214 lines
31 KiB
XML
214 lines
31 KiB
XML
<document id="6C300C549777EE76E65310558DFAF5F3" ID-DOI="10.2478/if-2019-0030" ID-ISSN="2533-4069" ID-Zenodo-Dep="4586737" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614972615996" checkinUser="felipe" docAuthor="Martin, Thomas, Averianov, Alexander O., Jäger, Kai R. K., Schwermann, Achim H. & Wings, Oliver" docDate="2019" docId="03F73338FF8C7657CEF190FCFD187642" docLanguage="en" docName="FossilImprint.75.3-4.504-509.pdf.imf" docOrigin="Fossil Imprint 75 (3 - 4)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Storchodon cingulatus Martin & Averianov & Jäger & Schwermann & Oliver & Wings 2019, sp. nov." docType="treatment" docVersion="3" lastPageNumber="507" masterDocId="FFCE4B40FF8D7654CE0F9737FFF8775C" masterDocTitle="A Large Morganucodontan Mammaliaform From The Late Jurassic Of Germany" masterLastPageNumber="509" masterPageNumber="504" pageNumber="505" updateTime="1698881601685" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:titleInfo id="23105B5D034335942190381E0FE026A8">
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<mods:title id="9405373C90B1FB044FBE93C73FD2239B">A Large Morganucodontan Mammaliaform From The Late Jurassic Of Germany</mods:title>
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<mods:name id="656BB515A6F4393B8FD41F43AFB775D9" type="personal">
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<mods:namePart id="FD7010E71B3E66E98A18695542CE4FC5">Martin, Thomas</mods:namePart>
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<mods:affiliation id="30B3CC6A13408E66CABE80815A1CA4A0">Section Paleontology, Institute of Geosciences, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany;</mods:affiliation>
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<mods:namePart id="5D94EC44144AF98A660971AB1E18BB2E">Averianov, Alexander O.</mods:namePart>
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<mods:affiliation id="577F6C1F5045C204BE6AC03AE6CD655D">Section Paleontology, Institute of Geosciences, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany; & Zoological Institute of the Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia. & Department of Sedimentary Geology, Institute of Earth Sciences, Saint Petersburg State University, 16 th Liniya V. O., 29, St. Petersburg,</mods:affiliation>
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<mods:roleTerm id="B1A68691B74517F25DDBBE8FD3A41030">Author</mods:roleTerm>
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<mods:namePart id="C8638E6C5BD3189B43FDEF8EF02BCA98">Jäger, Kai R. K.</mods:namePart>
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<mods:affiliation id="E5C05763123C1DCA1CFB364852FC8488">Section Paleontology, Institute of Geosciences, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany;</mods:affiliation>
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</mods:name>
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<mods:namePart id="654CD327590FCEDEF6C89CFB09F7ACFA">Schwermann, Achim H.</mods:namePart>
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<mods:affiliation id="28A0C524083DD48E1E52EBE6AB74E9CA">LWL-Museum fÜr Naturkunde, Westfälisches Landesmuseum mit Planetarium, Sentruper Strasse 285, 48161 MÜnster, Germany.</mods:affiliation>
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<mods:namePart id="732B08EBE61CC2DD84CA834C946ED6F7">Wings, Oliver</mods:namePart>
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<mods:affiliation id="BE2DFF5A89DC2F2E537167BA3F5C49DD">Martin-Luther-Universität Halle-Wittenberg, Zentralmagazin Naturwissenschaftlicher Sammlungen, Domplatz 4, 06108 Halle (Saale),</mods:affiliation>
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<mods:typeOfResource id="226BD776C005A55CE6CB2310F9359058">text</mods:typeOfResource>
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<mods:title id="7B52FE3E652C278BCCA54AE81F6B578C">Fossil Imprint</mods:title>
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<mods:part id="326E477D997F8F3A30E81E261B523A15">
