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<document id="0DB40A2CE69B1D5EE7DE489CFDCDCA3C" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629615386486" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2009" docId="1D214E2BF160FFE4DE8276452C39FBB6" docLanguage="en" docName="zt02038p119.pdf" docOrigin="Zootaxa 2038" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Parhydraena ora Perkins 2009, new species" docType="treatment" docVersion="3" lastPageNumber="30" masterDocId="E1183653F17CFFF9DE1575472E27FFB2" masterDocTitle="Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae)" masterLastPageNumber="119" masterPageNumber="1" pageNumber="29" updateTime="1699083612395" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="560477F803C437F6CEAE02E8C0AD5D6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A68501E81B7AE7973474D7FDABE7214E">
<mods:title id="516A6FF6B5A8ACF1C8A3DCB846CCB199">Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="D05204637AF341AF53489DA9F62ADCF1">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="7198C148A394D70D42514CCF9970A98A">text</mods:typeOfResource>
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<mods:title id="2F84EACC2EABD8C436077C1D0E57CF8D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="646E8A9074F94921991DF195FC31C7FB">
<mods:date id="B69E5308705933872BAA351A87D2CEA2">2009</mods:date>
<mods:detail id="445B4CCCB5085FFEF47E2973B167CFCF" type="pubDate">
<mods:number id="F6FE861B041D42959DE2BAC794712193">2009-03-16</mods:number>
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<mods:detail id="0004D06BFDE485F4894DE8DE61D8586B" type="volume">
<mods:number id="769D36386F9FEC76D33AFD803ECD7078">2038</mods:number>
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<mods:classification id="DF7064EDD3ECD4BAC3538CE5842E264E">journal article</mods:classification>
<mods:identifier id="4414CD844833F0847DDDEA536E9B1EBA" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="1D214E2BF160FFE4DE8276452C39FBB6" ID-DOI="http://doi.org/10.5281/zenodo.5321748" ID-Zenodo-Dep="5321748" LSID="urn:lsid:plazi:treatment:1D214E2BF160FFE4DE8276452C39FBB6" httpUri="http://treatment.plazi.org/id/1D214E2BF160FFE4DE8276452C39FBB6" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="DD92ACB6F160FFE5DE8276452FDBFCAE" box="[151,508,770,796]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="9537FF3DF160FFE5DE8276452FDBFCAE" blockId="28.[151,508,770,831]" box="[151,508,770,796]" pageId="28" pageNumber="29">
<heading id="CE7F4851F160FFE5DE8276452FDBFCAE" bold="true" box="[151,508,770,796]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="A7FC232FF160FFE5DE8276452FDBFCAE" bold="true" box="[151,508,770,796]" pageId="28" pageNumber="29">
<taxonomicName id="528884BEF160FFE5DE8276452F42FCAE" ID-CoL="4DLYW" authority="Perkins, 2009" authorityName="Perkins" authorityYear="2009" box="[151,357,770,796]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ora" status="sp. nov.">
<emphasis id="A7FC232FF160FFE5DE8276452F42FCAE" bold="true" box="[151,357,770,796]" italics="true" pageId="28" pageNumber="29">Parhydraena ora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCCF9E54F160FFE5DF7976452FDBFCAE" box="[364,508,770,796]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F160FFE5DE8276622F47FC8D" box="[151,352,805,831]" pageId="28" pageNumber="29" type="description">
<paragraph id="9537FF3DF160FFE5DE8276622F47FC8D" blockId="28.[151,508,770,831]" box="[151,352,805,831]" pageId="28" pageNumber="29">
(
<figureCitation id="0DB3E3B8F160FFE5DE8A76622F26FC8D" box="[159,257,805,831]" captionStart="FIGURE 54" captionStartId="85.[151,255,1730,1754]" captionTargetBox="[206,1377,211,1693]" captionTargetId="figure-25@85.[193,1394,194,1706]" captionTargetPageId="85" captionText="FIGURE 54. Parhydraena ora, dorsal, lateral and ventral habitus of holotype." pageId="28" pageNumber="29">Figs. 54</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DF1B76622F09FC8D" box="[270,302,805,831]" captionStart="FIGURES 5657" captionStartId="87.[151,269,1730,1754]" captionTargetBox="[358,1208,213,1691]" captionTargetId="figure-29@87.[326,1260,194,1706]" captionTargetPageId="87" captionText="FIGURES 5657. Parhydraena, aedeagi of holotypes. 56. P. ora. 57. P. semicostata." pageId="28" pageNumber="29">56</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DF2F76622F7FFC8D" box="[314,344,805,831]" captionStart="FIGURES 9697" captionStartId="116.[151,269,1730,1754]" captionTargetBox="[218,1369,194,1702]" captionTargetId="figure-21@116.[218,1369,194,1706]" captionTargetPageId="116" captionText="FIGURES 9697. Geographical distributions of Parhydraena species." pageId="28" pageNumber="29">96</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F160FFE5DE8276352D71FB95" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="9537FF3DF160FFE5DE8276352D71FB95" blockId="28.[151,1436,882,2030]" pageId="28" pageNumber="29">
<emphasis id="A7FC232FF160FFE5DE8276352F69FC3E" bold="true" box="[151,334,882,908]" pageId="28" pageNumber="29">Type material.</emphasis>
<materialsCitation id="25E0F560F160FFE5DF4276352D74FC01" country="South Africa" latitude="-33.916668" location="Town" longLatPrecision="1202" longitude="18.416666" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="4A33419FF160FFE5DF4276352FE3FC3E" box="[343,452,882,908]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="A7FC232FF160FFE5DC3F76352A20FC3E" bold="true" box="[554,1031,882,908]" pageId="28" pageNumber="29">
<collectingCountry id="ED9FBFADF160FFE5DC3F76352CF7FC3E" box="[554,720,882,908]" name="South Africa" pageId="28" pageNumber="29">South Africa</collectingCountry>
:
<collectingRegion id="574C31DFF160FFE5DCCB76352A20FC3E" box="[734,1031,882,908]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Western Cape Province</collectingRegion>
</emphasis>
,
<collectingRegion id="574C31DFF160FFE5DA0D76352A73FC3E" box="[1048,1108,882,908]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Cape</collectingRegion>
<location id="9057A9E6F160FFE5DA4876352A85FC3E" LSID="urn:lsid:plazi:treatment:1D214E2BF160FFE4DE8276452C39FBB6:9057A9E6F160FFE5DA4876352A85FC3E" box="[1117,1186,882,908]" country="South Africa" latitude="-33.916668" longLatPrecision="1202" longitude="18.416666" name="Town" pageId="28" pageNumber="29" stateProvince="Western Cape">Town</location>
,
<geoCoordinate id="F0BC99FAF160FFE5DABA76352B38FC3E" box="[1199,1311,882,908]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="-33.916668">33° 55' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF160FFE5DB3876352BBBFC3E" box="[1325,1436,882,908]" degrees="18" direction="east" minutes="25" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="18.416666">18° 25' E</geoCoordinate>
[no collector or date on label]. Deposited in the TMSA
</materialsCitation>
.
<materialsCitation id="25E0F560F160FFE5DD7176DF2CC5FBB2" collectingDate="1949-11-08" collectionCode="BMNH" collectorName="Du Toit's Kloof &amp; B. Malkin" country="South Africa" latitude="-33.716667" location="Du Toit's Kloof" longLatPrecision="1203" longitude="19.183332" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="A7FC232FF160FFE5DD7176DF2F22FC6B" bold="true" pageId="28" pageNumber="29">
<typeStatus id="4A33419FF160FFE5DD7176DF2DC6FC00" box="[868,993,920,946]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
(3):
<collectingCountry id="ED9FBFADF160FFE5DA3D76DF2AF5FC00" box="[1064,1234,920,946]" name="South Africa" pageId="28" pageNumber="29">South Africa</collectingCountry>
:
<collectingRegion id="574C31DFF160FFE5DAF076DF2F22FC6B" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Western Cape Province</collectingRegion>
</emphasis>
,
<collectingRegion id="574C31DFF160FFE5DF0076872F92FC68" box="[277,437,960,986]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Cape District</collectingRegion>
,
<collectorName id="387D9AEBF160FFE5DFD476872C5AFC68" box="[449,637,960,986]" pageId="28" pageNumber="29">
<location id="9057A9E6F160FFE5DFD476872C5AFC68" LSID="urn:lsid:plazi:treatment:1D214E2BF160FFE4DE8276452C39FBB6:9057A9E6F160FFE5DFD476872C5AFC68" box="[449,637,960,986]" country="South Africa" latitude="-33.716667" longLatPrecision="1203" longitude="19.183332" name="Du Toit's Kloof" pageId="28" pageNumber="29" stateProvince="Western Cape">Du Toit's Kloof</location>
</collectorName>
, under stones in tiny spring and among algae in bottom gravel,
<geoCoordinate id="F0BC99FAF160FFE5DB6676872EF3FBB2" degrees="33" direction="south" minutes="43" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="-33.716667">33° 43' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF160FFE5DEF776A12F76FBB2" box="[226,337,998,1024]" degrees="19" direction="east" minutes="11" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="19.183332">19° 11' E</geoCoordinate>
,
<date id="E136D9FDF160FFE5DF4B76A12FE8FBB2" box="[350,463,998,1024]" pageId="28" pageNumber="29" value="1949-11-08">
<collectingDate id="F1722015F160FFE5DF4B76A12FE8FBB2" box="[350,463,998,1024]" pageId="28" pageNumber="29" value="1949-11-08">8.xi.1949</collectingDate>
</date>
,
<collectorName id="387D9AEBF160FFE5DFC876A12C72FBB2" box="[477,597,998,1024]" pageId="28" pageNumber="29">B. Malkin</collectorName>
(1
<collectionCode id="F39967F8F160FFE5DC6876A12CFDFBB2" box="[637,730,998,1024]" pageId="28" pageNumber="29">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F160FFE5DCFA76A12B26FBB2" box="[751,1281,998,1024]" collectingDate="1949-11-22" collectionCode="BMNH" collectorName="B. Malkin" country="South Africa" location="As" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<location id="9057A9E6F160FFE5DCFA76A12D28FBB2" LSID="urn:lsid:plazi:treatment:1D214E2BF160FFE4DE8276452C39FBB6:9057A9E6F160FFE5DCFA76A12D28FBB2" box="[751,783,998,1024]" country="South Africa" name="As" pageId="28" pageNumber="29" stateProvince="Western Cape">As</location>
above,
<date id="E136D9FDF160FFE5DD7B76A12DCAFBB2" box="[878,1005,998,1024]" pageId="28" pageNumber="29" value="1949-11-22">
<collectingDate id="F1722015F160FFE5DD7B76A12DCAFBB2" box="[878,1005,998,1024]" pageId="28" pageNumber="29" value="1949-11-22">22.xi.1949</collectingDate>
</date>
,
<collectorName id="387D9AEBF160FFE5DDEE76A12A55FBB2" box="[1019,1138,998,1024]" pageId="28" pageNumber="29">B. Malkin</collectorName>
(1
<collectionCode id="F39967F8F160FFE5DA8976A12ADEFBB2" box="[1180,1273,998,1024]" pageId="28" pageNumber="29">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="25E0F560F160FFE5DB1B76A12D75FB95" collectionCode="MCZ" country="South Africa" latitude="-33.916668" location="Town" longLatPrecision="1202" longitude="18.416666" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<collectingRegion id="574C31DFF160FFE5DB1B76A12B6DFBB2" box="[1294,1354,998,1024]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Cape</collectingRegion>
<location id="9057A9E6F160FFE5DB4776A12BB0FBB2" LSID="urn:lsid:plazi:treatment:1D214E2BF160FFE4DE8276452C39FBB6:9057A9E6F160FFE5DB4776A12BB0FBB2" box="[1362,1431,998,1024]" country="South Africa" latitude="-33.916668" longLatPrecision="1202" longitude="18.416666" name="Town" pageId="28" pageNumber="29" stateProvince="Western Cape">Town</location>
,
<geoCoordinate id="F0BC99FAF160FFE5DE82714A2F24FB95" box="[151,259,1037,1063]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="-33.916668">33° 55' S</geoCoordinate>
,
<geoCoordinate id="F0BC99FAF160FFE5DF05714A2F5BFB95" box="[272,380,1037,1063]" degrees="18" direction="east" minutes="25" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="18.416666">18° 25' E</geoCoordinate>
[no collector or date on label] (1
<collectionCode id="F39967F8F160FFE5DD10714A2D6DFB95" box="[773,842,1037,1063]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F160FFE5DED371742BB0FB5A" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="9537FF3DF160FFE5DED371742BB0FB5A" blockId="28.[151,1436,882,2030]" pageId="28" pageNumber="29">
<emphasis id="A7FC232FF160FFE5DED371742FC6FBFF" bold="true" box="[198,481,1075,1101]" pageId="28" pageNumber="29">Differential diagnosis.</emphasis>
