treatments-xml/data/12/1F/87/121F87D60A4AFFABDA9B5B1CFC6A6808.xml

<document ID-DOI="10.25221/fee.450.1" ID-ISSN="2713-2196" approvalRequired="50" approvalRequired_for_textStreams="50" checkinTime="1665256513248" checkinUser="felipe" docAuthor="Ermilov, S. G. &amp; Salavatulin, V. M." docDate="2022" docId="121F87D60A4AFFABDA9B5B1CFC6A6808" docLanguage="en" docName="FarEastEntomol.450.1-8.pdf" docOrigin="Far Eastern Entomologist 450" docSource="http://dx.doi.org/10.25221/fee.450.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Epidamaeus hexatuberculatus Ermilov et Salavatulin 2022, sp. n." docType="treatment" docVersion="1" lastPageNumber="8" masterDocId="EE26FFAE0A48FFACDA785859FFC46936" masterDocTitle="A new species of the genus Epidamaeus Bulanova-Zachvatkina, 1957 (Acari: Oribatida: Damaeidae) from Western Siberia" masterLastPageNumber="8" masterPageNumber="1" pageNumber="3" updateTime="1665262495003" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of the genus Epidamaeus Bulanova-Zachvatkina, 1957 (Acari: Oribatida: Damaeidae) from Western Siberia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ermilov, S. G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salavatulin, V. M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>450</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>8</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.450.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.450.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:121F87D60A4AFFABDA9B5B1CFC6A6808" httpUri="http://treatment.plazi.org/id/121F87D60A4AFFABDA9B5B1CFC6A6808" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
<subSubSection pageId="2" pageNumber="3" type="nomenclature">
<paragraph blockId="2.[227,1056,837,894]" box="[227,897,837,862]" pageId="2" pageNumber="3">
<heading box="[227,897,837,862]" fontSize="9" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis bold="true" box="[227,897,837,862]" pageId="2" pageNumber="3">
<taxonomicName authority="Ermilov et Salavatulin" authorityName="Ermilov et Salavatulin" authorityYear="2022" box="[227,826,837,862]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis bold="true" box="[227,561,837,861]" italics="true" pageId="2" pageNumber="3">Epidamaeus hexatuberculatus</emphasis>
Ermilov et Salavatulin
</taxonomicName>
, 
<taxonomicNameLabel box="[837,897,838,862]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[227,1056,837,894]" box="[227,1056,872,894]" pageId="2" pageNumber="3">
http://zoobank.org/NomenclaturalActs/ 
<uuid box="[612,1056,872,894]" pageId="2" pageNumber="3">AB12E495-A525-4F71-93C4-4EECB7062682</uuid>
</paragraph>
</subSubSection>
<subSubSection box="[227,319,906,930]" pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[227,319,906,930]" box="[227,319,906,930]" pageId="2" pageNumber="3">
<figureCitation box="[227,319,906,930]" captionStart-0="Figs 3–5" captionStart-1="Figs 6–9" captionStartId-0="3.[265,306,1511,1533]" captionStartId-1="4.[265,306,1530,1552]" captionTargetBox-0="[238,1116,472,1467]" captionTargetBox-1="[249,1102,330,1455]" captionTargetId-0="figure-57@3.[227,1122,472,1478]" captionTargetId-1="figure-1@4.[227,1122,319,1469]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs 3–5. Epidamaeus hexatuberculatus sp. n., adult: 3 – dorsal view (legs not shown);" captionText-1="Figs 6–9. Epidamaeus hexatuberculatus sp. n., adult: 6 – leg I, right, antiaxial view; 7 –" pageId="2" pageNumber="3">Figs 3–9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="materials_examined">
<paragraph blockId="2.[227,1128,967,1605]" box="[265,1127,967,992]" pageId="2" pageNumber="3">
<materialsCitation box="[265,1127,967,992]" collectionCode="TYPE, MATERIAL" country="Russia" location="Western Siberia" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Tyumen'" typeStatus="holotype">
<collectionCode box="[265,482,967,991]" pageId="2" pageNumber="3">TYPE MATERIAL</collectionCode>
. 
<typeStatus box="[495,594,967,991]" pageId="2" pageNumber="3">Holotype</typeStatus>
– 
<specimenCount box="[626,646,968,991]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
, 
<emphasis bold="true" box="[662,738,967,991]" pageId="2" pageNumber="3">
<collectingCountry box="[662,738,967,991]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
</emphasis>
: 
<location LSID="urn:lsid:plazi:treatment:121F87D60A4AFFABDA9B5B1CFC6A6808:9F69601B0A4AFFAED88B5B91FC606AE9" box="[755,932,967,991]" country="Russia" name="Western Siberia" pageId="2" pageNumber="3" stateProvince="Tyumen'">Western Siberia</location>
, 
<collectingRegion box="[945,1123,967,992]" country="Russia" name="Tyumen'" pageId="2" pageNumber="3">Tyumen Region</collectingRegion>
,
</materialsCitation>
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
Tobolsk District, nearly Nadtsy Settlement, 
<geoCoordinate box="[704,856,993,1022]" degrees="58.631796" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="58.631798">
58.631796 
<superScript attach="left" box="[817,826,993,1007]" fontSize="6" pageId="2" pageNumber="3">o</superScript>
N
</geoCoordinate>
, 
<geoCoordinate box="[868,1017,993,1022]" degrees="68.566218" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="68.566216">
68.566218 
<superScript attach="left" box="[981,990,993,1007]" fontSize="6" pageId="2" pageNumber="3">o</superScript>
E
</geoCoordinate>
, soil-litter in mixed forest (birch-pine-cedar), 
<date box="[627,766,1028,1053]" pageId="2" pageNumber="3" value="2015-07-30">30.VII 2015</date>
, collected by 
<collectionCode box="[931,950,1029,1052]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
.M. 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[1000,1124,1028,1052]" pageId="2" pageNumber="3" rank="species" species="Salavatulin" status="sp. n.">Salavatulin</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" box="[227,1126,1059,1084]" pageId="2" pageNumber="3">
<typeStatus box="[227,335,1060,1084]" pageId="2" pageNumber="3">Paratypes</typeStatus>
: the same label as 
<typeStatus box="[546,642,1059,1083]" pageId="2" pageNumber="3">holotype</typeStatus>
, 
<specimenCount box="[656,691,1059,1083]" pageId="2" pageNumber="3" type="male">1♂</specimenCount>
, 
