KiB
XML
610 lines
55 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5872993" ID-GBIF-Dataset="6bc8f93f-8227-4361-9bef-0976c7102fe7" ID-GBIF-Taxon="192411994" ID-ISSN="1639-4798" ID-Zenodo-Dep="5872993" checkinTime="1629118158663" checkinUser="carolina" docAuthor="Rivero-Guerra, Aixa O." docDate="2012" docId="A6458955FFDD662AD244FF46FEB975ED" docLanguage="en" docName="Adansonia.34.1.133-154.pdf" docOrigin="Adansonia (3) 34 (1)" docSource="http://dx.doi.org/10.5252/a2012n1a16" docStyle="DocumentStyle:B98EC8A323C0F63E1C55336AD58D9273.1:Adansonia.2011-2014.journal_article" docStyleId="B98EC8A323C0F63E1C55336AD58D9273" docStyleName="Adansonia.2011-2014.journal_article" docStyleVersion="1" docTitle="Santolina oblongifolia var. ceratophylla Willk." docType="treatment" docVersion="7" lastPageId="11" lastPageNumber="144" masterDocId="5A7CF12DFFD86621D142FF9EFFD0734E" masterDocTitle="Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss." masterLastPageNumber="154" masterPageNumber="133" pageId="5" pageNumber="138" updateTime="1642515947841" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rivero-Guerra, Aixa O.</mods:namePart>
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<mods:affiliation>Universidad de Sevilla, Facultad de Biología, Departamento de Biología Vegetal y Ecología, avda. Reina Mercedes, E- 41012 Sevilla (Spain) & Centro de Investigação en Biodiversidade e Recursos Genéticos, Campus Agrario de Vairão, rua Padre Armando Quintas-Crasto, 4485 - 661 Vairão (Portugal)</mods:affiliation>
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<mods:nameIdentifier type="email">rivero-guerra@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Adansonia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="series">
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<mods:title>3</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2012-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>133</mods:start>
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<mods:end>154</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.5252/a2012n1a16</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/a2012n1a16</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6bc8f93f-8227-4361-9bef-0976c7102fe7</mods:identifier>
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<mods:identifier type="ISSN">1639-4798</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5205801</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5872993" ID-GBIF-Taxon="192411994" ID-Zenodo-Dep="5872993" LSID="urn:lsid:plazi:treatment:A6458955FFDD662AD244FF46FEB975ED" httpUri="http://treatment.plazi.org/id/A6458955FFDD662AD244FF46FEB975ED" lastPageId="11" lastPageNumber="144" pageId="5" pageNumber="138">
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<subSubSection pageId="5" pageNumber="138" type="nomenclature">
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<paragraph blockId="5.[774,1113,215,274]" pageId="5" pageNumber="138">
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<heading centered="true" fontSize="11" level="2" pageId="5" pageNumber="138" reason="3">
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4.
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<taxonomicName authority="Willk." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="ceratophylla">
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<emphasis bold="true" box="[802,1042,215,242]" italics="true" pageId="5" pageNumber="138">Santolina oblongifolia</emphasis>
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Boiss. var. β
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<emphasis bold="true" box="[873,1004,247,274]" italics="true" pageId="5" pageNumber="138">ceratophylla</emphasis>
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Willk.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="138" type="reference_group">
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<paragraph blockId="5.[675,1210,304,619]" pageId="5" pageNumber="138">
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<treatmentCitationGroup pageId="5" pageNumber="138">
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<treatmentCitation author="Willk." journal="Prodromus Florae Hispanicae" page="82" pageId="5" pageNumber="138" score="0.029850423456044287" volume="2" year="1865">
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<collectorName box="[699,829,304,328]" pageId="5" pageNumber="138">In Willkomm</collectorName>
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&
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<collectorName box="[858,918,305,329]" pageId="5" pageNumber="138">Lange</collectorName>
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,
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<emphasis box="[925,1187,304,328]" italics="true" pageId="5" pageNumber="138">Prodromus Florae Hispanicae</emphasis>
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2: 82 (1865)
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</treatmentCitation>
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</treatmentCitationGroup>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="138" type="materials_examined">
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<paragraph blockId="5.[675,1210,304,619]" pageId="5" pageNumber="138">
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—
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<materialsCitation ID-GBIF-Occurrence="3446332307" collectingDate="1863-07-08" collectingDateMax="1863-07-25" collectingDateMin="1863-07-08" collectionCode="COI-WILLK" collectorName="Bourgeau" country="Spain" location="Sierra de Gredos au Puerto del Pico" pageId="5" pageNumber="138" specimenCode="COI-WILLK 00035989" specimenCount="1" typeStatus="lectotype">
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Type (as given in protologue): “In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v.s.)”. —
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<typeStatus box="[1113,1209,437,461]" pageId="5" pageNumber="138">Lectotype</typeStatus>
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(here designated):
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<collectingCountry box="[859,916,463,487]" name="Spain" pageId="5" pageNumber="138">Spain</collectingCountry>
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, Rochers de la
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<location LSID="urn:lsid:plazi:treatment:A6458955FFDD662AD244FF46FEB975ED:2B336E98FFDD6624D56CFE51FC51714C" country="Spain" name="Sierra de Gredos au Puerto del Pico" pageId="5" pageNumber="138">Sierra de Gredos au Puerto del Pico</location>
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,
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<date box="[908,1053,489,513]" pageId="5" pageNumber="138" value="1863-07-08" valueMax="1863-07-25" valueMin="1863-07-08">
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<collectingDate box="[908,1053,489,513]" pageId="5" pageNumber="138" value="1863-07-08" valueMax="1863-07-25" valueMin="1863-07-08">8-25.VII.1863</collectingDate>
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</date>
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,
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<emphasis box="[1064,1210,490,514]" italics="true" pageId="5" pageNumber="138">
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<collectorName box="[1064,1152,490,513]" pageId="5" pageNumber="138">Bourgeau</collectorName>
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2540
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</emphasis>
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(
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<specimenCode box="[683,916,516,540]" collectionCode="COI-WILLK" pageId="5" pageNumber="138">COI-WILLK 00035989</specimenCode>
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[
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<figureCitation box="[927,983,516,540]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@6.[160,1163,215,1629]" captionTargetPageId="6" captionText="FIG 3. — Lectotype of Santolina oblongifolia var. α obtusifolia Willk. (COI-WILLK 00035955) and lectotype of S. oblongifolia var. β ceratophylla Willk. (COI-WILLK 00035989)." figureDoi="http://doi.org/10.5281/zenodo.5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="5" pageNumber="138">Fig. 3</figureCitation>