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<mods:date id="4E3844208AD985A3B5D8BBE397174CC5">2019</mods:date>
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<mods:number id="405957BA5EE09F169CE647561606EFFA">2019-12-30</mods:number>
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<mods:number id="6082B71081FC3AAB564162D974D2F185">75</mods:number>
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<mods:number id="E1E60D0C2528FF44B969809BBA955877">3 - 4</mods:number>
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<mods:identifier id="DA70D8E4976EA16DB4974BA9E7E83FD7" type="DOI">10.2478/if-2019-0030</mods:identifier>
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<mods:identifier id="5D64AA812D1996974F197F0662AF14C4" type="ISSN">2533-4069</mods:identifier>
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<treatment id="03F73338FF8C7657CEF190FCFD187642" LSID="urn:lsid:plazi:treatment:03F73338FF8C7657CEF190FCFD187642" httpUri="http://treatment.plazi.org/id/03F73338FF8C7657CEF190FCFD187642" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505">
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<subSubSection id="C344D1A5FF8C7655CEF190FCFDAC70B8" box="[254,596,1995,2020]" pageId="1" pageNumber="505" type="nomenclature">
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<paragraph id="8BE1822EFF8C7655CEF190FCFDAC70B8" blockId="1.[254,596,1995,2020]" box="[254,596,1995,2020]" pageId="1" pageNumber="505">
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<heading id="D0A93542FF8C7655CEF190FCFDAC70B8" box="[254,596,1995,2020]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="505" reason="2">
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<emphasis id="B92A5E3CFF8C7655CEF190FCFDAC70B8" bold="true" box="[254,596,1995,2020]" pageId="1" pageNumber="505">
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<taxonomicName id="4C5EF9ADFF8C7655CEF190FCFE0070BF" authority="Martin & Averianov & Jäger & Schwermann & Oliver & Wings, 2019" authorityName="Martin & Averianov & Jäger & Schwermann & Oliver & Wings" authorityYear="2019" box="[254,504,1995,2019]" class="Mammalia" family="Triconodontidae" genus="Storchodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="1" pageNumber="505" phylum="Chordata" rank="species" species="cingulatus" status="sp. nov.">
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<emphasis id="B92A5E3CFF8C7655CEF190FCFE0070BF" bold="true" box="[254,504,1995,2019]" italics="true" pageId="1" pageNumber="505">Storchodon cingulatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A219E347FF8C7655CFF090FBFDAC70B8" box="[511,596,1996,2020]" pageId="1" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C344D1A5FF8C7657CF6C90DBFD187642" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505" type="description">
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<paragraph id="8BE1822EFF8C7655CF6C90DBFE177F58" blockId="1.[355,495,2028,2052]" box="[355,495,2028,2052]" pageId="1" pageNumber="505">
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<figureCitation id="13659EABFF8C7655CF6C90DBFE2F7F58" box="[355,471,2028,2052]" captionStart="Text-fig" captionStartId="2.[147,228,606,627]" captionTargetBox="[157,1450,184,575]" captionTargetId="figure-621@2.[147,1462,58,587]" captionTargetPageId="2" captionText="Text-fig. 2. Storchodon cingulatus gen. et sp. nov., NLMH 105654, holotype, right upper molar, in buccal (a), occlusal (b, b’ stereopair), lingual (c), and basal (d) views. Langenberg Quarry, Lower Saxony, Germany; Süntel Formation, Late Jurassic (late Kimmeridgian) age. Abbreviations: A, B, C and D, main cusps of the crown; ic, interradical crest." figureDoi="http://doi.org/10.5281/zenodo.4586741" httpUri="https://zenodo.org/record/4586741/files/figure.png" pageId="1" pageNumber="505">Text-figs 2</figureCitation>
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,
