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<document id="B39F9524CF05C5540626CE429C1FB8CC" ID-CLB-Dataset="3979" ID-DOI="10.11646/zootaxa.5051.1.13" ID-GBIF-Dataset="a82cb38a-03d5-4d23-845c-436529542679" ID-ISSN="1175-5326" ID-Zenodo-Dep="5572417" ID-ZooBank="F94203E7-FCD1-4975-BAD3-0DF534806712" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634021882240" checkinUser="plazi" docAuthor="Huys, Rony &amp; Mu, Fanghong" docDate="2021" docId="951887EAFFFAFFA4FF51D099E05CFC05" docLanguage="en" docName="zootaxa.5051.1.13.pdf" docOrigin="Zootaxa 5051 (1)" docSource="https://doi.org/10.11646/zootaxa.5051.1.13" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Foweya Gee 2006" docType="treatment" docVersion="8" lastPageNumber="294" masterDocId="6921FF92FFC3FF9EFFC6D318E440FFDB" masterDocTitle="Johnwellsia, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera" masterLastPageNumber="318" masterPageNumber="236" pageNumber="293" updateTime="1699459569358" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="35B30B70EE21D31A87D0249008DEFDD8">Johnwellsia, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera</mods:title>
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<mods:affiliation id="40D0BFA6FD522EC9FCDA885616F17B35">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<mods:affiliation id="FE51C578385DEEE5756A7362F768B9A5">College of Marine Life Science, Ocean University of China, 5 Yushan Road, Qingdao 266003, China.</mods:affiliation>
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<mods:date id="558EBBB0004B769FC91EFD1A000FD1E1">2021</mods:date>
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<treatment id="951887EAFFFAFFA4FF51D099E05CFC05" ID-DOI="http://doi.org/10.5281/zenodo.5576523" ID-GBIF-Taxon="189091330" ID-Zenodo-Dep="5576523" LSID="urn:lsid:plazi:treatment:951887EAFFFAFFA4FF51D099E05CFC05" httpUri="http://treatment.plazi.org/id/951887EAFFFAFFA4FF51D099E05CFC05" lastPageId="58" lastPageNumber="294" pageId="57" pageNumber="293">
<subSubSection id="55AB6577FFFAFFA7FF51D099E538FC40" box="[151,376,897,923]" pageId="57" pageNumber="293" type="nomenclature">
<paragraph id="1D0E36FCFFFAFFA7FF51D099E538FC40" blockId="57.[151,376,897,923]" box="[151,376,897,923]" pageId="57" pageNumber="293">
<heading id="46468190FFFAFFA7FF51D099E538FC40" bold="true" box="[151,376,897,923]" fontSize="11" level="1" pageId="57" pageNumber="293" reason="1">
<taxonomicName id="DAB14D7FFFFAFFA7FF51D099E538FC40" authority="Gee, 2006" authorityName="Gee" authorityYear="2006" box="[151,376,897,923]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FF51D099E538FC40" bold="true" box="[151,376,897,923]" pageId="57" pageNumber="293">
<emphasis id="2FC5EAEEFFFAFFA7FF51D099E4B6FC40" bold="true" box="[151,246,897,923]" italics="true" pageId="57" pageNumber="293">Foweya</emphasis>
<bibRefCitation id="79204B0DFFFAFFA7FF3BD099E538FC40" author="Gee, J. M." box="[253,376,897,923]" pageId="57" pageNumber="293" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee, 2006</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="55AB6577FFFAFFA7FF51D0D1E68FFAB4" pageId="57" pageNumber="293" type="discussion">
<paragraph id="1D0E36FCFFFAFFA7FF51D0D1E68FFAB4" blockId="57.[151,1437,969,2039]" pageId="57" pageNumber="293">
<bibRefCitation id="79204B0DFFFAFFA7FF51D0D1E558FC3F" author="Gee, J. M." box="[151,280,969,996]" pageId="57" pageNumber="293" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
recognized the close resemblance between
<taxonomicName id="DAB14D7FFFFAFFA7FCCAD0D1E046FC38" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[780,1030,969,995]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FCCAD0D1E046FC38" box="[780,1030,969,995]" italics="true" pageId="57" pageNumber="293">Parastenhelia anglica</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFFAFFA7FB86D0D1E141FC38" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[1088,1281,969,995]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFFAFFA7FB86D0D1E141FC38" box="[1088,1281,969,995]" italics="true" pageId="57" pageNumber="293">Karllangia tertia</emphasis>
</taxonomicName>
and removed both species from their respective genera and placed them in a newly proposed genus
<taxonomicName id="DAB14D7FFFFAFFA7FB51D0F6E0B0FBDC" authorityName="Gee" authorityYear="2006" box="[1175,1264,1006,1031]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FB51D0F6E0B0FBDC" box="[1175,1264,1006,1031]" italics="true" pageId="57" pageNumber="293">Foweya</emphasis>