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]
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3446332312" pageId="5" pageNumber="138" specimenCount="1">isolecto-, A 00251664 (specimen to the left), HEID 702282 (specimen to the left), MPU (2 sheets), W (2 sheets, one of them with two specimen; the isolectotype is the specimen to the left]).</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="144" pageId="5" pageNumber="138" type="discussion">
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<paragraph blockId="5.[674,1212,660,1700]" box="[677,774,660,686]" pageId="5" pageNumber="138">REMARKS</paragraph>
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<paragraph blockId="5.[674,1212,660,1700]" pageId="5" pageNumber="138">
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These names were published by Willkomm
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<emphasis box="[1187,1210,692,717]" italics="true" pageId="5" pageNumber="138">in</emphasis>
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<bibRefCitation author="Willkomm & Lange" box="[679,994,722,749]" firstAuthor="Willkomm" pageId="5" pageNumber="138" refId="ref10035" refString="WILLKOMM M. & LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm & Lange (1865)</bibRefCitation>
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as
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="oblongifolia">
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<emphasis italics="true" pageId="5" pageNumber="138">S. oblongifolia α obtusifolia</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[844,1152,754,783]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="oblongifolia">
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<emphasis box="[844,1152,754,783]" italics="true" pageId="5" pageNumber="138">S. oblongifolia β ceratophylla</emphasis>
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</taxonomicName>
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. The author did not mentioned the rank of “
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<emphasis box="[1099,1112,788,815]" italics="true" pageId="5" pageNumber="138">α</emphasis>
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” and “
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<emphasis box="[1187,1200,788,815]" italics="true" pageId="5" pageNumber="138">β</emphasis>
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” in the introductory chapter of
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<emphasis italics="true" pageId="5" pageNumber="138">Prodromus Florae Hispanicae</emphasis>
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. However, Willkomm (
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<bibRefCitation author="Willkomm & Lange" firstAuthor="Willkomm" page="81" pageId="5" pageNumber="138" refId="ref10035" refString="WILLKOMM M. & LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm & Lange 1865: 81</bibRefCitation>
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) cited “
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<emphasis italics="true" pageId="5" pageNumber="138">
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<taxonomicName box="[925,1074,881,907]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="variety" variety="vulgaris">S. var. vulgaris</taxonomicName>
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="rosmarinifoliae">S. rosmarinifoliae</taxonomicName>
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</emphasis>
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”, indicating the rank of
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<taxonomicName box="[1005,1210,913,939]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="rosmarinifolia">
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<emphasis box="[1005,1179,913,939]" italics="true" pageId="5" pageNumber="138">S. rosmarinifolia</emphasis>
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L.
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</taxonomicName>
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<emphasis box="[677,777,944,973]" italics="true" pageId="5" pageNumber="138">α vulgaris</emphasis>
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Boiss.; therefore the rank of “
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<emphasis box="[1096,1109,946,973]" italics="true" pageId="5" pageNumber="138">α</emphasis>
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” and “
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<emphasis box="[1185,1198,946,973]" italics="true" pageId="5" pageNumber="138">β</emphasis>
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” can clearly be established as variety. In addition, Lange
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<emphasis box="[752,774,1009,1034]" italics="true" pageId="5" pageNumber="138">in</emphasis>
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<bibRefCitation author="Willkomm & Lange" box="[783,1098,1008,1034]" firstAuthor="Willkomm" pageId="5" pageNumber="138" refId="ref10063" refString="WILLKOMM M. & LANGE J. 1880. - Prodromus Florae Hispanicae III. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 1144 p." type="book" year="1880">Willkomm & Lange (1880)</bibRefCitation>
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, cited the varietal status of both names.
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</paragraph>
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<paragraph blockId="5.[674,1212,660,1700]" pageId="5" pageNumber="138">
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The varieties “
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<emphasis box="[853,960,1071,1097]" italics="true" pageId="5" pageNumber="138">obtusifolia</emphasis>
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” and “
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<emphasis box="[1038,1160,1071,1097]" italics="true" pageId="5" pageNumber="138">ceratophylla</emphasis>
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” are from the same
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<emphasis box="[831,966,1103,1129]" italics="true" pageId="5" pageNumber="138">locus classicus</emphasis>
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: “
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<emphasis italics="true" pageId="5" pageNumber="138">In rupestribus regionis montanae Hispanicae centralis, hucusque non nisi in Sierra de Gredos reperta (ad Serranillos, Isern, Cut.!. Puerto del Pico, Bourg.!) ubi utraque forma promiscue crescere videtur. – Julio (v.s.)</emphasis>
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” (Willkomm
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<emphasis box="[675,697,1262,1287]" italics="true" pageId="5" pageNumber="138">in</emphasis>
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<bibRefCitation author="Willkomm & Lange" box="[706,1011,1261,1288]" firstAuthor="Willkomm" pageId="5" pageNumber="138" refId="ref10035" refString="WILLKOMM M. & LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm & Lange 1865</bibRefCitation>
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). Willkomm also mentioned “
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<emphasis box="[811,917,1293,1319]" italics="true" pageId="5" pageNumber="138">Serranillos</emphasis>
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” in the protologue of
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<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="heterophylla">
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<emphasis box="[1147,1158,1293,1319]" italics="true" pageId="5" pageNumber="138">S</emphasis>
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.
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<emphasis italics="true" pageId="5" pageNumber="138">heterophylla</emphasis>
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</taxonomicName>
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. This indicates that Willkomm granted a varietal status to the intrapopulational variation.
