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<document id="9AAEB6F9FF2C353A7526C89146D50686" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFC116677A14FC70250E7A08" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="(Vietoderus) Fedorenko 2023, subg.n." docType="treatment" docVersion="3" lastPageNumber="26" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="26" updateTime="1712665435938" updateUser="ExternalLinkService">
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<mods:title id="806ABCD73927D59E7F139EA127737B71">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
</mods:titleInfo>
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<mods:namePart id="05E9876FF986BF833C6269196263D80D">Fedorenko, D. N.</mods:namePart>
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<mods:title id="C8494F9E3C31F616B7777E955AA3437C">Russian Entomological Journal</mods:title>
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<mods:date id="679A91A06102975EE86A2F94A28F6696">2023</mods:date>
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<mods:number id="2F343A516E42EF9C5A608940D521F0C5">2023-03-31</mods:number>
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<mods:identifier id="2DDAF970F5D9AD604E81F77D535530C7" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<subSubSection id="EFF2655EFFC116677A14FC702452760D" box="[882,1329,1001,1026]" pageId="10" pageNumber="26" type="nomenclature">
<paragraph id="A75736D5FFC116677A14FC702452760D" blockId="10.[812,1399,1001,2055]" box="[882,1329,1001,1026]" pageId="10" pageNumber="26">
<heading id="FC1F81B9FFC116677A14FC702452760D" box="[882,1329,1001,1026]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="26" reason="3">
Subgenus
<taxonomicName id="60E84D56FFC116677A87FC7325B7760D" ID-CoL="8MBT6" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[993,1236,1001,1026]" kingdom="Animalia" pageId="10" pageNumber="26" rank="subGenus" status="subgen. nov." subGenus="Vietoderus">
<emphasis id="959CEAC7FFC116677A87FC73253B760E" bold="true" box="[993,1112,1001,1025]" italics="true" pageId="10" pageNumber="26">Vietoderus</emphasis>
Fedorenko
</taxonomicName>
,
<emphasis id="959CEAC7FFC116677DB9FC702452760D" bold="true" box="[1247,1329,1002,1026]" pageId="10" pageNumber="26">
<taxonomicNameLabel id="8EAF57BCFFC116677DB9FC702452760D" box="[1247,1329,1002,1026]" pageId="10" pageNumber="26" rank="subGenus">subg.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC116677A34FB952455762D" box="[850,1334,1039,1058]" pageId="10" pageNumber="26" type="type_taxon">
<paragraph id="A75736D5FFC116677A34FB952455762D" blockId="10.[812,1399,1001,2055]" box="[850,1334,1039,1058]" pageId="10" pageNumber="26">
<typeStatus id="78538877FFC116677A34FB95221D762D" box="[850,894,1039,1058]" pageId="10" pageNumber="26">Type</typeStatus>
species:
<taxonomicName id="60E84D56FFC116677AB7FB9525F9762D" authorityName="Fedorenko" authorityYear="2023" box="[977,1178,1039,1058]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="species" species="laevibasis">
<emphasis id="959CEAC7FFC116677AB7FB9525F9762D" box="[977,1178,1039,1058]" italics="true" pageId="10" pageNumber="26">Pterostichus laevibasis</emphasis>
</taxonomicName>
Fedorenko,
<emphasis id="959CEAC7FFC116677C6DFB952455762D" bold="true" box="[1291,1334,1039,1058]" pageId="10" pageNumber="26">
<taxonomicNameLabel id="8EAF57BCFFC116677C6DFB952455762D" box="[1291,1334,1039,1058]" pageId="10" pageNumber="26" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC116677A34FBBD257077E3" pageId="10" pageNumber="26" type="diagnosis">
<paragraph id="A75736D5FFC116677A34FBBD257077E3" blockId="10.[812,1399,1001,2055]" pageId="10" pageNumber="26">
DIAGNOSIS. A monobasic subgenus of
<taxonomicName id="60E84D56FFC116677D9DFBBD24157633" authorityName="Bonelli" authorityYear="1810" box="[1275,1398,1063,1084]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="959CEAC7FFC116677D9DFBBD24157633" box="[1275,1398,1063,1084]" italics="true" pageId="10" pageNumber="26">Pterostichus</emphasis>
</taxonomicName>
different from
<taxonomicName id="60E84D56FFC116677ADBFBD825307658" authorityName="Tschitscherine" authorityYear="1898" box="[957,1107,1090,1111]" pageId="10" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
<emphasis id="959CEAC7FFC116677ADBFBD825307658" box="[957,1107,1090,1111]" italics="true" pageId="10" pageNumber="26">Neohaptoderus</emphasis>
</taxonomicName>
in the following points: head rather flat; mandibles smooth dorsally (
<emphasis id="959CEAC7FFC116677DD4FBC425A5767C" box="[1202,1222,1118,1139]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. densely obliquely striated); reflexed lateral margin of elytra dull from microsculpture much coarser than that on elytral intervals (
<emphasis id="959CEAC7FFC116677A53FB34222A76CC" box="[821,841,1198,1219]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. elytra subequally and superficially microsculptured throughout); humeral tooth nearly indistinct;
<emphasis id="959CEAC7FFC116677C63FB53247176D1" box="[1285,1298,1225,1246]" italics="true" pageId="10" pageNumber="26">ti</emphasis>
3 without (
<emphasis id="959CEAC7FFC116677A53FB7E222A76F6" box="[821,841,1252,1273]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. with) lateral setae; abdominal sternite VII with apical setae distant from (
<emphasis id="959CEAC7FFC116677A89FB652560771B" box="[1007,1027,1279,1300]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. proximate to) apical margin in both sexes; abdominal sternite VII modified into a large median tubercle (
<emphasis id="959CEAC7FFC116677AECFAAF22FD7745" box="[906,926,1333,1354]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. not or barely modified) in male; right paramere short; aedeagus in lateral view less curved; internal sac with a hypertrophied, transverse, preapical bulb having a few vesicles (
<emphasis id="959CEAC7FFC116677AE8FA1C22C17794" box="[910,930,1414,1435]" italics="true" pageId="10" pageNumber="26">vs</emphasis>
. rather simple, with rather small sclerotized vesicles only). Tergite VIII and sternite VIII wide (long) in female, sternite VIII glabrous apically (
<figureCitation id="3FD32A50FFC116677DDCFA21244C77DE" box="[1210,1327,1467,1489]" captionStart="Figs 5262" captionStartId="13.[226,263,1873,1892]" captionTargetBox="[193,1399,824,1838]" captionTargetId="figure-208@13.[189,1399,820,1839]" captionTargetPageId="13" captionText="Figs 5262. Abdominal urites VIIIIX in female: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 6162 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 5262. Уриты VIIIIX брюшка самки: 5253, 61 — Pterostichus austrinus sp.n.; 5455 — P. glabellus sp.n.; 5657 — P. unisetosus sp.n.; 5860 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 6162 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949112" httpUri="https://zenodo.org/record/10949112/files/figure.png" pageId="10" pageNumber="26">Figs 5859</figureCitation>
). Spermatheca differentiated.
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC116677A34FA6B24407433" pageId="10" pageNumber="26" type="etymology">
<paragraph id="A75736D5FFC116677A34FA6B24407433" blockId="10.[812,1399,1001,2055]" pageId="10" pageNumber="26">
NAME. An abbreviated combination of
<collectingCountry id="DFFF7645FFC116677D92FA6B24297408" box="[1268,1354,1521,1543]" name="Vietnam" pageId="10" pageNumber="26">Vietnam</collectingCountry>
, the country from where the
<typeStatus id="78538877FFC116677D72F996255D742E" box="[1044,1086,1548,1569]" pageId="10" pageNumber="26">type</typeStatus>
species originates, and
<taxonomicName id="60E84D56FFC116677C45F99622177433" authorityName="Tschitscherine" authorityYear="1898" pageId="10" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
<emphasis id="959CEAC7FFC116677C45F99622177433" italics="true" pageId="10" pageNumber="26">Neohaptoderus</emphasis>
</taxonomicName>
, the most similar subgenus of
<taxonomicName id="60E84D56FFC116677DC2F9BD247E7433" authorityName="Bonelli" authorityYear="1810" box="[1188,1309,1575,1596]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="959CEAC7FFC116677DC2F9BD247E7433" box="[1188,1309,1575,1596]" italics="true" pageId="10" pageNumber="26">Pterostichus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EFF2655EFFC116677A34F9D8250E7A08" pageId="10" pageNumber="26" type="discussion">
<paragraph id="A75736D5FFC116677A34F9D822DC756A" blockId="10.[812,1399,1001,2055]" pageId="10" pageNumber="26">
COMMENTS. In the
<collectingCountry id="DFFF7645FFC116677D55F9D825F47457" box="[1075,1175,1602,1624]" name="China" pageId="10" pageNumber="26">East Asia</collectingCountry>
, the new subgenus is similar to
<taxonomicName id="60E84D56FFC116677AE8F9C72542747D" authorityName="Tschitscherine" authorityYear="1898" box="[910,1057,1629,1650]" pageId="10" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
<emphasis id="959CEAC7FFC116677AE8F9C72542747D" box="[910,1057,1629,1650]" italics="true" pageId="10" pageNumber="26">Neohaptoderus</emphasis>
</taxonomicName>
and
<taxonomicName id="60E84D56FFC116677D37F9C72410747D" authority="Motschulsky, 1866" authorityName="Motschulsky" authorityYear="1866" box="[1105,1395,1628,1651]" class="Insecta" family="Carabidae" genus="Rhagadus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="959CEAC7FFC116677D37F9C725D2747D" box="[1105,1201,1629,1650]" italics="true" pageId="10" pageNumber="26">Rhagadus</emphasis>
<bibRefCitation id="C3794B24FFC116677DD1F9C72410747D" author="Motschulsky V." box="[1207,1395,1628,1651]" pageId="10" pageNumber="26" pagination="227 - 313" refId="ref20070" refString="Motschulsky V. 1866. Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. 4 - eme article (suite) // Bull. Soc. Nat. Mosc. T. 38 [1865]. P. 227 - 313." type="book chapter" year="1866">Motschulsky, 1866</bibRefCitation>
</taxonomicName>
, in having the apically beaded prosternal process, certainly plesiomorphic and rather rare character within
<taxonomicName id="60E84D56FFC116677D91F908241374A8" authorityName="Bonelli" authorityYear="1810" box="[1271,1392,1682,1703]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="959CEAC7FFC116677D91F908241374A8" box="[1271,1392,1682,1703]" italics="true" pageId="10" pageNumber="26">Pterostichus</emphasis>
</taxonomicName>
. The pronotum bisulcate on each side, the elytra without parascutellar striole in some species, and the interval 3 with two discal setae, d1 and d2, are characters the new subgenus shares with
<taxonomicName id="60E84D56FFC116677AC7F9642562751C" authorityName="Motschulsky" authorityYear="1866" box="[929,1025,1790,1811]" class="Insecta" family="Carabidae" genus="Rhagadus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="959CEAC7FFC116677AC7F9642562751C" box="[929,1025,1790,1811]" italics="true" pageId="10" pageNumber="26">Rhagadus</emphasis>
</taxonomicName>
. However, species of the latter subgenus have the body slenderer, the basolateral foveae punctate, the metepisterna long, and aedeagus peculiar, with right paramere long.
</paragraph>
<paragraph id="A75736D5FFC116677A34F8F0250E7A08" blockId="10.[812,1399,1001,2055]" pageId="10" pageNumber="26">
On the other hand, the only known species is more similar to some representatives of the subgenus
<taxonomicName id="60E84D56FFC116677DB5F81C243B7594" box="[1235,1368,1926,1947]" pageId="10" pageNumber="26" rank="subGenus" subGenus="Vietosteropus">
<emphasis id="959CEAC7FFC116677DB5F81C243B7594" box="[1235,1368,1926,1947]" italics="true" pageId="10" pageNumber="26">Vietosteropus</emphasis>
</taxonomicName>
in appearance, with which it also shares the elytra with coarsely microsculptured reflexed lateral margins, the mandibles smooth dorsally,
<emphasis id="959CEAC7FFC116677ABAF84C228A75E4" box="[988,1001,2006,2027]" italics="true" pageId="10" pageNumber="26">ti</emphasis>
3 without lateral setae, and the sternite VIII glabrous apically in female.
</paragraph>
</subSubSection>
</treatment>
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