<specimenCount box="[705,740,1059,1083]" pageId="2" pageNumber="3" type="female">2♀</specimenCount>
; 
<materialsCitation box="[755,1126,1059,1083]" country="Russia" location="Western Siberia" pageId="2" pageNumber="3" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[755,840,1059,1083]" pageId="2" pageNumber="3">
<collectingCountry box="[755,835,1059,1083]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:121F87D60A4AFFABDA9B5B1CFC6A6808:9F69601B0A4AFFAED9295C7DFBC66D0D" box="[849,1026,1059,1083]" country="Russia" name="Western Siberia" pageId="2" pageNumber="3">Western Siberia</location>
, Khanty-
</materialsCitation>
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
Mansiysk Autonomous Region, Nephteyugansk District, nearly Kut-Yakh Settlement, 
<geoCoordinate box="[295,448,1116,1144]" degrees="60.351376" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="60.351376">
60.351376 
<superScript attach="left" box="[409,418,1116,1130]" fontSize="6" pageId="2" pageNumber="3">o</superScript>
N
</geoCoordinate>
, 
<geoCoordinate box="[460,610,1116,1144]" degrees="72.099673" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="72.09967">
72.099673 
<superScript attach="left" box="[574,583,1116,1130]" fontSize="6" pageId="2" pageNumber="3">o</superScript>
E
</geoCoordinate>
, soil-litter in mixed forest (spruce-birch-cedar),
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" box="[227,777,1151,1175]" pageId="2" pageNumber="3">
<materialsCitation box="[227,774,1151,1175]" collectingDate="2015-07-23" collectionCode="V" pageId="2" pageNumber="3" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<date box="[227,360,1151,1175]" pageId="2" pageNumber="3" value="2015-07-23">
<collectingDate box="[227,360,1151,1175]" pageId="2" pageNumber="3" value="2015-07-23">23.VII 2015</collectingDate>
</date>
, 
<specimenCount box="[371,406,1151,1175]" pageId="2" pageNumber="3" type="male">2♂</specimenCount>
, 
<specimenCount box="[417,437,1152,1175]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
, collected by 
<collectionCode box="[587,609,1152,1175]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="3" type="Museum">V</collectionCode>
.M. 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[650,774,1151,1175]" pageId="2" pageNumber="3" rank="species" species="Salavatulin" status="sp. n.">Salavatulin</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" box="[265,1126,1182,1206]" pageId="2" pageNumber="3">
<materialsCitation box="[265,1126,1182,1206]" collectionCode="TYPE, DEPOSITION" pageId="2" pageNumber="3" specimenCount="1" typeStatus="holotype">
<collectionCode box="[265,509,1182,1206]" pageId="2" pageNumber="3">TYPE DEPOSITION</collectionCode>
. The 
<typeStatus box="[586,679,1182,1206]" pageId="2" pageNumber="3">holotype</typeStatus>
is deposited in the collection of the
</materialsCitation>
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
Senckenberg Museum of Natural History, Görlitz, 
<collectingCountry box="[792,895,1213,1237]" name="Germany" pageId="2" pageNumber="3">Germany</collectingCountry>
; 
<specimenCount box="[910,1083,1213,1237]" pageId="2" pageNumber="3" type="generic" typeStatus="paratypes">seven paratypes</specimenCount>
are deposited in the collection of the Tyumen State University Museum of Zoology,
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
<collectingRegion box="[227,317,1274,1298]" country="Russia" name="Tyumen'" pageId="2" pageNumber="3">Tyumen</collectingRegion>
, 
<collectingCountry box="[328,401,1274,1298]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
DIAGNOSIS. Body length: 531–630. Cerotegument filamentous. Exuvial scalps usually present on notogaster. Propodolateral apophysis absent. Three pairs of simple prodorsobasal tubercles (
<emphasis box="[597,630,1397,1420]" italics="true" pageId="2" pageNumber="3">Da</emphasis>
, 
<emphasis box="[651,681,1397,1420]" italics="true" pageId="2" pageNumber="3">Ba</emphasis>
, 
<emphasis box="[701,731,1397,1420]" italics="true" pageId="2" pageNumber="3">Bp</emphasis>
); 
<emphasis box="[762,795,1397,1420]" italics="true" pageId="2" pageNumber="3">Da</emphasis>
located anteromedial to 
<emphasis box="[1090,1120,1397,1420]" italics="true" pageId="2" pageNumber="3">Ba</emphasis>
.
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" box="[227,1127,1427,1451]" pageId="2" pageNumber="3">Rostral and lamellar setae setiform, barbed. Interlamellar seta narrowly phylliform.</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" pageId="2" pageNumber="3">
Bothridial seta rod-like, barbed. 
<taxonomicName authorityName="Kaup" authorityYear="1829" box="[584,645,1458,1482]" class="Aves" genus="Spina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">Spina</taxonomicName>
adnata of medium length, thorn-like. Notogastral setae of 
<emphasis box="[395,407,1489,1512]" italics="true" pageId="2" pageNumber="3">c</emphasis>
-, 
<emphasis box="[430,437,1488,1512]" italics="true" pageId="2" pageNumber="3">l</emphasis>
- and 
<emphasis box="[499,512,1488,1512]" italics="true" pageId="2" pageNumber="3">h</emphasis>
-series phylliform, 
<emphasis box="[717,730,1489,1512]" italics="true" pageId="2" pageNumber="3">p</emphasis>
<subScript attach="right" box="[731,740,1500,1514]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
, 
<emphasis box="[753,766,1489,1512]" italics="true" pageId="2" pageNumber="3">p</emphasis>
<subScript attach="left" box="[767,776,1500,1514]" fontSize="6" pageId="2" pageNumber="3">2</subScript>
and 
<emphasis box="[828,841,1489,1512]" italics="true" pageId="2" pageNumber="3">p</emphasis>
<subScript attach="left" box="[842,851,1500,1514]" fontSize="6" pageId="2" pageNumber="3">3</subScript>
setiform, barbed. Ventral enantiophyses 
<emphasis box="[381,411,1519,1543]" italics="true" pageId="2" pageNumber="3">E2</emphasis>
and 
<emphasis box="[462,478,1520,1543]" italics="true" pageId="2" pageNumber="3">V</emphasis>
absent; mediosejugal enantiophysis well-developed. Parastigmatic tubercle 
<emphasis box="[385,412,1550,1573]" italics="true" pageId="2" pageNumber="3">Sa</emphasis>
long, thorn-like, 
<emphasis box="[598,625,1550,1573]" italics="true" pageId="2" pageNumber="3">Sp</emphasis>
tubercle-like. Epimeral setal formula: 3-1-3-4.