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</paragraph>
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<paragraph blockId="5.[674,1212,660,1700]" lastBlockId="7.[111,649,216,1699]" lastPageId="7" lastPageNumber="140" pageId="5" pageNumber="138">
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The specimens COI-WILLK 00035955 and COI-WILLK 00035989 of Willkomm’s herbarium are on the same sheet (
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<figureCitation box="[840,905,1451,1477]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@6.[160,1163,215,1629]" captionTargetPageId="6" captionText="FIG 3. — Lectotype of Santolina oblongifolia var. α obtusifolia Willk. (COI-WILLK 00035955) and lectotype of S. oblongifolia var. β ceratophylla Willk. (COI-WILLK 00035989)." figureDoi="http://doi.org/10.5281/zenodo.5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="5" pageNumber="138">Fig. 3</figureCitation>
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). This sheet has two labels in the lower left-hand margin with the following notation in black ink:
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<emphasis bold="true" box="[863,942,1514,1541]" pageId="5" pageNumber="138">Label 1</emphasis>
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(Bourgeau’s handwritten notation): “E. Bourgeau
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<emphasis box="[946,961,1548,1575]" italics="true" pageId="5" pageNumber="138">–</emphasis>
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Pl. d’Espagne 1863 // 2540.
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<taxonomicName authority="Boiss. Diagn." authorityName="Boiss. Diagn." box="[745,1119,1578,1604]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="oblongifolia">
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<emphasis box="[745,971,1578,1604]" italics="true" pageId="5" pageNumber="138">Santolina oblongifolia</emphasis>
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Boiss. Diagn.
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</taxonomicName>
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/ Pl. Or. Ser.2, fasc. 3, p. 18 (Planta herbaria Paroniani, cujus patria nodium involucrate) // (J. Gay) // Rochers de la Sierra de Gredos au Puerto del Pico // 8 et 25 juill”;
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<emphasis bold="true" box="[214,295,216,242]" pageId="7" pageNumber="140">Label 2</emphasis>
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(Willkomm’s handwritten notation): “
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<taxonomicName authority="Boiss." authorityName="Boiss." box="[190,484,247,273]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
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<emphasis box="[190,415,247,273]" italics="true" pageId="7" pageNumber="140">Santolina oblongifolia</emphasis>
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Boiss.
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</taxonomicName>
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[underlined] // 1. var.
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<emphasis box="[180,306,279,308]" italics="true" pageId="7" pageNumber="140">α obtusifolia</emphasis>
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/ 2. var.
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<emphasis box="[395,534,279,308]" italics="true" pageId="7" pageNumber="140">β ceratophylla</emphasis>
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”. Voucher specimens from the
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<emphasis box="[345,491,311,337]" italics="true" pageId="7" pageNumber="140">locus classicus</emphasis>
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and with the number “
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<emphasis box="[222,279,343,369]" italics="true" pageId="7" pageNumber="140">2540</emphasis>
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” were found in A 00251664, G, HEID 702282, LY (2 sheets),
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<accessionNumber box="[440,576,374,401]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MA720935" pageId="7" pageNumber="140">MA 720935</accessionNumber>
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, MPU (3 sheets), P (3 sheets) and W (3 sheets), but they do not include a revision label of Willkomm with the epithets “
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<emphasis box="[257,364,469,495]" italics="true" pageId="7" pageNumber="140">obtusifolia</emphasis>
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” and “
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<emphasis box="[441,563,469,495]" italics="true" pageId="7" pageNumber="140">ceratophylla</emphasis>
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”.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5205807" ID-Zenodo-Dep="5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="6" pageNumber="139" startId="6.[113,124,1657,1674]" targetBox="[160,1163,215,1629]" targetPageId="6">
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<paragraph blockId="6.[113,1209,1657,1697]" pageId="6" pageNumber="139">
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FIG 3. — Lectotype of
|
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<taxonomicName box="[305,485,1657,1674]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="139" phylum="Tracheophyta" rank="species" species="oblongifolia">
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<emphasis box="[305,485,1657,1674]" italics="true" pageId="6" pageNumber="139">Santolina oblongifolia</emphasis>
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</taxonomicName>
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var. α
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<emphasis box="[546,632,1657,1674]" italics="true" pageId="6" pageNumber="139">obtusifolia</emphasis>
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Willk. (COI-WILLK 00035955) and lectotype of
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<taxonomicName box="[1035,1155,1657,1674]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="139" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[1035,1155,1657,1674]" italics="true" pageId="6" pageNumber="139">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
var. β
|
||
<emphasis box="[113,214,1680,1697]" italics="true" pageId="6" pageNumber="139">ceratophylla</emphasis>
|
||
Willk. (COI-WILLK 00035989).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[111,649,216,1699]" pageId="7" pageNumber="140">
|
||
The collection to the left, COI-WILLK 00035955, comprises two samples from the same gathering, labelled with the name “
|
||
<taxonomicName box="[366,587,564,590]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[366,587,564,590]" italics="true" pageId="7" pageNumber="140">Santolina oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
var.
|
||
<emphasis italics="true" pageId="7" pageNumber="140">α obtusifolia</emphasis>
|
||
” (
|
||
<typeStatus box="[242,340,596,622]" pageId="7" pageNumber="140">lectotype</typeStatus>
|
||
here designated;
|
||
<figureCitation box="[519,580,596,622]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@6.[160,1163,215,1629]" captionTargetPageId="6" captionText="FIG 3. — Lectotype of Santolina oblongifolia var. α obtusifolia Willk. (COI-WILLK 00035955) and lectotype of S. oblongifolia var. β ceratophylla Willk. (COI-WILLK 00035989)." figureDoi="http://doi.org/10.5281/zenodo.5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="7" pageNumber="140">Fig. 3</figureCitation>
|
||
). One of them (sample to the right) has three flowering stems and a small sterile stem, and the other (sample to the left) is a robust sample with four flowering stems and several vegetative stems. The
|
||
<typeStatus box="[514,608,723,749]" pageId="7" pageNumber="140">lectotype</typeStatus>
|
||
has two revision labels by Sánchez-Mata and Sardinero with the following notation in black ink:
|
||
<emphasis bold="true" box="[556,637,786,812]" pageId="7" pageNumber="140">Label 1</emphasis>
|
||
: “
|
||
<taxonomicName authority="Willk." authorityName="Willk." box="[125,647,818,844]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="obtusifolia">
|
||
<emphasis box="[125,351,818,844]" italics="true" pageId="7" pageNumber="140">Santolina oblongifolia</emphasis>
|
||
Boiss. var.