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<figureCitation id="13659EABFF8C7655CFED90DBFE177F58" box="[482,495,2028,2052]" captionStart="Text-fig" captionStartId="2.[129,210,1960,1981]" captionTargetBox="[141,751,1426,1921]" captionTargetId="figure-722@2.[400,673,1589,1760]" captionTargetPageId="2" captionText="Text-fig. 3. Right upper molars of Morganucodon watsoni KÜHNE, 1949, UMZC_Eo.M(Ur).27 (cast) (a) and Storchodon cingulatus gen. et sp. nov., NLMH 105654, holotype (b) in lingual and occlusal views. Abbreviations: A, B, C, D, E and F, main cusps of the crown. Specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.4586743" httpUri="https://zenodo.org/record/4586743/files/figure.png" pageId="1" pageNumber="505">3</figureCitation>
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</paragraph>
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<paragraph id="8BE1822EFF8C7655CE9F9F2BFD497F68" blockId="1.[144,689,2076,2100]" box="[144,689,2076,2100]" pageId="1" pageNumber="505">H o l o t y p e. NLMH 105654, a right upper molar.</paragraph>
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<paragraph id="8BE1822EFF8C7655CD559126FB2E7115" blockId="1.[820,1459,1552,1609]" pageId="1" pageNumber="505">E t y m o l o g y. The species name refers to the welldeveloped cingulum around the crown.</paragraph>
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<paragraph id="8BE1822EFF8C7655CD559155FAC17125" blockId="1.[858,1337,1633,1657]" box="[858,1337,1633,1657]" pageId="1" pageNumber="505">
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<materialsCitation id="3B368873FF8C7655CD559155FAC17125" box="[858,1337,1633,1657]" collectionCode="R" pageId="1" pageNumber="505" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED4F1AEBFF8C7655CD559155FC947125" box="[858,876,1634,1657]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="505">R</collectionCode>
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e f e r r e d s p e c i m e n s.
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<typeStatus id="54E53C8CFF8C7655CA959156FB007125" box="[1178,1272,1633,1657]" pageId="1" pageNumber="505" type="holotype">Holotype</typeStatus>
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only.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE1822EFF8C7655CD5591A5FAB7702D" blockId="1.[820,1459,1681,1905]" pageId="1" pageNumber="505">
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<materialsCitation id="3B368873FF8C7655CD5591A5FAB3702D" collectionCode="T" country="Germany" latitude="51.901833" location="Langenberg Quarry" longLatPrecision="1" longitude="10.508333" municipality="Goslar" pageId="1" pageNumber="505" specimenCount="1" stateProvince="Niedersachsen">
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<collectionCode id="ED4F1AEBFF8C7655CD5591A5FC9271F5" box="[858,874,1682,1705]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="505">T</collectionCode>
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y p e l o c a l i t y a n d h o r i z o n.
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<location id="8E81D4F5FF8C7655CB3B91A6FC787197" LSID="urn:lsid:plazi:treatment:03F73338FF8C7657CEF190FCFD187642:8E81D4F5FF8C7655CB3B91A6FC787197" country="Germany" latitude="51.901833" longLatPrecision="1" longitude="10.508333" municipality="Goslar" name="Langenberg Quarry" pageId="1" pageNumber="505" stateProvince="Niedersachsen">Langenberg Quarry</location>
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near the town of
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<collectingMunicipality id="6B851854FF8C7655CA5F9184FB617197" box="[1104,1177,1715,1739]" pageId="1" pageNumber="505">Goslar</collectingMunicipality>
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,
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<collectingRegion id="499A4CCCFF8C7655CAA79184FAB27197" box="[1192,1354,1715,1739]" country="Germany" name="Niedersachsen" pageId="1" pageNumber="505">Lower Saxony</collectingRegion>
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, northern