</paragraph>
<paragraph blockId="2.[227,1128,967,1605]" box="[227,1125,1580,1605]" pageId="2" pageNumber="3">Epimeral and anogenital setae setiform, slightly barbed. Discidium tubercle-like. Legs</paragraph>
<paragraph blockId="3.[227,1127,264,441]" box="[227,1127,264,288]" pageId="3" pageNumber="4">I and IV longer than body length; leg IV longest. Formulas of leg segments (I-IV):</paragraph>
<paragraph blockId="3.[227,1127,264,441]" pageId="3" pageNumber="4">
femur 7-6-4-4; genu and tibia 4-4-3-3; tarsus 21-18-18-15. Seta 
<emphasis box="[938,950,295,318]" italics="true" pageId="3" pageNumber="4">v</emphasis>
<subScript attach="left" box="[950,959,306,320]" fontSize="6" pageId="3" pageNumber="4">2</subScript>
<emphasis box="[958,967,295,318]" italics="true" pageId="3" pageNumber="4">’</emphasis>
present on all tarsi.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph blockId="3.[227,1127,264,441]" pageId="3" pageNumber="4">
<materialsCitation box="[265,939,356,380]" collectionCode="DESCRIPTION" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
<collectionCode box="[265,442,356,380]" pageId="3" pageNumber="4">DESCRIPTION</collectionCode>
. Measurements. Body length: 581 (
<typeStatus box="[833,932,356,380]" pageId="3" pageNumber="4">holotype</typeStatus>
)
</materialsCitation>
, 498 (male 
<typeStatus pageId="3" pageNumber="4">paratypes</typeStatus>
); 531–630 (female 
<typeStatus box="[513,620,387,411]" pageId="3" pageNumber="4">paratypes</typeStatus>
); notogaster width: 381 (
<typeStatus box="[910,1009,386,410]" pageId="3" pageNumber="4">holotype</typeStatus>
), 315–332
</paragraph>
<paragraph blockId="3.[227,1127,264,441]" box="[227,726,417,441]" pageId="3" pageNumber="4">
(male 
<typeStatus box="[294,401,417,441]" pageId="3" pageNumber="4">paratypes</typeStatus>
); 332–381 (female 
<typeStatus box="[607,714,417,441]" pageId="3" pageNumber="4">paratypes</typeStatus>
).
</paragraph>
<caption box="[265,1127,1511,1533]" pageId="3" pageNumber="4" startId="3.[265,306,1511,1533]" targetBox="[238,1116,472,1467]" targetPageId="3">
<paragraph blockId="3.[227,1127,1511,1588]" box="[265,1127,1511,1533]" pageId="3" pageNumber="4">
Figs 3–5. 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[364,656,1511,1532]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis box="[364,656,1511,1532]" italics="true" pageId="3" pageNumber="4">Epidamaeus hexatuberculatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[664,720,1511,1532]" pageId="3" pageNumber="4">
<taxonomicNameLabel box="[664,720,1511,1532]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult: 3 – dorsal view (legs not shown);
</paragraph>
</caption>
<paragraph blockId="3.[227,1127,1511,1588]" box="[227,1127,1538,1560]" pageId="3" pageNumber="4">4 – ventral view (gnathosoma and legs not shown); 5 – notogaster and basal part of prodorsum,</paragraph>
<paragraph blockId="3.[227,1127,1511,1588]" box="[227,527,1566,1588]" pageId="3" pageNumber="4">lateral view. Scale bar 100 μm.</paragraph>
<caption box="[265,1127,1530,1552]" pageId="4" pageNumber="5" startId="4.[265,306,1530,1552]" targetBox="[249,1102,330,1455]" targetPageId="4">
<paragraph blockId="4.[227,1127,1530,1607]" box="[265,1127,1530,1552]" pageId="4" pageNumber="5">
Figs 6–9. 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[363,653,1530,1551]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis box="[363,653,1530,1551]" italics="true" pageId="4" pageNumber="5">Epidamaeus hexatuberculatus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[661,716,1530,1551]" pageId="4" pageNumber="5">
<taxonomicNameLabel box="[661,716,1530,1551]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult: 6 – leg I, right, antiaxial view; 7 –
</paragraph>
</caption>
<paragraph blockId="4.[227,1127,1530,1607]" box="[227,1127,1557,1579]" pageId="4" pageNumber="5">leg II, without tarsus, right, dorsoantiaxial view; 8 – leg III, without tarsus, trochanter turned,</paragraph>
<paragraph blockId="4.[227,1127,1530,1607]" box="[227,882,1585,1607]" pageId="4" pageNumber="5">left, antiaxial view; 9 – leg IV, left, antiaxial view. Scale bar 50 μm.</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">Integument. Color medium to dark brown in preserved mature adults. Cerotegument conspicuous, excrescences varying in length but mostly filamentous on body and legs; setae usually without cerotegument. Procuticle mostly smooth.</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,356,380]" pageId="5" pageNumber="6">Usually with full stack of juvenile exuvial scalps, often with other adherent structures,</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">particularly eggs or eggshells and fecal pellets, but without compacted organic debris.</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
Prodorsum. Rostrum rounded. Propodolateral apophysis absent. Prodorsobasal tubercle 
<emphasis box="[322,355,479,502]" italics="true" pageId="5" pageNumber="6">Da</emphasis>
and postbothridial enantiophysis (
<emphasis box="[733,763,479,502]" italics="true" pageId="5" pageNumber="6">Ba</emphasis>
, 
<emphasis box="[778,808,479,502]" italics="true" pageId="5" pageNumber="6">Bp</emphasis>
) well developed, subconical,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
distally rounded, similar in size; 
<emphasis box="[575,607,510,533]" italics="true" pageId="5" pageNumber="6">Da</emphasis>
located clearly anteromedial to 
<emphasis box="[949,978,510,533]" italics="true" pageId="5" pageNumber="6">Ba</emphasis>
; laterosejugal enantiophysis absent. Rostral (77–82) and lamellar (102–106) setae setiform, barbed;
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,570,594]" pageId="5" pageNumber="6">
<emphasis box="[227,246,570,594]" italics="true" pageId="5" pageNumber="6">le</emphasis>
thicker than 
<emphasis box="[390,414,571,594]" italics="true" pageId="5" pageNumber="6">ro</emphasis>
. Indistinct transverse furrow between insertions of lamellar setae.