|
||
<emphasis box="[466,573,818,844]" italics="true" pageId="7" pageNumber="140">obtusifolia</emphasis>
|
||
Willk.
|
||
</taxonomicName>
|
||
<emphasis box="[113,135,850,875]" italics="true" pageId="7" pageNumber="140">in</emphasis>
|
||
Willk. & Lange //
|
||
<typeStatus box="[353,459,850,876]" pageId="7" pageNumber="140">Lectotype</typeStatus>
|
||
[handwritten] //
|
||
<collectingRegion box="[115,212,881,907]" country="Portugal" name="Coimbra" pageId="7" pageNumber="140">Coimbra</collectingRegion>
|
||
[printed]
|
||
<date box="[320,408,881,908]" pageId="7" pageNumber="140" value="1993-09">1993 IX</date>
|
||
[handwritten]”;
|
||
<emphasis bold="true" pageId="7" pageNumber="140">Label 2</emphasis>
|
||
: “
|
||
<taxonomicName authority="Boiss." authorityName="Boiss." box="[155,459,913,939]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[155,386,913,939]" italics="true" pageId="7" pageNumber="140">Santolina oblongifolia</emphasis>
|
||
Boiss.
|
||
</taxonomicName>
|
||
[handwritten] //
|
||
<collectingRegion box="[115,212,944,970]" country="Portugal" name="Coimbra" pageId="7" pageNumber="140">Coimbra</collectingRegion>
|
||
[printed]
|
||
<date box="[323,412,945,971]" pageId="7" pageNumber="140" value="1993-09">1993 IX</date>
|
||
[handwritten]”. Both samples have spatulate leaves, but the flowering stems are strongly thickened above and the capitula are big (
|
||
<figureCitation box="[201,267,1039,1066]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@6.[160,1163,215,1629]" captionTargetPageId="6" captionText="FIG 3. — Lectotype of Santolina oblongifolia var. α obtusifolia Willk. (COI-WILLK 00035955) and lectotype of S. oblongifolia var. β ceratophylla Willk. (COI-WILLK 00035989)." figureDoi="http://doi.org/10.5281/zenodo.5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="7" pageNumber="140">Fig. 3</figureCitation>
|
||
). The literature does not show the publication of this
|
||
<typeStatus box="[323,421,1071,1097]" pageId="7" pageNumber="140">lectotype</typeStatus>
|
||
by Sánchez-Mata & Sardinero. There is another specimen with the name “
|
||
<taxonomicName box="[194,605,1134,1164]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[194,424,1134,1160]" italics="true" pageId="7" pageNumber="140">Santolina oblongifolia</emphasis>
|
||
var.
|
||
<emphasis box="[478,605,1134,1164]" italics="true" pageId="7" pageNumber="140">α obtusifolia</emphasis>
|
||
</taxonomicName>
|
||
” in W herbarium. The sheet (W 0026403;
|
||
<figureCitation box="[535,597,1166,1192]" captionStart="FIG" captionStartId="8.[113,124,1644,1661]" captionTargetBox="[161,1162,216,1630]" captionTargetId="figure-26@8.[161,1162,216,1630]" captionTargetPageId="8" captionText="FIG 4. — Specimen labelled by Willkomm as Santolina oblongifolia var. obtusifolia Willk. The sheet has a different collection date for the type material (Sierra de Gredos, Leg. Bourgeau 3.VII.1863, Cour. Willkomm), and the number “2540” is not shown." figureDoi="http://doi.org/10.5281/zenodo.5205809" httpUri="https://zenodo.org/record/5205809/files/figure.png" pageId="7" pageNumber="140">Fig. 4</figureCitation>
|
||
) has a different collection date (Sierra de Gredos, Leg. Bourgeau
|
||
<date box="[223,347,1229,1255]" pageId="7" pageNumber="140" value="1863-07-03">3.VII.1863</date>
|
||
, Cour. Willkomm), and the number “2540” is not shown. This specimen does not show intermediate characteristics between
|
||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis italics="true" pageId="7" pageNumber="140">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="subsp. rosmarinifolia" box="[256,640,1325,1351]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="subSpecies" species="rosmarinifolia" subSpecies="rosmarinifolia">
|
||
<emphasis box="[256,423,1325,1351]" italics="true" pageId="7" pageNumber="140">S. rosmarinifolia</emphasis>
|
||
subsp.
|
||
<emphasis box="[498,640,1325,1351]" italics="true" pageId="7" pageNumber="140">rosmarinifolia</emphasis>
|
||
</taxonomicName>
|
||
. All the characteristics fit with those listed in the protologue (Appendix 1).
|
||
</paragraph>
|
||
<paragraph blockId="7.[111,649,216,1699]" lastBlockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
<bibRefCitation author="Jordan & Fourreau" box="[136,410,1420,1447]" firstAuthor="Jordan" pageId="7" pageNumber="140" refId="ref9516" refString="JORDAN A. & FOURREAU J. 1869. - Icones ad Floram Europae novo fundamento instaurandam spectantes. II. Apud Savy, Bibliopolam, Paris, 25 p., tabs 201 - 280." type="book" year="1869">
|
||
<collectingCountry box="[136,206,1420,1446]" name="Jordan" pageId="7" pageNumber="140">Jordan</collectingCountry>
|
||
& Fourreau (1869)
|
||
</bibRefCitation>
|
||
published a new taxon (
|
||
<taxonomicName box="[120,214,1452,1478]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[120,214,1452,1478]" italics="true" pageId="7" pageNumber="140">S. sericea</emphasis>
|
||
</taxonomicName>
|
||
) based on plants from the
|
||
<emphasis box="[508,646,1451,1477]" italics="true" pageId="7" pageNumber="140">locus classicus</emphasis>
|
||
of the varieties
|
||
<emphasis box="[282,389,1483,1509]" italics="true" pageId="7" pageNumber="140">obtusifolia</emphasis>
|
||
and
|
||
<emphasis box="[446,568,1483,1509]" italics="true" pageId="7" pageNumber="140">ceratophylla</emphasis>
|
||
. However, they did not cite it in the protologue. They also published a detailed diagnosis of
|
||
<taxonomicName box="[549,648,1547,1573]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[549,648,1547,1573]" italics="true" pageId="7" pageNumber="140">S. sericea</emphasis>
|
||
</taxonomicName>
|
||
(Appendix 1). Furthermore, figure 321, table 240 (
|
||
<bibRefCitation author="Jordan & Fourreau" box="[121,392,1610,1637]" firstAuthor="Jordan" pageId="7" pageNumber="140" refId="ref9516" refString="JORDAN A. & FOURREAU J. 1869. - Icones ad Floram Europae novo fundamento instaurandam spectantes. II. Apud Savy, Bibliopolam, Paris, 25 p., tabs 201 - 280." type="book" year="1869">
|
||
<collectingCountry box="[121,195,1610,1636]" name="Jordan" pageId="7" pageNumber="140">Jordan</collectingCountry>
|
||
& Fourreau 1869
|
||
</bibRefCitation>
|
||
) shows a robust sample of
|
||
<taxonomicName box="[144,239,1642,1668]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[144,157,1642,1668]" italics="true" pageId="7" pageNumber="140">S</emphasis>
|
||
.