</taxonomicName>
. Both
<taxonomicName id="DAB14D7FFFFAFFA7FA82D0F6E642FBF7" authority="(Norman &amp; Scott, 1905)" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FA82D0F6E4ABFBF0" italics="true" pageId="57" pageNumber="293">Foweya anglica</emphasis>
(
<bibRefCitation id="79204B0DFFFAFFA7FF3FD709E5BAFBF7" author="Norman, A. M. &amp; Scott, T." box="[249,506,1041,1068]" pageId="57" pageNumber="293" pagination="284 - 300" refId="ref76779" refString="Norman, A. M. &amp; Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284 - 300. https: // doi. org / 10.1080 / 03745480509443044" type="journal article" year="1905">Norman &amp; Scott, 1905</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFFAFFA7FDFED70AE76BFBF7" authority="(Kunz, 1975)" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[568,811,1041,1068]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFFAFFA7FDFED70AE6D0FBF0" box="[568,656,1042,1067]" italics="true" pageId="57" pageNumber="293">F. tertia</emphasis>
(
<bibRefCitation id="79204B0DFFFAFFA7FD58D709E762FBF7" author="Kunz, H." box="[670,802,1041,1068]" pageId="57" pageNumber="293" pagination="179 - 212" refId="ref75043" refString="Kunz, H. (1975) Copepoda Harpacticoida aus dem Litoral des sudlichen Afrika. I. Teil. Kieler Meeresforschungen, 31, 179 - 212, pls. 1 - 30 (unpaginated)." type="journal article" year="1975">Kunz, 1975</bibRefCitation>
)
</taxonomicName>
exhibit the narrowing of the outer cuticle of P1 enp-1 at around the insertion level of the inner seta (shared with
<taxonomicName id="DAB14D7FFFFAFFA7FD38D72DE7C4FB94" authorityName="Monard" authorityYear="1935" box="[766,900,1077,1103]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FD38D72DE7C4FB94" box="[766,900,1077,1103]" italics="true" pageId="57" pageNumber="293">Thalestrella</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFFAFFA7FC7FD72DE0FEFB94" authorityName="Mielke" authorityYear="1994" box="[953,1214,1077,1103]" class="Hexanauplia" family="Parastenheliidae" genus="Psammoleptomesochra" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FC7FD72DE0FEFB94" box="[953,1214,1077,1103]" italics="true" pageId="57" pageNumber="293">Psammoleptomesochra</emphasis>
</taxonomicName>
), the bifid, sexually dimorphic inner spine on the P1 basis in the male (shared with
<taxonomicName id="DAB14D7FFFFAFFA7FC8BD741E06AFBA8" authorityName="Wells" authorityYear="1967" box="[845,1066,1113,1139]" class="Hexanauplia" family="Parastenheliidae" genus="Paraleptomesochra" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FC8BD741E06AFBA8" box="[845,1066,1113,1139]" italics="true" pageId="57" pageNumber="293">Paraleptomesochra</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFFAFFA7FB98D741E123FBA8" authorityName="Mielke" authorityYear="1994" box="[1118,1379,1113,1139]" class="Hexanauplia" family="Parastenheliidae" genus="Psammoleptomesochra" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FB98D741E123FBA8" box="[1118,1379,1113,1139]" italics="true" pageId="57" pageNumber="293">Psammoleptomesochra</emphasis>
</taxonomicName>
), the 10-segmented male antennule with only three segments distal to the articulation, the absence of sexual dimorphism in the endopod of P3 (shared with
<taxonomicName id="DAB14D7FFFFAFFA7FDF5D7B9E6F9FB60" authorityName="Monard" authorityYear="1935" box="[563,697,1185,1211]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FDF5D7B9E6F9FB60" box="[563,697,1185,1211]" italics="true" pageId="57" pageNumber="293">Thalestrella</emphasis>
</taxonomicName>
,
<taxonomicName id="DAB14D7FFFFAFFA7FD0DD7B9E7E8FB60" authorityName="Wells" authorityYear="1967" box="[715,936,1185,1211]" class="Hexanauplia" family="Parastenheliidae" genus="Paraleptomesochra" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FD0DD7B9E7E8FB60" box="[715,936,1185,1211]" italics="true" pageId="57" pageNumber="293">Paraleptomesochra</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFFAFFA7FC20D7B9E0ABFB60" authorityName="Mielke" authorityYear="1994" box="[998,1259,1185,1211]" class="Hexanauplia" family="Parastenheliidae" genus="Psammoleptomesochra" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFFAFFA7FC20D7B9E0ABFB60" box="[998,1259,1185,1211]" italics="true" pageId="57" pageNumber="293">Psammoleptomesochra</emphasis>
</taxonomicName>
), and the most primitive swimming leg armature formula within the family, including two inner setae on P2 exp-3 and enp-3 and three inner setae on P3 enp-3.