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1128,601,625]" pageId="5" pageNumber="6">Interlamellar seta (69–73) narrowly phylliform, directed mediad or posteromediad.</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,632,656]" pageId="5" pageNumber="6">Bothridial seta (221–225) rod-like, barbed. Bothridium with projecting part goblet-</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
shaped, with vertical banding; mutual distance of openings slightly greater than that of tubercles 
<emphasis box="[360,389,694,717]" italics="true" pageId="5" pageNumber="6">Ba</emphasis>
. Anterobothridial ridge present. Exobothridial seta (49–57) setiform,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
slightly barbed. Alveolar vestige of second exobothridial seta present posterior to seta 
<emphasis box="[275,299,755,778]" italics="true" pageId="5" pageNumber="6">ex</emphasis>
.
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
Notogaster. Nearly circular in outline; uniformly curved, hemispherical with maximum height in middle. 
<taxonomicName authorityName="Kaup" authorityYear="1829" box="[534,595,816,840]" class="Aves" genus="Spina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">Spina</taxonomicName>
adnata (41–53) shorter than a mutual distance of pair, thorn-like, directed anterolaterad, reaching or surpassing level of tubercle 
<emphasis box="[1097,1127,847,870]" italics="true" pageId="5" pageNumber="6">Ba</emphasis>
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,877,901]" pageId="5" pageNumber="6">
(sometimes 
<emphasis box="[363,396,878,901]" italics="true" pageId="5" pageNumber="6">Da</emphasis>
). Dorsal notogastral setae (
<emphasis box="[714,726,878,901]" italics="true" pageId="5" pageNumber="6">c</emphasis>
-, 
<emphasis box="[755,762,877,901]" italics="true" pageId="5" pageNumber="6">l</emphasis>
- and 
<emphasis box="[835,848,877,901]" italics="true" pageId="5" pageNumber="6">h</emphasis>
-series) directed radially,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,908,933]" pageId="5" pageNumber="6">
inserted in uniformly curving, nearly semicircular row; 
<emphasis box="[825,837,909,932]" italics="true" pageId="5" pageNumber="6">c</emphasis>
<subScript attach="left" box="[836,845,919,933]" fontSize="6" pageId="5" pageNumber="6">1</subScript>
, 
<emphasis box="[859,871,909,932]" italics="true" pageId="5" pageNumber="6">c</emphasis>
<subScript attach="left" box="[871,880,919,933]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
(73–82) and 
<emphasis box="[1024,1045,908,932]" italics="true" pageId="5" pageNumber="6">la</emphasis>
, 
<emphasis box="[1059,1086,908,932]" italics="true" pageId="5" pageNumber="6">lm</emphasis>
, 
<emphasis box="[1099,1120,908,932]" italics="true" pageId="5" pageNumber="6">lp</emphasis>
,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,938,964]" pageId="5" pageNumber="6">
<emphasis box="[227,240,938,962]" italics="true" pageId="5" pageNumber="6">h</emphasis>
<subScript attach="left" box="[240,249,950,964]" fontSize="6" pageId="5" pageNumber="6">3</subScript>
(57–61) relatively broadly phylliform, 
<emphasis box="[671,684,938,962]" italics="true" pageId="5" pageNumber="6">h</emphasis>
<subScript attach="left" box="[685,694,950,964]" fontSize="6" pageId="5" pageNumber="6">1</subScript>
and 
<emphasis box="[746,759,938,962]" italics="true" pageId="5" pageNumber="6">h</emphasis>
<subScript attach="left" box="[760,769,950,964]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
(41–45) narrowly phylliform, 
<emphasis box="[1098,1111,939,962]" italics="true" pageId="5" pageNumber="6">p</emphasis>
<subScript attach="left" box="[1111,1120,950,964]" fontSize="6" pageId="5" pageNumber="6">1</subScript>
,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
<emphasis box="[227,240,970,993]" italics="true" pageId="5" pageNumber="6">p</emphasis>
<subScript attach="left" box="[240,249,980,994]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
and 
<emphasis box="[321,334,970,993]" italics="true" pageId="5" pageNumber="6">p</emphasis>
<subScript attach="left" box="[334,343,980,994]" fontSize="6" pageId="5" pageNumber="6">3</subScript>
(57–61) setiform, barbed. Opisthonotal gland opening, notogastral lyrifissures 
<emphasis box="[353,374,1000,1023]" italics="true" pageId="5" pageNumber="6">ia</emphasis>
, 
<emphasis box="[388,415,1000,1023]" italics="true" pageId="5" pageNumber="6">im</emphasis>
, 
<emphasis box="[429,450,1000,1023]" italics="true" pageId="5" pageNumber="6">ip</emphasis>
, 
<emphasis box="[464,485,1000,1024]" italics="true" pageId="5" pageNumber="6">ih</emphasis>
and 
<emphasis box="[539,570,1000,1023]" italics="true" pageId="5" pageNumber="6">ips</emphasis>
and circumgastric sigillar band all distinct in trans-
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,358,1030,1054]" pageId="5" pageNumber="6">mitted light.</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
Gnathosoma. Typical for 
<taxonomicName authority="(Norton &amp; Ermilov 2021)" baseAuthorityName="Norton &amp; Ermilov" baseAuthorityYear="2021" box="[544,967,1061,1085]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis box="[544,676,1061,1085]" italics="true" pageId="5" pageNumber="6">Epidamaeus</emphasis>
(Norton &amp; Ermilov 2021)
</taxonomicName>
. Subcapitulum size: 110–123 × 73–77. Subcapitular setae (
<emphasis box="[718,731,1093,1116]" italics="true" pageId="5" pageNumber="6">a</emphasis>
: 28–32; 
<emphasis box="[834,853,1093,1116]" italics="true" pageId="5" pageNumber="6">m</emphasis>
and 
<emphasis box="[913,926,1092,1116]" italics="true" pageId="5" pageNumber="6">h</emphasis>
: 49–53) setiform,
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
slightly barbed. Adoral seta (12) setiform, straight, roughened. Palp (94–102) with the typical formula: 0-2-1-3-9(+ω); ω baculiform, appressed to surface. Postpalpal seta (8) spiniform, roughened. Chelicera (110–123) with two setiform setae; 
<emphasis box="[1088,1127,1184,1208]" italics="true" pageId="5" pageNumber="6">cha</emphasis>
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,868,1214,1238]" pageId="5" pageNumber="6">
(41) barbed, 
<emphasis box="[364,402,1214,1238]" italics="true" pageId="5" pageNumber="6">chb</emphasis>
(28) unilaterally ciliate in mediodistal part.
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
Epimeral and lateral podosomal regions. Tectum of podocephalic fossa normally developed, without lateral tooth-like projection. Medial coxisternal pit present on epimere I, shallow, with vaguely defined margins. Enantiophyses 
<emphasis box="[974,1003,1306,1330]" italics="true" pageId="5" pageNumber="6">E2</emphasis>
and 
<emphasis box="[1066,1082,1307,1330]" italics="true" pageId="5" pageNumber="6">V</emphasis>
are absent. Parastigmatic enantiophysis strongly dimorphic: 
<emphasis box="[866,893,1337,1360]" italics="true" pageId="5" pageNumber="6">Sa</emphasis>
long, thorn-like; 
<emphasis box="[1100,1127,1337,1360]" italics="true" pageId="5" pageNumber="6">Sp</emphasis>
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
tubercle-like, distally rounded, directed nearly perpendicular to 
<emphasis box="[937,964,1368,1391]" italics="true" pageId="5" pageNumber="6">Sa</emphasis>
. Mediosejugal enantiophysis distinct, just anterior to genital aperture; tubercles simple, 
<emphasis box="[1020,1056,1399,1422]" italics="true" pageId="5" pageNumber="6">Ma</emphasis>
larger than 
<emphasis box="[281,316,1430,1453]" italics="true" pageId="5" pageNumber="6">Mp</emphasis>
. Epimeral setal formula: 3-1-3-4. All setae (
<emphasis box="[803,830,1429,1453]" italics="true" pageId="5" pageNumber="6">1a</emphasis>
, 
<emphasis box="[845,872,1429,1453]" italics="true" pageId="5" pageNumber="6">2a</emphasis>
, 
<emphasis box="[887,914,1429,1453]" italics="true" pageId="5" pageNumber="6">3a</emphasis>
: 32–36; 
<emphasis box="[1012,1037,1429,1453]" italics="true" pageId="5" pageNumber="6">1c</emphasis>
: 24–32;
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,1127,1460,1484]" pageId="5" pageNumber="6">
others: 57–61) setiform, slightly barbed; 
<emphasis box="[674,699,1460,1484]" italics="true" pageId="5" pageNumber="6">1c</emphasis>
inserted on small but distinct tubercle;
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" pageId="5" pageNumber="6">
<emphasis box="[227,252,1490,1514]" italics="true" pageId="5" pageNumber="6">3c</emphasis>
removed medially from tubercle 
<emphasis box="[617,644,1491,1514]" italics="true" pageId="5" pageNumber="6">Sp</emphasis>
and acetabulum III. Discidium poorly developed, tubercle-like.
</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[265,1127,1552,1576]" pageId="5" pageNumber="6">Anogenital region. Genital, aggenital, anal and adanal setae similar in length (41–</paragraph>
<paragraph blockId="5.[227,1128,264,1606]" box="[227,963,1582,1606]" pageId="5" pageNumber="6">45), setiform, slightly barbed. Adanal lyrifissure oriented diagonally.</paragraph>
<caption pageId="6" pageNumber="7" startId="6.[227,288,264,288]">
<paragraph blockId="6.[227,1123,264,319]" pageId="6" pageNumber="7">
Table 1. Leg mean lengths (micrometers) and their ratio to body mean length of one paratype 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[363,686,294,318]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis box="[363,686,294,318]" italics="true" pageId="6" pageNumber="7">Epidamaeus hexatuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[692,749,295,318]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="7">
<table box="[227,1106,342,479]" gridcols="8" gridrows="5" pageId="6" pageNumber="7" rowsContinueIn="6.[227,1091,870,1119]">
<tr box="[227,1106,342,363]" gridrow="0" pageId="6" pageNumber="7">
<th box="[227,266,342,363]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[227,266,342,363]" pageId="6" pageNumber="7">Leg</emphasis>
</th>
<th box="[313,349,342,363]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[319,343,342,363]" italics="true" pageId="6" pageNumber="7">Tr</emphasis>
</th>
<th box="[407,443,342,363]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[412,439,342,363]" italics="true" pageId="6" pageNumber="7">Fe</emphasis>
</th>
<th box="[506,534,342,363]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[506,534,342,363]" italics="true" pageId="6" pageNumber="7">Ge</emphasis>
</th>
<th box="[596,633,342,363]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[604,625,342,363]" italics="true" pageId="6" pageNumber="7">Ti</emphasis>
</th>
<th box="[691,727,342,363]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[696,723,342,363]" italics="true" pageId="6" pageNumber="7">Ta</emphasis>
</th>
<th box="[785,821,342,363]" gridcol="6" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[788,819,342,363]" pageId="6" pageNumber="7">All</emphasis>
</th>
<th box="[865,1106,342,363]" gridcol="7" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[865,1106,342,363]" pageId="6" pageNumber="7">Leg: body mean length</emphasis>
</th>
</tr>
<tr box="[227,1106,370,392]" gridrow="1" pageId="6" pageNumber="7">
<th box="[227,266,370,392]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">I</th>
<td box="[313,349,370,392]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">20</td>
<td box="[407,443,370,392]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">184</td>
<td box="[506,534,370,392]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">69</td>
<td box="[596,633,370,392]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">102</td>
<td box="[691,727,370,392]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">192</td>
<td box="[785,821,370,392]" gridcol="6" gridrow="1" pageId="6" pageNumber="7">567</td>
<td box="[865,1106,370,392]" gridcol="7" gridrow="1" pageId="6" pageNumber="7">1.13</td>
</tr>
<tr box="[227,1106,399,421]" gridrow="2" pageId="6" pageNumber="7">
<th box="[227,266,399,421]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">II</th>
<td box="[313,349,399,421]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">20</td>
<td box="[407,443,399,421]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">139</td>
<td box="[506,534,399,421]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">61</td>
<td box="[596,633,399,421]" gridcol="4" gridrow="2" pageId="6" pageNumber="7">82</td>
<td box="[691,727,399,421]" gridcol="5" gridrow="2" pageId="6" pageNumber="7">164</td>
<td box="[785,821,399,421]" gridcol="6" gridrow="2" pageId="6" pageNumber="7">466</td>
<td box="[865,1106,399,421]" gridcol="7" gridrow="2" pageId="6" pageNumber="7">0.93</td>
</tr>
<tr box="[227,1106,428,450]" gridrow="3" pageId="6" pageNumber="7">
<th box="[227,266,428,450]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">III</th>
<td box="[313,349,428,450]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">94</td>
<td box="[407,443,428,450]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">114</td>
<td box="[506,534,428,450]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">61</td>
<td box="[596,633,428,450]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">82</td>
<td box="[691,727,428,450]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">151</td>
<td box="[785,821,428,450]" gridcol="6" gridrow="3" pageId="6" pageNumber="7">502</td>
<td box="[865,1106,428,450]" gridcol="7" gridrow="3" pageId="6" pageNumber="7">1.00</td>
</tr>
<tr box="[227,1106,457,479]" gridrow="4" pageId="6" pageNumber="7">
<th box="[227,266,457,479]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">IV</th>
<td box="[313,349,457,479]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">114</td>
<td box="[407,443,457,479]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">127</td>
<td box="[506,534,457,479]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">73</td>
<td box="[596,633,457,479]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">114</td>
<td box="[691,727,457,479]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">205</td>
<td box="[785,821,457,479]" gridcol="6" gridrow="4" pageId="6" pageNumber="7">633</td>
<td box="[865,1106,457,479]" gridcol="7" gridrow="4" pageId="6" pageNumber="7">1.27</td>
</tr>
</table>
</paragraph>
<paragraph blockId="6.[227,997,502,524]" box="[227,997,502,524]" pageId="6" pageNumber="7">
<tableNote box="[227,997,502,524]" pageId="6" pageNumber="7" targetBox="[227,1106,342,479]" targetPageId="6">Note: Basal part of each segment included in previous segment is not measured.</tableNote>
</paragraph>
<paragraph blockId="6.[227,1127,547,787]" pageId="6" pageNumber="7">
Legs. Leg III similar to body in length, leg II shorter than body length, legs I and IV longer than body length, leg IV longest; approximate length of leg segments shown in 
<tableCitation box="[334,417,608,632]" captionStart="Table 1" captionStartId="6.[227,288,264,288]" captionText="Table 1. Leg mean lengths (micrometers) and their ratio to body mean length of one paratype Epidamaeus hexatuberculatus sp. n." pageId="6" pageNumber="7">Table 1</tableCitation>
. Claw of each tarsus smooth. Porose area on femora I-IV and on trochanters III and IV distinctly visible. Formulas of leg setation and solenidia: I (1-
</paragraph>
<paragraph blockId="6.[227,1127,547,787]" box="[227,1127,669,693]" pageId="6" pageNumber="7">7-4-4-21) [1-2-2], II (1-6-4-4-18) [1-1-2], III (2-4-3-3-18) [1-1-0], IV (1-4-3-3-15)</paragraph>
<paragraph blockId="6.[227,1127,547,787]" box="[227,1127,700,725]" pageId="6" pageNumber="7">
[0-1-0]; homology of setae and solenidia indicated in 
<tableCitation box="[809,893,700,724]" captionStart="Table" captionStartId="6.[227,291,823,847]" captionText="Table. 2. Leg setation and solenidia of adult Epidamaeus hexatuberculatus sp. n." pageId="6" pageNumber="7">Table 2</tableCitation>
. Solenidion on genua
</paragraph>
<paragraph blockId="6.[227,1127,547,787]" pageId="6" pageNumber="7">
I-III shorter than coupled setae 
<emphasis box="[592,605,731,755]" italics="true" pageId="6" pageNumber="7">d</emphasis>
. Famulus of tarsus I short, setiform, inserted between solenidia ω 
<subScript attach="left" box="[443,452,773,787]" fontSize="6" pageId="6" pageNumber="7">1</subScript>
and ω 
<subScript attach="left" box="[521,530,773,787]" fontSize="6" pageId="6" pageNumber="7">2</subScript>
. Seta 
<emphasis box="[596,608,762,785]" italics="true" pageId="6" pageNumber="7">v</emphasis>
<subScript attach="left" box="[607,616,773,787]" fontSize="6" pageId="6" pageNumber="7">2</subScript>
<emphasis box="[616,625,762,785]" italics="true" pageId="6" pageNumber="7">’</emphasis>
present on all tarsi.