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<collectingCountry id="F349C2BEFF8C7655CD3B91E3FC6F71B0" box="[820,919,1748,1772]" name="Germany" pageId="1" pageNumber="505">Germany</collectingCountry>
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(
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<geoCoordinate id="EE6AE4E9FF8C7655CDA391E3FBA871B7" box="[940,1104,1746,1772]" degrees="51" direction="north" minutes="54.110" orientation="latitude" pageId="1" pageNumber="505" precision="1" value="51.901833">N 51° 54.110′</geoCoordinate>
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,
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<geoCoordinate id="EE6AE4E9FF8C7655CA6F91E3FAFB71B7" box="[1120,1283,1746,1772]" degrees="10" direction="east" minutes="30.500" orientation="longitude" pageId="1" pageNumber="505" precision="1" value="10.508333">E 10° 30.500′</geoCoordinate>
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). Bed 83 (after
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<bibRefCitation id="EFCFFFDFFF8C7655CD3B91C2FC3E7051" author="Fischer, R." box="[820,966,1781,1805]" pageId="1" pageNumber="505" pagination="21 - 36" refId="ref3873" refString="Fischer, R. (1991): Die Oberjura-Schichtenfolge des Lan- genbergs bei Oker. - Arbeitskreis Palaontologie Hanno- ver, 19: 21 - 36." type="journal article" year="1991">Fischer 1991</bibRefCitation>
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). The sample was collected in a temporarily exposed dark grey lens of marl within the normally light grey-greenish marly limestone bed. It belongs to the Süntel Formation, Late Jurassic (late Kimmeridgian) age
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE1822EFF8C7655CD5590BEFA8170FD" blockId="1.[858,1401,1928,1953]" box="[858,1401,1928,1953]" pageId="1" pageNumber="505">D i f f e r e n t i a l d i a g n o s i s.As for the genus.</paragraph>
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<paragraph id="8BE1822EFF8C7657CD55908EFD9677B3" blockId="1.[820,1460,1976,2100]" lastBlockId="3.[106,745,151,240]" lastPageId="3" lastPageNumber="507" pageId="1" pageNumber="505">
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D e s c r i p t i o n. The presence of a continuous cingulum around the crown identifies the tooth as an upper molar (
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<figureCitation id="13659EABFF8C7655CD8E90CCFC0C7F4F" box="[897,1012,2043,2067]" captionStart="Text-fig" captionStartId="2.[147,228,606,627]" captionTargetBox="[157,1450,184,575]" captionTargetId="figure-621@2.[147,1462,58,587]" captionTargetPageId="2" captionText="Text-fig. 2. Storchodon cingulatus gen. et sp. nov., NLMH 105654, holotype, right upper molar, in buccal (a), occlusal (b, b’ stereopair), lingual (c), and basal (d) views. Langenberg Quarry, Lower Saxony, Germany; Süntel Formation, Late Jurassic (late Kimmeridgian) age. Abbreviations: A, B, C and D, main cusps of the crown; ic, interradical crest." figureDoi="http://doi.org/10.5281/zenodo.4586741" httpUri="https://zenodo.org/record/4586741/files/figure.png" pageId="1" pageNumber="505">Text-fig. 2</figureCitation>
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); in lower molars of
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<taxonomicName id="4C5EF9ADFF8C7655CAD990CCFA747F4F" authorityName="KERMACK, MUSSET et RIGNEY" authorityYear="1973" box="[1238,1420,2043,2067]" class="Mammalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="1" pageNumber="505" phylum="Chordata" rank="order">Morganucodonta</taxonomicName>
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the buccal cingulid is absent (
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<bibRefCitation id="EFCFFFDFFF8C7655CA5E9F2BFA507F68" author="Kielan-Jaworowska, Z. & Cifelli, R. L. & Luo, Z. - X." box="[1105,1448,2076,2100]" pageId="1" pageNumber="505" refId="ref4360" refString="Kielan-Jaworowska, Z., Cifelli, R. L., Luo, Z. - X. (2004): Mammals from the Age of Dinosaurs: Origins, Evolu- tion, and Structure. Columbia University Press, New York, 630 pp. https: // doi. org / 10.7312 / kiel 11918" type="book" year="2004">Kielan-Jaworowska et al. 2004</bibRefCitation>
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). The three main cusps A, B, and C are aligned serially. Cusp D is set off this line and sits more lingually compared to cusp C. Thus, the line A-B-C-D is slightly convexly curved in a lingual direction. Cusp A is not completely preserved; its base is only slightly larger than that of cusp C. Cusp A was probably only little higher than cusp C and vertically directed, whereas cusp C is slightly distally oriented. Cusp B has about two thirds the size of cusp C and is slightly mesially directed. The main cusps are connected by a sharp crest which extends to their apices (incompletely preserved on cusp A). This crest is extended on the mesial side of cusp B, as well as on the distal side of cusp C and on both sides of cusp D. The crown is surrounded by a continuous cingulum bearing numerous cusps. There is a distinct valley between the cingulum and the bases of the main cusps on the buccal, lingual, and mesial sides of the crown. Cusp D (incomplete) is more distally directed than cusp C. Cusp D is not part of the cingulum (a unique character of the new taxon within
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<taxonomicName id="4C5EF9ADFF8F7656CD4495C2FBFD7451" authorityName="KERMACK, MUSSET et RIGNEY" authorityYear="1973" box="[843,1029,757,781]" class="Mammalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="order">Morganucodonta</taxonomicName>
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), but is connected distally with a rather large cingular cusp. Only in
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<taxonomicName id="4C5EF9ADFF8F7656CA9F9422FACD7471" authorityName="CROMPTON" authorityYear="1964" box="[1168,1333,789,813]" class="Mammalia" family="Morganucodontidae" genus="Erythrotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
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<emphasis id="B92A5E3CFF8F7656CA9F9422FACD7471" box="[1168,1333,789,813]" italics="true" pageId="2" pageNumber="506">Erythrotherium</emphasis>
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</taxonomicName>
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and in some individuals of
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<taxonomicName id="4C5EF9ADFF8F7656CDE69402FB747411" authorityName="KUHNE" authorityYear="1949" box="[1001,1164,821,845]" class="Mammalia" family="Morganucodontidae" genus="Morganucodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
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<emphasis id="B92A5E3CFF8F7656CDE69402FB747411" box="[1001,1164,821,845]" italics="true" pageId="2" pageNumber="506">Morganucodon</emphasis>
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</taxonomicName>
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is cusp D equally prominent (
|
||
<figureCitation id="13659EABFF8F7656CD5D9461FC3E7432" box="[850,966,854,878]" captionStart="Text-fig" captionStartId="2.[129,210,1960,1981]" captionTargetBox="[141,751,1426,1921]" captionTargetId="figure-722@2.[400,673,1589,1760]" captionTargetPageId="2" captionText="Text-fig. 3. Right upper molars of Morganucodon watsoni KÜHNE, 1949, UMZC_Eo.M(Ur).27 (cast) (a) and Storchodon cingulatus gen. et sp. nov., NLMH 105654, holotype (b) in lingual and occlusal views. Abbreviations: A, B, C, D, E and F, main cusps of the crown. Specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.4586743" httpUri="https://zenodo.org/record/4586743/files/figure.png" pageId="2" pageNumber="506">Text-fig. 3</figureCitation>
|
||
). A relatively large cingular cusp is situated on the mesial end of the crown; it is separated by a groove from cusp B. This mesial cingular cusp possibly corresponds to cusp E of other morganucodontans. Two large cingular cusps are located on the buccal cingulum opposite to cusp C. Distally and mesially to these cusps, the size of the cingular cusps decreases. Altogether there are 13 cusps on the buccal cingulumandatleast 14 cuspsonthepreservedlingualportion of the cingulum (part of the lingual cingulum between cusps B and A is missing due to dental wear). The size variation between the lingual cingular cusps is smaller than that of the buccal cingular cusps. The lingual cingulum is elevated opposite to cusp A and between cusps A and C, similar to the “protocone” in the “symmetrodontan” mammal
|
||
<emphasis id="B92A5E3CFF8F7656CB5593CFFA32724C" box="[1370,1482,1272,1296]" italics="true" pageId="2" pageNumber="506">Yermackia</emphasis>
|
||
from the Early Cretaceous of West Siberia (
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CB25922FFC27720C" author="Averianov, A. O. & Lopatin, A. V." pageId="2" pageNumber="506" pagination="548 - 552" refId="ref3263" refString="Averianov, A. O., Lopatin, A. V. (2008): " Protocone " in a pretribosphenic mammal and upper dentition of tinodontid " symmetrodontans ". - Journal of Vertebrate Paleon- tology, 28: 548 - 552. https: // doi. org / 10.1671 / 0272 - 4634 (2008) 28 [548: PIAP-" type="journal article" year="2008">Averianov and Lopatin 2008</bibRefCitation>
|
||
). The mesial and distal sides of the crown are rhomboidal with pointed ends. This, and the lack of mesial cingular cusps E and F with an embayment between them, indicates that the new taxon lacked an upper molar interlock between cusps D and E-F, which is also seen in
|
||
<taxonomicName id="4C5EF9ADFF8F7656CB30928EFA3B728D" authorityName="YADAGIRI" authorityYear="1984" box="[1343,1475,1465,1489]" class="Mammalia" family="Morganucodontidae" genus="Indotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92A5E3CFF8F7656CB30928EFA3B728D" box="[1343,1475,1465,1489]" italics="true" pageId="2" pageNumber="506">Indotherium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5EF9ADFF8F7656CD4492EEFC1C72AD" authorityName="DATTA et DAS" authorityYear="2001" box="[843,996,1497,1521]" class="Mammalia" family="Morganucodontidae" genus="Indozostrodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92A5E3CFF8F7656CD4492EEFC1C72AD" box="[843,996,1497,1521]" italics="true" pageId="2" pageNumber="506">Indozostrodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5EF9ADFF8F7656CA1792EEFB5E72AD" authorityName="CLEMENS" authorityYear="2011" box="[1048,1190,1497,1521]" class="Mammalia" family="Cynodontia" genus="Bridetherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92A5E3CFF8F7656CA1792EEFB5E72AD" box="[1048,1190,1497,1521]" italics="true" pageId="2" pageNumber="506">Bridetherium</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CABA92EEFA3E72AD" author="Prasad, G. V. R. & Manhas, B. K." box="[1205,1478,1497,1521]" pageId="2" pageNumber="506" pagination="563 - 572" refId="ref5120" refString="Prasad, G. V. R., Manhas, B. K. (1997): A new symmetrodont mammal from the Lower Jurassic Kota Formation, Pran- hita-Godavari Valley, India. - Geobios, 30: 563 - 572. https: // doi. org / 10.1016 / S 0016 - 6995 (97) 80122 - 2" type="journal article" year="1997">Prasad and Manhas 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CD4492CEFBDF714D" author="Datta, P. M. & Das, D. P." box="[843,1063,1529,1553]" pageId="2" pageNumber="506" pagination="528 - 534" refId="ref3761" refString="Datta, P. M., Das, D. P. (2001): Indozostrodon simpsoni, gen. et sp. nov., an Early Jurassic megazostrodontid mammal from India. - Journal of Vertebrate Paleontology, 21: 528 - 534. https: // doi. org / 10.1671 / 0272 - 4634 (2001) 021 [0528: IS-" type="journal article" year="2001">Datta and Das 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CA3B92CEFB2B714D" author="Clemens, W. A." box="[1076,1235,1529,1553]" pageId="2" pageNumber="506" pagination="1139 - 1156" refId="ref3548" refString="Clemens, W. A. (2011): New morganucodontans from an Early Jurassic fissure filling in Wales (United Kingdom). - Palaeontology, 54: 1139 - 1156. https: // doi. org / 10.1111 / j. 1475 - 4983.2011.01094. x" type="journal article" year="2011">Clemens 2011</bibRefCitation>
|
||