|
||
<emphasis box="[173,239,1642,1667]" italics="true" pageId="7" pageNumber="140">sericea</emphasis>
|
||
</taxonomicName>
|
||
. Two sheets are preserved in the Jordan’s herbarium (LY) with the epithets “
|
||
<taxonomicName box="[561,628,1674,1699]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[561,628,1674,1699]" italics="true" pageId="7" pageNumber="140">sericea</emphasis>
|
||
</taxonomicName>
|
||
”. The records indicate that the sample
|
||
<collectingCountry box="[1081,1153,216,242]" name="Jordan" pageId="7" pageNumber="140">Jordan</collectingCountry>
|
||
used to illustrate this name was split into two units and mounted on two sheets. They were labelled as “
|
||
<emphasis italics="true" pageId="7" pageNumber="140">
|
||
1
|
||
<superScript attach="none" box="[1190,1210,281,297]" fontSize="7" pageId="7" pageNumber="140">ère</superScript>
|
||
feuille
|
||
</emphasis>
|
||
” (
|
||
<figureCitation box="[764,826,311,338]" captionStart="FIG" captionStartId="9.[274,285,1680,1697]" captionTargetBox="[218,1104,215,1629]" captionTargetId="figure-17@9.[218,1104,215,1629]" captionTargetPageId="9" captionText="FIG 5. — Lectotype of Santolina sericea Jord. & Fourr. (LY, “1ère feuille”, specimen to the left)." figureDoi="http://doi.org/10.5281/zenodo.5205811" httpUri="https://zenodo.org/record/5205811/files/figure.png" pageId="7" pageNumber="140">Fig. 5</figureCitation>
|
||
) and “
|
||
<emphasis box="[899,1008,311,337]" italics="true" pageId="7" pageNumber="140">
|
||
2
|
||
<superScript attach="left" box="[913,941,313,329]" fontSize="7" pageId="7" pageNumber="140">ème</superScript>
|
||
feuille
|
||
</emphasis>
|
||
” (
|
||
<figureCitation box="[1034,1098,311,337]" captionStart="FIG" captionStartId="10.[257,268,1680,1697]" captionTargetBox="[222,1101,215,1629]" captionTargetId="figure-25@10.[222,1101,215,1629]" captionTargetPageId="10" captionText="FIG 6. — Isolectotype of Santolina sericea Jord. & Fourr. (LY, “2ème feuille”, specimen to the right)." figureDoi="http://doi.org/10.5281/zenodo.5205813" httpUri="https://zenodo.org/record/5205813/files/figure.png" pageId="7" pageNumber="140">Fig. 6</figureCitation>
|
||
, specimen to the right).
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
The sheet designated “
|
||
<emphasis box="[933,1033,374,400]" italics="true" pageId="7" pageNumber="140">
|
||
1
|
||
<superScript attach="left" box="[947,967,376,392]" fontSize="7" pageId="7" pageNumber="140">ère</superScript>
|
||
feuille
|
||
</emphasis>
|
||
” (Jordan’s handwritten notation in graphite in the lower left margin) has a sample, in the centre of the sheet, with three flowering stems, two fragments of flowering stems and three sterile stems. There is also a large, isolated sterile stem to the right of the sample (
|
||
<figureCitation box="[1083,1145,532,559]" captionStart="FIG" captionStartId="9.[274,285,1680,1697]" captionTargetBox="[218,1104,215,1629]" captionTargetId="figure-17@9.[218,1104,215,1629]" captionTargetPageId="9" captionText="FIG 5. — Lectotype of Santolina sericea Jord. & Fourr. (LY, “1ère feuille”, specimen to the left)." figureDoi="http://doi.org/10.5281/zenodo.5205811" httpUri="https://zenodo.org/record/5205811/files/figure.png" pageId="7" pageNumber="140">Fig. 5</figureCitation>
|
||
) [this sample is the portion to the right-hand of the original specimen].