<bibRefCitation id="79204B0DFFFAFFA7FE3AD7F1E63DFADF" author="Gee, J. M." box="[508,637,1257,1284]" pageId="57" pageNumber="293" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
confirmed that the specimens, previously identified as
<taxonomicName id="DAB14D7FFFFAFFA7FB39D7F1E4ABFAFC" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FB39D7F1E4ABFAFC" italics="true" pageId="57" pageNumber="293">Parastenhelia anglica</emphasis>
</taxonomicName>
from St. Agnes, St. Mary’s and Tresco (Isles of Scilly) by
<bibRefCitation id="79204B0DFFFAFFA7FC43D615E050FAFC" author="Wells, J. B. J." box="[901,1040,1293,1319]" pageId="57" pageNumber="293" pagination="262 - 274" refId="ref82063" refString="Wells, J. B. J. (1961) Interstitial copepods from the Isles of Scilly. Crustaceana, 2, 262 - 274. https: // doi. org / 10.1163 / 156854061 X 00392" type="journal article" year="1961">Wells (1961</bibRefCitation>
,
<bibRefCitation id="79204B0DFFFAFFA7FBDAD615E015FAFC" author="Wells, J. B. J." box="[1052,1109,1293,1319]" pageId="57" pageNumber="293" pagination="255 - 268" refId="ref82252" refString="Wells, J. B. J. (1970) The marine flora and fauna of the Isles of Scilly. Crustacea: Copepoda: Harpacticoida. Journal of natural History, 4, 255 - 268. https: // doi. org / 10.1080 / 00222937000770251" type="journal article" year="1970">1970</bibRefCitation>
) and from
<collectingCountry id="65A6766CFFFAFFA7FB12D615E174FAFC" box="[1236,1332,1293,1319]" name="Namibia" pageId="57" pageNumber="293">Namibia</collectingCountry>
by
<bibRefCitation id="79204B0DFFFAFFA7FAA6D615E4A4FA97" author="Kunz, H." pageId="57" pageNumber="293" pagination="33 - 51" refId="ref75017" refString="Kunz, H. (1963) Weitere Harpacticoiden (Crustacea Copepoda) von der sudwestafrikanischen Kuste. Zoologischer Anzeiger, 171, 33 - 51" type="journal article" year="1963">Kunz (1963)</bibRefCitation>
, are conspecific with
<taxonomicName id="DAB14D7FFFFAFFA7FE25D629E6B7FA90" authorityName="Monard." authorityYear="1935" baseAuthorityName="sensu Por" baseAuthorityYear="1935" box="[483,759,1329,1355]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFFAFFA7FE25D629E6B7FA90" box="[483,759,1329,1355]" italics="true" pageId="57" pageNumber="293">Thalestrella ornatissima</emphasis>
</taxonomicName>
(
<typeStatus id="C20A885EFFFAFFA7FCCED629E77AFA90" box="[776,826,1329,1355]" pageId="57" pageNumber="293">type</typeStatus>
locality: Roscoff, Brittany), and
<taxonomicName id="DAB14D7FFFFAFFA7FB7CD62AE119FA90" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[1210,1369,1330,1355]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFFAFFA7FB7CD62AE119FA90" box="[1210,1369,1330,1355]" italics="true" pageId="57" pageNumber="293">Foweya tertia</emphasis>
</taxonomicName>
(
<typeStatus id="C20A885EFFFAFFA7FAACD629E1DCFA90" box="[1386,1436,1329,1355]" pageId="57" pageNumber="293">type</typeStatus>
locality:
<collectingRegion id="DF75F81EFFFAFFA7FF3CD64DE5D1FAB4" box="[250,401,1365,1391]" country="South Africa" name="Eastern Cape" pageId="57" pageNumber="293">Eastern Cape</collectingRegion>
,
<collectingCountry id="65A6766CFFFAFFA7FE5AD64DE66FFAAB" box="[412,559,1365,1392]" name="South Africa" pageId="57" pageNumber="293">South Africa</collectingCountry>
), respectively.