). The molar contact in the maxilla was likely imbricating. The tips of cusps B and C have small apical abrasion facets, but otherwise the cusps are unworn. There is a large wear facet on the lingual side between cusps A and B which was caused by cusp a of the lower antagonist, and a small facet between cusps A and C (caused by cusp c of the antagonist). According to
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CB9E918DFC6C71AE" author="Mills, J. R. E." pageId="2" pageNumber="506" pagination="29 - 63" refId="ref4956" refString="Mills, J. R. E. (1971): The dentition of Morganucodon. - In: Kermack, D. M., Kermack, K. A. (eds), Early Mammals. Zoological Journal of the Linnean Society, London, 50 (Supplement 1): 29 - 63." type="journal article" year="1971">Mills (1971)</bibRefCitation>
|
||
, the cingulum wears rapidly on the lingual side in
|
||
<taxonomicName id="4C5EF9ADFF8F7656CD4491CDFC16704E" authorityName="KUHNE" authorityYear="1949" box="[843,1006,1786,1810]" class="Mammalia" family="Morganucodontidae" genus="Morganucodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92A5E3CFF8F7656CD4491CDFC16704E" box="[843,1006,1786,1810]" italics="true" pageId="2" pageNumber="506">Morganucodon</emphasis>
|
||
</taxonomicName>
|
||
. The insignificant apical wear and the only partially worn lingual cingulum suggest that NLMH 105654 was only recently erupted. This occlusal mode is identical with the one described for
|
||
<taxonomicName id="4C5EF9ADFF8F7656CA78906CFC1070CF" authority="(Crompton and Jenkins 1968)" baseAuthorityName="Crompton and Jenkins" baseAuthorityYear="1968" class="Mammalia" family="Morganucodontidae" genus="Morganucodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92A5E3CFF8F7656CA78906CFAE2702F" box="[1143,1306,1883,1907]" italics="true" pageId="2" pageNumber="506">Morganucodon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CB24906CFC2770CF" author="Crompton, A. W. & Jenkins, F. A., Jr." pageId="2" pageNumber="506" pagination="427 - 458" refId="ref3702" refString="Crompton, A. W., Jenkins, F. A., Jr. (1968): Molar occlusion in Late Triassic mammals. - Biological Reviews, 43: 427 - 458. https: // doi. org / 10.1111 / j. 1469 - 185 X. 1968. tb 00966. x" type="journal article" year="1968">Crompton and Jenkins 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. There are two roots connected basally by an interradical crest (
|
||
<figureCitation id="13659EABFF8F7656CA3790ACFB4570EF" box="[1080,1213,1947,1971]" captionStart="Text-fig" captionStartId="2.[147,228,606,627]" captionTargetBox="[157,1450,184,575]" captionTargetId="figure-621@2.[147,1462,58,587]" captionTargetPageId="2" captionText="Text-fig. 2. Storchodon cingulatus gen. et sp. nov., NLMH 105654, holotype, right upper molar, in buccal (a), occlusal (b, b’ stereopair), lingual (c), and basal (d) views. Langenberg Quarry, Lower Saxony, Germany; Süntel Formation, Late Jurassic (late Kimmeridgian) age. Abbreviations: A, B, C and D, main cusps of the crown; ic, interradical crest." figureDoi="http://doi.org/10.5281/zenodo.4586741" httpUri="https://zenodo.org/record/4586741/files/figure.png" pageId="2" pageNumber="506">Text-fig. 2d</figureCitation>
|
||
). The mesial root has a normal position while the distal root is set obliquely to the longitudinal axis of the crown. This apparently reflects an oblique position of cusp D relative to cusp C (both cusps are supported by the distal root). The pulp cavity of the distal root is smaller compared with that of the mesial root. The distal root is oval in cross section, with the mesiodistal diameter being larger than the buccolingual one.
|
||
</paragraph>
|
||
<caption id="DF21D2A6FF8F7656CE9C9569FB8175F2" ID-DOI="http://doi.org/10.5281/zenodo.4586741" ID-Zenodo-Dep="4586741" httpUri="https://zenodo.org/record/4586741/files/figure.png" pageId="2" pageNumber="506" startId="2.[147,228,606,627]" targetBox="[157,1450,184,575]" targetPageId="2">
|
||
<paragraph id="8BE1822EFF8F7656CE9C9569FB8175F2" blockId="2.[147,1463,606,686]" pageId="2" pageNumber="506">
|
||
Text-fig. 2.