Similar in dorsal habitus to
<taxonomicName id="528884BEF160FFE5DD4B71732DE0FBFF" authorityName="Perkins" authorityYear="2009" box="[862,967,1076,1101]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="A7FC232FF160FFE5DD4B71732DE0FBFF" box="[862,967,1076,1101]" italics="true" pageId="28" pageNumber="29">P. parva</emphasis>
</taxonomicName>
and
<taxonomicName id="528884BEF160FFE5DA0471732A85FBFF" authorityName="Perkins" authorityYear="2009" box="[1041,1186,1076,1101]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="namaqua">
<emphasis id="A7FC232FF160FFE5DA0471732A85FBFF" box="[1041,1186,1076,1101]" italics="true" pageId="28" pageNumber="29">P. namaqua</emphasis>
</taxonomicName>
(
<figureCitation id="0DB3E3B8F160FFE5DAAE71732B02FBFC" box="[1211,1317,1076,1102]" captionStart="FIGURE 42" captionStartId="76.[151,255,1730,1754]" captionTargetBox="[226,1363,214,1689]" captionTargetId="figure-26@76.[206,1381,194,1706]" captionTargetPageId="76" captionText="FIGURE 42. Parhydraena parva, dorsal and lateral habitus of holotype." pageId="28" pageNumber="29">Figs. 42</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DB2C71732B7EFBFC" box="[1337,1369,1076,1102]" captionStart="FIGURE 51" captionStartId="83.[151,255,1730,1754]" captionTargetBox="[203,1384,194,1706]" captionTargetId="figure-24@83.[203,1384,194,1706]" captionTargetPageId="83" captionText="FIGURE 51. Parhydraena namaqua, dorsal and lateral habitus of holotype." pageId="28" pageNumber="29">51</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DB7B71732BA8FBFC" box="[1390,1423,1076,1102]" captionStart="FIGURE 54" captionStartId="85.[151,255,1730,1754]" captionTargetBox="[206,1377,211,1693]" captionTargetId="figure-25@85.[193,1394,194,1706]" captionTargetPageId="85" captionText="FIGURE 54. Parhydraena ora, dorsal, lateral and ventral habitus of holotype." pageId="28" pageNumber="29">54</figureCitation>
). Distinguished from
<taxonomicName id="528884BEF160FFE5DF9E711C2FD6FBC6" authorityName="Perkins" authorityYear="2009" box="[395,497,1115,1140]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="A7FC232FF160FFE5DF9E711C2FD6FBC6" box="[395,497,1115,1140]" italics="true" pageId="28" pageNumber="29">P. parva</emphasis>
</taxonomicName>
by the larger size (length ca. 1.77 vs.
<quantity id="527052D8F160FFE5DDC9711D2A69FBC6" box="[988,1102,1114,1140]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="28" pageNumber="29" unit="mm" value="1.64">1.64 mm</quantity>
), the comparatively shorter maxillary palpi (palpus length/head width ca.
<quantity id="527052D8F160FFE5DCCF71C62D69FB29" box="[730,846,1153,1179]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="28" pageNumber="29" unit="mm" value="0.78">0.78 mm</quantity>
), and the different shape, in males, of the 6th abdominal ventrite. Males have the 6th ventrite modified similar to that of
<taxonomicName id="528884BEF160FFE5DA3571EF2A8BFB73" authorityName="Perkins" authorityYear="2009" box="[1056,1196,1192,1217]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="namaqua">
<emphasis id="A7FC232FF160FFE5DA3571EF2A8BFB73" box="[1056,1196,1192,1217]" italics="true" pageId="28" pageNumber="29">P. namaqua</emphasis>
</taxonomicName>
. The male genitalia distinctively differ in the three species, and should be examined for reliable determinations (
<figureCitation id="0DB3E3B8F160FFE5DADA71892B16FB5A" box="[1231,1329,1230,1256]" captionStart="FIGURES 4445" captionStartId="78.[151,269,1730,1754]" captionText="FIGURES 4445. Parhydraena, aedeagi of holotypes. 44. P. parva. 45. P. seriata." pageId="28" pageNumber="29">Figs. 44</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DB2871892B7AFB5A" box="[1341,1373,1230,1256]" captionStart="FIGURES 5253" captionStartId="84.[151,269,1730,1754]" captionTargetBox="[246,1333,222,1670]" captionTargetId="figure-31@84.[228,1360,194,1706]" captionTargetPageId="84" captionText="FIGURES 5253. Parhydraena, aedeagi of holotypes. 52. P. lancicula. 53. P. namaqua." pageId="28" pageNumber="29">53</figureCitation>