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFFAFFA7FF01D661E752F99C" pageId="57" pageNumber="293" type="description">
<paragraph id="1D0E36FCFFFAFFA7FF01D661E752F99C" blockId="57.[151,1437,969,2039]" pageId="57" pageNumber="293">
Males of the two known
<taxonomicName id="DAB14D7FFFFAFFA7FE36D662E6E5FA48" box="[496,677,1401,1427]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2FC5EAEEFFFAFFA7FE36D662E609FA48" box="[496,585,1402,1427]" italics="true" pageId="57" pageNumber="293">Foweya</emphasis>
species
</taxonomicName>
can be differentiated by the segmentation of the P2 endopod (3- segmented in
<taxonomicName id="DAB14D7FFFFAFFA7FEF5D686E5E2FA6C" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[307,418,1437,1463]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FEF5D686E5E2FA6C" box="[307,418,1437,1463]" italics="true" pageId="57" pageNumber="293">F. anglica</emphasis>
</taxonomicName>
, 2-segmented in
<taxonomicName id="DAB14D7FFFFAFFA7FDA5D686E6FBFA6C" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[611,699,1438,1463]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFFAFFA7FDA5D686E6FBFA6C" box="[611,699,1438,1463]" italics="true" pageId="57" pageNumber="293">F. tertia</emphasis>
</taxonomicName>
) and the morphology of the fifth legs (with five exopodal setae in
<emphasis id="2FC5EAEEFFFAFFA7FF51D6DAE54EFA00" box="[151,270,1474,1499]" italics="true" pageId="57" pageNumber="293">
<taxonomicName id="DAB14D7FFFFAFFA7FF51D6DAE4B0FA00" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[151,240,1474,1499]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">F. tertia</taxonomicName>
vs
</emphasis>
six in
<taxonomicName id="DAB14D7FFFFAFFA7FE9CD6DAE589FA00" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[346,457,1473,1499]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FE9CD6DAE589FA00" box="[346,457,1473,1499]" italics="true" pageId="57" pageNumber="293">F. anglica</emphasis>
</taxonomicName>
; endopodal lobe relatively weakly developed and both setae equally long in
<emphasis id="2FC5EAEEFFFAFFA7FAE3D6DAE1DCFA00" box="[1317,1436,1474,1499]" italics="true" pageId="57" pageNumber="293">
<taxonomicName id="DAB14D7FFFFAFFA7FAE3D6DAE13EFA00" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[1317,1406,1474,1499]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">F. tertia</taxonomicName>
vs
</emphasis>
well developed lobe and outer endopodal seta distinctly longer than inner one in
<taxonomicName id="DAB14D7FFFFAFFA7FBECD6FEE0DBFA24" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[1066,1179,1509,1535]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFFAFFA7FBECD6FEE0DBFA24" box="[1066,1179,1509,1535]" italics="true" pageId="57" pageNumber="293">F. anglica</emphasis>
</taxonomicName>
). Females (and males) can be distinguished by the ornamentation of the anal operculum (with approximately 30 small, closely set denticles in
<emphasis id="2FC5EAEEFFFAFFA7FF72D536E501F99C" box="[180,321,1581,1607]" italics="true" pageId="57" pageNumber="293">
<taxonomicName id="DAB14D7FFFFAFFA7FF72D536E563F99C" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[180,291,1581,1607]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="anglica">F. anglica</taxonomicName>
vs
</emphasis>
numerous very fine spinules in