</paragraph>
</subSubSection>
<caption box="[227,1089,823,847]" pageId="6" pageNumber="7" startId="6.[227,291,823,847]">
<paragraph blockId="6.[227,1089,823,847]" box="[227,1089,823,847]" pageId="6" pageNumber="7">
Table. 2. Leg setation and solenidia of adult 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[702,1025,823,847]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis box="[702,1025,823,847]" italics="true" pageId="6" pageNumber="7">Epidamaeus hexatuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1031,1089,823,846]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="7">
<table box="[227,1091,870,1119]" gridcols="6" gridrows="5" pageId="6" pageNumber="7" rowsContinueFrom="6.[227,1106,342,479]">
<tr box="[227,1091,870,892]" gridrow="0" pageId="6" pageNumber="7">
<th box="[227,266,870,892]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[227,266,870,891]" pageId="6" pageNumber="7">Leg</emphasis>
</th>
<th box="[294,340,870,892]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[294,318,871,892]" italics="true" pageId="6" pageNumber="7">Tr</emphasis>
</th>
<th box="[371,482,870,892]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[371,398,871,892]" italics="true" pageId="6" pageNumber="7">Fe</emphasis>
</th>
<th box="[527,618,870,892]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[527,555,870,891]" italics="true" pageId="6" pageNumber="7">Ge</emphasis>
</th>
<th box="[662,791,870,892]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[662,683,870,891]" italics="true" pageId="6" pageNumber="7">Ti</emphasis>
</th>
<th box="[821,1091,870,892]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">
<emphasis bold="true" box="[821,848,871,892]" italics="true" pageId="6" pageNumber="7">Ta</emphasis>
</th>
</tr>
<tr box="[227,1091,899,950]" gridrow="1" pageId="6" pageNumber="7">
<th box="[227,266,899,950]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">I</th>
<td box="[294,340,899,950]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">
<emphasis box="[294,313,913,934]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
</td>
<td box="[371,482,899,950]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">d, (l), bv”, (v</emphasis>
1
<emphasis box="[397,428,927,948]" italics="true" pageId="6" pageNumber="7">), v</emphasis>
2 
<emphasis box="[436,444,927,948]" italics="true" pageId="6" pageNumber="7">’</emphasis>
</td>
<td box="[527,618,899,950]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">
<emphasis box="[527,605,912,934]" italics="true" pageId="6" pageNumber="7">(l), v’, d</emphasis>
σ
</td>
<td box="[662,791,899,950]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">
<emphasis box="[662,729,912,934]" italics="true" pageId="6" pageNumber="7">(l), (v),</emphasis>
φ1, φ2
</td>
<td box="[821,1091,899,950]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">(ft), (tc), (it), (p), (u), (a), s, (pv), (pl), (v</emphasis>
1
<emphasis box="[945,976,927,948]" italics="true" pageId="6" pageNumber="7">), v</emphasis>
2 
<emphasis box="[984,998,927,948]" italics="true" pageId="6" pageNumber="7">’,</emphasis>
ɛ, ω1, ω2
</td>
</tr>
<tr box="[227,1091,955,1006]" gridrow="2" pageId="6" pageNumber="7">
<th box="[227,266,955,1006]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">II</th>
<td box="[294,340,955,1006]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">
<emphasis box="[294,313,969,990]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
</td>
<td box="[371,482,955,1006]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">d, (l), bv”, (v)</emphasis>
</td>
<td box="[527,618,955,1006]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">
<emphasis box="[527,605,969,990]" italics="true" pageId="6" pageNumber="7">(l), v’, d</emphasis>
σ
</td>
<td box="[662,791,955,1006]" gridcol="4" gridrow="2" pageId="6" pageNumber="7">
<emphasis box="[662,729,969,990]" italics="true" pageId="6" pageNumber="7">(l), (v),</emphasis>
φ
</td>
<td box="[821,1091,955,1006]" gridcol="5" gridrow="2" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">(ft), (tc), (it), (p), (u), (a), s, (pv), (v</emphasis>
1
<emphasis box="[898,929,983,1004]" italics="true" pageId="6" pageNumber="7">), v</emphasis>
2 
<emphasis box="[937,951,983,1004]" italics="true" pageId="6" pageNumber="7">’,</emphasis>
ω1, ω2
</td>
</tr>
<tr box="[227,1091,1012,1063]" gridrow="3" pageId="6" pageNumber="7">
<th box="[227,266,1012,1063]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">III</th>
<td box="[294,340,1012,1063]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">
<emphasis box="[294,340,1025,1047]" italics="true" pageId="6" pageNumber="7">l’, v’</emphasis>
</td>
<td box="[371,482,1012,1063]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">
<emphasis box="[371,482,1025,1047]" italics="true" pageId="6" pageNumber="7">d, l’, ev’, v’</emphasis>
</td>
<td box="[527,618,1012,1063]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">
<emphasis box="[527,597,1025,1047]" italics="true" pageId="6" pageNumber="7">l’, v’, d</emphasis>
σ
</td>
<td box="[662,791,1012,1063]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">
<emphasis box="[662,721,1025,1047]" italics="true" pageId="6" pageNumber="7">l’, (v),</emphasis>
φ
</td>
<td box="[821,1091,1012,1063]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">(ft), (tc), (it), (p), (u), (a), s, (pv), (v</emphasis>
1
<emphasis box="[898,929,1040,1061]" italics="true" pageId="6" pageNumber="7">), v</emphasis>
2 
<emphasis box="[937,945,1040,1061]" italics="true" pageId="6" pageNumber="7">’</emphasis>
</td>
</tr>
<tr box="[227,1091,1068,1119]" gridrow="4" pageId="6" pageNumber="7">
<th box="[227,266,1068,1119]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">IV</th>
<td box="[294,340,1068,1119]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">
<emphasis box="[294,313,1082,1103]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
</td>
<td box="[371,482,1068,1119]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">
<emphasis box="[371,482,1082,1103]" italics="true" pageId="6" pageNumber="7">d, l’, ev’, v’</emphasis>
</td>
<td box="[527,618,1068,1119]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">
<emphasis box="[527,597,1082,1103]" italics="true" pageId="6" pageNumber="7">d, l’, v’</emphasis>
</td>
<td box="[662,791,1068,1119]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">
<emphasis box="[662,721,1082,1103]" italics="true" pageId="6" pageNumber="7">l’, (v),</emphasis>
φ
</td>
<td box="[821,1091,1068,1119]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">
<emphasis italics="true" pageId="6" pageNumber="7">ft”, (tc), (p), (u), (a), s, (pv), (v</emphasis>
1
<emphasis box="[848,879,1096,1117]" italics="true" pageId="6" pageNumber="7">), v</emphasis>
2’
</td>
</tr>
</table>
</paragraph>
<tableNote pageId="6" pageNumber="7" targetBox="[227,1091,870,1119]" targetPageId="6">
<paragraph blockId="6.[227,1127,1141,1245]" box="[265,1126,1141,1163]" pageId="6" pageNumber="7">Note: Roman letters refer to normal setae, Greek letters to solenidia (except ɛ = famulus),</paragraph>
<paragraph blockId="6.[227,1127,1141,1245]" pageId="6" pageNumber="7">
<emphasis box="[227,239,1168,1189]" italics="true" pageId="6" pageNumber="7">d</emphasis>
σ – seta and solenidion coupled. Single prime (
<emphasis box="[714,722,1169,1190]" italics="true" pageId="6" pageNumber="7">’</emphasis>
) marks setae on the anterior and double prime (
<emphasis box="[299,312,1196,1217]" italics="true" pageId="6" pageNumber="7">”</emphasis>
) setae on the posterior side of a given leg segment. Parentheses refer to a pair of setae.
</paragraph>
</tableNote>
<subSubSection pageId="6" pageNumber="7" type="etymology">
<paragraph blockId="6.[227,1127,1268,1599]" pageId="6" pageNumber="7">
ETYMOLOGY. The species name refers to the presence of six prodorsobasal tubercles (paired 
<emphasis box="[412,445,1300,1323]" italics="true" pageId="6" pageNumber="7">Da</emphasis>
, 
<emphasis box="[458,488,1300,1323]" italics="true" pageId="6" pageNumber="7">Ba</emphasis>
and 
<emphasis box="[539,569,1300,1323]" italics="true" pageId="6" pageNumber="7">Bp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph blockId="6.[227,1127,1268,1599]" pageId="6" pageNumber="7">
REMARKS. 
<taxonomicName authority="Ermilov &amp; Salavatulin, 2022" authorityName="Ermilov &amp; Salavatulin" authorityYear="2022" box="[417,753,1329,1353]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hexatuberculatus" status="sp. n.">
<emphasis box="[417,753,1329,1353]" italics="true" pageId="6" pageNumber="7">Epidamaeus hexatuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[767,833,1330,1353]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
is morphologically most similar to 
<emphasis box="[341,562,1360,1384]" italics="true" pageId="6" pageNumber="7">
<taxonomicName baseAuthorityName="Norton &amp; Ermilov" baseAuthorityYear="2021" box="[341,473,1360,1384]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Epidamaeus</taxonomicName>
nortoni
</emphasis>
Bayartogtokh, 2004 from 
<collectingCountry box="[857,961,1360,1384]" name="Mongolia" pageId="6" pageNumber="7">Mongolia</collectingCountry>
and 
<taxonomicName box="[1018,1127,1360,1384]" pageId="6" pageNumber="7">
<emphasis box="[1018,1127,1360,1384]" italics="true" pageId="6" pageNumber="7">E. tritylos</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[227,1127,1268,1599]" pageId="6" pageNumber="7">
Behan-Pelletier et Norton, 1983 from the Palaearctic region in having three pairs of prodorsobasal tubercles and phylliform dorsal notogastral setae. However, the new species differs from both by the absence (
<emphasis box="[662,684,1453,1476]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
. presence) of enantiophyses 
<emphasis box="[985,1015,1452,1476]" italics="true" pageId="6" pageNumber="7">E2</emphasis>
and 
<emphasis box="[1066,1082,1453,1476]" italics="true" pageId="6" pageNumber="7">V</emphasis>
, the presence (
<emphasis box="[336,358,1484,1507]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
. absence) of mediosejugal enantiophysis, narrowly phylliform (
<emphasis box="[1045,1067,1484,1507]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
. setiform) interlamellar seta and tubercle-like, rounded distally (
<emphasis box="[890,912,1514,1537]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
. triangular, pointed distally) discidium, and the localization of prodorsobasal tubercle 
<emphasis box="[974,1007,1545,1568]" italics="true" pageId="6" pageNumber="7">Da</emphasis>
(distinctly anteromedial to 
<emphasis box="[404,462,1576,1599]" italics="true" pageId="6" pageNumber="7">Ba vs</emphasis>
. medial to 
<emphasis box="[585,615,1576,1599]" italics="true" pageId="6" pageNumber="7">Ba</emphasis>
). Also, 
<emphasis box="[703,812,1575,1599]" italics="true" pageId="6" pageNumber="7">E. nortoni</emphasis>
has four pairs (
<emphasis box="[983,1006,1576,1599]" italics="true" pageId="6" pageNumber="7">vs</emphasis>
. one pair)
</paragraph>
<paragraph blockId="7.[227,1127,264,319]" pageId="7" pageNumber="8">
of setae on the epimere II and does not have a parastigmatic tubercle 
<emphasis box="[998,1025,264,287]" italics="true" pageId="7" pageNumber="8">Sa</emphasis>
(
<emphasis box="[1043,1065,265,288]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. well developed); 
<taxonomicName authorityName="Behan-Pelletier et Norton" authorityYear="1983" box="[360,464,294,318]" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tritylos">
<emphasis box="[360,464,294,318]" italics="true" pageId="7" pageNumber="8">E. tritylos</emphasis>
</taxonomicName>
has bacilliform (
<emphasis box="[647,669,295,318]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. rod-like) bothridial seta.
</paragraph>
</subSubSection>
</treatment>
</document>