,
<figureCitation id="0DB3E3B8F160FFE5DB7C71892BADFB5A" box="[1385,1418,1230,1256]" captionStart="FIGURES 5657" captionStartId="87.[151,269,1730,1754]" captionTargetBox="[358,1208,213,1691]" captionTargetId="figure-29@87.[326,1260,194,1706]" captionTargetPageId="87" captionText="FIGURES 5657. Parhydraena, aedeagi of holotypes. 56. P. ora. 57. P. semicostata." pageId="28" pageNumber="29">56</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F160FFE4DED371B22D13FC22" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="9537FF3DF160FFE5DED371B22A5BFAEF" blockId="28.[151,1436,882,2030]" pageId="28" pageNumber="29">
<emphasis id="A7FC232FF160FFE5DED371B22F79FABD" bold="true" box="[198,350,1269,1295]" pageId="28" pageNumber="29">Description.</emphasis>
Size (length/width, mm)
<typeStatus id="4A33419FF160FFE5DC9E71B22CD1FABD" box="[651,758,1269,1295]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
: body (length to elytral apices) 1.77/0.73; head 0.28/0.49; pronotum 0.40/0.66, PA 0.54, PB 0.58; elytra 1.09/0.73. Color dark brown except head and disc of pronotum piceous; apex of distal mere of maxillary palpus not darker than remainder of palpus.
</paragraph>
<paragraph id="9537FF3DF160FFE5DED3702E2ED2F920" blockId="28.[151,1436,882,2030]" pageId="28" pageNumber="29">Head with eyes comparatively small, in dorsal aspect 7 rather coarse, convex facets in longest series. Ocelli distinct round small shining, each located at base of moderately deep oblique interocular fovea. Frons microreticulate, dull, disc indistinctly finely punctate, ca. 1xef; area between interocular foveae and eye flat or very weakly convex. Clypeus markedly microreticulate, dull, except anterior margin shining. Labrum dull, apicomedially emarginate to form rounded lobe on each side; laterally fringed with short arcuate setae. Maxillary palpi elongate, combined lengths of meres less than width of head (ratio ca. 25/32); ratios of meres 24 ca. 9/5/11. Mentum and submentum sparsely finely punctulate, effacedly microreticulate, interstices shining.</paragraph>
<paragraph id="9537FF3DF160FFE5DED373D92CF7F875" blockId="28.[151,1436,882,2030]" pageId="28" pageNumber="29">Pronotum with anterior margin straight or nearly so, median 1/3 with extremely narrow hyaline border. Anterior angles obtuse, moderately angulate. Sides rounded, widest just before middle, behind middle gradually attenuate, straight. Posterior angle obtusangulate. Pronotum coarsely and very densely punctate, rugulose, dull, except discal relief much more finely punctate, ca. 1xef, interstices smooth and shining. Discal impressions deep, broadly U-shaped, many punctures confluent; rather deep, wide impression at each posterior angle. Each puncture with anterior margin extended posteriorly as narrow ridge, dividing puncture into a pair of two smaller punctures, most punctures with indistinct, short recumbent or decumbent seta arising from posterior extreme of dividing ridge.</paragraph>
<paragraph id="9537FF3DF160FFE4DED372932C14FEFE" blockId="28.[151,1436,882,2030]" lastBlockId="29.[151,1436,152,1028]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">Elytra elongate-ovate, sides weakly arcuate, margins distinct from shoulder for 4/5 of length. Ten-seriate punctate, only first series striate-impressed, in distal 1/2; punctures ca. 1xef, longitudinally separated by ca. 1xpd, becoming finer and shallower on apical declivity, each puncture with minute granule at anterior margin. Interval width ca. 2xpd, shining but with irregular reflections, each with unilinear or slightly irregular row of minute granules, each granule at its posterior side with short recumbent seta, setae very similar in length and density to setae of serial punctures.</paragraph>
<paragraph id="9537FF3DF161FFE4DED3741E2F2BFD30" blockId="29.[151,1436,152,1028]" pageId="29" pageNumber="30">Metaventrite with moderately shallow median elongate oval impression in basal 1/2, midlongitudinal shining carina between anterior extreme of oval impression and mesoventral intercoxal process. Metaventrite and abdominal ventrites 14 hydrofuge pubescent. Ventrite 5 hydrofuge pubescent in basal 1/3 laterally, posterior margin of pubescent area produced in middle to 4/5 length of ventrite; shining part of ventrite 5 very sparsely pubescent. Ventrite 6 very weakly, effacedly microreticulate, shining, with prominent tubercle in midline, area posterior to tubercle markedly concave. Last ventrite asymmetrical, apex lobiform and setose; last tergite with tuft of setae on each side of apicomedian notch into which fits lobe of last ventrite. Female unknown.</paragraph>
<paragraph id="9537FF3DF161FFE4DED377C92C2EFD44" blockId="29.[151,1436,152,1028]" pageId="29" pageNumber="30">Legs moderately long and slender, combined lengths of metatibia and metatarsus greater than width of head (ratio ca. 49/32), ratio of lengths of metatibia/metatarsus ca. 28/21. Basal 3 pro- and mesotarsomeres, in male, with pad of suction setae.</paragraph>
<paragraph id="9537FF3DF161FFE4DED376452C43FCAE" blockId="29.[151,1436,152,1028]" box="[198,612,770,796]" pageId="29" pageNumber="30">
Wings fully developed in
<typeStatus id="4A33419FF161FFE4DFE076452C78FCAE" box="[501,607,770,796]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9537FF3DF161FFE4DED3766E2D13FC22" blockId="29.[151,1436,152,1028]" pageId="29" pageNumber="30">
Aedeagus length ca.
<quantity id="527052D8F161FFE4DFD1766E2C16FCF1" box="[452,561,809,835]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="29" pageNumber="30" unit="mm" value="0.41">0.41 mm</quantity>
, slightly sinuate in ventral view; main-piece widened apically in ventral view, with large pointed process in left lateral view; distal piece with small rounded lobe and much larger hood-like lobe, latter with bifurcate left margin (
<figureCitation id="0DB3E3B8F161FFE4DCDA76312D01FC22" box="[719,806,886,912]" captionStart="FIGURES 5657" captionStartId="87.[151,269,1730,1754]" captionTargetBox="[358,1208,213,1691]" captionTargetId="figure-29@87.[326,1260,194,1706]" captionTargetPageId="87" captionText="FIGURES 5657. Parhydraena, aedeagi of holotypes. 56. P. ora. 57. P. semicostata." pageId="29" pageNumber="30">Fig. 56</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F161FFE4DED376DA2A74FC05" box="[198,1107,925,951]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="9537FF3DF161FFE4DED376DA2A74FC05" blockId="29.[151,1436,152,1028]" box="[198,1107,925,951]" pageId="29" pageNumber="30">
<emphasis id="A7FC232FF161FFE4DED376DA2F74FC05" bold="true" box="[198,339,925,951]" pageId="29" pageNumber="30">Etymology.</emphasis>
Named in reference to the complicated left edge of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="DD92ACB6F161FFE4DED376842C39FBB6" pageId="29" pageNumber="30" type="distribution">
<paragraph id="9537FF3DF161FFE4DED376842C39FBB6" blockId="29.[151,1436,152,1028]" pageId="29" pageNumber="30">
<emphasis id="A7FC232FF161FFE4DED376842F40FC6F" bold="true" box="[198,359,963,989]" pageId="29" pageNumber="30">Distribution.</emphasis>
Currently known only from &quot;
<collectingRegion id="574C31DFF161FFE4DCF076832D02FC6C" box="[741,805,964,990]" country="South Africa" name="Western Cape" pageId="29" pageNumber="30">Cape</collectingRegion>
Town&quot; and Du Toit's Kloof, in southwesternmost
<collectingRegion id="574C31DFF161FFE4DE8276AD2F8EFBB6" box="[151,425,1002,1028]" country="South Africa" name="Western Cape" pageId="29" pageNumber="30">Western Cape Province</collectingRegion>
(
<figureCitation id="0DB3E3B8F161FFE4DFAC76AD2C36FBB6" box="[441,529,1002,1028]" captionStart="FIGURES 9697" captionStartId="116.[151,269,1730,1754]" captionTargetBox="[218,1369,194,1702]" captionTargetId="figure-21@116.[218,1369,194,1706]" captionTargetPageId="116" captionText="FIGURES 9697. Geographical distributions of Parhydraena species." pageId="29" pageNumber="30">Fig. 96</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>