|
||
<collectionCode box="[839,856,596,622]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="140" type="Herbarium">A</collectionCode>
|
||
label in the central lower margin indicates: “
|
||
<taxonomicName box="[793,963,628,654]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[793,963,628,654]" italics="true" pageId="7" pageNumber="140">Santolina sericea</emphasis>
|
||
</taxonomicName>
|
||
[underlined] // Jord. et Fourr. //
|
||
<typeStatus box="[776,894,660,686]" pageId="7" pageNumber="140">Lectotypus</typeStatus>
|
||
[underlined] //
|
||
<collectorName box="[1074,1181,659,686]" pageId="7" pageNumber="140">D. Rivera</collectorName>
|
||
//
|
||
<date box="[677,806,691,717]" pageId="7" pageNumber="140" value="1984-10-18">
|
||
<collectingDate box="[677,806,691,717]" pageId="7" pageNumber="140" value="1984-10-18">18/10/1984</collectingDate>
|
||
</date>
|
||
[
|
||
<collectorName box="[823,941,691,718]" pageId="7" pageNumber="140">D. Rivera’s</collectorName>
|
||
handwritten notation]”. This
|
||
<typeStatus box="[730,828,723,749]" pageId="7" pageNumber="140">lectotype</typeStatus>
|
||
was not published by
|
||
<collectorName box="[1069,1178,723,749]" pageId="7" pageNumber="140">D. Rivera</collectorName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
The sheet designated “
|
||
<emphasis box="[930,1035,754,781]" italics="true" pageId="7" pageNumber="140">
|
||
2
|
||
<superScript attach="left" box="[943,970,756,772]" fontSize="7" pageId="7" pageNumber="140">ème</superScript>
|
||
feuille
|
||
</emphasis>
|
||
” (Jordan’s handwritten notation in the central lower margin in graphite) has two samples (
|
||
<figureCitation box="[972,1038,818,845]" captionStart="FIG" captionStartId="10.[257,268,1680,1697]" captionTargetBox="[222,1101,215,1629]" captionTargetId="figure-25@10.[222,1101,215,1629]" captionTargetPageId="10" captionText="FIG 6. — Isolectotype of Santolina sericea Jord. & Fourr. (LY, “2ème feuille”, specimen to the right)." figureDoi="http://doi.org/10.5281/zenodo.5205813" httpUri="https://zenodo.org/record/5205813/files/figure.png" pageId="7" pageNumber="140">Fig. 6</figureCitation>
|
||
). The sample to the right is the portion to the left-hand-side of the original material. It has a decumbent branch with eight flowering stems. The sample to the left (representing a different gathering) has a small branch with three flowering stems. This sample shows clear intermediate characteristics (linear and narrowly spatulate leaves, peduncles strongly thickened above and umbilicate capitula) between
|
||
<taxonomicName box="[1021,1165,1071,1098]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[1021,1034,1072,1098]" italics="true" pageId="7" pageNumber="140">S</emphasis>
|
||
.
|
||
<emphasis box="[1048,1165,1071,1097]" italics="true" pageId="7" pageNumber="140">oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="subsp. rosmarinifolia" box="[675,1075,1103,1129]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="subSpecies" species="rosmarinifolia" subSpecies="rosmarinifolia">
|
||
<emphasis box="[675,688,1103,1129]" italics="true" pageId="7" pageNumber="140">S</emphasis>
|
||
.
|
||
<emphasis box="[702,848,1103,1129]" italics="true" pageId="7" pageNumber="140">rosmarinifolia</emphasis>
|
||
subsp.
|
||
<emphasis box="[929,1075,1103,1129]" italics="true" pageId="7" pageNumber="140">rosmarinifolia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
The
|
||
<typeStatus box="[745,842,1134,1160]" pageId="7" pageNumber="140">lectotype</typeStatus>
|
||
is the specimen from the sheet designated “1ère feuille” (
|
||
<figureCitation box="[918,981,1166,1193]" captionStart="FIG" captionStartId="9.[274,285,1680,1697]" captionTargetBox="[218,1104,215,1629]" captionTargetId="figure-17@9.[218,1104,215,1629]" captionTargetPageId="9" captionText="FIG 5. — Lectotype of Santolina sericea Jord. & Fourr. (LY, “1ère feuille”, specimen to the left)." figureDoi="http://doi.org/10.5281/zenodo.5205811" httpUri="https://zenodo.org/record/5205811/files/figure.png" pageId="7" pageNumber="140">Fig. 5</figureCitation>
|
||
[sample to the left]). It has three flowering stems and three other short vegetative stems. Furthermore,
|
||
<bibRefCitation author="Nyman" box="[998,1155,1230,1256]" firstAuthor="Nyman" pageId="7" pageNumber="140" refId="ref9712" refString="NYMAN C. F. 1879. - Conspectus florae Europaeae. Suecia typis Officinae Bohlinianae, Orebro, 856 p." type="book" year="1879">Nyman (1879)</bibRefCitation>
|
||
cited the present Bourgeau’s collection as
|
||
<taxonomicName box="[1056,1202,1261,1288]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[1056,1202,1261,1288]" italics="true" pageId="7" pageNumber="140">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
Willkomm
|
||
<emphasis box="[824,846,1294,1319]" italics="true" pageId="7" pageNumber="140">in</emphasis>
|
||
Willkomm & Lage (1865), in the protologue of the
|
||
<taxonomicName authorityName="Willk. (Willkomm & Lange" authorityYear="1865" box="[866,1077,1325,1351]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="ceratophylla">variety ceratophylla</taxonomicName>
|
||
, cited
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis italics="true" pageId="7" pageNumber="140">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of this name.
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" pageId="7" pageNumber="140">
|
||
There are two possibilities regarding
|
||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[1069,1209,1388,1414]" italics="true" pageId="7" pageNumber="140">S. oblongifolia</emphasis>
|
||
var.
|
||
<emphasis box="[721,864,1420,1449]" italics="true" pageId="7" pageNumber="140">β ceratophylla</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
<paragraph blockId="7.[674,1212,216,1700]" lastBlockId="11.[111,648,216,1700]" lastPageId="11" lastPageNumber="144" pageId="7" pageNumber="140">
|
||
– Option 1: Willkomm created a new name, at varietal rank for
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[855,1007,1483,1509]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[855,1007,1483,1509]" italics="true" pageId="7" pageNumber="140">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
(even if he did not use the same final epithet). This theory is supported by the following evidences: 1) Willkomm & Cutanda in
|
||
<bibRefCitation author="Willkomm" box="[805,1010,1578,1605]" firstAuthor="Willkomm" pageId="7" pageNumber="140" pagination="106 - 107" refId="ref9993" refString="WILLKOMM M. 1859. - Pugillus plantarum novarum peninsulae pyrenaicae. Linnaea 30: 106 - 107." type="journal article" year="1859">Willkomm (1859)</bibRefCitation>
|
||
in the protologue of
|
||
<taxonomicName box="[708,860,1610,1636]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="heteroplylla">
|
||
<emphasis box="[708,860,1610,1636]" italics="true" pageId="7" pageNumber="140">S. heteroplylla</emphasis>
|
||
</taxonomicName>
|
||
wrote “An varietas S. oblongifoliae?”, indicating that the authors had doubts about the species status of
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[963,1116,1673,1699]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="140" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[963,1116,1673,1699]" italics="true" pageId="7" pageNumber="140">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
; 2) both names have the same
|
||
<emphasis box="[349,493,216,242]" italics="true" pageId="11" pageNumber="144">locus classicus</emphasis>
|
||
; 3) the single specimen cited in the protologue of
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
(COI-WILLK 0035956) bears the both names; 4)
|
||
<bibRefCitation author="Willkomm" box="[225,427,311,338]" firstAuthor="Willkomm" pageId="11" pageNumber="144" refId="ref10035" refString="WILLKOMM M. & LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm (1865)</bibRefCitation>
|
||
cited
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[492,646,311,337]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[492,646,311,337]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
as synonym of the variety “
|
||
<emphasis box="[422,549,342,368]" italics="true" pageId="11" pageNumber="144">ceratophylla</emphasis>
|
||
”. If this is the case, the two names (
|
||
<taxonomicName box="[438,594,374,401]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heteroplylla">
|
||
<emphasis box="[438,594,374,401]" italics="true" pageId="11" pageNumber="144">S. heteroplylla</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[113,269,406,432]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[113,269,406,432]" italics="true" pageId="11" pageNumber="144">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
var.
|
||
<emphasis box="[324,472,406,435]" italics="true" pageId="11" pageNumber="144">β ceratophylla</emphasis>
|
||
) have the same
|
||
<typeStatus box="[114,218,437,463]" pageId="11" pageNumber="144">lectotype</typeStatus>
|
||
(COI-WILLK 0035956, here designated; see ICBN: art. 7.3 and 7.4;
|
||
<bibRefCitation author="McNeill" etAl="et al." firstAuthor="McNeill" pageId="11" pageNumber="144" refId="ref9601" refString="MCNEILL J., BARRIE F. R., BURDET H. M., DE- MOULIN V., HAWKSWORTH D. L., MARHOLD K., NICOLSON D. H., PRADO J., SILVA P. C., SKOG J. E., WIERSEMA J. H. & TURLAND N. J. 2006. - International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress Vienna. Austria, July 2005. A. R." type="proceedings" year="2006">
|
||
McNeill
|
||
<emphasis box="[594,647,469,495]" italics="true" pageId="11" pageNumber="144">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
), therefore the both names are homotypic synonyms.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5205809" ID-Zenodo-Dep="5205809" httpUri="https://zenodo.org/record/5205809/files/figure.png" pageId="8" pageNumber="141" startId="8.[113,124,1644,1661]" targetBox="[161,1162,216,1630]" targetPageId="8">
|
||
<paragraph blockId="8.[113,1209,1644,1684]" pageId="8" pageNumber="141">
|
||
FIG 4. — Specimen labelled by Willkomm as
|
||
<taxonomicName authority="Willk. The" authorityName="Willk. The" box="[487,878,1644,1661]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="141" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="obtusifolia">
|
||
<emphasis box="[487,665,1644,1661]" italics="true" pageId="8" pageNumber="141">Santolina oblongifolia</emphasis>
|
||
var.
|
||
<emphasis box="[706,791,1644,1661]" italics="true" pageId="8" pageNumber="141">obtusifolia</emphasis>
|
||
Willk. The
|
||
</taxonomicName>
|
||
sheet has a different collection date for the type material (Sierra de Gredos, Leg. Bourgeau 3.VII.1863, Cour. Willkomm), and the number “2540” is not shown.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5205811" ID-Zenodo-Dep="5205811" box="[274,1051,1680,1697]" httpUri="https://zenodo.org/record/5205811/files/figure.png" pageId="9" pageNumber="142" startId="9.[274,285,1680,1697]" targetBox="[218,1104,215,1629]" targetPageId="9">
|
||
<paragraph blockId="9.[274,1051,1680,1697]" box="[274,1051,1680,1697]" pageId="9" pageNumber="142">
|
||
FIG 5. — Lectotype of
|
||
<taxonomicName authority="Jord. & Fourr." authorityName="Jord. & Fourr." box="[458,719,1680,1697]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="142" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[458,600,1680,1697]" italics="true" pageId="9" pageNumber="142">Santolina sericea</emphasis>
|
||
Jord. & Fourr.
|
||
</taxonomicName>
|
||
(LY, “1ère feuille”, specimen to the left).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5205813" ID-Zenodo-Dep="5205813" box="[257,1067,1680,1697]" httpUri="https://zenodo.org/record/5205813/files/figure.png" pageId="10" pageNumber="143" startId="10.[257,268,1680,1697]" targetBox="[222,1101,215,1629]" targetPageId="10">
|
||
<paragraph blockId="10.[257,1067,1680,1697]" box="[257,1067,1680,1697]" pageId="10" pageNumber="143">
|
||
FIG 6. — Isolectotype of
|
||
<taxonomicName authority="Jord. & Fourr." authorityName="Jord. & Fourr." box="[461,722,1680,1697]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="143" phylum="Tracheophyta" rank="species" species="sericea">
|
||
<emphasis box="[461,602,1680,1697]" italics="true" pageId="10" pageNumber="143">Santolina sericea</emphasis>
|
||
Jord. & Fourr.
|
||
</taxonomicName>
|
||
(LY, “2ème feuille”, specimen to the right).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[111,648,216,1700]" pageId="11" pageNumber="144">
|
||
– Option 2: Willkomm created a new taxon at varietal rank for
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[308,469,596,622]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[308,469,596,622]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
. This theory is supported by the following evidences: 1) the author published a more accurate diagnosis with elements of the diagnosis of
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[422,576,691,717]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[422,576,691,717]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
; 2) he changed “
|
||
<emphasis box="[223,451,723,749]" italics="true" pageId="11" pageNumber="144">spathulato-linearibus</emphasis>
|
||
” (
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[479,636,723,749]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[479,636,723,749]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
) to “
|
||
<emphasis box="[155,334,754,780]" italics="true" pageId="11" pageNumber="144">cuneato-linearia</emphasis>
|
||
” (
|
||
<taxonomicName box="[363,520,754,781]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[363,520,754,781]" italics="true" pageId="11" pageNumber="144">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
var.
|
||
<emphasis italics="true" pageId="11" pageNumber="144">β ceratophylla</emphasis>
|
||
); 3) the author included new elements in the diagnosis: “
|
||
<emphasis italics="true" pageId="11" pageNumber="144">folia superiora aut summa integerrima, omnia virentia</emphasis>
|
||
”; and 4) Willkomm studied more specimens and redefined the concept of
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[196,354,913,939]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[196,354,913,939]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
. If the option 2 is correct, the specimen to the right (
|
||
<figureCitation box="[414,482,944,971]" captionStart="FIG" captionStartId="6.[113,124,1657,1674]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@6.[160,1163,215,1629]" captionTargetPageId="6" captionText="FIG 3. — Lectotype of Santolina oblongifolia var. α obtusifolia Willk. (COI-WILLK 00035955) and lectotype of S. oblongifolia var. β ceratophylla Willk. (COI-WILLK 00035989)." figureDoi="http://doi.org/10.5281/zenodo.5205807" httpUri="https://zenodo.org/record/5205807/files/figure.png" pageId="11" pageNumber="144">Fig. 3</figureCitation>
|
||
; COI-WILLK 00035989) is the
|
||
<typeStatus box="[313,416,976,1002]" pageId="11" pageNumber="144">lectotype</typeStatus>
|
||
(here designated) of
|
||
<taxonomicName box="[113,275,1008,1034]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[113,275,1008,1034]" italics="true" pageId="11" pageNumber="144">S. oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
var.
|
||
<emphasis box="[337,493,1008,1037]" italics="true" pageId="11" pageNumber="144">β ceratophylla</emphasis>
|
||
. The sample has two flowering stems and a sterile stem.This specimen has two revision labels by Sánchez-Mata & Sardinero with the following notation in black ink (the labels are placed under one of the specimens of the variety “
|
||
<emphasis box="[508,627,1166,1192]" italics="true" pageId="11" pageNumber="144">obtusifolia</emphasis>
|
||
”, probably a handling error):
|
||
<emphasis bold="true" box="[429,513,1198,1224]" pageId="11" pageNumber="144">Label 1</emphasis>
|
||
: “
|
||
<taxonomicName authority="Willk." authorityName="Willk. (Willkomm & Lange" authorityYear="1865" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="ceratophylla">
|
||
<emphasis italics="true" pageId="11" pageNumber="144">Santolina oblongifolia</emphasis>
|
||
Boiss. var.
|
||
<emphasis box="[392,527,1229,1255]" italics="true" pageId="11" pageNumber="144">ceratophylla</emphasis>
|
||
Willk.
|
||
</taxonomicName>
|
||
<emphasis box="[623,646,1230,1255]" italics="true" pageId="11" pageNumber="144">in</emphasis>
|
||
Willk. & Lange // Typus [handwritten] //
|
||
<collectingRegion country="Portugal" name="Coimbra" pageId="11" pageNumber="144">Coimbra</collectingRegion>
|
||
[printed]
|
||
<date box="[297,390,1293,1319]" pageId="11" pageNumber="144" value="1993-09">1993 IX</date>
|
||
[handwritten]”;
|
||
<emphasis bold="true" pageId="11" pageNumber="144">Label 2</emphasis>
|
||
: “
|
||
<taxonomicName authority="Boiss." authorityName="Boiss." box="[157,478,1325,1351]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[157,402,1325,1351]" italics="true" pageId="11" pageNumber="144">Santolina oblongifolia</emphasis>
|
||
Boiss.
|
||
</taxonomicName>
|
||
[handwritten] //
|
||
<collectingRegion box="[141,242,1356,1382]" country="Portugal" name="Coimbra" pageId="11" pageNumber="144">Coimbra</collectingRegion>
|
||
[printed]
|
||
<date box="[363,458,1356,1383]" pageId="11" pageNumber="144" value="1993-09">1993 IX</date>
|
||
[handwritten]”. However, the specimen COI-WILLK 00035956 (
|
||
<figureCitation box="[120,191,1420,1446]" captionStart="FIG" captionStartId="4.[351,362,1680,1697]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@4.[160,1163,215,1629]" captionTargetPageId="4" captionText="FIG 2. — Lectotype of Santolina heterophylla Willk. (COI-WILLK 00035956)." figureDoi="http://doi.org/10.5281/zenodo.5205805" httpUri="https://zenodo.org/record/5205805/files/figure.png" pageId="11" pageNumber="144">Fig. 2</figureCitation>
|
||
, see lectotypification of
|
||
<taxonomicName authorityName="Willk. & Cut. (Willkomm" authorityYear="1859" box="[474,635,1420,1446]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="heterophylla">
|
||
<emphasis box="[474,635,1420,1446]" italics="true" pageId="11" pageNumber="144">S. heterophylla</emphasis>
|
||
</taxonomicName>
|
||
) was also selected by Sánchez-Mata & Sardinero as
|
||
<emphasis box="[140,193,1484,1509]" italics="true" pageId="11" pageNumber="144">typus</emphasis>
|
||
of
|
||
<taxonomicName authorityName="Willk. (Willkomm & Lange" authorityYear="1865" box="[228,639,1483,1510]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="ceratophylla">
|
||
<emphasis box="[228,462,1483,1509]" italics="true" pageId="11" pageNumber="144">Santolina oblongifolia</emphasis>
|
||
var.
|
||
<emphasis box="[513,639,1483,1509]" italics="true" pageId="11" pageNumber="144">ceratophylla</emphasis>
|
||
</taxonomicName>
|
||
. The specimens display characteristics consistent with those listed in the protologue (Appendix 1) and show characteristics intermediate between
|
||
<taxonomicName box="[113,273,1610,1636]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="species" species="oblongifolia">
|
||
<emphasis box="[113,126,1610,1636]" italics="true" pageId="11" pageNumber="144">S</emphasis>
|
||
.
|
||
<emphasis box="[144,273,1610,1636]" italics="true" pageId="11" pageNumber="144">oblongifolia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="subsp. rosmarinifolia" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="144" phylum="Tracheophyta" rank="subSpecies" species="rosmarinifolia" subSpecies="rosmarinifolia">
|
||
<emphasis box="[333,346,1610,1636]" italics="true" pageId="11" pageNumber="144">S</emphasis>
|
||
.
|
||
<emphasis box="[363,518,1610,1636]" italics="true" pageId="11" pageNumber="144">rosmarinifolia</emphasis>
|
||
subsp.
|
||
<emphasis italics="true" pageId="11" pageNumber="144">rosmarinifolia</emphasis>
|
||
</taxonomicName>
|
||
. AORG considers that option 2 is the most appropriate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |