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<mods:title id="60C503DBB47C6F4317859494F69192AB">Tanypus urszulae, a new Tanypodinae (Diptera: Chironomidae) from the Neotropical Region</mods:title>
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<paragraph id="8BAA3699D918FFC0FF278ED704D38AF7" blockId="1.[151,426,221,247]" box="[151,426,221,247]" pageId="1" pageNumber="594">
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<taxonomicName id="4C154D1AD918FFC0FF278ED704D38AF7" ID-CoL="9CJWX" authority="Meigen, 1803" authorityName="Meigen" authorityYear="1803" box="[151,426,221,247]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8BD918FFC0FF278ED704D38AF7" bold="true" box="[151,426,221,247]" pageId="1" pageNumber="594">
<emphasis id="B961EA8BD918FFC0FF278ED705838AF7" bold="true" box="[151,250,221,247]" italics="true" pageId="1" pageNumber="594">Tanypus</emphasis>
<bibRefCitation id="EF844B68D918FFC0FEB48ED704D38AF7" author="Meigen" box="[260,426,221,247]" pageId="1" pageNumber="594" refString="Meigen, J. W. (1803) Versuch einer neuen Gattungs Eintheilung der europaischen zweiflugeligen Insekten. Illigers Magazin fur Insektenkunde, 2, 259 - 281." type="journal article" year="1803">Meigen, 1803</bibRefCitation>
</emphasis>
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<taxonomicName id="4C154D1AD918FFC0FF278F2F04CD8B3A" ID-CoL="9CJWX" authority="Meigen, 1803: 261" authorityName="Meigen" authorityPageNumber="261" authorityYear="1803" box="[151,436,292,314]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8BD918FFC0FF278F2F05928B3A" box="[151,235,293,314]" italics="true" pageId="1" pageNumber="594">Tanypus</emphasis>
<treatmentCitation id="0AB41088D918FFC0FF418F2E04CD8B3A" author="Meigen" box="[241,436,292,314]" page="261" pageId="1" pageNumber="594" year="1803">
<bibRefCitation id="EF844B68D918FFC0FF418F2E04F88B3A" author="Meigen" box="[241,385,292,314]" pageId="1" pageNumber="594" refString="Meigen, J. W. (1803) Versuch einer neuen Gattungs Eintheilung der europaischen zweiflugeligen Insekten. Illigers Magazin fur Insektenkunde, 2, 259 - 281." type="journal article" year="1803">Meigen, 1803</bibRefCitation>
: 261
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.
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</paragraph>
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<paragraph id="8BAA3699D918FFC0FF278F6C005D8B7F" blockId="1.[151,1436,358,779]" box="[151,1316,358,383]" pageId="1" pageNumber="594">
<emphasis id="B961EA8BD918FFC0FF278F6C044B8B7F" bold="true" box="[151,306,358,383]" pageId="1" pageNumber="594">Type species.</emphasis>
<taxonomicName id="4C154D1AD918FFC0FE898F6C07C48B7F" authority="Latreille, 1810" authorityName="Latreille" authorityYear="1810" box="[313,701,358,383]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B961EA8BD918FFC0FE898F6C04B58B7E" box="[313,460,358,383]" italics="true" pageId="1" pageNumber="594">Tipula cincta</emphasis>
sensu
<bibRefCitation id="EF844B68D918FFC0FDA88F6C07C48B7F" author="Latreille" box="[536,701,358,383]" pageId="1" pageNumber="594" refString="Latreille, P. A. (1810) Considerations generales sur l'ordre naturel des animaux composant les classes des crustaces, des arachnides et des insects; avec un tableau methodique de leurs genres, disposes en familles. Chez F. Schoell, Paris, 444 pp." type="book" year="1810">Latreille, 1810</bibRefCitation>
</taxonomicName>
[misidentified
<emphasis id="B961EA8BD918FFC0FCD88F6C06018B7F" bold="true" box="[872,888,358,383]" pageId="1" pageNumber="594">=</emphasis>
<taxonomicName id="4C154D1AD918FFC0FCCF8F6D00638B7F" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" box="[895,1306,358,383]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B961EA8BD918FFC0FCCF8F6D01088B7E" box="[895,1137,359,382]" italics="true" pageId="1" pageNumber="594">Tanypus punctipennis</emphasis>
<bibRefCitation id="EF844B68D918FFC0FBC88F6C00638B7F" author="Meigen" box="[1144,1306,358,383]" pageId="1" pageNumber="594" refString="Meigen, J. W. (1818) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Part 1. Bei Friedrich Wilhelm Forstmann, Aachen, 332 pp." type="book" year="1818">Meigen, 1818</bibRefCitation>
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].
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<emphasis id="B961EA8BD918FFC0FF778F8004CC8BA3" bold="true" box="[199,437,394,419]" pageId="1" pageNumber="594">Emended diagnosis.</emphasis>
<emphasis id="B961EA8BD918FFC0FE0C8F8604868BA3" box="[444,511,396,419]" italics="true" pageId="1" pageNumber="594">Larva</emphasis>
: ligula with 56 teeth.
</paragraph>
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<subSubSection id="C30F6512D918FFC0FF778FA40042890B" pageId="1" pageNumber="594" type="discussion">
<paragraph id="8BAA3699D918FFC0FF778FA40042890B" blockId="1.[151,1436,358,779]" pageId="1" pageNumber="594">
<emphasis id="B961EA8BD918FFC0FF778FA404428BC7" bold="true" box="[199,315,430,455]" pageId="1" pageNumber="594">Remarks.</emphasis>
Generic diagnoses are given by
<bibRefCitation id="EF844B68D918FFC0FD0D8FA4061E8BC7" author="Roback" box="[701,871,430,455]" pageId="1" pageNumber="594" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback (1971)</bibRefCitation>
and
<bibRefCitation id="EF844B68D918FFC0FC148FA501BE8BC7" author="Murray" box="[932,1223,430,455]" pageId="1" pageNumber="594" refString="Murray, D. A. &amp; Fittkau, E. J. (1989) The adult males of Tanypodinae (Diptera: Chironomidae) of the Holarctic Region - Keys and diagnoses. Entomologica scandinavica supplement, 34, 37 - 123." type="journal article" year="1989">Murray &amp; Fittkau (1989)</bibRefCitation>
for the adult; and
<bibRefCitation id="EF844B68D918FFC0FF278FD804388BEB" author="Roback" box="[151,321,466,491]" pageId="1" pageNumber="594" refString="Roback, S. S. (1977) The immature chironomids of the Eastern United States II. Tanypodinae - Tanypodini. Proceedings of the Academy of Natural Sciences of Philadelphia, 128, 55 - 88." type="journal article" year="1977">Roback (1977)</bibRefCitation>
and
<bibRefCitation id="EF844B68D918FFC0FECD8FD807E78BEB" author="Fittkau" box="[381,670,466,491]" pageId="1" pageNumber="594" refString="Fittkau, E. J. &amp; Murray, D. A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses. Entomologica Scandinavica supplement, 28, 31 - 113." type="journal article" year="1986">Fittkau &amp; Murray (1986)</bibRefCitation>
for the pupa. The diagnoses in
<bibRefCitation id="EF844B68D918FFC0FBA38FD801C68BEB" author="Roback" box="[1043,1215,466,491]" pageId="1" pageNumber="594" refString="Roback, S. S. (1977) The immature chironomids of the Eastern United States II. Tanypodinae - Tanypodini. Proceedings of the Academy of Natural Sciences of Philadelphia, 128, 55 - 88." type="journal article" year="1977">Roback (1977)</bibRefCitation>
,
<bibRefCitation id="EF844B68D918FFC0FB7E8FD80598880F" author="Fittkau" pageId="1" pageNumber="594" refString="Fittkau, E. J. &amp; Roback, S. S. (1983) The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic Region - Keys and diagnoses. Entomologica scandinavica supplement, 19, 33 - 110." type="journal article" year="1983">Fittkau &amp; Roback (1983)</bibRefCitation>
and
<bibRefCitation id="EF844B68D918FFC0FEA78FFD0754880F" author="Cranston" box="[279,557,502,527]" pageId="1" pageNumber="594" refString="Cranston, P. S. &amp; Epler, J. H. (2013) The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. Insect Systematics &amp; Evolution, 66, 39 - 136." type="journal article" year="2013">Cranston &amp; Epler (2013)</bibRefCitation>
for the larva should be emended to accommodate the Neotropical species with an unusual feature for the genus. Concerning the
<taxonomicName id="4C154D1AD918FFC0FD598C16063C8833" box="[745,837,540,563]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8BD918FFC0FD598C16063C8833" box="[745,837,540,563]" italics="true" pageId="1" pageNumber="594">Tanypus</emphasis>
</taxonomicName>
pupa,
<bibRefCitation id="EF844B68D918FFC0FC2C8C1001BC8833" author="Fittkau" box="[924,1221,538,563]" pageId="1" pageNumber="594" refString="Fittkau, E. J. &amp; Murray, D. A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses. Entomologica Scandinavica supplement, 28, 31 - 113." type="journal article" year="1986">Fittkau &amp; Murray (1986)</bibRefCitation>
wrote, “anal lobe reduced, generally slightly broader than long and shorter than segment VIII.” The pupa of the species treated here, however, has well-developed anal lobe, which is much longer than the abdominal segment VIII. The male genital sac is shorter than the anal lobe, and the relative length is much shorter than in
<taxonomicName id="4C154D1AD918FFC0FB8D8C8D001C889F" authority="Coquillett" authorityName="Coquillett" box="[1085,1381,646,671]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="stellatus" subGenus="T.">
<emphasis id="B961EA8BD918FFC0FB8D8C8D0133889E" box="[1085,1098,647,670]" italics="true" pageId="1" pageNumber="594">T</emphasis>
. (
<emphasis id="B961EA8BD918FFC0FBD58C8D010B889E" box="[1125,1138,647,670]" italics="true" pageId="1" pageNumber="594">T</emphasis>
.)
<emphasis id="B961EA8BD918FFC0FB3D8C8C0193889F" box="[1165,1258,646,671]" italics="true" pageId="1" pageNumber="594">stellatus</emphasis>
Coquillett
</taxonomicName>
(see
<bibRefCitation id="EF844B68D918FFC0FF278CA0044888C3" author="Roback" box="[151,305,682,707]" pageId="1" pageNumber="594" refString="Roback, S. S. (1977) The immature chironomids of the Eastern United States II. Tanypodinae - Tanypodini. Proceedings of the Academy of Natural Sciences of Philadelphia, 128, 55 - 88." type="journal article" year="1977">Roback 1977</bibRefCitation>
, figure 70;
<bibRefCitation id="EF844B68D918FFC0FE098CA007B288C3" author="Fittkau" box="[441,715,682,707]" pageId="1" pageNumber="594" refString="Fittkau, E. J. &amp; Murray, D. A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses. Entomologica Scandinavica supplement, 28, 31 - 113." type="journal article" year="1986">Fittkau &amp; Murray 1986</bibRefCitation>
, figure 5.38 F). These features may change the concept of the
<taxonomicName id="4C154D1AD918FFC0FF278CC5058D88E6" box="[151,244,719,742]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8BD918FFC0FF278CC5058D88E6" box="[151,244,719,742]" italics="true" pageId="1" pageNumber="594">Tanypus</emphasis>
</taxonomicName>
pupa. This genus is divided into two subgenera,
<taxonomicName id="4C154D1AD918FFC0FC8D8CC4017B88E7" authority="Roback" authorityName="Roback" box="[829,1026,718,743]" class="Insecta" family="Chironomidae" genus="Apelopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA8BD918FFC0FC8D8CC406DB88E7" box="[829,930,718,743]" italics="true" pageId="1" pageNumber="594">Apelopia</emphasis>
Roback
</taxonomicName>
and
<emphasis id="B961EA8BD918FFC0FBF28CC5019188E6" box="[1090,1256,719,742]" italics="true" pageId="1" pageNumber="594">
<taxonomicName id="4C154D1AD918FFC0FBF28CC501E688E6" box="[1090,1183,719,742]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus" sensu="stricto">Tanypus</taxonomicName>
<taxonomicNameLabel id="A25257F0D918FFC0FB198CC5019188E6" box="[1193,1256,719,742]" pageId="1" pageNumber="594" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EF844B68D918FFC0FB4A8CC400EE88E7" author="Roback" box="[1274,1431,718,743]" pageId="1" pageNumber="594" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback 1971</bibRefCitation>
,
<bibRefCitation id="EF844B68D918FFC0FF278CF905AA890B" author="Roback" box="[151,211,755,779]" pageId="1" pageNumber="594" refString="Roback, S. S. (1977) The immature chironomids of the Eastern United States II. Tanypodinae - Tanypodini. Proceedings of the Academy of Natural Sciences of Philadelphia, 128, 55 - 88." type="journal article" year="1977">1977</bibRefCitation>
).
<bibRefCitation id="EF844B68D918FFC0FF568CF90486890B" author="Cranston" box="[230,511,754,779]" pageId="1" pageNumber="594" refString="Cranston, P. S. &amp; Epler, J. H. (2013) The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region. Keys and diagnoses. Insect Systematics &amp; Evolution, 66, 39 - 136." type="journal article" year="2013">Cranston &amp; Epler (2013)</bibRefCitation>
, however, noted that the genus cannot be separated yet in the larval stage.
</paragraph>
</subSubSection>
</treatment>
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