<taxonomicName id="DAB14D7FFFFAFFA7FD6FD536E742F99C" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[681,770,1582,1607]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFFAFFA7FD6FD536E742F99C" box="[681,770,1582,1607]" italics="true" pageId="57" pageNumber="293">F. tertia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFFAFFA4FF01D549E613FCA9" lastPageId="58" lastPageNumber="294" pageId="57" pageNumber="293" type="diagnosis">
<paragraph id="1D0E36FCFFFAFFA7FF01D549E162F8C4" blockId="57.[151,1437,969,2039]" pageId="57" pageNumber="293">
<emphasis id="2FC5EAEEFFFAFFA7FF01D549E502F9B0" bold="true" box="[199,322,1617,1643]" pageId="57" pageNumber="293">Diagnosis.</emphasis>
<materialsCitation id="ADD93CA1FFFAFFA7FE88D549E15BF8C4" ID-GBIF-Occurrence="3387337341" collectionCode="V" pageId="57" pageNumber="293" specimenCount="1" specimenCount-female="1">
<taxonomicName id="DAB14D7FFFFAFFA7FE88D549E649F9B0" baseAuthorityName="Willen" baseAuthorityYear="2000" box="[334,521,1617,1643]" class="Hexanauplia" family="Parastenheliidae" kingdom="Animalia" order="Harpacticoida" pageId="57" pageNumber="293" phylum="Arthropoda" rank="family">Parastenheliidae</taxonomicName>
. Sexual dimorphism in antennule, P1 inner basal spine, P5–P6 and urosomal segmentation; occasionally in segmentation of P2 endopod. Body subcylindrical, short; posterior margin of cephalothorax and somites bearing P2–P5 with plain hyaline frills; remaining urosomites (except anal somite) with denticulodigitate hyaline frills. Rostrum defined at base, of moderate size, reaching at most to end of second segment of antennule; linguiform. Anal operculum semicircular, bordered with fine spinules or denticles. Caudal ramus wider than long, with dorsal spinular row; with seven setae, setae IV–
<collectionCode id="7BA0AE39FFFAFFA7FC36D41DE044F8C4" box="[1008,1028,1797,1823]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="57" pageNumber="293" type="Museum">V</collectionCode>
not basally inflated in
<specimenCount id="0BB7FD75FFFAFFA7FAC3D41DE15BF8C4" box="[1285,1307,1797,1823]" pageId="57" pageNumber="293" type="female">♀</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="1D0E36FCFFFAFFA4FF01D431E50BFE9A" blockId="57.[151,1437,969,2039]" lastBlockId="58.[151,1436,151,538]" lastPageId="58" lastPageNumber="294" pageId="57" pageNumber="293">
<materialsCitation id="ADD93CA1FFFAFFA4FF01D431E507FE9A" ID-GBIF-Occurrence="3387337343" collectionCode="VIII, IX, XII, XIII, XIV, XVI, VII, XVIII, XIX, XX, XXI, XXIII, XXIV, XXV, XXVI, XXVIII" lastPageId="58" lastPageNumber="294" pageId="57" pageNumber="293" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
Antennule elongate and 9-segmented in
<specimenCount id="0BB7FD75FFFAFFA7FD4DD431E6E0F898" box="[651,672,1833,1859]" pageId="57" pageNumber="293" type="female">♀</specimenCount>
, segment 1 not elongate, segments 5–8 shortest, with aesthetascs on segments 4 and 9; haplocer and 10-segmented with three segments distal to geniculation in
<specimenCount id="0BB7FD75FFFAFFA7FB74D455E089F8BC" box="[1202,1225,1869,1895]" pageId="57" pageNumber="293" type="male">♂</specimenCount>
, with geniculation between segments 7 and 8, segments 5–7 swollen and with aesthetasc on segment 5; segmental homologies in
<specimenCount id="0BB7FD75FFFAFFA7FAB9D469E1D7F850" box="[1407,1431,1905,1931]" pageId="57" pageNumber="293" type="male">♂</specimenCount>
: I, II–
<collectionCode id="7BA0AE39FFFAFFA7FF17D48DE543F874" box="[209,259,1941,1967]" pageId="57" pageNumber="293">VIII</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FEC8D48DE56CF874" box="[270,300,1941,1967]" pageId="57" pageNumber="293">IX</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FEFDD48DE523F874" box="[315,355,1941,1967]" pageId="57" pageNumber="293">XII</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FEA9D48DE5E2F874" box="[367,418,1941,1967]" pageId="57" pageNumber="293">XIII</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FE6BD48DE5A0F874" box="[429,480,1941,1967]" pageId="57" pageNumber="293">XIV</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FE29D48DE662F874" box="[495,546,1941,1967]" pageId="57" pageNumber="293">XVI</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FDEBD48DE614F874" box="[557,596,1941,1967]" pageId="57" pageNumber="293">VII</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FDA5D48DE6E9F874" box="[611,681,1941,1967]" pageId="57" pageNumber="293">XVIII</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FD72D48DE6A7F874" box="[692,743,1941,1967]" pageId="57" pageNumber="293">XIX</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FD30D48DE75FF874" box="[758,799,1941,1967]" pageId="57" pageNumber="293">XX</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FCEDD48DE71EF874" box="[811,862,1941,1967]" pageId="57" pageNumber="293">XXI</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FCAAD48DE7F2F874" box="[876,946,1941,1967]" pageId="57" pageNumber="293">XXIII</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FC7BD48DE043F874" box="[957,1027,1941,1967]" pageId="57" pageNumber="293">XXIV</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FBD7D48DE00DF874" box="[1041,1101,1941,1967]" pageId="57" pageNumber="293">XXV</collectionCode>
,
<collectionCode id="7BA0AE39FFFAFFA7FB9FD48DE0E0F874" box="[1113,1184,1941,1967]" pageId="57" pageNumber="293">XXVI</collectionCode>
–
<collectionCode id="7BA0AE39FFFAFFA7FB69D48DE149F874" box="[1199,1289,1941,1967]" pageId="57" pageNumber="293">XXVIII</collectionCode>
. Antenna not sexually dimorphic; proximal endopodal segment largely fused to basis forming allobasis, with pinnate seta on abexopodal margin; exopod 2-segmented, proximal segment with two setae, distal segment with two lateral and three apical elements; distal endopodal segment without penicillate elements. Mandible with three elements on basis; endopod with eight setae; exopod 1-segmented, with four setae. Maxillulary coxal epipodite represented by one seta. Maxilla with three endites on syncoxa; endopod discrete, with 4–5 setae. Maxilliped with three setae on syncoxa; basis with two setae on palmar margin; endopod represented by pinnate claw, accompanied by two accessory setae
</materialsCitation>
.
</paragraph>
<paragraph id="1D0E36FCFFF9FFA4FF01D253E0F0FDC2" blockId="58.[151,1436,151,538]" pageId="58" pageNumber="294">
P1 inner basal spine sexually dimorphic (with bifid tip in
<specimenCount id="0BB7FD75FFF9FFA4FCABD253E7C4FEBE" box="[877,900,331,357]" pageId="58" pageNumber="294" type="male">♂</specimenCount>
). P1 exopod 3-segmented; exp-2 only slightly longer than exp-1, with inner seta; exp-3 small, with two pinnate spines and two geniculate setae. P1 endopod 2- segmented; enp-1 elongate, almost twice as long as exopod, with moderately long, pinnate inner seta originating from proximal third (at 35% of ramus length in a region of reduced chitinization of segment wall); enp-2 very small, with one naked minute seta and two unipinnate claws. P2–P4 rami 3-segmented (except P2 endopod
<specimenCount id="0BB7FD75FFF9FFA4FB33D2C3E14AFE2E" box="[1269,1290,475,501]" pageId="58" pageNumber="294" type="male">♂</specimenCount>
occasionally 2-segmented); P3 endopod
<specimenCount id="0BB7FD75FFF9FFA4FE0CD2E7E59FFDC2" box="[458,479,511,537]" pageId="58" pageNumber="294" type="male">♂</specimenCount>
not sexually dimorphic. Armature formula of P2–P4 as follows:
</paragraph>
<paragraph id="1D0E36FCFFF9FFA4FD8BD154E00DFD3B" pageId="58" pageNumber="294">
<table id="6FB1C45CFFF90061FEA6D154E103FD3B" box="[352,1347,588,736]" gridcols="3" gridrows="4" pageId="58" pageNumber="294" rowsContinueFrom="57.[352,1113,162,310]" rowsContinueIn="60.[352,1303,162,310]">
<tr id="A38134BEFFF90061FEA6D154E103FDBD" box="[352,1347,588,614]" gridrow="0" pageId="58" pageNumber="294" rowspan-0="1">
<th id="E0505DC2FFF90061FD8BD154E6E1FDBD" box="[589,673,588,614]" gridcol="1" gridrow="0" pageId="58" pageNumber="294">exopod</th>
<th id="E0505DC2FFF90061FC3FD154E103FDBD" box="[1017,1347,588,614]" gridcol="2" gridrow="0" pageId="58" pageNumber="294">endopod</th>
</tr>
<tr id="A38134BEFFF90061FEA6D16CE103FD54" box="[352,1347,628,655]" gridrow="1" pageId="58" pageNumber="294">
<th id="E0505DC2FFF90061FEA6D16CE53AFD54" box="[352,378,628,655]" gridcol="0" gridrow="1" pageId="58" pageNumber="294">P2</th>
<td id="E0505DC2FFF90061FD8BD16CE6E1FD54" box="[589,673,628,655]" gridcol="1" gridrow="1" pageId="58" pageNumber="294">1.1.223</td>
<td id="E0505DC2FFF90061FC3FD16CE103FD54" box="[1017,1347,628,655]" gridcol="2" gridrow="1" pageId="58" pageNumber="294">1.1.221 [♂: 1.1.221 or 1.321]</td>
</tr>
<tr id="A38134BEFFF90061FEA6D186E103FD63" box="[352,1347,670,696]" gridrow="2" pageId="58" pageNumber="294">
<th id="E0505DC2FFF90061FEA6D186E53AFD63" box="[352,378,670,696]" gridcol="0" gridrow="2" pageId="58" pageNumber="294">P3</th>
<td id="E0505DC2FFF90061FD8BD186E6E1FD63" box="[589,673,670,696]" gridcol="1" gridrow="2" pageId="58" pageNumber="294">1.1.323</td>
<td id="E0505DC2FFF90061FC3FD186E103FD63" box="[1017,1347,670,696]" gridcol="2" gridrow="2" pageId="58" pageNumber="294">1.1.321</td>
</tr>
<tr id="A38134BEFFF90061FEA6D1DEE103FD3B" box="[352,1347,710,736]" gridrow="3" pageId="58" pageNumber="294">
<th id="E0505DC2FFF90061FEA6D1DEE53AFD3B" box="[352,378,710,736]" gridcol="0" gridrow="3" pageId="58" pageNumber="294">P4</th>
<td id="E0505DC2FFF90061FD8BD1DEE6E1FD3B" box="[589,673,710,736]" gridcol="1" gridrow="3" pageId="58" pageNumber="294">1.1.323</td>
<td id="E0505DC2FFF90061FC3FD1DEE103FD3B" box="[1017,1347,710,736]" gridcol="2" gridrow="3" pageId="58" pageNumber="294">1.1.221</td>
</tr>
</table>
</paragraph>
<paragraph id="1D0E36FCFFF9FFA4FF01D017E613FCA9" blockId="58.[151,1437,783,990]" pageId="58" pageNumber="294">
P
<specimenCount id="0BB7FD75FFF9FFA4FF1ED017E4BFFCF1" box="[216,255,783,810]" pageId="58" pageNumber="294" type="female">5 ♀</specimenCount>
endopodal lobe with five setae; inner margin without transverse striae; exopod elongate, with six elements. P
<specimenCount id="0BB7FD75FFF9FFA4FF6ED02BE492FC95" box="[168,210,819,846]" pageId="58" pageNumber="294" type="male">5 ♂</specimenCount>
endopodal lobe with two elements; exopod 1-segmented, with 5–6 elements. Vestigial P
<specimenCount id="0BB7FD75FFF9FFA4FB02D02BE0AEFC95" box="[1220,1262,819,846]" pageId="58" pageNumber="294" type="female">6 ♀</specimenCount>
represented by three short setae. P
<specimenCount id="0BB7FD75FFF9FFA4FEADD04FE5D5FCA9" box="[363,405,855,882]" pageId="58" pageNumber="294" type="male">6 ♂</specimenCount>
with three setae.
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFF9FFA4FF01D063E05CFC05" pageId="58" pageNumber="294" type="type_taxon">
<paragraph id="1D0E36FCFFF9FFA4FF01D063E5C7FC61" blockId="58.[151,1437,783,990]" pageId="58" pageNumber="294">
<emphasis id="2FC5EAEEFFF9FFA4FF01D063E524FC4E" bold="true" box="[199,356,891,917]" pageId="58" pageNumber="294">
<typeStatus id="C20A885EFFF9FFA4FF01D063E542FC4E" box="[199,258,891,917]" pageId="58" pageNumber="294">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="DAB14D7FFFF9FFA4FEABD063E736FC4D" authority="Norman &amp; Scott, 1905" authorityName="Norman &amp; Scott" authorityYear="1905" box="[365,886,891,918]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="294" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFF9FFA4FEABD063E627FC4E" box="[365,615,891,917]" italics="true" pageId="58" pageNumber="294">Parastenhelia anglica</emphasis>
<bibRefCitation id="79204B0DFFF9FFA4FDB6D064E736FC4D" author="Norman, A. M. &amp; Scott, T." box="[624,886,891,918]" pageId="58" pageNumber="294" pagination="284 - 300" refId="ref76779" refString="Norman, A. M. &amp; Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284 - 300. https: // doi. org / 10.1080 / 03745480509443044" type="journal article" year="1905">Norman &amp; Scott, 1905</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF9FFA4FC51D064E12EFC4D" authority="(Norman &amp; Scott, 1905)" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[919,1390,891,918]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="294" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFF9FFA4FC51D064E00DFC4E" box="[919,1101,891,917]" italics="true" pageId="58" pageNumber="294">Foweya anglica</emphasis>
(
<bibRefCitation id="79204B0DFFF9FFA4FB9BD063E126FC4D" author="Norman, A. M. &amp; Scott, T." box="[1117,1382,891,918]" pageId="58" pageNumber="294" pagination="284 - 300" refId="ref76779" refString="Norman, A. M. &amp; Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284 - 300. https: // doi. org / 10.1080 / 03745480509443044" type="journal article" year="1905">Norman &amp; Scott, 1905</bibRefCitation>
)
</taxonomicName>
(by original designation).
</paragraph>
<paragraph id="1D0E36FCFFF9FFA4FF01D0DBE05CFC05" blockId="58.[151,1437,783,990]" box="[199,1052,963,990]" pageId="58" pageNumber="294">
<emphasis id="2FC5EAEEFFF9FFA4FF01D0DBE52FFC06" bold="true" box="[199,367,963,989]" pageId="58" pageNumber="294">Other species.</emphasis>
<taxonomicName id="DAB14D7FFFF9FFA4FEB0D0DBE6FEFC05" authority="Kunz, 1975" authorityName="Kunz" authorityYear="1975" box="[374,702,963,990]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="294" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFF9FFA4FEB0D0DBE675FC06" box="[374,565,963,989]" italics="true" pageId="58" pageNumber="294">Karllangia tertia</emphasis>
<bibRefCitation id="79204B0DFFF9FFA4FDFDD0DCE6FEFC05" author="Kunz, H." box="[571,702,963,990]" pageId="58" pageNumber="294" pagination="179 - 212" refId="ref75043" refString="Kunz, H. (1975) Copepoda Harpacticoida aus dem Litoral des sudlichen Afrika. I. Teil. Kieler Meeresforschungen, 31, 179 - 212, pls. 1 - 30 (unpaginated)." type="journal article" year="1975">Kunz, 1975</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF9FFA4FD1AD0DCE057FC05" authority="(Kunz, 1975)" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[732,1047,963,990]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="58" pageNumber="294" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFF9FFA4FD1AD0DCE738FC06" box="[732,888,964,989]" italics="true" pageId="58" pageNumber="294">Foweya tertia</emphasis>
(
<bibRefCitation id="79204B0DFFF9FFA4FC41D0DBE04FFC05" author="Kunz, H." box="[903,1039,963,990]" pageId="58" pageNumber="294" pagination="179 - 212" refId="ref75043" refString="Kunz, H. (1975) Copepoda Harpacticoida aus dem Litoral des sudlichen Afrika. I. Teil. Kieler Meeresforschungen, 31, 179 - 212, pls. 1 - 30 (unpaginated)." type="journal article" year="1975">Kunz, 1975</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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