|
||
<taxonomicName id="4C5EF9ADFF8F7656CF019568FE0A7528" authority="Martin & Averianov & Jäger & Schwermann & Oliver & Wings, 2019" authorityName="Martin & Averianov & Jäger & Schwermann & Oliver & Wings" authorityYear="2019" box="[270,498,607,628]" class="Mammalia" family="Triconodontidae" genus="Storchodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="2" pageNumber="506" phylum="Chordata" rank="species" species="cingulatus" status="gen. et sp. nov.">
|
||
<emphasis id="B92A5E3CFF8F7656CF019568FE0A7528" bold="true" box="[270,498,607,628]" italics="true" pageId="2" pageNumber="506">Storchodon cingulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219E347FF8F7656CFF39569FD65752F" box="[508,669,606,627]" pageId="2" pageNumber="506" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, NLMH 105654, holotype, right upper molar, in buccal (a), occlusal (b, b’ stereopair), lingual (c), and basal (d) views. Langenberg Quarry, Lower Saxony, Germany; Süntel Formation, Late Jurassic (late Kimmeridgian) age. Abbreviations: A, B, C and D, main cusps of the crown; ic, interradical crest.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF21D2A6FF8F7656CE8E909FFDBF7F6F" ID-DOI="http://doi.org/10.5281/zenodo.4586743" ID-Zenodo-Dep="4586743" httpUri="https://zenodo.org/record/4586743/files/figure.png" pageId="2" pageNumber="506" startId="2.[129,210,1960,1981]" targetBox="[141,751,1426,1921]" targetPageId="2">
|
||
<paragraph id="8BE1822EFF8F7656CE8E909FFDBF7F6F" blockId="2.[128,764,1960,2099]" pageId="2" pageNumber="506">
|
||
<emphasis id="B92A5E3CFF8F7656CE8E909FFF3C7086" bold="true" pageId="2" pageNumber="506">
|
||
Text-fig. 3. Right upper molars of
|
||
<emphasis id="B92A5E3CFF8F7656CC06909FFD0370E1" bold="true" box="[521,763,1960,1981]" italics="true" pageId="2" pageNumber="506">
|
||
<taxonomicName id="4C5EF9ADFF8F7656CC06909FFD5A70E1" baseAuthorityName="Crompton and Jenkins" baseAuthorityYear="1968" box="[521,674,1960,1981]" class="Mammalia" family="Morganucodontidae" genus="Morganucodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Morganucodonta" pageId="2" pageNumber="506" phylum="Chordata" rank="genus">Morganucodon</taxonomicName>
|
||
watsoni
|
||
</emphasis>
|
||
<bibRefCitation id="EFCFFFDFFF8F7656CE8E90F1FF3C7086" author="Kuhne, W. G." box="[129,196,1990,2011]" pageId="2" pageNumber="506" pagination="345 - 350" refId="ref4419" refString="Kuhne, W. G. (1949): On a triconodont tooth of a new pattern from a fissure-filling in South Glamorgan. - Proceedings of the Zoological Society of London, 119: 345 - 350. https: // doi. org / 10.1111 / j. 1096 - 3642.1949. tb 00883. x" type="journal article" year="1949">KÜHNE</bibRefCitation>
|
||
</emphasis>
|
||
, 1949, UMZC_Eo.M(Ur).27 (cast) (a) and
|
||
<taxonomicName id="4C5EF9ADFF8F7656CC8690F1FF1070A4" authority="Martin & Averianov & Jäger & Schwermann & Oliver & Wings, 2019" authorityName="Martin & Averianov & Jäger & Schwermann & Oliver & Wings" authorityYear="2019" class="Mammalia" family="Triconodontidae" genus="Storchodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Multituberculata" pageId="2" pageNumber="506" phylum="Chordata" rank="species" species="cingulatus" status="gen. et sp. nov.">
|
||
<emphasis id="B92A5E3CFF8F7656CC8690F1FF1070A4" bold="true" italics="true" pageId="2" pageNumber="506">Storchodon cingulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A219E347FF8F7656CEFB90D4FE6370A4" box="[244,411,2019,2040]" pageId="2" pageNumber="506" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, NLMH 105654, holotype (b) in lingual and occlusal views. Abbreviations: A, B, C, D, E and F, main cusps of the crown. Specimens to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1822EFF8E7657CE9F9630FD187642" blockId="3.[144,736,263,287]" box="[144,736,263,287]" pageId="3" pageNumber="507">
|
||
M e a s u r e m e n t s. Length
|
||
<quantity id="4CA62FCBFF8E7657CFC49630FDD47642" box="[459,556,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="3.07" pageId="3" pageNumber="507" unit="mm" value="3.07">3.07 mm</quantity>
|
||
, width
|
||
<quantity id="4CA62FCBFF8E7657CC759630FD257642" box="[634,733,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="3" pageNumber="507" unit="mm" value="1.44">1.44 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |