treatments-xml/data/83/09/87/8309879FF93A2035B9AEF8CDFCB8FD36.xml

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<mods:title id="6269D482C55C9E0A507C7FCFE12FDC08">Rediscovery and redescription of Laelaps lignicola G. &amp; R. Canestrini, a remarkable myrmecophilous mite of the genus Cosmolaelaps Berlese (Acari: Mesostigmata: Laelapidae) from Italy</mods:title>
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<heading id="505781E5F93A2024B9AEF8CDFC5AF8FB" bold="true" box="[151,920,1866,1893]" fontSize="11" level="1" pageId="1" pageNumber="22" reason="1">
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<taxonomicName id="CCA04D0AF93A2024B9AEF8CDFE7FF8FA" box="[151,445,1866,1892]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF93A2024B9AEF8CDFE7FF8FA" bold="true" box="[151,445,1866,1892]" italics="true" pageId="1" pageNumber="22">Cosmolaelaps lignicolus</emphasis>
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(G. &amp; R. Canestrini, 1882) comb. nov.
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(
<figureCitation id="939B2A0CF93A2024B9A6F8EAFECDF818" box="[159,271,1901,1926]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 7" captionStart-2="FIGURE 8" captionStart-3="FIGURES 9 – 12" captionStart-4="FIGURE 13 – 15" captionStart-5="FIGURES 16 – 21" captionStart-6="FIGURES 22 – 24" captionStart-7="FIGURES 25 – 26" captionStart-8="FIGURES 27 – 28" captionStart-9="FIGURES 29 – 34" captionStartId-0="4.[151,264,1939,1961]" captionStartId-1="5.[151,264,1778,1800]" captionStartId-2="6.[151,250,1772,1794]" captionStartId-3="7.[151,264,1183,1205]" captionStartId-4="8.[151,250,1851,1873]" captionStartId-5="9.[151,264,1586,1608]" captionStartId-6="10.[151,264,1580,1602]" captionStartId-7="12.[151,264,1969,1991]" captionStartId-8="13.[151,264,1437,1459]" captionStartId-9="15.[151,264,1915,1937]" captionTargetBox-0="[155,1421,421,1907]" captionTargetBox-1="[164,1424,214,1739]" captionTargetBox-2="[171,1416,210,1728]" captionTargetBox-3="[160,1424,207,1150]" captionTargetBox-4="[161,1420,221,1818]" captionTargetBox-5="[164,1418,210,1548]" captionTargetBox-6="[174,1419,211,1540]" captionTargetBox-7="[231,1335,206,1928]" captionTargetBox-8="[183,1388,193,1405]" captionTargetBox-9="[251,1331,223,1864]" captionTargetId-0="figure@4.[150,1436,409,1918]" captionTargetId-1="figure@5.[151,1436,193,1756]" captionTargetId-2="figure@6.[151,1436,193,1751]" captionTargetId-3="figure@7.[151,1436,193,1162]" captionTargetId-4="figure@8.[151,1436,193,1829]" captionTargetId-5="figure@9.[151,1436,193,1565]" captionTargetId-6="figure@10.[151,1436,193,1558]" captionTargetId-7="figure@12.[225,1345,193,1942]" captionTargetId-8="figure@13.[173,1411,181,1417]" captionTargetId-9="figure@15.[234,1351,205,1881]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="15" captionText-0="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." captionText-1="FIGURES 3 – 7. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4 – 7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm." captionText-2="FIGURE 8. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma (partial view; setae Zv 5 and glands gv 2 not discernible in this specimen). Scale bar = 100 µm." captionText-3="FIGURES 9 – 12. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma, four different specimens (partial views). Scale bars: 100 µm." captionText-4="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." captionText-5="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." captionText-6="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." captionText-7="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." captionText-8="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." captionText-9="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri-0="https://zenodo.org/record/292759/files/figure.png" httpUri-1="https://zenodo.org/record/292760/files/figure.png" httpUri-2="https://zenodo.org/record/292761/files/figure.png" httpUri-3="https://zenodo.org/record/292762/files/figure.png" httpUri-4="https://zenodo.org/record/292763/files/figure.png" httpUri-5="https://zenodo.org/record/292881/files/figure.png" httpUri-6="https://zenodo.org/record/292765/files/figure.png" httpUri-7="https://zenodo.org/record/292766/files/figure.png" httpUri-8="https://zenodo.org/record/292767/files/figure.png" httpUri-9="https://zenodo.org/record/292768/files/figure.png" pageId="1" pageNumber="22">Figs 1–31</figureCitation>
)
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<taxonomicName id="CCA04D0AF93A2024B9AEF832FE85F855" box="[151,327,1973,1995]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF93A2024B9AEF832FE85F855" box="[151,327,1973,1995]" italics="true" pageId="1" pageNumber="22">Laelaps lignicola</emphasis>
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G. &amp; 
<treatmentCitation id="8A011098F93A2024B8BAF831FDBFF855" author="Canestrini" box="[387,637,1973,1995]" page="74" pageId="1" pageNumber="22" year="1882">
<bibRefCitation id="6F314B78F93A2024B8BAF831FD95F855" author="Canestrini" box="[387,599,1973,1995]" pageId="1" pageNumber="22" refString="Canestrini, G. &amp; Canestrini, R. (1882 a) I Gamasi Italiani. Memoria. Atti della Societa Veneto - Trentina di Scienze Naturali Residente in Padova, 8, 3 - 82, pls. I - VII." type="journal article" year="1882" yearSuffix="a">R. Canestrini, 1882a</bibRefCitation>
: 74
</treatmentCitation>
. 
<taxonomicName id="CCA04D0AF93A2024B9AEF853FE85F874" box="[151,327,2004,2026]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF93A2024B9AEF853FE85F874" box="[151,327,2004,2026]" italics="true" pageId="1" pageNumber="22">Laelaps lignicola</emphasis>
</taxonomicName>
G. &amp; 
<treatmentCitation id="8A011098F93A2024B8BAF852FDBDF874" author="Canestrini" box="[387,639,2004,2026]" page="72" pageId="1" pageNumber="22" year="1882">
<bibRefCitation id="6F314B78F93A2024B8BAF852FD9AF874" author="Canestrini" box="[387,600,2004,2026]" pageId="1" pageNumber="22" refString="Canestrini, G. &amp; Canestrini, R. (1882 b) I Gamasi Italiani. Monografia. Proserpini, Padova, 80 pp., pls. I - VII." type="book" year="1882" yearSuffix="b">R. Canestrini, 1882b</bibRefCitation>
: 72
</treatmentCitation>
.
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</paragraph>
<paragraph id="0B1F3689F9392027B9AEFF1EFD40FF52" blockId="2.[151,1268,152,266]" pageId="2" pageNumber="23">
<treatmentCitationGroup id="2BB011A7F9392027B9AEFF1EFD40FF52" pageId="2" pageNumber="23">
<taxonomicName id="CCA04D0AF9392027B9AEFF1EFEA0FF30" box="[151,354,152,174]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027B9AEFF1EFEA0FF30" box="[151,354,152,174]" italics="true" pageId="2" pageNumber="23">Hypoaspis lignicola</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="8A011098F9392027B8B9FF1FFD90FF30" author="Canestrini" box="[384,594,152,174]" page="89" pageId="2" pageNumber="23" year="1885">
<bibRefCitation id="6F314B78F9392027B8B9FF1FFDEEFF30" author="Canestrini" box="[384,556,152,174]" pageId="2" pageNumber="23" refString="Canestrini, G. (1885) Prospetto dell'Acarofauna Italiana, Volume 1, Famiglie: Oribatini, Gamasini, Hoplopini, Trombidini, Rhyncholophini. Proserpini, Padova, 163 pp., pls. I - IX." type="book" year="1885">Canestrini, 1885</bibRefCitation>
: 89
</treatmentCitation>
; 
<treatmentCitation id="8A011098F9392027BB64FF1FFCF0FF30" author="Oudemans" box="[605,818,152,174]" page="24" pageId="2" pageNumber="23" year="1902">
<bibRefCitation id="6F314B78F9392027BB64FF1FFCCFFF30" author="Oudemans" box="[605,781,152,174]" pageId="2" pageNumber="23" refString="Oudemans, A. C. (1902) New list of Dutch Acari. Second part. Tijdschrift voor Entomologie, 45, 1 - 52, pls. 1 - 6." type="journal article" year="1902">Oudemans, 1902</bibRefCitation>
: 24
</treatmentCitation>
; 
<treatmentCitation id="8A011098F9392027BA05FF1FFBDCFF30" author="Oudemans" box="[828,1054,152,174]" page="129" pageId="2" pageNumber="23" year="1903">
<bibRefCitation id="6F314B78F9392027BA05FF1FFC29FF30" author="Oudemans" box="[828,1003,152,174]" pageId="2" pageNumber="23" refString="Oudemans, A. C. (1903) Notes on Acari. Fifth series. Tijdschrift voor Entomologie, 45, 123 - 150." type="journal article" year="1903">Oudemans, 1903</bibRefCitation>
: 129
</treatmentCitation>
; Moreira, 2014: 409. 
<taxonomicName id="CCA04D0AF9392027B9AEFF31FE53FF52" box="[151,401,182,204]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola" subGenus="Iphis">
<emphasis id="39D4EA9BF9392027B9AEFF31FF25FF52" box="[151,231,182,204]" italics="true" pageId="2" pageNumber="23">Laelaps</emphasis>
(
<emphasis id="39D4EA9BF9392027B9CFFF31FEEBFF52" box="[246,297,182,204]" italics="true" pageId="2" pageNumber="23">Iphis</emphasis>
) 
<emphasis id="39D4EA9BF9392027B80EFF31FE53FF52" box="[311,401,182,204]" italics="true" pageId="2" pageNumber="23">lignicola</emphasis>
</taxonomicName>
.—Berlese, 1892: 70, 2.
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</paragraph>
<paragraph id="0B1F3689F9392027B9AEFF52FDE1FF75" blockId="2.[151,1268,152,266]" box="[151,547,213,235]" pageId="2" pageNumber="23">
<treatmentCitationGroup id="2BB011A7F9392027B9AEFF52FDE1FF75" box="[151,547,213,235]" pageId="2" pageNumber="23">
<taxonomicName id="CCA04D0AF9392027B9AEFF52FE85FF75" box="[151,327,213,235]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027B9AEFF52FE85FF75" box="[151,327,213,235]" italics="true" pageId="2" pageNumber="23">Laelaps lignicola</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="8A011098F9392027B85CFF52FDDDFF75" author="Tipton" box="[357,543,213,235]" page="299" pageId="2" pageNumber="23" year="1960">
<bibRefCitation id="6F314B78F9392027B85CFF52FE2FFF75" author="Tipton" box="[357,493,213,235]" pageId="2" pageNumber="23" refString="Tipton, V. J. (1960) The genus Laelaps with a review of the Laelaptinae and a new subfamily Alphalaelaptinae (Acarina: Laelaptidae). University of California Publications in Entomology, Technical Bulletin, Berkeley, 16 (6), 233 - 356." type="journal article" year="1960">Tipton, 1960</bibRefCitation>
: 299
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="0B1F3689F9392027B9AEFF73FD58FE94" blockId="2.[151,1268,152,266]" box="[151,666,244,266]" pageId="2" pageNumber="23">
<treatmentCitationGroup id="2BB011A7F9392027B9AEFF73FD58FE94" box="[151,666,244,266]" pageId="2" pageNumber="23">
<taxonomicName id="CCA04D0AF9392027B9AEFF73FE4BFE94" box="[151,393,244,266]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027B9AEFF73FE4BFE94" box="[151,393,244,266]" italics="true" pageId="2" pageNumber="23">Gymnolaelaps lignicola</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="8A011098F9392027B89EFF73FD54FE94" author="Bernini" box="[423,662,244,266]" page="27" pageId="2" pageNumber="23" year="1995">
<bibRefCitation id="6F314B78F9392027B89EFF73FDB2FE94" author="Bernini" box="[423,624,244,266]" pageId="2" pageNumber="23" refString="Bernini, E., Castagnoli, M. &amp; Nannelli, R. (1995) Arachnida Acari. In: Minelli, A., Ruffo, S. &amp; La Posta, S. (Eds.), Checklist delle specie della fauna italiana, 24, pp. 1 - 131. [Calderini, Bologna]" type="journal article" year="1995">
Bernini 
<emphasis id="39D4EA9BF9392027B8C2FF72FDEAFE94" box="[507,552,244,266]" italics="true" pageId="2" pageNumber="23">et al</emphasis>
., 1995
</bibRefCitation>
: 27
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="43BA6502F9392027B9AEFEB1FEE4FA19" pageId="2" pageNumber="23" type="materials_examined">
<paragraph id="0B1F3689F9392027B9AEFEB1FC35FE09" blockId="2.[151,1437,310,2027]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B9AEFEB1FE45FED1" bold="true" box="[151,391,310,335]" pageId="2" pageNumber="23">Material examined.</emphasis>
The material of 
<taxonomicName id="CCA04D0AF9392027BB67FEB0FD37FED0" box="[606,757,310,334]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF9392027BB67FEB0FDB3FED0" box="[606,625,311,334]" italics="true" pageId="2" pageNumber="23">C</emphasis>
. 
<emphasis id="39D4EA9BF9392027BBBFFEB1FD37FED0" box="[646,757,310,334]" italics="true" pageId="2" pageNumber="23">lignicolus</emphasis>
</taxonomicName>
studied here was collected mostly by the senior author exclusively from nests of 
<taxonomicName id="CCA04D0AF9392027B885FEDCFCC7FEED" authority="Olivier" authorityName="Olivier" box="[444,773,346,371]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="emarginatus">
<emphasis id="39D4EA9BF9392027B885FEDCFD5BFEEC" box="[444,665,347,370]" italics="true" pageId="2" pageNumber="23">Lasius emarginatus</emphasis>
(Olivier)
</taxonomicName>
, and where it is not indicated, this material is deposited in the Collection of Lentate sul Seveso Civic Museum, Monza &amp; Brianza, 
<collectingCountry id="73B77619F9392027BA84FEF9FC30FE09" box="[957,1010,382,407]" name="Italy" pageId="2" pageNumber="23">Italy</collectingCountry>
.
</paragraph>
<paragraph id="0B1F3689F9392027B9FEFE24FEE4FA19" blockId="2.[151,1437,310,2027]" pageId="2" pageNumber="23">
<materialsCitation id="BBC83CD4F9392027B9FEFE24FADAFD9D" ID-GBIF-Occurrence="1434654789" collectionCode="COLL" collectorName="G. R. Can. &amp; Padova" county="Padova" location="Centro" municipality="Berlese Acaroteca" pageId="2" pageNumber="23" specimenCount="3" specimenCount-female="3" stateProvince="Veneto">
<emphasis id="39D4EA9BF9392027B9FEFE24FE74FE25" box="[199,438,419,443]" italics="true" pageId="2" pageNumber="23">Specimens examined</emphasis>
: 
<specimenCount id="1DA6FD00F9392027B8F5FE24FD80FE25" box="[460,578,418,443]" pageId="2" pageNumber="23" type="female">2 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BB6DFE24FD67FE24" box="[596,677,419,442]" country="Italy" name="Veneto" pageId="2" pageNumber="23">Veneto</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027BB8EFE25FCCEFE25" box="[695,780,418,443]" pageId="2" pageNumber="23">Padova</collectingCounty>
, rotting wood, other collecting data unknown, 
<collectingMunicipality id="EB7BACF3F9392027BC71FE25FECBFE41" pageId="2" pageNumber="23">Berlese Acaroteca</collectingMunicipality>
(
<location id="0E7F6052F9392027B824FE40FEA9FE41" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B824FE40FEA9FE41" box="[285,363,455,479]" county="Padova" municipality="Berlese Acaroteca" name="Centro" pageId="2" pageNumber="23" stateProvince="Veneto">Centro</location>
di ricerca per l'agrobiologia e la pedologia, 
<location id="0E7F6052F9392027BABEFE41FC2FFE41" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BABEFE41FC2FFE41" box="[903,1005,454,479]" county="Padova" municipality="Berlese Acaroteca" name="Florence" pageId="2" pageNumber="23" stateProvince="Veneto">Florence</location>
), slide 44/38 [original label: 
<collectionCode id="6DB1AE4CF9392027BC75FE40FA5BFE41" box="[1356,1433,455,479]" pageId="2" pageNumber="23">COLL</collectionCode>
. BERLESE— 
<taxonomicName id="CCA04D0AF9392027B815FE6CFE60FD9C" box="[300,418,491,514]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027B815FE6CFE60FD9C" box="[300,418,491,514]" italics="true" pageId="2" pageNumber="23">Hypoaspis</emphasis>
</taxonomicName>
<specimenCount id="1DA6FD00F9392027B895FE6CFE79FD9D" box="[428,443,491,515]" pageId="2" pageNumber="23" type="female">♀</specimenCount>
—(
<taxonomicName id="CCA04D0AF9392027B8DFFE6CFD4AFD9D" box="[486,648,491,515]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027B8DFFE6CFD4AFD9D" box="[486,648,491,515]" italics="true" pageId="2" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
)— 
<taxonomicName id="CCA04D0AF9392027BB94FE6CFCD3FD9D" box="[685,785,491,515]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027BB94FE6CFCD3FD9D" box="[685,785,491,515]" italics="true" pageId="2" pageNumber="23">lignicola</emphasis>
</taxonomicName>
—cotipi (
<collectorName id="A655535FF9392027BA46FE6CFC35FD9D" box="[895,1015,490,515]" pageId="2" pageNumber="23">G. R. Can.</collectorName>
)— 
<collectorName id="A655535FF9392027BD22FE6DFBB2FD9D" box="[1051,1136,490,515]" pageId="2" pageNumber="23">Padova</collectorName>
legno guasto]
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BC1FFE6CFB80FDD5" ID-GBIF-Occurrence="1434654796" collectionCode="COLL" collectorName="A. Dodero &amp; Berlese Acaroteca &amp; G. R. Can. &amp; Dodero" location="Genova" pageId="2" pageNumber="23" specimenCount="3" specimenCount-female="3" stateProvince="Liguria">
<specimenCount id="1DA6FD00F9392027BC1FFE6CFA5AFD9D" box="[1318,1432,490,515]" pageId="2" pageNumber="23" type="female">2 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027B9AEFD89FF2EFDB9" box="[151,236,526,551]" country="Italy" name="Liguria" pageId="2" pageNumber="23">Liguria</collectingRegion>
, 
<location id="0E7F6052F9392027B838FD88FE99FDB9" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B838FD88FE99FDB9" box="[257,347,527,551]" name="Genova" pageId="2" pageNumber="23" stateProvince="Liguria">Genova</location>
, date unknown, 
<collectorName id="A655535FF9392027BB16FD88FD6DFDB9" box="[559,687,526,551]" pageId="2" pageNumber="23">A. Dodero</collectorName>
coll., 
<collectorName id="A655535FF9392027BA31FD89FC23FDB9" box="[776,993,526,551]" pageId="2" pageNumber="23">Berlese Acaroteca</collectorName>
, slide 50/34 [original label: 
<collectionCode id="6DB1AE4CF9392027BC75FD88FA5BFDB9" box="[1356,1433,527,551]" pageId="2" pageNumber="23">COLL</collectionCode>
. BERLESE— 
<taxonomicName id="CCA04D0AF9392027B815FDB4FE0CFDD5" box="[300,462,563,587]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027B815FDB4FE0CFDD5" box="[300,462,563,587]" italics="true" pageId="2" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
<specimenCount id="1DA6FD00F9392027B8E0FDB4FE39FDD4" box="[473,507,563,586]" pageId="2" pageNumber="23" type="female">♀♀</specimenCount>
— 
<taxonomicName id="CCA04D0AF9392027BB26FDB4FD41FDD5" box="[543,643,563,587]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027BB26FDB4FD41FDD5" box="[543,643,563,587]" italics="true" pageId="2" pageNumber="23">lignicola</emphasis>
</taxonomicName>
—(
<collectorName id="A655535FF9392027BB91FDB4FCE0FDD5" box="[680,802,562,587]" pageId="2" pageNumber="23">G. R. Can.</collectorName>
)—Genova—(
<collectorName id="A655535FF9392027BAFDFDB5FBD9FDD5" box="[964,1051,562,587]" pageId="2" pageNumber="23">Dodero</collectorName>
!)]
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BD6FFDB4FC64FD0D" ID-GBIF-Occurrence="1434654794" collectionCode="COLL" collectorName="A. Dodero &amp; Berlese Acaroteca &amp; G. R. Can. &amp; Dodero" location="Genova" pageId="2" pageNumber="23" specimenCount="3" specimenCount-female="3" stateProvince="Liguria">
<specimenCount id="1DA6FD00F9392027BD6FFDB4FB0BFDD5" box="[1110,1225,562,587]" pageId="2" pageNumber="23" type="female">2 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BDE0FDB5FAECFDD5" box="[1241,1326,562,587]" country="Italy" name="Liguria" pageId="2" pageNumber="23">Liguria</collectingRegion>
, 
<location id="0E7F6052F9392027BC07FDB4FA5AFDD5" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BC07FDB4FA5AFDD5" box="[1342,1432,563,587]" name="Genova" pageId="2" pageNumber="23" stateProvince="Liguria">Genova</location>
, date unknown, 
<collectorName id="A655535FF9392027B867FDD0FE21FDF1" box="[350,483,598,623]" pageId="2" pageNumber="23">A. Dodero</collectorName>
coll., 
<collectorName id="A655535FF9392027BB7DFDD1FCE0FDF1" box="[580,802,598,623]" pageId="2" pageNumber="23">Berlese Acaroteca</collectorName>
, slide 50/35 [original label: 
<collectionCode id="6DB1AE4CF9392027BD9BFDD0FB2DFDF1" box="[1186,1263,599,623]" pageId="2" pageNumber="23">COLL</collectionCode>
. BERLESE— 
<emphasis id="39D4EA9BF9392027B9AEFDFCFE7AFD0D" box="[151,440,635,659]" italics="true" pageId="2" pageNumber="23">
<taxonomicName id="CCA04D0AF9392027B9AEFDFCFEFBFD0D" box="[151,313,635,659]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">Gymnolaelaps</taxonomicName>
— 
<taxonomicName id="CCA04D0AF9392027B86DFDFCFE7AFD0D" box="[340,440,635,659]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">lignicola</taxonomicName>
</emphasis>
<specimenCount id="1DA6FD00F9392027B8FBFDFCFE20FD0D" box="[450,482,635,659]" pageId="2" pageNumber="23" type="female">♀♀</specimenCount>
—(
<collectorName id="A655535FF9392027BB34FDFCFD45FD0D" box="[525,647,635,659]" pageId="2" pageNumber="23">G. R. Can.</collectorName>
)—Genova—(
<collectorName id="A655535FF9392027BA10FDFCFC42FD0D" box="[809,896,635,659]" pageId="2" pageNumber="23">Dodero</collectorName>
!)]
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BA8FFDFCFC75FD45" ID-GBIF-Occurrence="1434654784" collectionCode="COLL" collectorName="A. Dodero &amp; Berlese Acaroteca &amp; G. R. Can. &amp; Dodero" location="Genova" pageId="2" pageNumber="23" specimenCount="2" specimenCount-female="2" stateProvince="Liguria">
<specimenCount id="1DA6FD00F9392027BA8FFDFCFBDEFD0D" box="[950,1052,634,659]" pageId="2" pageNumber="23" type="female">1 female</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BD13FDFDFBBDFD0D" box="[1066,1151,634,659]" country="Italy" name="Liguria" pageId="2" pageNumber="23">Liguria</collectingRegion>
, 
<location id="0E7F6052F9392027BDB4FDFCFB25FD0D" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BDB4FDFCFB25FD0D" box="[1165,1255,635,659]" name="Genova" pageId="2" pageNumber="23" stateProvince="Liguria">Genova</location>
, date unknown, 
<collectorName id="A655535FF9392027B9AEFD18FED9FD29" box="[151,283,670,695]" pageId="2" pageNumber="23">A. Dodero</collectorName>
coll., 
<collectorName id="A655535FF9392027B845FD19FD9BFD29" box="[380,601,670,695]" pageId="2" pageNumber="23">Berlese Acaroteca</collectorName>
, slide 208/50 [original label: 
<collectionCode id="6DB1AE4CF9392027BADCFD18FBF0FD29" box="[997,1074,671,695]" pageId="2" pageNumber="23">COLL</collectionCode>
. BERLESE— 
<taxonomicName id="CCA04D0AF9392027BDD9FD18FA94FD28" box="[1248,1366,671,694]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027BDD9FD18FA94FD28" box="[1248,1366,671,694]" italics="true" pageId="2" pageNumber="23">Hypoaspis</emphasis>
</taxonomicName>
<specimenCount id="1DA6FD00F9392027BC52FD18FAB8FD28" box="[1387,1402,671,694]" pageId="2" pageNumber="23" type="female">♀</specimenCount>
— (
<taxonomicName id="CCA04D0AF9392027B999FD44FE80FD45" box="[160,322,707,731]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027B999FD44FE80FD45" box="[160,322,707,731]" italics="true" pageId="2" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
)— 
<taxonomicName id="CCA04D0AF9392027B85EFD44FE09FD45" box="[359,459,707,731]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027B85EFD44FE09FD45" box="[359,459,707,731]" italics="true" pageId="2" pageNumber="23">lignicola</emphasis>
</taxonomicName>
—rotta (
<collectorName id="A655535FF9392027BB10FD44FD59FD45" box="[553,667,706,731]" pageId="2" pageNumber="23">G. R. Can.</collectorName>
)—Genova—(
<collectorName id="A655535FF9392027BA05FD45FC51FD45" box="[828,915,706,731]" pageId="2" pageNumber="23">Dodero</collectorName>
!)]
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BAFDFD44FC58FCBD" ID-GBIF-Occurrence="1434654788" collectionCode="COLL" collectorName="A. Dodero &amp; Berlese Acaroteca &amp; G. R. Can. &amp; Dodero" location="Genova" pageId="2" pageNumber="23" specimenCount="2" specimenCount-female="2" stateProvince="Liguria">
<specimenCount id="1DA6FD00F9392027BAFDFD44FBEAFD45" box="[964,1064,706,731]" pageId="2" pageNumber="23" type="female">1 female</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BD0AFD45FB4AFD45" box="[1075,1160,706,731]" country="Italy" name="Liguria" pageId="2" pageNumber="23">Liguria</collectingRegion>
, 
<location id="0E7F6052F9392027BDAAFD44FB2FFD45" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BDAAFD44FB2FFD45" box="[1171,1261,707,731]" name="Genova" pageId="2" pageNumber="23" stateProvince="Liguria">Genova</location>
, date unknown, 
<collectorName id="A655535FF9392027B9AEFD60FEDEFD61" box="[151,284,742,767]" pageId="2" pageNumber="23">A. Dodero</collectorName>
coll., 
<collectorName id="A655535FF9392027B8B9FD61FD9CFD61" box="[384,606,742,767]" pageId="2" pageNumber="23">Berlese Acaroteca</collectorName>
, slide 209/2 [original label: 
<collectionCode id="6DB1AE4CF9392027BADAFD60FBF2FD61" box="[995,1072,743,767]" pageId="2" pageNumber="23">COLL</collectionCode>
. BERLESE— 
<taxonomicName id="CCA04D0AF9392027BDE6FD60FA97FD60" box="[1247,1365,743,766]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027BDE6FD60FA97FD60" box="[1247,1365,743,766]" italics="true" pageId="2" pageNumber="23">Hypoaspis</emphasis>
</taxonomicName>
<specimenCount id="1DA6FD00F9392027BC52FD60FAB8FD60" box="[1387,1402,743,766]" pageId="2" pageNumber="23" type="female">♀</specimenCount>
— (
<taxonomicName id="CCA04D0AF9392027B999FC8CFE80FCBD" box="[160,322,779,803]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF9392027B999FC8CFE80FCBD" box="[160,322,779,803]" italics="true" pageId="2" pageNumber="23">Gymnolaelaps</emphasis>
</taxonomicName>
)— 
<taxonomicName id="CCA04D0AF9392027B85EFC8CFE09FCBD" box="[359,459,779,803]" class="Arachnida" family="Laelapidae" genus="Gymnolaelaps" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lignicola">
<emphasis id="39D4EA9BF9392027B85EFC8CFE09FCBD" box="[359,459,779,803]" italics="true" pageId="2" pageNumber="23">lignicola</emphasis>
</taxonomicName>
—(
<collectorName id="A655535FF9392027B8C9FC8CFDB6FCBD" box="[496,628,778,803]" pageId="2" pageNumber="23">G. R. Can.</collectorName>
)—Genova—(
<collectorName id="A655535FF9392027BA2FFC8DFCAFFCBD" box="[790,877,778,803]" pageId="2" pageNumber="23">Dodero</collectorName>
!)]
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BA88FC8CFCC1FCD9" ID-GBIF-Occurrence="1434654785" collectingDate="2009-10-04" location="Solaro" municipality="Milano province" pageId="2" pageNumber="23" specimenCount="11" specimenCount-female="10" specimenCount-male="1" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BA88FC8CFBF5FCBD" box="[945,1079,778,803]" pageId="2" pageNumber="23" type="female">10 females</specimenCount>
, 
<specimenCount id="1DA6FD00F9392027BD75FC8CFB63FCBD" box="[1100,1185,778,803]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
and 1 deutonymph, 
<collectingRegion id="C964F86BF9392027B9AEFCA9FED7FCD9" box="[151,277,814,839]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027B81AFCA9FE21FCD9" box="[291,483,814,839]" pageId="2" pageNumber="23">Milano province</collectingMunicipality>
, 
<location id="0E7F6052F9392027B8C8FCA9FDFFFCD9" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B8C8FCA9FDFFFCD9" box="[497,573,814,839]" municipality="Milano province" name="Solaro" pageId="2" pageNumber="23" stateProvince="Lombardia">Solaro</location>
, 
<collectingDate id="6F5AE9A1F9392027BB72FCA8FCC1FCD9" box="[587,771,814,839]" pageId="2" pageNumber="23" value="2009-10-04">
<date id="7F1E1049F9392027BB72FCA8FCC1FCD9" box="[587,771,814,839]" pageId="2" pageNumber="23" value="2009-10-04">4 October 2009</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BA2BFCA9FDC6FCF5" ID-GBIF-Occurrence="1434654786" collectingDate="2013-08-20" county="Foggia province" location="Foresta Umbra" municipality="Vieste" pageId="2" pageNumber="23" specimenCount="17" specimenCount-female="16" specimenCount-male="1" stateProvince="Puglia">
<specimenCount id="1DA6FD00F9392027BA2BFCA9FC4DFCD9" box="[786,911,814,839]" pageId="2" pageNumber="23" type="female">16 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BAF5FCA8FBDFFCD9" box="[972,1053,814,839]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BD12FCA9FBB4FCD9" box="[1067,1142,814,839]" country="Italy" name="Puglia" pageId="2" pageNumber="23">Puglia</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027BDBDFCA9FA83FCD9" box="[1156,1345,814,839]" pageId="2" pageNumber="23">Foggia province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027BC76FCA9FA5AFCD9" box="[1359,1432,814,839]" pageId="2" pageNumber="23">Vieste</collectingMunicipality>
, 
<location id="0E7F6052F9392027B9AEFCD4FE80FCF5" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B9AEFCD4FE80FCF5" box="[151,322,850,875]" county="Foggia province" municipality="Vieste" name="Foresta Umbra" pageId="2" pageNumber="23" stateProvince="Puglia">Foresta Umbra</location>
, 
<collectingDate id="6F5AE9A1F9392027B874FCD4FDC6FCF5" box="[333,516,851,875]" pageId="2" pageNumber="23" value="2013-08-20">
<date id="7F1E1049F9392027B874FCD4FDC6FCF5" box="[333,516,851,875]" pageId="2" pageNumber="23" value="2013-08-20">20 August 2013</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BB29FCD5FCE4FC49" ID-GBIF-Occurrence="1434654792" collectingDate="2013-11-10" collectorName="Arthropods Collection &amp; Natural History Museum &amp; Athens &amp; United States" country="Iran" county="Como province" location="Alpe" municipality="Albavilla" pageId="2" pageNumber="23" specimenCount="104" specimenCount-female="99" specimenCount-male="5" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BB29FCD5FD49FCF5" box="[528,651,850,875]" pageId="2" pageNumber="23" type="female">85 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BBFBFCD4FCDEFCF5" box="[706,796,850,875]" pageId="2" pageNumber="23" type="male">4 males</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BA11FCD5FC64FCF5" box="[808,934,850,875]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027BA88FCD4FBA0FCF5" box="[945,1122,850,875]" pageId="2" pageNumber="23">Como province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027BD57FCD5FB1BFCF5" box="[1134,1241,850,875]" pageId="2" pageNumber="23">Albavilla</collectingMunicipality>
, 
<location id="0E7F6052F9392027BDDDFCD5FAD8FCF5" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BDDDFCD5FAD8FCF5" box="[1252,1306,850,875]" country="Iran" county="Como province" municipality="Albavilla" name="Alpe" pageId="2" pageNumber="23" stateProvince="Lombardia">Alpe</location>
del 
<location id="0E7F6052F9392027BC75FCD5FA5AFCF5" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BC75FCD5FA5AFCF5" box="[1356,1432,850,875]" country="Iran" county="Como province" municipality="Albavilla" name="Vicere" pageId="2" pageNumber="23" stateProvince="Lombardia">Vicerè</location>
, 
<collectingDate id="6F5AE9A1F9392027B9AEFCF0FEBEFC11" box="[151,380,886,911]" pageId="2" pageNumber="23" value="2013-11-10">
<date id="7F1E1049F9392027B9AEFCF0FEBEFC11" box="[151,380,886,911]" pageId="2" pageNumber="23" value="2013-11-10">10 November 2013</date>
</collectingDate>
(
<specimenCount id="1DA6FD00F9392027B8AAFCF0FDD6FC11" box="[403,532,886,911]" pageId="2" pageNumber="23" type="female">10 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BB61FCF0FD68FC11" box="[600,682,886,911]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
deposited in the Acarological Collection, Department of Plant Protection, 
<collectingRegion id="C964F86BF9392027B824FC1DFE97FC2D" box="[285,341,922,947]" country="Iran" name="Yazd" pageId="2" pageNumber="23">Yazd</collectingRegion>
Branch, Islamic Azad University, 
<collectingCountry id="73B77619F9392027BBCAFC1CFCE6FC2C" box="[755,804,923,946]" name="Iran" pageId="2" pageNumber="23">Iran</collectingCountry>
; 
<specimenCount id="1DA6FD00F9392027BA0DFC1CFC67FC2D" box="[820,933,922,947]" pageId="2" pageNumber="23" type="female">4 females</specimenCount>
deposited in the 
<collectorName id="A655535FF9392027BD40FC1DFABBFC2D" box="[1145,1401,922,947]" pageId="2" pageNumber="23">Arthropods Collection</collectorName>
of 
<collectorName id="A655535FF9392027B9AEFC39FE70FC48" box="[151,434,958,983]" pageId="2" pageNumber="23">Natural History Museum</collectorName>
, 
<collectorName id="A655535FF9392027B887FC39FDD2FC49" box="[446,528,958,983]" pageId="2" pageNumber="23">Athens</collectorName>
, 
<collectingCountry id="73B77619F9392027BB25FC39FDBBFC49" box="[540,633,958,983]" name="Georgia" pageId="2" pageNumber="23">Georgia</collectingCountry>
, 
<collectorName id="A655535FF9392027BBBDFC39FCDCFC49" box="[644,798,958,983]" pageId="2" pageNumber="23">United States</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BA0BFC39FE19FC65" ID-GBIF-Occurrence="1434654798" collectingDate="2014-03-09" country="Iran" county="Como province" location="Monte Cornizzolo" municipality="Eupilio" pageId="2" pageNumber="23" specimenCount="6" specimenCount-female="6" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BA0BFC39FC60FC49" box="[818,930,958,983]" pageId="2" pageNumber="23" type="female">6 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BA94FC39FBE9FC49" box="[941,1067,958,983]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027BD0FFC38FB25FC49" box="[1078,1255,958,983]" pageId="2" pageNumber="23">Como province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027BDCBFC39FA85FC49" box="[1266,1351,958,983]" pageId="2" pageNumber="23">Eupilio</collectingMunicipality>
, 
<location id="0E7F6052F9392027BC6AFC38FED4FC65" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BC6AFC38FED4FC65" country="Iran" county="Como province" municipality="Eupilio" name="Monte Cornizzolo" pageId="2" pageNumber="23" stateProvince="Lombardia">Monte Cornizzolo</location>
, 
<collectingDate id="6F5AE9A1F9392027B813FC64FE19FC65" box="[298,475,994,1019]" pageId="2" pageNumber="23" value="2014-03-09">
<date id="7F1E1049F9392027B813FC64FE19FC65" box="[298,475,994,1019]" pageId="2" pageNumber="23" value="2014-03-09">9 March 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027B8D6FC65FACFFC65" ID-GBIF-Occurrence="1434654787" box="[495,1293,994,1019]" collectingDate="2014-03-23" country="Iran" location="Senago" municipality="Milano province" pageId="2" pageNumber="23" specimenCount="5" specimenCount-female="5" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027B8D6FC65FDA5FC65" box="[495,615,994,1019]" pageId="2" pageNumber="23" type="female">5 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BB43FC65FD3AFC65" box="[634,760,994,1019]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027BA32FC65FC13FC65" box="[779,977,994,1019]" pageId="2" pageNumber="23">Milano province</collectingMunicipality>
, 
<location id="0E7F6052F9392027BADDFC64FBFBFC65" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BADDFC64FBFBFC65" box="[996,1081,995,1019]" country="Iran" municipality="Milano province" name="Senago" pageId="2" pageNumber="23" stateProvince="Lombardia">Senago</location>
, 
<collectingDate id="6F5AE9A1F9392027BD74FC64FACFFC65" box="[1101,1293,994,1019]" pageId="2" pageNumber="23" value="2014-03-23">
<date id="7F1E1049F9392027BD74FC64FACFFC65" box="[1101,1293,994,1019]" pageId="2" pageNumber="23" value="2014-03-23">23 March 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BC18FC64FC44FB81" ID-GBIF-Occurrence="1434654797" collectingDate="2014-03-30" country="Iran" county="Milano province" location="Castellazzo" municipality="Bollate" pageId="2" pageNumber="23" specimenCount="3" specimenCount-female="3" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BC18FC64FA5AFC65" box="[1313,1432,994,1019]" pageId="2" pageNumber="23" type="female">3 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027B9AEFB81FED7FB81" box="[151,277,1030,1055]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027B819FB81FE1FFB81" box="[288,477,1030,1055]" pageId="2" pageNumber="23">Milano province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027B8D1FB81FDFEFB81" box="[488,572,1030,1055]" pageId="2" pageNumber="23">Bollate</collectingMunicipality>
, 
<location id="0E7F6052F9392027BB71FB81FD0EFB81" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BB71FB81FD0EFB81" box="[584,716,1030,1055]" country="Iran" county="Milano province" municipality="Bollate" name="Castellazzo" pageId="2" pageNumber="23" stateProvince="Lombardia">Castellazzo</location>
, 
<collectingDate id="6F5AE9A1F9392027BBEEFB80FC44FB81" box="[727,902,1030,1055]" pageId="2" pageNumber="23" value="2014-03-30">
<date id="7F1E1049F9392027BBEEFB80FC44FB81" box="[727,902,1030,1055]" pageId="2" pageNumber="23" value="2014-03-30">30 March 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BAABFB81FC8CFBDD" ID-GBIF-Occurrence="1434654795" collectingDate="2014-05-25" country="Italy" location="Limido Comasco" municipality="Como province" pageId="2" pageNumber="23" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BAABFB81FBC3FB81" box="[914,1025,1030,1055]" pageId="2" pageNumber="23" type="female">6 females</specimenCount>
, 
<specimenCount id="1DA6FD00F9392027BD34FB80FB9AFB81" box="[1037,1112,1030,1055]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
and 2 deutonymphs, 
<collectingCountry id="73B77619F9392027BC73FB80FA5BFB81" box="[1354,1433,1031,1055]" name="Italy" pageId="2" pageNumber="23">ITALY</collectingCountry>
, 
<collectingRegion id="C964F86BF9392027B9AEFBADFED7FBDD" box="[151,277,1066,1091]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027B818FBACFE11FBDD" box="[289,467,1066,1091]" pageId="2" pageNumber="23">Como province</collectingMunicipality>
, 
<location id="0E7F6052F9392027B8D9FBADFD64FBDD" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B8D9FBADFD64FBDD" box="[480,678,1066,1091]" country="Italy" municipality="Como province" name="Limido Comasco" pageId="2" pageNumber="23" stateProvince="Lombardia">Limido Comasco</location>
, 
<collectingDate id="6F5AE9A1F9392027BB8AFBADFC8CFBDD" box="[691,846,1066,1091]" pageId="2" pageNumber="23" value="2014-05-25">
<date id="7F1E1049F9392027BB8AFBADFC8CFBDD" box="[691,846,1066,1091]" pageId="2" pageNumber="23" value="2014-05-25">25 May 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BA65FBACFB4FFB4D" ID-GBIF-Occurrence="1434654800" collectingDate="2014-06-01" collectorName="Lido di Pomposa &amp; Arthropods Collection &amp; Natural History Museum &amp; Athens &amp; United States" country="Iran" county="Ferrara province" location="Lido di Pomposa" municipality="Comacchio" pageId="2" pageNumber="23" specimenCount="92" specimenCount-female="49" specimenCount-male="43" stateProvince="Emilia Romagna">
<specimenCount id="1DA6FD00F9392027BA65FBACFC18FBDD" box="[860,986,1066,1091]" pageId="2" pageNumber="23" type="female">39 females</specimenCount>
, 
<specimenCount id="1DA6FD00F9392027BADEFBADFB8EFBDD" box="[999,1100,1066,1091]" pageId="2" pageNumber="23" type="male">36 males</specimenCount>
and 14 deutonymphs, 
<collectingRegion id="C964F86BF9392027BC6BFBADFEC7FBF9" country="Italy" name="Emilia-Romagna" pageId="2" pageNumber="23">Emilia Romagna</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027B82BFBC8FE13FBF9" box="[274,465,1102,1127]" pageId="2" pageNumber="23">Ferrara province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027B8E7FBC9FDA3FBF9" box="[478,609,1102,1127]" pageId="2" pageNumber="23">Comacchio</collectingMunicipality>
, 
<collectorName id="A655535FF9392027BB54FBC9FCF1FBF8" box="[621,819,1102,1127]" pageId="2" pageNumber="23">
<location id="0E7F6052F9392027BB54FBC9FCF1FBF8" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BB54FBC9FCF1FBF8" box="[621,819,1102,1127]" country="Iran" county="Ferrara province" municipality="Comacchio" name="Lido di Pomposa" pageId="2" pageNumber="23" stateProvince="Emilia Romagna">Lido di Pomposa</location>
</collectorName>
, 
<collectingDate id="6F5AE9A1F9392027BA79FBC8FC09FBF9" box="[832,971,1103,1127]" pageId="2" pageNumber="23" value="2014-06-01">
<date id="7F1E1049F9392027BA79FBC8FC09FBF9" box="[832,971,1103,1127]" pageId="2" pageNumber="23" value="2014-06-01">1 June 2014</date>
</collectingDate>
(
<specimenCount id="1DA6FD00F9392027BAE7FBC8FB8EFBF9" box="[990,1100,1102,1127]" pageId="2" pageNumber="23" type="female">8 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BDBFFBC8FB1CFBF9" box="[1158,1246,1102,1127]" pageId="2" pageNumber="23" type="male">4 males</specimenCount>
deposited in the Acarological Collection, Department of Plant Protection, 
<collectingRegion id="C964F86BF9392027BA10FBF5FCA3FB15" box="[809,865,1138,1163]" country="Iran" name="Yazd" pageId="2" pageNumber="23">Yazd</collectingRegion>
Branch, Islamic Azad University, 
<collectingCountry id="73B77619F9392027BDC9FBF4FAE2FB14" box="[1264,1312,1139,1162]" name="Iran" pageId="2" pageNumber="23">Iran</collectingCountry>
; 
<specimenCount id="1DA6FD00F9392027BC17FBF4FA5EFB15" box="[1326,1436,1138,1163]" pageId="2" pageNumber="23" type="female">2 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027B9FFFB10FED3FB31" box="[198,273,1174,1199]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
deposited in Canestrini collection at Zoological Museum, University of Padua, 
<collectingCountry id="73B77619F9392027BDA8FB11FB0AFB31" box="[1169,1224,1174,1199]" name="Italy" pageId="2" pageNumber="23">Italy</collectingCountry>
; 
<specimenCount id="1DA6FD00F9392027BDEAFB10FAE8FB31" box="[1235,1322,1174,1199]" pageId="2" pageNumber="23" type="male">2 males</specimenCount>
deposited in the 
<collectorName id="A655535FF9392027B9E4FB3DFE1BFB4D" box="[221,473,1210,1235]" pageId="2" pageNumber="23">Arthropods Collection</collectorName>
of 
<collectorName id="A655535FF9392027B8C7FB3DFCD8FB4C" box="[510,794,1210,1235]" pageId="2" pageNumber="23">Natural History Museum</collectorName>
, 
<collectorName id="A655535FF9392027BA1CFB3DFCB5FB4D" box="[805,887,1210,1235]" pageId="2" pageNumber="23">Athens</collectorName>
, 
<collectingCountry id="73B77619F9392027BABBFB3DFC1DFB4D" box="[898,991,1210,1235]" name="Georgia" pageId="2" pageNumber="23">Georgia</collectingCountry>
, 
<collectorName id="A655535FF9392027BAD3FB3DFB47FB4D" box="[1002,1157,1210,1235]" pageId="2" pageNumber="23">United States</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BDA1FB3CFBB8FB69" ID-GBIF-Occurrence="1434654793" collectingDate="2014-07-21" collectorName="San Vincenzo, S." country="Iran" location="Livorno province" pageId="2" pageNumber="23" specimenCount="14" specimenCount-female="13" specimenCount-male="1" stateProvince="Toscana">
<specimenCount id="1DA6FD00F9392027BDA1FB3CFAD4FB4D" box="[1176,1302,1210,1235]" pageId="2" pageNumber="23" type="female">13 females</specimenCount>
, 
<specimenCount id="1DA6FD00F9392027BC18FB3CFAAEFB4D" box="[1313,1388,1210,1235]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
and 3 deutonymphs, 
<collectingRegion id="C964F86BF9392027B867FB58FE7FFB68" box="[350,445,1247,1270]" country="Italy" name="Toscana" pageId="2" pageNumber="23">Toscana</collectingRegion>
, 
<location id="0E7F6052F9392027B8F7FB59FD59FB69" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B8F7FB59FD59FB69" box="[462,667,1246,1271]" country="Iran" name="Livorno province" pageId="2" pageNumber="23" stateProvince="Toscana">Livorno province</location>
, 
<collectorName id="A655535FF9392027BB95FB58FCBAFB69" box="[684,888,1246,1271]" pageId="2" pageNumber="23">San Vincenzo, S.</collectorName>
Carlo, 
<collectingDate id="6F5AE9A1F9392027BAE0FB58FBB8FB69" box="[985,1146,1246,1271]" pageId="2" pageNumber="23" value="2014-07-21">
<date id="7F1E1049F9392027BAE0FB58FBB8FB69" box="[985,1146,1246,1271]" pageId="2" pageNumber="23" value="2014-07-21">21 July 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BDB4FB58FC7DFA85" ID-GBIF-Occurrence="1434654801" collectingDate="2014-08-03" country="Iran" location="Castelnuovo Scrivia" municipality="Alessandria province" pageId="2" pageNumber="23" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Piemonte">
<specimenCount id="1DA6FD00F9392027BDB4FB58FB3DFB69" box="[1165,1279,1246,1271]" pageId="2" pageNumber="23" type="female">4 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BC7AFB58FA5AFB69" box="[1347,1432,1246,1271]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
, 
<collectingRegion id="C964F86BF9392027B9AEFA85FEC6FA85" box="[151,260,1282,1307]" country="Italy" name="Piemonte" pageId="2" pageNumber="23">Piemonte</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027B82AFA85FDC5FA85" box="[275,519,1282,1307]" pageId="2" pageNumber="23">Alessandria province</collectingMunicipality>
, 
<location id="0E7F6052F9392027BB2FFA85FD3DFA85" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BB2FFA85FD3DFA85" box="[534,767,1282,1307]" country="Iran" municipality="Alessandria province" name="Castelnuovo Scrivia" pageId="2" pageNumber="23" stateProvince="Piemonte">Castelnuovo Scrivia</location>
, 
<collectingDate id="6F5AE9A1F9392027BA36FA84FC7DFA85" box="[783,959,1283,1307]" pageId="2" pageNumber="23" value="2014-08-03">
<date id="7F1E1049F9392027BA36FA84FC7DFA85" box="[783,959,1283,1307]" pageId="2" pageNumber="23" value="2014-08-03">3 August 2014</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BAF7FA84FD21FAA1" ID-GBIF-Occurrence="1434654799" collectingDate="2014-03-30" collectorName="M. Galuppi" country="Iran" location="Lonate Pozzolo" municipality="Varese province" pageId="2" pageNumber="23" specimenCount="9" specimenCount-female="9" stateProvince="Lombardia">
<specimenCount id="1DA6FD00F9392027BAF7FA84FB80FA85" box="[974,1090,1282,1307]" pageId="2" pageNumber="23" type="female">9 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BD68FA85FB0DFA85" box="[1105,1231,1282,1307]" country="Italy" name="Lombardia" pageId="2" pageNumber="23">Lombardia</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027BDE4FA84FA5AFA85" box="[1245,1432,1282,1307]" pageId="2" pageNumber="23">Varese province</collectingMunicipality>
, 
<location id="0E7F6052F9392027B9AEFAA0FE8EFAA1" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027B9AEFAA0FE8EFAA1" box="[151,332,1318,1343]" country="Iran" municipality="Varese province" name="Lonate Pozzolo" pageId="2" pageNumber="23" stateProvince="Lombardia">Lonate Pozzolo</location>
, 
<collectingDate id="6F5AE9A1F9392027B862FAA0FDD3FAA1" box="[347,529,1318,1343]" pageId="2" pageNumber="23" value="2014-03-30">
<date id="7F1E1049F9392027B862FAA0FDD3FAA1" box="[347,529,1318,1343]" pageId="2" pageNumber="23" value="2014-03-30">30 March 2014</date>
</collectingDate>
, 
<collectorName id="A655535FF9392027BB26FAA0FD66FAA1" box="[543,676,1318,1343]" pageId="2" pageNumber="23">M. Galuppi</collectorName>
coll.
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027BBCAFAA0FE53FAFD" ID-GBIF-Occurrence="1434654790" collectingDate="2015-07-27" country="Iceland" location="Monte Perone" municipality="Livorno province" pageId="2" pageNumber="23" specimenCount="4" specimenCount-female="4" stateProvince="Toscana">
<specimenCount id="1DA6FD00F9392027BBCAFAA0FCA7FAA1" box="[755,869,1318,1343]" pageId="2" pageNumber="23" type="female">4 females</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BA4DFAA0FC11FAA0" box="[884,979,1319,1342]" country="Italy" name="Toscana" pageId="2" pageNumber="23">Toscana</collectingRegion>
, 
<collectingMunicipality id="EB7BACF3F9392027BAD8FAA1FB69FAA1" box="[993,1195,1318,1343]" pageId="2" pageNumber="23">Livorno province</collectingMunicipality>
, 
<collectingCountry id="73B77619F9392027BD83FAA1FB32FAA1" box="[1210,1264,1318,1343]" name="Iceland" pageId="2" pageNumber="23">Isola</collectingCountry>
d'Elba, 
<location id="0E7F6052F9392027BC6AFAA0FF2AFAFC" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BC6AFAA0FF2AFAFC" country="Iceland" municipality="Livorno province" name="Monte Perone" pageId="2" pageNumber="23" stateProvince="Toscana">Monte Perone</location>
, 
<collectingDate id="6F5AE9A1F9392027B9CFFACCFE53FAFD" box="[246,401,1354,1379]" pageId="2" pageNumber="23" value="2015-07-27">
<date id="7F1E1049F9392027B9CFFACCFE53FAFD" box="[246,401,1354,1379]" pageId="2" pageNumber="23" value="2015-07-27">27 July 2015</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="BBC83CD4F9392027B899FACCFEE1FA19" ID-GBIF-Occurrence="1434654791" collectingDate="2016-03-20" country="Iceland" county="Alessandria province" location="Salada" municipality="Mongiardino Ligure" pageId="2" pageNumber="23" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Piemonte">
<specimenCount id="1DA6FD00F9392027B899FACCFDD2FAFD" box="[416,528,1354,1379]" pageId="2" pageNumber="23" type="female">3 females</specimenCount>
and 
<specimenCount id="1DA6FD00F9392027BB74FACCFD5DFAFD" box="[589,671,1354,1379]" pageId="2" pageNumber="23" type="male">1 male</specimenCount>
, 
<collectingRegion id="C964F86BF9392027BB97FACDFCD9FAFD" box="[686,795,1354,1379]" country="Italy" name="Piemonte" pageId="2" pageNumber="23">Piemonte</collectingRegion>
, 
<collectingCounty id="E27E4E05F9392027BA10FACDFBDFFAFD" box="[809,1053,1354,1379]" pageId="2" pageNumber="23">Alessandria province</collectingCounty>
, 
<collectingMunicipality id="EB7BACF3F9392027BD12FACDFAD7FAFD" box="[1067,1301,1354,1379]" pageId="2" pageNumber="23">Mongiardino Ligure</collectingMunicipality>
, 
<location id="0E7F6052F9392027BC1DFACDFAB3FAFD" LSID="urn:lsid:plazi:treatment:8309879FF93A2035B9AEF8CDFCB8FD36:0E7F6052F9392027BC1DFACDFAB3FAFD" box="[1316,1393,1354,1379]" country="Iceland" county="Alessandria province" municipality="Mongiardino Ligure" name="Salada" pageId="2" pageNumber="23" stateProvince="Piemonte">Salada</location>
, 
<collectingDate id="6F5AE9A1F9392027BCB9FACCFEE1FA19" pageId="2" pageNumber="23" value="2016-03-20">
<date id="7F1E1049F9392027BCB9FACCFEE1FA19" pageId="2" pageNumber="23" value="2016-03-20">20 March 2016</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43BA6502F9392027B9FEFA15FD78F8A9" pageId="2" pageNumber="23" type="diagnosis">
<paragraph id="0B1F3689F9392027B9FEFA15FD78F8A9" blockId="2.[151,1437,310,2027]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B9FEFA15FDBBFA35" bold="true" box="[199,633,1426,1451]" pageId="2" pageNumber="23">Diagnosis (adults and deutonymph).</emphasis>
Idiosoma prolonged posteriorly, with only tarsal segment of the fourth legs protruding beyond its posterior margin; all idiosomatic setae simple, without a discernible asymmetric swelling. Dorsal shield highly convex in adults, with some setae in marginal and submarginal position (
<emphasis id="39D4EA9BF9392027BC1EFA5CFAA4FA6C" box="[1319,1382,1499,1523]" italics="true" pageId="2" pageNumber="23">r2–r5</emphasis>
, 
<emphasis id="39D4EA9BF9392027BC4CFA5CFF72F989" italics="true" pageId="2" pageNumber="23">s1– s3</emphasis>
, 
<emphasis id="39D4EA9BF9392027B986FA79FF1AF988" box="[191,216,1534,1558]" italics="true" pageId="2" pageNumber="23">s6</emphasis>
, 
<emphasis id="39D4EA9BF9392027B9DEFA78FEEFF988" box="[231,301,1535,1559]" italics="true" pageId="2" pageNumber="23">S1–S5</emphasis>
and extra pairs 
<date id="7F1E1049F9392027B8DDFA78FDE5F989" box="[484,551,1535,1559]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B8DDFA78FDE5F989" box="[484,551,1535,1559]" italics="true" pageId="2" pageNumber="23">x1–x3</emphasis>
</date>
) barely discernible in dorsal view (even in partially pressed specimens); most dorsal shield setae relatively short, slightly decreasing in length from anterior to posterior positions in opisthonotal region; opisthonotum slightly hypertrichous in deutonymphs; setae 
<emphasis id="39D4EA9BF9392027BAADF9C0FC73F9C0" box="[916,945,1607,1630]" italics="true" pageId="2" pageNumber="23">Z5</emphasis>
thickened, blade-like, barbed in distal end; lyrifissures 
<emphasis id="39D4EA9BF9392027B819F9ECFE92F91D" box="[288,336,1643,1667]" italics="true" pageId="2" pageNumber="23">idJ2</emphasis>
and 
<emphasis id="39D4EA9BF9392027B8A8F9ECFE00F91D" box="[401,450,1643,1667]" italics="true" pageId="2" pageNumber="23">idJ4</emphasis>
not discernible in adults; three (
<date id="7F1E1049F9392027BA74F9ECFC4DF91D" box="[845,911,1643,1667]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027BA74F9ECFC4DF91D" box="[845,911,1643,1667]" italics="true" pageId="2" pageNumber="23">x1–x3</emphasis>
</date>
) and one (
<date id="7F1E1049F9392027BD20F9ECFBF1F91D" box="[1049,1075,1643,1667]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027BD20F9ECFBF1F91D" box="[1049,1075,1643,1667]" italics="true" pageId="2" pageNumber="23">x1</emphasis>
</date>
) extra pairs of marginal setae respectively in adults and deutonymph. Opisthogastric setae short. Sternal shield distinctly wider than long in female, with irregular lobes arising anteriorly to 
<emphasis id="39D4EA9BF9392027BB8DF934FD16F955" box="[692,724,1715,1739]" italics="true" pageId="2" pageNumber="23">
st 
<quantity id="CC589B6CF9392027BBFFF934FD16F955" box="[710,724,1715,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="23" unit="in" value="4.0">4</quantity>
</emphasis>
in deutonymph. Genital shield distinctly enlarged behind coxae IV. In the male: setae 
<emphasis id="39D4EA9BF9392027B8A8F950FE6CF970" box="[401,430,1751,1774]" italics="true" pageId="2" pageNumber="23">Px</emphasis>
generally absent; outer lobes of internal malae without fimbriae; seta 
<emphasis id="39D4EA9BF9392027BDF7F950FB2BF970" box="[1230,1257,1751,1774]" italics="true" pageId="2" pageNumber="23">av</emphasis>
on telefemur II inserted on a well-developed apophysis, seta 
<emphasis id="39D4EA9BF9392027BB90F97CFD01F88C" box="[681,707,1787,1810]" italics="true" pageId="2" pageNumber="23">av</emphasis>
on genu and tibia II, and setae 
<emphasis id="39D4EA9BF9392027BD7AF97CFBA0F88C" box="[1091,1122,1787,1810]" italics="true" pageId="2" pageNumber="23">mv</emphasis>
, 
<emphasis id="39D4EA9BF9392027BD4DF97CFB5EF88D" box="[1140,1180,1787,1811]" italics="true" pageId="2" pageNumber="23">pv2</emphasis>
and 
<emphasis id="39D4EA9BF9392027BDE2F97CFAC6F88D" box="[1243,1284,1787,1811]" italics="true" pageId="2" pageNumber="23">av2</emphasis>
on tarsus II, thickened, spine-like, and shorter than in female.
</paragraph>
</subSubSection>
<subSubSection id="43BA6502F939202BB9FEF8C5FC22FDA3" lastPageId="14" lastPageNumber="35" pageId="2" pageNumber="23" type="description">
<paragraph id="0B1F3689F9392027B9FEF8C5FDB5F8C5" blockId="2.[151,1437,310,2027]" box="[199,631,1858,1883]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B9FEF8C5FE90F8C5" bold="true" box="[199,338,1858,1883]" pageId="2" pageNumber="23">Description</emphasis>
(adults and deutonymph).
</paragraph>
<paragraph id="0B1F3689F9392027B9FEF8E1FC39F859" blockId="2.[151,1437,310,2027]" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B9FEF8E1FEDCF8E1" bold="true" box="[199,286,1894,1919]" italics="true" pageId="2" pageNumber="23">Female</emphasis>
. 
<figureCitation id="939B2A0CF9392027B80DF8E1FE6DF8E1" box="[308,431,1894,1919]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 7" captionStart-2="FIGURE 8" captionStart-3="FIGURES 9 – 12" captionStartId-0="4.[151,264,1939,1961]" captionStartId-1="5.[151,264,1778,1800]" captionStartId-2="6.[151,250,1772,1794]" captionStartId-3="7.[151,264,1183,1205]" captionTargetBox-0="[155,1421,421,1907]" captionTargetBox-1="[164,1424,214,1739]" captionTargetBox-2="[171,1416,210,1728]" captionTargetBox-3="[160,1424,207,1150]" captionTargetId-0="figure@4.[150,1436,409,1918]" captionTargetId-1="figure@5.[151,1436,193,1756]" captionTargetId-2="figure@6.[151,1436,193,1751]" captionTargetId-3="figure@7.[151,1436,193,1162]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." captionText-1="FIGURES 3 – 7. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4 – 7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm." captionText-2="FIGURE 8. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma (partial view; setae Zv 5 and glands gv 2 not discernible in this specimen). Scale bar = 100 µm." captionText-3="FIGURES 9 – 12. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma, four different specimens (partial views). Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/292759/files/figure.png" httpUri-1="https://zenodo.org/record/292760/files/figure.png" httpUri-2="https://zenodo.org/record/292761/files/figure.png" httpUri-3="https://zenodo.org/record/292762/files/figure.png" pageId="2" pageNumber="23">Figs 1–12</figureCitation>
, 
<figureCitation id="939B2A0CF9392027B8FAF8E1FDC9F8E1" box="[451,523,1894,1919]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="2" pageNumber="23">16–18</figureCitation>
, 
<figureCitation id="939B2A0CF9392027BB19F8E0FDB3F8E1" box="[544,625,1895,1919]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="2" pageNumber="23">22, 24</figureCitation>
, 
<figureCitation id="939B2A0CF9392027BBBCF8E0FD0DF8E1" box="[645,719,1895,1919]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="2" pageNumber="23">29–30</figureCitation>
; 
<specimenCount id="1DA6FD00F9392027BBDAF8E0FC41F8E1" box="[739,899,1894,1919]" pageId="2" pageNumber="23" type="generic">10 specimens</specimenCount>
measured. Idiosoma oval-shaped, prolonged posteriorly, with only tarsal segment of the fourth legs protruding beyond its posterior margin; 617 ± 13 (600–640) long and 474 ± 8 (460–490) wide, ratio length/width 1.30 ± 0.03 (1.24–1.35).
</paragraph>
<paragraph id="0B1F3689F9392026B9FEF854FF23FCA6" blockId="2.[151,1437,310,2027]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
<emphasis id="39D4EA9BF9392027B9FEF854FE46F875" box="[199,388,2003,2027]" italics="true" pageId="2" pageNumber="23">Dorsal idiosoma</emphasis>
(
<figureCitation id="939B2A0CF9392027B8ACF855FE38F875" box="[405,506,2002,2027]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 7" captionStartId-0="4.[151,264,1939,1961]" captionStartId-1="5.[151,264,1778,1800]" captionTargetBox-0="[155,1421,421,1907]" captionTargetBox-1="[164,1424,214,1739]" captionTargetId-0="figure@4.[150,1436,409,1918]" captionTargetId-1="figure@5.[151,1436,193,1756]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." captionText-1="FIGURES 3 – 7. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4 – 7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/292759/files/figure.png" httpUri-1="https://zenodo.org/record/292760/files/figure.png" pageId="2" pageNumber="23">Figs 1–7</figureCitation>
, 
<figureCitation id="939B2A0CF9392027BB31F854FD96F875" box="[520,596,2003,2027]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="2" pageNumber="23">29–30</figureCitation>
). Dorsal shield entire, completely covering dorsal surface, highly convex, with some setae in marginal and submarginal position (
<emphasis id="39D4EA9BF9382026BA1CFF1FFCA7FF31" box="[805,869,152,176]" italics="true" pageId="3" pageNumber="24">r2–r5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BA4EFF1FFC75FF2E" box="[887,951,152,176]" italics="true" pageId="3" pageNumber="24">s1–s3</emphasis>
, 
<emphasis id="39D4EA9BF9382026BAF0FF1FFC20FF2E" box="[969,994,152,176]" italics="true" pageId="3" pageNumber="24">s6</emphasis>
, 
<emphasis id="39D4EA9BF9382026BACDFF1FFBF8FF31" box="[1012,1082,152,176]" italics="true" pageId="3" pageNumber="24">S1–S5</emphasis>
and extra pairs 
<date id="7F1E1049F9382026BDC7FF1FFA83FF2E" box="[1278,1345,152,176]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BDC7FF1FFA83FF2E" box="[1278,1345,152,176]" italics="true" pageId="3" pageNumber="24">x1–x3</emphasis>
</date>
) barely discernible in dorsal view (even in pressed specimens) (
<figureCitation id="939B2A0CF9382026BA33FF3BFC8EFF4A" box="[778,844,188,213]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,264,1939,1961]" captionTargetBox="[155,1421,421,1907]" captionTargetId="figure@4.[150,1436,409,1918]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." httpUri="https://zenodo.org/record/292759/files/figure.png" pageId="3" pageNumber="24">Fig. 1</figureCitation>
); with 43 pairs of setae, 23 pairs on podonotum (
<emphasis id="39D4EA9BF9382026BC41FF3BFF6EFF66" italics="true" pageId="3" pageNumber="24">j1– j6</emphasis>
, 
<emphasis id="39D4EA9BF9382026B983FF67FF38FF66" box="[186,250,224,248]" italics="true" pageId="3" pageNumber="24">z1–z6</emphasis>
, 
<emphasis id="39D4EA9BF9382026B831FF67FE8AFF66" box="[264,328,224,248]" italics="true" pageId="3" pageNumber="24">s1–s6</emphasis>
, 
<emphasis id="39D4EA9BF9382026B86CFF67FE57FF69" box="[341,405,224,248]" italics="true" pageId="3" pageNumber="24">r2–r5</emphasis>
; 
<emphasis id="39D4EA9BF9382026B89AFF67FE7EFF66" box="[419,444,224,248]" italics="true" pageId="3" pageNumber="24">r6</emphasis>
ventrally placed) including a marginal supernumerary pair (
<date id="7F1E1049F9382026BD64FF67FBB5FF66" box="[1117,1143,224,248]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BD64FF67FBB5FF66" box="[1117,1143,224,248]" italics="true" pageId="3" pageNumber="24">x1</emphasis>
</date>
) posterior to setae 
<emphasis id="39D4EA9BF9382026BC74FF67FAA4FF66" box="[1357,1382,224,248]" italics="true" pageId="3" pageNumber="24">s6</emphasis>
, and 20 pairs on opisthonotum (
<emphasis id="39D4EA9BF9382026B8FFFE83FDCBFE85" box="[454,521,260,284]" italics="true" pageId="3" pageNumber="24">J1–J5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB2EFE83FDA2FE85" box="[535,608,260,284]" italics="true" pageId="3" pageNumber="24">Z1–Z5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB56FE83FD77FE85" box="[623,693,260,284]" italics="true" pageId="3" pageNumber="24">S1–S5</emphasis>
) including three pairs of extra setae (
<emphasis id="39D4EA9BF9382026BD62FE83FB02FE82" box="[1115,1216,260,284]" italics="true" pageId="3" pageNumber="24">Px1–Px3</emphasis>
) (
<emphasis id="39D4EA9BF9382026BDE3FE83FAFEFE82" box="[1242,1340,260,284]" italics="true" pageId="3" pageNumber="24">Zx1–Zx3</emphasis>
of other authors) between 
<emphasis id="39D4EA9BF9382026B867FEAEFEA8FEDE" box="[350,362,297,320]" italics="true" pageId="3" pageNumber="24">J</emphasis>
and 
<emphasis id="39D4EA9BF9382026B89BFEAEFE73FEDE" box="[418,433,297,320]" italics="true" pageId="3" pageNumber="24">Z</emphasis>
series and two marginal supernumerary pairs, at level of setae 
<emphasis id="39D4EA9BF9382026BD4EFEAFFB51FEDE" box="[1143,1171,296,320]" italics="true" pageId="3" pageNumber="24">S1</emphasis>
(
<date id="7F1E1049F9382026BD9DFEAFFB7DFEDE" box="[1188,1215,296,320]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BD9DFEAFFB7DFEDE" box="[1188,1215,296,320]" italics="true" pageId="3" pageNumber="24">x2</emphasis>
</date>
) and between setae 
<emphasis id="39D4EA9BF9382026B9AEFECBFF71FEFA" box="[151,179,332,356]" italics="true" pageId="3" pageNumber="24">S1</emphasis>
and 
<emphasis id="39D4EA9BF9382026B9D4FECBFECBFEFA" box="[237,265,332,356]" italics="true" pageId="3" pageNumber="24">S2</emphasis>
(
<date id="7F1E1049F9382026B822FECBFEF4FEFA" box="[283,310,332,356]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026B822FECBFEF4FEFA" box="[283,310,332,356]" italics="true" pageId="3" pageNumber="24">x3</emphasis>
</date>
); setae 
<emphasis id="39D4EA9BF9382026B8A8FECBFE11FEFA" box="[401,467,332,356]" italics="true" pageId="3" pageNumber="24">J2–J4</emphasis>
very unstable in position (in some cases, one or both setae of these pairs apparently absent); with a variable number (4–7) of unpaired setae (
<emphasis id="39D4EA9BF9382026BA25FEF6FCF7FE16" box="[796,821,369,392]" italics="true" pageId="3" pageNumber="24">Jx</emphasis>
) always present between 
<emphasis id="39D4EA9BF9382026BD63FEF6FBA4FE16" box="[1114,1126,369,392]" italics="true" pageId="3" pageNumber="24">J</emphasis>
series (in some specimens, setae 
<emphasis id="39D4EA9BF9382026B9EFFE12FF2DFE32" box="[214,239,405,428]" italics="true" pageId="3" pageNumber="24">Jx</emphasis>
between 
<emphasis id="39D4EA9BF9382026B865FE13FEB4FE32" box="[348,374,404,428]" italics="true" pageId="3" pageNumber="24">J4</emphasis>
and 
<emphasis id="39D4EA9BF9382026B897FE13FE0AFE35" box="[430,456,404,427]" italics="true" pageId="3" pageNumber="24">J5</emphasis>
appearing as a supplementary pair); all setae simple, without a discernible asymmetric swelling, setiform and smooth (
<figureCitation id="939B2A0CF9382026BB3FFE30FD50FE4E" box="[518,658,439,464]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="3" pageNumber="24">Figs. 29–30</figureCitation>
), except 
<emphasis id="39D4EA9BF9382026BBC2FE3FFCD3FE4E" box="[763,785,440,464]" italics="true" pageId="3" pageNumber="24">j1</emphasis>
, 
<emphasis id="39D4EA9BF9382026BA1BFE3FFCF9FE4E" box="[802,827,440,464]" italics="true" pageId="3" pageNumber="24">z1</emphasis>
and 
<emphasis id="39D4EA9BF9382026BA4EFE3FFC57FE51" box="[887,917,440,463]" italics="true" pageId="3" pageNumber="24">Z5</emphasis>
, thickened, 
<emphasis id="39D4EA9BF9382026BD1DFE3FFBFBFE4E" box="[1060,1081,440,464]" italics="true" pageId="3" pageNumber="24">j1</emphasis>
and 
<emphasis id="39D4EA9BF9382026BD4FFE3FFB4DFE4E" box="[1142,1167,440,464]" italics="true" pageId="3" pageNumber="24">z1</emphasis>
acuminate, 
<emphasis id="39D4EA9BF9382026BC24FE3FFAF9FE51" box="[1309,1339,440,463]" italics="true" pageId="3" pageNumber="24">Z5</emphasis>
(
<figureCitation id="939B2A0CF9382026BC74FE30FA51FE4E" box="[1357,1427,439,464]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,264,1939,1961]" captionTargetBox="[155,1421,421,1907]" captionTargetId="figure@4.[150,1436,409,1918]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." httpUri="https://zenodo.org/record/292759/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
) blade-like and barbed in distal end (in some specimens setae 
<emphasis id="39D4EA9BF9382026BA06FE5BFC9BFE6A" box="[831,857,476,500]" italics="true" pageId="3" pageNumber="24">J4</emphasis>
, 
<emphasis id="39D4EA9BF9382026BA5EFE5BFC43FE6D" box="[871,897,476,499]" italics="true" pageId="3" pageNumber="24">J5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BAB6FE5BFC6EFE6A" box="[911,940,476,500]" italics="true" pageId="3" pageNumber="24">Z4</emphasis>
and 
<emphasis id="39D4EA9BF9382026BADBFE5AFC38FE6A" box="[994,1018,477,500]" italics="true" pageId="3" pageNumber="24">Jx</emphasis>
, between 
<emphasis id="39D4EA9BF9382026BD55FE5BFB44FE6A" box="[1132,1158,476,500]" italics="true" pageId="3" pageNumber="24">J4</emphasis>
and 
<emphasis id="39D4EA9BF9382026BD85FE5BFB14FE6D" box="[1212,1238,476,499]" italics="true" pageId="3" pageNumber="24">J5</emphasis>
, slightly barbed); most setae not long enough to reach base of next posterior seta, 
<emphasis id="39D4EA9BF9382026BAACFD87FC6EFD86" box="[917,940,512,536]" italics="true" pageId="3" pageNumber="24">j1</emphasis>
and 
<emphasis id="39D4EA9BF9382026BAD0FD87FBC0FD86" box="[1001,1026,512,536]" italics="true" pageId="3" pageNumber="24">z1</emphasis>
the shortest, other podonotal setae subequal in length, opisthonotal setae slightly decreasing in length from anterior to posterior positions, with 
<emphasis id="39D4EA9BF9382026BC6CFDA3FAB1FDA5" box="[1365,1395,548,571]" italics="true" pageId="3" pageNumber="24">Z5</emphasis>
the shortest: 
<emphasis id="39D4EA9BF9382026B9C4FDCFFED6FDFE" box="[253,276,584,608]" italics="true" pageId="3" pageNumber="24">
j 
<date id="7F1E1049F9382026B830FDCFFED6FDFE" box="[265,276,584,608]" pageId="3" pageNumber="24" value="1918-01-19">1</date>
</emphasis>
18–19, 
<emphasis id="39D4EA9BF9382026B857FDCFFE41FDFE" box="[366,387,584,608]" italics="true" pageId="3" pageNumber="24">j2</emphasis>
34–38, 
<emphasis id="39D4EA9BF9382026B8D9FDCFFE34FDC1" box="[480,502,584,607]" italics="true" pageId="3" pageNumber="24">j5</emphasis>
39–45, 
<emphasis id="39D4EA9BF9382026BB68FDCFFDA8FDFE" box="[593,618,584,608]" italics="true" pageId="3" pageNumber="24">
z 
<date id="7F1E1049F9382026BB67FDCFFDA8FDFE" box="[606,618,584,608]" pageId="3" pageNumber="24" value="2014-01-16">1</date>
</emphasis>
14–16, 
<emphasis id="39D4EA9BF9382026BBFCFDCFFD1CFDFE" box="[709,734,584,608]" italics="true" pageId="3" pageNumber="24">z2</emphasis>
38–43, 
<emphasis id="39D4EA9BF9382026BA03FDCFFC91FDFE" box="[826,851,584,608]" italics="true" pageId="3" pageNumber="24">z3</emphasis>
35–41, 
<emphasis id="39D4EA9BF9382026BA97FDCFFC05FDFE" box="[942,967,584,608]" italics="true" pageId="3" pageNumber="24">z4</emphasis>
44–49, 
<emphasis id="39D4EA9BF9382026BD1AFDCFFBFEFDC1" box="[1059,1084,584,607]" italics="true" pageId="3" pageNumber="24">z5</emphasis>
43–46, 
<geoCoordinate id="6E94504EF9382026BDAEFDCFFB17FDFE" box="[1175,1237,583,608]" direction="south" orientation="latitude" pageId="3" pageNumber="24" precision="925" value="-5.7666664">
<emphasis id="39D4EA9BF9382026BDAEFDCFFB72FDC1" box="[1175,1200,584,607]" italics="true" pageId="3" pageNumber="24">s5</emphasis>
46
</geoCoordinate>
–51, 
<date id="7F1E1049F9382026BC35FDCFFA8BFDFE" box="[1292,1353,583,608]" pageId="3" pageNumber="24" value="1936-10-01">
<emphasis id="39D4EA9BF9382026BC35FDCFFAE4FDFE" box="[1292,1318,584,608]" italics="true" pageId="3" pageNumber="24">x1</emphasis>
36
</date>
–40, 
<emphasis id="39D4EA9BF9382026BCB8FDCFFA5EFDFE" box="[1409,1436,584,608]" italics="true" pageId="3" pageNumber="24">
x 
<date id="7F1E1049F9382026BCB6FDCFFA5EFDFE" box="[1423,1436,584,608]" pageId="3" pageNumber="24" value="1934-03-30">3</date>
</emphasis>
30–34, 
<emphasis id="39D4EA9BF9382026B9D5FDEBFEC4FD1A" box="[236,262,620,644]" italics="true" pageId="3" pageNumber="24">J1</emphasis>
38–43, 
<emphasis id="39D4EA9BF9382026B85AFDEBFEBFFD1A" box="[355,381,620,644]" italics="true" pageId="3" pageNumber="24">J2</emphasis>
39–41, 
<emphasis id="39D4EA9BF9382026B8E2FDEBFE37FD1A" box="[475,501,620,644]" italics="true" pageId="3" pageNumber="24">J4</emphasis>
34–38, 
<emphasis id="39D4EA9BF9382026BB6BFDEAFDA9FD1A" box="[594,619,621,644]" italics="true" pageId="3" pageNumber="24">Jx</emphasis>
33–35, 
<emphasis id="39D4EA9BF9382026BBF1FDEBFD20FD1D" box="[712,738,620,643]" italics="true" pageId="3" pageNumber="24">
J 
<date id="7F1E1049F9382026BBECFDEBFD20FD1D" box="[725,738,620,643]" pageId="3" pageNumber="24" value="1939-05-31">5</date>
</emphasis>
31–39, 
<emphasis id="39D4EA9BF9382026BA06FDEBFC9FFD1A" box="[831,861,620,644]" italics="true" pageId="3" pageNumber="24">Z1</emphasis>
43–46, 
<emphasis id="39D4EA9BF9382026BA83FDEBFC1AFD1A" box="[954,984,620,644]" italics="true" pageId="3" pageNumber="24">Z2</emphasis>
38–41, 
<emphasis id="39D4EA9BF9382026BD0DFDEBFB91FD1A" box="[1076,1107,620,644]" italics="true" pageId="3" pageNumber="24">Z3</emphasis>
34–38, 
<emphasis id="39D4EA9BF9382026BD96FDEBFB0FFD1A" box="[1199,1229,620,644]" italics="true" pageId="3" pageNumber="24">Z4</emphasis>
39–41, 
<emphasis id="39D4EA9BF9382026BC10FDEBFA85FD1D" box="[1321,1351,620,643]" italics="true" pageId="3" pageNumber="24">
Z 
<date id="7F1E1049F9382026BC03FDEBFA85FD1D" box="[1338,1351,620,643]" pageId="3" pageNumber="24" value="1920-05-23">5</date>
</emphasis>
20–23, 
<emphasis id="39D4EA9BF9382026B9AEFD17FF01FD36" box="[151,195,656,680]" italics="true" pageId="3" pageNumber="24">Px1</emphasis>
39–46, 
<emphasis id="39D4EA9BF9382026B826FD17FE88FD36" box="[287,330,656,680]" italics="true" pageId="3" pageNumber="24">Px2</emphasis>
36–46, 
<emphasis id="39D4EA9BF9382026B89EFD17FE10FD36" box="[423,466,656,680]" italics="true" pageId="3" pageNumber="24">Px3</emphasis>
34–40; with 23 pairs of discernible pore-like structures, 12 on podonotum (
<emphasis id="39D4EA9BF9382026BC16FD17FA98FD36" box="[1327,1370,656,680]" italics="true" pageId="3" pageNumber="24">idj1</emphasis>
, 
<emphasis id="39D4EA9BF9382026BC50FD17FA57FD36" box="[1385,1429,656,680]" italics="true" pageId="3" pageNumber="24">idj3</emphasis>
, 
<emphasis id="39D4EA9BF9382026B9AEFD33FF04FD52" box="[151,198,692,716]" italics="true" pageId="3" pageNumber="24">idz3</emphasis>
, 
<emphasis id="39D4EA9BF9382026B9EDFD33FEC0FD52" box="[212,258,692,716]" italics="true" pageId="3" pageNumber="24">idr5</emphasis>
, 
<emphasis id="39D4EA9BF9382026B829FD33FEFEFD52" box="[272,316,692,716]" italics="true" pageId="3" pageNumber="24">idj6</emphasis>
, 
<emphasis id="39D4EA9BF9382026B873FD33FEBBFD52" box="[330,377,692,716]" italics="true" pageId="3" pageNumber="24">ids6</emphasis>
, 
<emphasis id="39D4EA9BF9382026B8BEFD33FE7AFD52" box="[391,440,692,716]" italics="true" pageId="3" pageNumber="24">idx1</emphasis>
, 
<emphasis id="39D4EA9BF9382026B8FCFD33FE3AFD52" box="[453,504,692,716]" italics="true" pageId="3" pageNumber="24">gdj2</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB3CFD33FDFAFD52" box="[517,568,692,716]" italics="true" pageId="3" pageNumber="24">gdj4</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB7CFD33FDB8FD52" box="[581,634,692,716]" italics="true" pageId="3" pageNumber="24">gdr4</emphasis>
, 
<emphasis id="39D4EA9BF9382026BBB0FD33FD7CFD52" box="[649,702,692,716]" italics="true" pageId="3" pageNumber="24">gdz5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BBF5FD33FCC2FD52" box="[716,768,692,716]" italics="true" pageId="3" pageNumber="24">gdz6</emphasis>
; 
<emphasis id="39D4EA9BF9382026BA29FD33FC86FD52" box="[784,836,692,716]" italics="true" pageId="3" pageNumber="24">gds4</emphasis>
not discernible) and 11 on opisthonotum (
<emphasis id="39D4EA9BF9382026BC1BFD33FA97FD52" box="[1314,1365,692,716]" italics="true" pageId="3" pageNumber="24">idZ1</emphasis>
, 
<emphasis id="39D4EA9BF9382026BC5AFD33FA56FD52" box="[1379,1428,692,716]" italics="true" pageId="3" pageNumber="24">idS1</emphasis>
, 
<emphasis id="39D4EA9BF9382026B9AEFD5FFF0AFD6E" box="[151,200,728,752]" italics="true" pageId="3" pageNumber="24">idS2</emphasis>
, 
<emphasis id="39D4EA9BF9382026B9E1FD5FFECAFD6E" box="[216,264,728,752]" italics="true" pageId="3" pageNumber="24">idJ1</emphasis>
, 
<emphasis id="39D4EA9BF9382026B821FD5FFE89FD6E" box="[280,331,728,752]" italics="true" pageId="3" pageNumber="24">idPx</emphasis>
, 
<emphasis id="39D4EA9BF9382026B862FD5FFE4FFD6E" box="[347,397,728,752]" italics="true" pageId="3" pageNumber="24">idS3</emphasis>
, 
<emphasis id="39D4EA9BF9382026B8A5FD5FFE0DFD6E" box="[412,463,728,752]" italics="true" pageId="3" pageNumber="24">idZ4</emphasis>
, 
<emphasis id="39D4EA9BF9382026B8E6FD5FFDCDFD6E" box="[479,527,728,752]" italics="true" pageId="3" pageNumber="24">idJ5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB27FD5FFD90FD6E" box="[542,594,728,752]" italics="true" pageId="3" pageNumber="24">idZ5</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB58FD5FFD5AFD6E" box="[609,664,728,752]" italics="true" pageId="3" pageNumber="24">gdx2</emphasis>
, 
<emphasis id="39D4EA9BF9382026BB9EFD5FFD1DFD6E" box="[679,735,728,752]" italics="true" pageId="3" pageNumber="24">gdS3</emphasis>
; 
<emphasis id="39D4EA9BF9382026BBD6FD5FFCEBFD6E" box="[751,809,728,752]" italics="true" pageId="3" pageNumber="24">gdZ3</emphasis>
not discernible), with 
<emphasis id="39D4EA9BF9382026BD15FD5FFB9EFD6E" box="[1068,1116,728,752]" italics="true" pageId="3" pageNumber="24">idJ2</emphasis>
and 
<emphasis id="39D4EA9BF9382026BDACFD5FFB04FD6E" box="[1173,1222,728,752]" italics="true" pageId="3" pageNumber="24">idJ4</emphasis>
apparently absent; dorsal surface smooth, ornamented throughout with irregular polygonal pattern, less regular in the central region of shield.
</paragraph>
<paragraph id="0B1F3689F9382021B9FEFCC3FAFAFEFA" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026B9FEFCC3FE4AFCC2" box="[199,392,836,860]" italics="true" pageId="3" pageNumber="24">Ventral idiosoma</emphasis>
(
<figureCitation id="939B2A0CF9382026B8A1FCC3FDCCFCC2" box="[408,526,836,861]" captionStart-0="FIGURE 8" captionStart-1="FIGURES 9 – 12" captionStartId-0="6.[151,250,1772,1794]" captionStartId-1="7.[151,264,1183,1205]" captionTargetBox-0="[171,1416,210,1728]" captionTargetBox-1="[160,1424,207,1150]" captionTargetId-0="figure@6.[151,1436,193,1751]" captionTargetId-1="figure@7.[151,1436,193,1162]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 8. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma (partial view; setae Zv 5 and glands gv 2 not discernible in this specimen). Scale bar = 100 µm." captionText-1="FIGURES 9 – 12. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma, four different specimens (partial views). Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/292761/files/figure.png" httpUri-1="https://zenodo.org/record/292762/files/figure.png" pageId="3" pageNumber="24">Figs 8–12</figureCitation>
). Tritosternum with subrectangular base well separated from anterior margin of sternal shield, base longer 39 ± 3 (35–44) than basal width 20 ± 1 (16–21), and two free pilose laciniae. Presternal area with faint lines of sclerotisation, without platelets well-developed. Sternal shield well sclerotised, wider than long, 67 ± 3 (63–70) long, 126 ± 3 (123–131) wide; with anterior margin well defined, medially almost straight, and antero-lateral corners pointed, concave on posterior margin; with three pairs of smooth setae (
<emphasis id="39D4EA9BF9382026BDC5FC53FADEFC72" box="[1276,1308,980,1004]" italics="true" pageId="3" pageNumber="24">st1</emphasis>
40–45, 
<emphasis id="39D4EA9BF9382026BC42FC52FA4CFC72" box="[1403,1422,981,1004]" italics="true" pageId="3" pageNumber="24">st</emphasis>
2 45–50, 
<emphasis id="39D4EA9BF9382026B9D7FC7EFEC3FB8E" box="[238,257,1017,1040]" italics="true" pageId="3" pageNumber="24">st</emphasis>
3 40–43) and two pairs of slit-like lyrifissures, adjacent to 
<emphasis id="39D4EA9BF9382026BA8BFC7FFC11FB8E" box="[946,979,1016,1040]" italics="true" pageId="3" pageNumber="24">st1</emphasis>
and between 
<emphasis id="39D4EA9BF9382026BD41FC7EFB49FB8E" box="[1144,1163,1017,1040]" italics="true" pageId="3" pageNumber="24">st</emphasis>
2 and 
<emphasis id="39D4EA9BF9382026BDEFFC7EFB2BFB8E" box="[1238,1257,1017,1040]" italics="true" pageId="3" pageNumber="24">st</emphasis>
3 setae; surface delicately ornamented with irregular polygonal pattern. Endopodals II–III fused with sternal shield, endopodal platelets III–IV distinct and well-developed, boomerang-shaped, not fused anteriorly to sternal shield, with deeper adaxial portion, posteriorly connected with sclerites of the sperm access system. Metasternal platelets absent; setae 
<emphasis id="39D4EA9BF9382026B9AEFB0EFF68FB3E" box="[151,170,1161,1184]" italics="true" pageId="3" pageNumber="24">st</emphasis>
4 (30–33) situated on soft cuticle covering adaxial portion of the endopodal platelets III–IV, poroids 
<date id="7F1E1049F9382026BC1AFB0FFA84FB3E" box="[1315,1350,1160,1184]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BC1AFB0FFA84FB3E" box="[1315,1350,1160,1184]" italics="true" pageId="3" pageNumber="24">iv3</emphasis>
</date>
on soft integument, well separated from endopodal platelets. Genital shield flask-shaped, enlarged behind coxae IV, 317 ± 7 (305–323) long, 201 ± 3 (198–205) wide; widely separated from anal shield and extending posteriorly to level of 
<emphasis id="39D4EA9BF9382026B9AEFB73FF7FFA92" box="[151,189,1268,1292]" italics="true" pageId="3" pageNumber="24">Jv1</emphasis>
setae; with rounded posterior margin and hyaline anterior margin, anteriorly reaching beyond posterior margin of sternal shield; bearing only setae 
<emphasis id="39D4EA9BF9382026BB15FA9FFD8EFAB1" box="[556,588,1304,1327]" italics="true" pageId="3" pageNumber="24">st5</emphasis>
(34–35); setae 
<emphasis id="39D4EA9BF9382026BBC3FA9FFCE6FAAE" box="[762,804,1304,1328]" italics="true" pageId="3" pageNumber="24">Zv1</emphasis>
(15–16) and 
<emphasis id="39D4EA9BF9382026BA83FA9FFC23FAAE" box="[954,993,1304,1328]" italics="true" pageId="3" pageNumber="24">Jv1</emphasis>
(18–21) close to the edge of the shield, on unsclerotised cuticle; genital lyrifissures 
<date id="7F1E1049F9382026BBA4FABBFD7DFACD" box="[669,703,1340,1363]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BBA4FABBFD7DFACD" box="[669,703,1340,1363]" italics="true" pageId="3" pageNumber="24">iv5</emphasis>
</date>
on soft integument beside genital setae; surface of the shield ornamented with irregular polygonal pattern, basally composed of large cells, the outer edges of the upper cells forming an inverted u- or v-shaped line. Peritrematal shields only with antiaxial and post-stigmatic sections developed, not extending behind coxa IV, free from exopodals and fused to dorsal shield only in a very small section anterior to 
<emphasis id="39D4EA9BF9382026B854FA4BFE44FA7A" box="[365,390,1484,1508]" italics="true" pageId="3" pageNumber="24">z1</emphasis>
; peritreme extending anteriorly beyond coxae I, up to level between 
<emphasis id="39D4EA9BF9382026BD99FA4BFB78FA7A" box="[1184,1210,1484,1508]" italics="true" pageId="3" pageNumber="24">s1</emphasis>
and 
<emphasis id="39D4EA9BF9382026BDCAFA4BFACEFA7A" box="[1267,1292,1484,1508]" italics="true" pageId="3" pageNumber="24">z1</emphasis>
; each shield bearing five discernible pore-like structures (a lyrifissure 
<emphasis id="39D4EA9BF9382026BA22FA77FCF3F999" box="[795,817,1520,1543]" italics="true" pageId="3" pageNumber="24">ip</emphasis>
and a gland pore 
<emphasis id="39D4EA9BF9382026BAC1FA76FBD6F996" box="[1016,1044,1521,1544]" italics="true" pageId="3" pageNumber="24">gp</emphasis>
at level of coxa II, two lyrifissures 
<emphasis id="39D4EA9BF9382026B9AEF992FF6FF9B2" box="[151,173,1557,1580]" italics="true" pageId="3" pageNumber="24">ip</emphasis>
and a gland pore 
<emphasis id="39D4EA9BF9382026B847F992FE58F9B2" box="[382,410,1557,1580]" italics="true" pageId="3" pageNumber="24">gp</emphasis>
on post-stigmatic section); a small pore-like structure within peritreme at level of coxa I. Setae 
<emphasis id="39D4EA9BF9382026B9E4F9BFFF34F9CE" box="[221,246,1592,1616]" italics="true" pageId="3" pageNumber="24">r6</emphasis>
(12–15) on unsclerotised integument between peritrematal shields and antero-lateral margins of dorsal shield, about at level of stigmata. Exopodal platelets I–II absent, exopodals II–III present, although weakly sclerotised, generally posteriorly not connected with well-developed exopodal platelets III–IV, the latter ones extending along posterior margin of coxae IV, but not expanded posteriorly in podal elements; glands 
<emphasis id="39D4EA9BF9382026BC22F923FA81F922" box="[1307,1347,1700,1724]" italics="true" pageId="3" pageNumber="24">gv2</emphasis>
present on posterior rim of exopodals III–IV. Anal shield subtriangular, about as long as wide, 106 ± 5 (100–113) long, 105 ± 4 (100–110) wide; anal opening situated about at mid-level of shield; para-anal setae (23–25) longer than unpaired post-anal seta (20–21); pores 
<emphasis id="39D4EA9BF9382026BB72F897FDB1F8B6" box="[587,627,1808,1832]" italics="true" pageId="3" pageNumber="24">gv3</emphasis>
on lateral margins of the anal shield, situated from the mid-level of anal opening to just behind its posterior margin; cribrum extending anteriorly along lateral margins of shield beyond insertion of post-anal seta; with some sculptural lines on anterior half. Opisthogastric unsclerotised integument with 15 pairs of setae, excluding a variable number of setae 
<emphasis id="39D4EA9BF9382026BA34F8FAFCF0F80A" box="[781,818,1917,1940]" italics="true" pageId="3" pageNumber="24">UR</emphasis>
(chaetotaxy with some variation; 
<figureCitation id="939B2A0CF9382026BD8EF8FBFAE8F80A" box="[1207,1322,1916,1941]" captionStart="FIGURES 9 – 12" captionStartId="7.[151,264,1183,1205]" captionTargetBox="[160,1424,207,1150]" captionTargetId="figure@7.[151,1436,193,1162]" captionTargetPageId="7" captionText="FIGURES 9 – 12. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma, four different specimens (partial views). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292762/files/figure.png" pageId="3" pageNumber="24">Figs 9–12</figureCitation>
), and five pairs of lyrifissures 
<emphasis id="39D4EA9BF9382026B84AF826FE57F826" box="[371,405,1953,1976]" italics="true" pageId="3" pageNumber="24">ivp</emphasis>
(excluding genital pores 
<date id="7F1E1049F9382026BB97F827FD12F829" box="[686,720,1952,1975]" pageId="3" pageNumber="24">
<emphasis id="39D4EA9BF9382026BB97F827FD12F829" box="[686,720,1952,1975]" italics="true" pageId="3" pageNumber="24">iv5</emphasis>
</date>
); 
<emphasis id="39D4EA9BF9382026BBD1F827FCCCF826" box="[744,782,1952,1976]" italics="true" pageId="3" pageNumber="24">Jv1</emphasis>
and 
<emphasis id="39D4EA9BF9382026BA7DF827FCA9F826" box="[836,875,1952,1976]" italics="true" pageId="3" pageNumber="24">Jv2</emphasis>
pairs with setae widely separated from each other, displaced posteriorly (
<emphasis id="39D4EA9BF9382026B8B4F843FE71F842" box="[397,435,1988,2012]" italics="true" pageId="3" pageNumber="24">Jv1</emphasis>
about at level of 
<emphasis id="39D4EA9BF9382026BB48F843FD59F842" box="[625,667,1988,2012]" italics="true" pageId="3" pageNumber="24">Zv2</emphasis>
); all setae smooth and needle-like, subequal in length (except 
<emphasis id="39D4EA9BF9382026BC7FF843FAAFF842" box="[1350,1389,1988,2012]" italics="true" pageId="3" pageNumber="24">Jv2</emphasis>
, the longest), not reaching base of next posterior seta (
<emphasis id="39D4EA9BF93F2021BB86FF1FFD24FF2E" box="[703,742,152,176]" italics="true" pageId="4" pageNumber="25">
Jv 
<date id="7F1E1049F93F2021BBE2FF1FFD24FF2E" box="[731,742,152,176]" pageId="4" pageNumber="25" value="1918-01-21">1</date>
</emphasis>
18 
<emphasis id="39D4EA9BF93F2021BA31FF1EFCD4FF2E" box="[776,790,153,176]" italics="true" pageId="4" pageNumber="25">–</emphasis>
21, 
<emphasis id="39D4EA9BF93F2021BA79FF1FFCA5FF2E" box="[832,871,152,176]" italics="true" pageId="4" pageNumber="25">
Jv 
<date id="7F1E1049F93F2021BA63FF1FFCA5FF2E" box="[858,871,152,176]" pageId="4" pageNumber="25" value="1923-02-25">2</date>
</emphasis>
23–25, 
<emphasis id="39D4EA9BF93F2021BAF9FF1FFC25FF2E" box="[960,999,152,176]" italics="true" pageId="4" pageNumber="25">
Jv 
<date id="7F1E1049F93F2021BAE3FF1FFC25FF2E" box="[986,999,152,176]" pageId="4" pageNumber="25" value="1918-03-21">3</date>
</emphasis>
18–21, 
<emphasis id="39D4EA9BF93F2021BD78FF1FFBAAFF2E" box="[1089,1128,152,176]" italics="true" pageId="4" pageNumber="25">
Jv 
<date id="7F1E1049F93F2021BD63FF1FFBAAFF2E" box="[1114,1128,152,176]" pageId="4" pageNumber="25" value="2013-04-15">4</date>
</emphasis>
13–15, 
<emphasis id="39D4EA9BF93F2021BDF8FF1FFB25FF31" box="[1217,1255,152,175]" italics="true" pageId="4" pageNumber="25">
Jv 
<date id="7F1E1049F93F2021BDE3FF1FFB25FF31" box="[1242,1255,152,175]" pageId="4" pageNumber="25" value="2013-05-15">5</date>
</emphasis>
13–15, 
<emphasis id="39D4EA9BF93F2021BC7BFF1FFAA9FF2E" box="[1346,1387,152,176]" italics="true" pageId="4" pageNumber="25">
Zv 
<date id="7F1E1049F93F2021BC59FF1FFAA9FF2E" box="[1376,1387,152,176]" pageId="4" pageNumber="25" value="2015-01-16">1</date>
</emphasis>
15– 16, 
<emphasis id="39D4EA9BF93F2021B9F8FF3BFF29FF4A" box="[193,235,188,212]" italics="true" pageId="4" pageNumber="25">
Zv 
<date id="7F1E1049F93F2021B9E7FF3BFF29FF4A" box="[222,235,188,212]" pageId="4" pageNumber="25" value="2015-02-18">2</date>
</emphasis>
15–18, 
<emphasis id="39D4EA9BF93F2021B87EFF3BFEB0FF4A" box="[327,370,188,212]" italics="true" pageId="4" pageNumber="25">
Zv 
<date id="7F1E1049F93F2021B85CFF3BFEB0FF4A" box="[357,370,188,212]" pageId="4" pageNumber="25" value="2015-11-03">3</date>
</emphasis>
11–15, 
<emphasis id="39D4EA9BF93F2021B8F5FF3BFE34FF4A" box="[460,502,188,212]" italics="true" pageId="4" pageNumber="25">
Zv 
<date id="7F1E1049F93F2021B8D0FF3BFE34FF4A" box="[489,502,188,212]" pageId="4" pageNumber="25" value="2013-11-04">4</date>
</emphasis>
11–13, 
<emphasis id="39D4EA9BF93F2021BB68FF3BFDB9FF4D" box="[593,635,188,211]" italics="true" pageId="4" pageNumber="25">
Zv 
<date id="7F1E1049F93F2021BB57FF3BFDB9FF4D" box="[622,635,188,211]" pageId="4" pageNumber="25" value="2013-10-05">5</date>
</emphasis>
10–13, 
<emphasis id="39D4EA9BF93F2021BBEEFF3BFD34FF4A" box="[727,758,188,212]" italics="true" pageId="4" pageNumber="25">
R 
<date id="7F1E1049F93F2021BBD3FF3BFD34FF4A" box="[746,758,188,212]" pageId="4" pageNumber="25" value="2015-11-01">1</date>
</emphasis>
11–15, 
<emphasis id="39D4EA9BF93F2021BA68FF3BFCB2FF4A" box="[849,880,188,212]" italics="true" pageId="4" pageNumber="25">
R 
<date id="7F1E1049F93F2021BA5AFF3BFCB2FF4A" box="[867,880,188,212]" pageId="4" pageNumber="25" value="2013-10-02">2</date>
</emphasis>
10–13, 
<emphasis id="39D4EA9BF93F2021BAF5FF3BFC29FF4A" box="[972,1003,188,212]" italics="true" pageId="4" pageNumber="25">
R 
<date id="7F1E1049F93F2021BAE7FF3BFC29FF4A" box="[990,1003,188,212]" pageId="4" pageNumber="25" value="2013-09-03">3</date>
</emphasis>
9–13, 
<emphasis id="39D4EA9BF93F2021BD03FF3BFB9BFF4A" box="[1082,1113,188,212]" italics="true" pageId="4" pageNumber="25">
R 
<date id="7F1E1049F93F2021BD72FF3BFB9BFF4A" box="[1099,1113,188,212]" pageId="4" pageNumber="25" value="2013-10-04">4</date>
</emphasis>
10–13, 
<emphasis id="39D4EA9BF93F2021BD8CFF3BFB16FF4D" box="[1205,1236,188,211]" italics="true" pageId="4" pageNumber="25">
R 
<date id="7F1E1049F93F2021BDFFFF3BFB16FF4D" box="[1222,1236,188,211]" pageId="4" pageNumber="25" value="2013-10-05">5</date>
</emphasis>
10–13, 
<emphasis id="39D4EA9BF93F2021BC09FF3AFA97FF4A" box="[1328,1365,189,212]" italics="true" pageId="4" pageNumber="25">UR</emphasis>
9–13, except seta 
<emphasis id="39D4EA9BF93F2021B824FF66FE80FF66" box="[285,322,225,248]" italics="true" pageId="4" pageNumber="25">UR</emphasis>
between 
<emphasis id="39D4EA9BF93F2021B888FF67FE12FF66" box="[433,464,224,248]" italics="true" pageId="4" pageNumber="25">R1</emphasis>
and post-stigmatic section of peritrematal shield, 15–19, when present always longer than setae 
<emphasis id="39D4EA9BF93F2021B834FE82FEDCFE82" box="[269,286,261,284]" italics="true" pageId="4" pageNumber="25">R</emphasis>
and other setae 
<emphasis id="39D4EA9BF93F2021B8EDFE82FE3BFE82" box="[468,505,261,284]" italics="true" pageId="4" pageNumber="25">UR</emphasis>
); metapodal platelets elongate, oriented obliquely, 42 ± 5 (35–49) long; one pair of small parapodal elements variable in shape; two pairs of rod-shaped platelets close to 
<emphasis id="39D4EA9BF93F2021BD60FEAFFB40FEDE" box="[1113,1154,296,320]" italics="true" pageId="4" pageNumber="25">Zv1</emphasis>
, one larger, between 
<emphasis id="39D4EA9BF93F2021BC4BFEAFFA5EFEDE" box="[1394,1436,296,320]" italics="true" pageId="4" pageNumber="25">Zv1</emphasis>
and lateral margin of the genital shield (generally not fused with shield), the other one tiny, external to 
<emphasis id="39D4EA9BF93F2021BC3EFECBFAF3FEFA" box="[1287,1329,332,356]" italics="true" pageId="4" pageNumber="25">Zv1</emphasis>
.
</paragraph>
<caption id="5FDF6601F93F2021B9AEF814FB4CF878" httpUri="https://zenodo.org/record/292759/files/figure.png" pageId="4" pageNumber="25" targetBox="[155,1421,421,1907]" targetPageId="4">
<paragraph id="0B1F3689F93F2021B9AEF814FB4CF878" blockId="4.[151,1436,1939,2022]" pageId="4" pageNumber="25">
<emphasis id="39D4EA9BF93F2021B9AEF814FEF7F837" bold="true" box="[151,309,1939,1961]" pageId="4" pageNumber="25">FIGURES 1–2</emphasis>
. 
<taxonomicName id="CCA04D0AF93F2021B87DF814FDFDF837" box="[324,575,1939,1961]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF93F2021B87DF814FDFDF837" box="[324,575,1939,1961]" italics="true" pageId="4" pageNumber="25">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae 
<emphasis id="39D4EA9BF93F2021B807F835FE97F856" box="[318,341,1970,1992]" italics="true" pageId="4" pageNumber="25">x2</emphasis>
and 
<emphasis id="39D4EA9BF93F2021B8B1F835FE63F856" box="[392,417,1970,1992]" italics="true" pageId="4" pageNumber="25">x3</emphasis>
, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta 
<emphasis id="39D4EA9BF93F2021B81DF856FEFDF878" box="[292,319,2001,2022]" italics="true" pageId="4" pageNumber="25">Z5</emphasis>
and lyrifissure 
<emphasis id="39D4EA9BF93F2021B8E4F857FDC9F878" box="[477,523,2000,2022]" italics="true" pageId="4" pageNumber="25">idZ5</emphasis>
(ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2.
</paragraph>
</caption>
<caption id="5FDF6601F93E2020B9AEF975FBA6F8B8" httpUri="https://zenodo.org/record/292760/files/figure.png" pageId="5" pageNumber="26" targetBox="[164,1424,214,1739]" targetPageId="5">
<paragraph id="0B1F3689F93E2020B9AEF975FBA6F8B8" blockId="5.[151,1435,1778,1830]" pageId="5" pageNumber="26">
<emphasis id="39D4EA9BF93E2020B9AEF975FEF7F896" bold="true" box="[151,309,1778,1800]" pageId="5" pageNumber="26">FIGURES 3–7</emphasis>
. 
<taxonomicName id="CCA04D0AF93E2020B87DF975FD83F896" box="[324,577,1778,1800]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF93E2020B87DF975FD83F896" box="[324,577,1778,1800]" italics="true" pageId="5" pageNumber="26">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4–7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm.
</paragraph>
</caption>
<caption id="5FDF6601F93D2023B9AEF96BFD51F8BF" httpUri="https://zenodo.org/record/292761/files/figure.png" pageId="6" pageNumber="27" targetBox="[171,1416,210,1728]" targetPageId="6">
<paragraph id="0B1F3689F93D2023B9AEF96BFD51F8BF" blockId="6.[151,1436,1772,1825]" pageId="6" pageNumber="27">
<emphasis id="39D4EA9BF93D2023B9AEF96BFECEF89C" bold="true" box="[151,268,1772,1794]" pageId="6" pageNumber="27">FIGURE 8</emphasis>
. 
<taxonomicName id="CCA04D0AF93D2023B821F96BFDD3F89C" box="[280,529,1772,1794]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF93D2023B821F96BFDD3F89C" box="[280,529,1772,1794]" italics="true" pageId="6" pageNumber="27">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), female, ventral idiosoma (partial view; setae 
<emphasis id="39D4EA9BF93D2023BD8AF96AFB1BF89C" box="[1203,1241,1773,1794]" italics="true" pageId="6" pageNumber="27">Zv5</emphasis>
and glands 
<emphasis id="39D4EA9BF93D2023BC68F96BFAB7F89C" box="[1361,1397,1772,1794]" italics="true" pageId="6" pageNumber="27">gv2</emphasis>
not discernible in this specimen). Scale bar = 100 µm.
</paragraph>
</caption>
<caption id="5FDF6601F93C2022B9AEFB18FE72FB4A" httpUri="https://zenodo.org/record/292762/files/figure.png" pageId="7" pageNumber="28" targetBox="[160,1424,207,1150]" targetPageId="7">
<paragraph id="0B1F3689F93C2022B9AEFB18FE72FB4A" blockId="7.[151,1435,1183,1236]" pageId="7" pageNumber="28">
<emphasis id="39D4EA9BF93C2022B9AEFB18FE81FB2B" bold="true" box="[151,323,1183,1205]" pageId="7" pageNumber="28">FIGURES 9–12</emphasis>
. 
<taxonomicName id="CCA04D0AF93C2022B86AFB18FD8DFB2B" box="[339,591,1183,1205]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF93C2022B86AFB18FD8DFB2B" box="[339,591,1183,1205]" italics="true" pageId="7" pageNumber="28">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), female, ventral idiosoma, four different specimens (partial views). Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="0B1F3689F93C2022B9FEFA86FA57F957" blockId="7.[151,1437,1281,2026]" pageId="7" pageNumber="28">
<taxonomicName id="CCA04D0AF93C2022B9FEFA86FE94FA87" box="[199,342,1281,1305]" class="Arachnida" family="Podocinidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF93C2022B9FEFA86FE94FA87" box="[199,342,1281,1305]" italics="true" pageId="7" pageNumber="28">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="939B2A0CF93C2022B850FA86FE33FA84" box="[361,497,1281,1306]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Figs 16–18</figureCitation>
). Epistome triangular, with anterior margin mostly smooth, with sparse isolated denticles (
<figureCitation id="939B2A0CF93C2022B835FAA3FEA0FAA3" box="[268,354,1316,1341]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Fig. 16</figureCitation>
). Hypostome with three pairs of smooth and needle-like setae, 
<emphasis id="39D4EA9BF93C2022BD05FAA2FB95FAA3" box="[1084,1111,1317,1341]" italics="true" pageId="7" pageNumber="28">h1</emphasis>
(43–46) shorter than 
<emphasis id="39D4EA9BF93C2022BC6DFAA2FAB2FAA3" box="[1364,1392,1317,1341]" italics="true" pageId="7" pageNumber="28">h3</emphasis>
, 
<emphasis id="39D4EA9BF93C2022BCB9FAA2FA59FAA3" box="[1408,1435,1317,1341]" italics="true" pageId="7" pageNumber="28">h2</emphasis>
(21–26) the shortest, 
<emphasis id="39D4EA9BF93C2022B8BFFACEFE61FAFF" box="[390,419,1353,1377]" italics="true" pageId="7" pageNumber="28">h3</emphasis>
(60–64) the longest; palpcoxal setae (33–38) smooth and needle-like; deutosternal groove usually with six-seven rows of denticles, each bearing 4-10 small teeth; corniculi horn-shaped, parallel, reaching about middle of palp-femur and about four times as long as basal width (
<figureCitation id="939B2A0CF93C2022BA87FA16FBD2FA37" box="[958,1040,1425,1450]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Fig. 18</figureCitation>
). Internal malae bilobed, inner lobe narrow, pointed and serrated, outer lobe wide, bearing well-developed fimbriae with rounded tips (
<figureCitation id="939B2A0CF93C2022BC0CFA32FA4DFA53" box="[1333,1423,1461,1486]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Fig. 18</figureCitation>
). Labrum blade-like, pilose, considerably longer than corniculi and internal malae. Palp chaetotaxy normal for the genus; apotele 2-tined, tines of unequal length; all setae smooth and needle-like, except setae 
<emphasis id="39D4EA9BF93C2022BD85FA7AFB10F98B" box="[1212,1234,1533,1557]" italics="true" pageId="7" pageNumber="28">al</emphasis>
on femur and 
<emphasis id="39D4EA9BF93C2022BC41FA7AFA5EF98B" box="[1400,1436,1533,1557]" italics="true" pageId="7" pageNumber="28">al1</emphasis>
on genu, both stout and spur-like; 
<emphasis id="39D4EA9BF93C2022BB02F9A6FD9CF9A7" box="[571,606,1569,1593]" italics="true" pageId="7" pageNumber="28">al2</emphasis>
about twice length of 
<emphasis id="39D4EA9BF93C2022BA40F9A6FC5FF9A7" box="[889,925,1569,1593]" italics="true" pageId="7" pageNumber="28">al1</emphasis>
(
<figureCitation id="939B2A0CF93C2022BA8BF9A6FBCEF9A7" box="[946,1036,1569,1594]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Fig. 18</figureCitation>
). Chelate-dentate chelicerae welldeveloped, movable digit bidentate, fixed digit multidentate generally with four robust teeth approximately equally-sized (some of which can be partially or totally fused together) and three small subapical teeth; dorsal seta short, arthrodial membrane extended into a brush-like structure; inner face of chelicera with an irregular window of weakly sclerotised integument, connected with a narrow scissure arising dorsally from base of fixed digit (
<figureCitation id="939B2A0CF93C2022BC0FF936FA4AF957" box="[1334,1416,1713,1738]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="7" pageNumber="28">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="0B1F3689F93C2022B9FEF951FBDEF874" blockId="7.[151,1437,1281,2026]" pageId="7" pageNumber="28">
<emphasis id="39D4EA9BF93C2022B9FEF951FF3EF973" box="[199,252,1750,1773]" italics="true" pageId="7" pageNumber="28">Legs</emphasis>
(
<figureCitation id="939B2A0CF93C2022B835F952FE53F973" box="[268,401,1749,1774]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="7" pageNumber="28">Figs 22, 24</figureCitation>
). Leg II stouter than other legs (
<figureCitation id="939B2A0CF93C2022BA39F952FC97F973" box="[768,853,1749,1774]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="7" pageNumber="28">Fig. 24</figureCitation>
). Chaetotaxy normal for free-living 
<taxonomicName id="CCA04D0AF93C2022BDC9F952FAA9F970" box="[1264,1387,1749,1774]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="28" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
and genus 
<taxonomicName id="CCA04D0AF93C2022B9E6F97EFEBCF88F" box="[223,382,1785,1809]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF93C2022B9E6F97EFEBCF88F" box="[223,382,1785,1809]" italics="true" pageId="7" pageNumber="28">Cosmolaelaps</emphasis>
</taxonomicName>
; leg I: coxa 0 0/1 0/1 0, trochanter 1 0/2 1/1 1, femur 2 3/1 2/3 2, genu 2 3/2 3/1 2, tibia 2 3/2 3/1 2; leg II (
<figureCitation id="939B2A0CF93C2022B810F89AFEBFF8AB" box="[297,381,1821,1846]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="7" pageNumber="28">Fig. 24</figureCitation>
): coxa 0 0/1 0/1 0, trochanter 1 0/1 0/2 1, femur 2 3/1 2/2 1, genu 2 3/1 2/1 2, tibia 2 2/1 2/1 2; leg III: coxa 0 0/1 0/1 0, trochanter 1 0/2 0/1 1, femur 1 2/1 1/0 1, genu 2 2/1 2/1 1, tibia 2 1/1 2/1 1; leg IV (
<figureCitation id="939B2A0CF93C2022BC57F8C6FF75F8E3" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="7" pageNumber="28">Fig. 22</figureCitation>
): coxa 0 0/1 0/0 0, trochanter 1 0/2 0/1 1, femur 1 2/0 1/1 1 (
<emphasis id="39D4EA9BF93C2022BA4EF8E2FC51F8E3" box="[887,915,1893,1917]" italics="true" pageId="7" pageNumber="28">pd</emphasis>
about half as long as length of 
<emphasis id="39D4EA9BF93C2022BDC7F8E2FAEAF8E3" box="[1278,1320,1893,1917]" italics="true" pageId="7" pageNumber="28">ad1</emphasis>
and 
<emphasis id="39D4EA9BF93C2022BC5BF8E2FA4EF8E3" box="[1378,1420,1893,1917]" italics="true" pageId="7" pageNumber="28">ad2</emphasis>
), genu 2 2/1 3/0 1, tibia 2 1/1 3/1 2 (
<emphasis id="39D4EA9BF93C2022BB25F80DFDF4F83F" box="[540,566,1930,1953]" italics="true" pageId="7" pageNumber="28">av</emphasis>
and 
<emphasis id="39D4EA9BF93C2022BB54F80DFD4AF83F" box="[621,648,1930,1953]" italics="true" pageId="7" pageNumber="28">pv</emphasis>
thicker and longer than other setae); tarsi II–IV with 18 setae, 3 3/2 3/ 2 3 + setae 
<emphasis id="39D4EA9BF93C2022B82EF829FEFAF85B" box="[279,312,1966,1989]" italics="true" pageId="7" pageNumber="28">mv</emphasis>
and 
<emphasis id="39D4EA9BF93C2022B857F82AFE52F85B" box="[366,400,1965,1989]" italics="true" pageId="7" pageNumber="28">md</emphasis>
; all setae simple and needle-like; all pre-tarsi with a well-developed ambulacral stalk, a pair of claws and three rounded pulvillar lobes, extensively projecting beyond claws.
</paragraph>
<caption id="5FDF6601F933202DB9AEF8BCFEACF810" httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="8" pageNumber="29" targetBox="[161,1420,221,1818]" targetPageId="8">
<paragraph id="0B1F3689F933202DB9AEF8BCFEACF810" blockId="8.[151,1437,1851,1934]" pageId="8" pageNumber="29">
<emphasis id="39D4EA9BF933202DB9AEF8BCFE82F8CF" bold="true" box="[151,320,1851,1873]" pageId="8" pageNumber="29">FIGURE 13–15</emphasis>
. 
<taxonomicName id="CCA04D0AF933202DB874F8BCFD85F8CF" box="[333,583,1851,1873]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF933202DB874F8BCFD85F8CF" box="[333,583,1851,1873]" italics="true" pageId="8" pageNumber="29">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="5FDF6601F932202CB9AEF9B5FABFF918" httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="9" pageNumber="30" targetBox="[164,1418,210,1548]" targetPageId="9">
<paragraph id="0B1F3689F932202CB9AEF9B5FABFF918" blockId="9.[151,1436,1586,1670]" pageId="9" pageNumber="30">
<emphasis id="39D4EA9BF932202CB9AEF9B5FE8FF9D6" bold="true" box="[151,333,1586,1608]" pageId="9" pageNumber="30">FIGURES 16–21</emphasis>
. 
<taxonomicName id="CCA04D0AF932202CB862F9B5FD97F9D6" box="[347,597,1586,1608]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF932202CB862F9B5FD97F9D6" box="[347,597,1586,1608]" italics="true" pageId="9" pageNumber="30">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, 
<taxonomicName id="CCA04D0AF932202CB8BBF9D6FE3EF9F9" box="[386,508,1617,1639]" class="Arachnida" family="Podocinidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="35" phylum="Arthropoda" rank="genus">gnathosoma</taxonomicName>
(ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18–21.
</paragraph>
</caption>
<paragraph id="0B1F3689F932202CB9FEF92AFE64F890" blockId="9.[151,1436,1709,1986]" pageId="9" pageNumber="30">
<emphasis id="39D4EA9BF932202CB9FEF92AFE18F95B" box="[199,474,1709,1733]" italics="true" pageId="9" pageNumber="30">Spermathecal apparatus</emphasis>
(
<figureCitation id="939B2A0CF932202CB8D1F92AFDEEF958" box="[488,556,1709,1734]" captionStart="FIGURE 8" captionStartId="6.[151,250,1772,1794]" captionTargetBox="[171,1416,210,1728]" captionTargetId="figure@6.[151,1436,193,1751]" captionTargetPageId="6" captionText="FIGURE 8. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female, ventral idiosoma (partial view; setae Zv 5 and glands gv 2 not discernible in this specimen). Scale bar = 100 µm." httpUri="https://zenodo.org/record/292761/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
). Sperm access system composed of a complex of sclerites, situated at level of coxa IV and continuous with endopodals III–IV; proximal section of tubulus sclerotised and enlarged; other structures unsclerotised.
</paragraph>
<paragraph id="0B1F3689F932202CB9FEF89EFBF7F85C" blockId="9.[151,1436,1709,1986]" pageId="9" pageNumber="30">
<emphasis id="39D4EA9BF932202CB9FEF89EFEC1F8AC" bold="true" box="[199,259,1817,1842]" italics="true" pageId="9" pageNumber="30">Male</emphasis>
(
<figureCitation id="939B2A0CF932202CB82FF89EFE5FF8AC" box="[278,413,1817,1842]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="9" pageNumber="30">Figs 13–15</figureCitation>
, 
<figureCitation id="939B2A0CF932202CB894F89DFE34F8AC" box="[429,502,1818,1842]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="9" pageNumber="30">19–21</figureCitation>
, 
<figureCitation id="939B2A0CF932202CBB3EF89DFDE7F8AC" box="[519,549,1818,1842]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="9" pageNumber="30">23</figureCitation>
, 
<figureCitation id="939B2A0CF932202CBB0FF89DFD97F8AC" box="[566,597,1818,1842]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="9" pageNumber="30">31</figureCitation>
; 
<specimenCount id="1DA6FD00F932202CBB5FF89EFD34F8AC" box="[614,758,1817,1842]" pageId="9" pageNumber="30" type="generic">5 specimens</specimenCount>
measured). Idiosoma slightly more oval-elongate than in female; 575 ± 33 (530–615) long and 415 ± 19 (395–435) wide, ratio length/width 1.39 ± 0.10 (1.28–1.51). 
<emphasis id="39D4EA9BF932202CB9FEF8E6FE47F8E7" box="[199,389,1889,1913]" italics="true" pageId="9" pageNumber="30">Dorsal idiosoma</emphasis>
(
<figureCitation id="939B2A0CF932202CB8ACF8E6FDD8F8E4" box="[405,538,1889,1914]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="9" pageNumber="30">Figs 13–14</figureCitation>
). Dorsal shield narrower than in female, with setae and pore-like structures of some series noticeably closer than in female (i.e. 
<emphasis id="39D4EA9BF932202CBBF1F802FCCBF803" box="[712,777,1925,1949]" italics="true" pageId="9" pageNumber="30">idPx2</emphasis>
, 
<emphasis id="39D4EA9BF932202CBA21F802FCF4F803" box="[792,822,1925,1949]" italics="true" pageId="9" pageNumber="30">Z4</emphasis>
and 
<emphasis id="39D4EA9BF932202CBA56F802FC61F803" box="[879,931,1925,1949]" italics="true" pageId="9" pageNumber="30">idZ4</emphasis>
); extension, shape, chaetotaxy and pore-like structures of dorsal shield as in female, except for setae 
<emphasis id="39D4EA9BF932202CBA33F82EFCADF85F" box="[778,879,1961,1985]" italics="true" pageId="9" pageNumber="30">Px1–Px3</emphasis>
generally absent.
</paragraph>
<caption id="5FDF6601F931202FB9AEF9ABFEE1F9E1" httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="10" pageNumber="31" targetBox="[174,1419,211,1540]" targetPageId="10">
<paragraph id="0B1F3689F931202FB9AEF9ABFEE1F9E1" blockId="10.[151,1436,1580,1663]" pageId="10" pageNumber="31">
<emphasis id="39D4EA9BF931202FB9AEF9ABFE8CF9DC" bold="true" box="[151,334,1580,1602]" pageId="10" pageNumber="31">FIGURES 22–24</emphasis>
. 
<taxonomicName id="CCA04D0AF931202FB865F9ABFD94F9DC" box="[348,598,1580,1602]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF931202FB865F9ABFD94F9DC" box="[348,598,1580,1602]" italics="true" pageId="10" pageNumber="31">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="0B1F3689F931202EB9FEF921FDD3FE6B" blockId="10.[151,1436,1702,2015]" lastBlockId="11.[151,1437,151,501]" lastPageId="11" lastPageNumber="32" pageId="10" pageNumber="31">
<emphasis id="39D4EA9BF931202FB9FEF921FE45F920" box="[199,391,1702,1726]" italics="true" pageId="10" pageNumber="31">Ventral idiosoma</emphasis>
(
<figureCitation id="939B2A0CF931202FB8AEF921FDD8F921" box="[407,538,1702,1727]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="10" pageNumber="31">Figs 14–15</figureCitation>
). Tritosternum and presternal area as in female. Sterno-genital, ventro-anal and endopodal shields fused in a well-developed holoventral shield, with anterior margin well defined, prominent at level of genital opening, and posterior portion of variable extension and shape; shield 475 ± 24 (455–510) long and 254 ± 16 (230–275) wide (behind coxae IV), bearing 10 pairs of simple and needle-like setae, excluding circumanal setae (
<emphasis id="39D4EA9BF931202FB82DF8B0FEA0F8D1" box="[276,354,1847,1871]" italics="true" pageId="10" pageNumber="31">st1–st5</emphasis>
, 
<emphasis id="39D4EA9BF931202FB849F8B0FE11F8D1" box="[368,467,1847,1871]" italics="true" pageId="10" pageNumber="31">Zv1–Zv2</emphasis>
, 
<emphasis id="39D4EA9BF931202FB8D9F8B0FDF9F8D1" box="[480,571,1847,1871]" italics="true" pageId="10" pageNumber="31">Jv1–Jv3</emphasis>
), 5 pairs of lyrifissures (
<date id="7F1E1049F931202FBA77F8B0FCB2F8D1" box="[846,880,1847,1871]" pageId="10" pageNumber="31">
<emphasis id="39D4EA9BF931202FBA77F8B0FCB2F8D1" box="[846,880,1847,1871]" italics="true" pageId="10" pageNumber="31">iv1</emphasis>
</date>
slit-like; a pair of lyrifissures 
<emphasis id="39D4EA9BF931202FBDF9F8B0FB20F8D0" box="[1216,1250,1847,1870]" italics="true" pageId="10" pageNumber="31">ivp</emphasis>
about at level of 
<emphasis id="39D4EA9BF931202FB9AEF8DDFF7FF8EC" box="[151,189,1882,1906]" italics="true" pageId="10" pageNumber="31">Jv1</emphasis>
) and glands 
<emphasis id="39D4EA9BF931202FB875F8DDFEB6F8EC" box="[332,372,1882,1906]" italics="true" pageId="10" pageNumber="31">gv3</emphasis>
, situated about at mid-level of anal opening; setae 
<emphasis id="39D4EA9BF931202FBA95F8DDFC15F8EC" box="[940,983,1882,1906]" italics="true" pageId="10" pageNumber="31">Zv2</emphasis>
and 
<emphasis id="39D4EA9BF931202FBD35F8DDFBF0F8EC" box="[1036,1074,1882,1906]" italics="true" pageId="10" pageNumber="31">Jv3</emphasis>
on or well removed from lateral margins of shield or on unsclerotised integument (depending on the shape and lateral extension of shield at level of these setae); 
<emphasis id="39D4EA9BF931202FB813F825FE92F824" box="[298,336,1954,1978]" italics="true" pageId="10" pageNumber="31">Jv1</emphasis>
and 
<emphasis id="39D4EA9BF931202FB8B0F825FE72F824" box="[393,432,1954,1978]" italics="true" pageId="10" pageNumber="31">Jv2</emphasis>
more anteriorly and medially located than in female; 
<emphasis id="39D4EA9BF931202FBD2DF825FBA6F824" box="[1044,1124,1954,1978]" italics="true" pageId="10" pageNumber="31">st1–st3</emphasis>
and 
<emphasis id="39D4EA9BF931202FBDA5F824FB7FF824" box="[1180,1213,1955,1978]" italics="true" pageId="10" pageNumber="31">st5</emphasis>
subequal in length, 
<emphasis id="39D4EA9BF931202FB9AEF840FF75F841" box="[151,183,1991,2015]" italics="true" pageId="10" pageNumber="31">st4</emphasis>
shorter than other sternal setae (
<tableCitation id="46220332F931202FBB08F841FD4AF841" box="[561,648,1990,2015]" captionStart="TABLE 1" captionStartId="11.[151,239,548,571]" captionTargetBox="[159,1372,598,827]" captionTargetPageId="11" captionText="TABLE 1. Length of holoventral shield setae in five males of Cosmolaelaps lignicolus." httpUri="http://table.plazi.org/id/5FDF6601F930202EB9AEFDA3FBFFFDA5" pageId="10" pageNumber="31" tableUuid="5FDF6601F930202EB9AEFDA3FBFFFDA5">Table 1</tableCitation>
); relative length of setae 
<emphasis id="39D4EA9BF931202FBA8BF840FBD6F841" box="[946,1044,1991,2015]" italics="true" pageId="10" pageNumber="31">Zv1–Zv2</emphasis>
and 
<emphasis id="39D4EA9BF931202FBD76F840FB69F841" box="[1103,1195,1991,2015]" italics="true" pageId="10" pageNumber="31">Jv1–Jv3</emphasis>
highly variable (
<emphasis id="39D4EA9BF931202FBC4BF840FA5EF841" box="[1394,1436,1991,2015]" italics="true" pageId="10" pageNumber="31">Zv1</emphasis>
with greater variation; 
<tableCitation id="46220332F930202EB8A4FF10FE32FF2E" box="[413,496,151,176]" captionStart="TABLE 1" captionStartId="11.[151,239,548,571]" captionTargetBox="[159,1372,598,827]" captionTargetPageId="11" captionText="TABLE 1. Length of holoventral shield setae in five males of Cosmolaelaps lignicolus." httpUri="http://table.plazi.org/id/5FDF6601F930202EB9AEFDA3FBFFFDA5" pageId="11" pageNumber="32" tableUuid="5FDF6601F930202EB9AEFDA3FBFFFDA5">Table 1</tableCitation>
); para-anal setae longer than unpaired post-anal seta (as in female); cribrum as in female; shield throughout with polygonal ornamentation, except posteriorly to anal opening. Peritrematal shields only with antiaxial and post-stigmatic sections developed, not extending behind coxa IV and free from exopodals (as in female), broadly fused to dorsal shield in their anterior portion, about to level of 
<emphasis id="39D4EA9BF930202EBDAFFE83FB6DFE82" box="[1174,1199,260,284]" italics="true" pageId="11" pageNumber="32">r2</emphasis>
(
<figureCitation id="939B2A0CF930202EBDFBFE83FADAFE82" box="[1218,1304,260,285]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="11" pageNumber="32">Fig. 14</figureCitation>
); peritreme extending anteriorly beyond coxae I, up to level of 
<emphasis id="39D4EA9BF930202EBBE8FEAFFD29FEDE" box="[721,747,296,320]" italics="true" pageId="11" pageNumber="32">s1</emphasis>
(
<figureCitation id="939B2A0CF930202EBBC0FEA0FC8EFEDE" box="[761,844,295,320]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="11" pageNumber="32">Fig. 14</figureCitation>
); peritrematal and post-stigmatic pore-like structures as in female. Setae 
<emphasis id="39D4EA9BF930202EB841FECBFE52FEFA" box="[376,400,332,356]" italics="true" pageId="11" pageNumber="32">r6</emphasis>
as in female. Exopodal shields as in female; exopodals III–IV connected with holoventral shield. Opisthogastric unsclerotised integument with 10 pairs of setae, excluding a variable number of setae 
<emphasis id="39D4EA9BF930202EBC56FEF6FA56FE16" box="[1391,1428,369,392]" italics="true" pageId="11" pageNumber="32">UR</emphasis>
, and 5 pairs of lyrifissures 
<emphasis id="39D4EA9BF930202EB887FE12FE22FE32" box="[446,480,405,428]" italics="true" pageId="11" pageNumber="32">ivp</emphasis>
; all setae smooth and needle-like, subequal in length (
<emphasis id="39D4EA9BF930202EBD73FE13FB71FE35" box="[1098,1203,404,428]" italics="true" pageId="11" pageNumber="32">
Zv3– Zv 
<date id="7F1E1049F930202EBD9FFE13FB71FE35" box="[1190,1203,404,427]" pageId="11" pageNumber="32" value="2016-11-05">5</date>
</emphasis>
11–16, 
<emphasis id="39D4EA9BF930202EBC36FE13FAAEFE35" box="[1295,1388,404,428]" italics="true" pageId="11" pageNumber="32">
Jv4–Jv 
<date id="7F1E1049F930202EBC67FE13FAAEFE35" box="[1374,1388,404,427]" pageId="11" pageNumber="32" value="1918-11-05">5</date>
</emphasis>
11– 18; seta 
<emphasis id="39D4EA9BF930202EB9CDFE3EFEDBFE4E" box="[244,281,441,464]" italics="true" pageId="11" pageNumber="32">UR</emphasis>
between 
<emphasis id="39D4EA9BF930202EB8BCFE3FFE67FE4E" box="[389,421,440,464]" italics="true" pageId="11" pageNumber="32">R1</emphasis>
and post-stigmatic section of peritrematal shield, when present always longer than setae 
<emphasis id="39D4EA9BF930202EBCB2FE3EFA5EFE4E" box="[1419,1436,441,464]" italics="true" pageId="11" pageNumber="32">R</emphasis>
and other setae 
<emphasis id="39D4EA9BF930202EB87FFE5AFEA9FE6A" box="[326,363,477,500]" italics="true" pageId="11" pageNumber="32">UR</emphasis>
, as in female).
</paragraph>
<caption id="5FDF6601F930202EB9AEFDA3FBFFFDA5" ID-Table-UUID="5FDF6601F930202EB9AEFDA3FBFFFDA5" box="[151,1085,547,571]" httpUri="http://table.plazi.org/id/5FDF6601F930202EB9AEFDA3FBFFFDA5" pageId="11" pageNumber="32" targetBox="[159,1372,598,827]" targetIsTable="true" targetPageId="11">
<paragraph id="0B1F3689F930202EB9AEFDA3FBFFFDA5" blockId="11.[151,1085,547,571]" box="[151,1085,547,571]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB9AEFDA3FEC1FDA5" bold="true" box="[151,259,548,571]" pageId="11" pageNumber="32">TABLE 1</emphasis>
. Length of holoventral shield setae in five males of 
<taxonomicName id="CCA04D0AF930202EBA16FDA4FBF4FDA5" box="[815,1078,547,571]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF930202EBA16FDA4FBF4FDA5" box="[815,1078,547,571]" italics="true" pageId="11" pageNumber="32">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="0B1F3689F930202EB83CFDD1FA91FCA5" pageId="11" pageNumber="32">
<table id="79A0C429F930DFDAB9A6FDD1FA9EFCA5" box="[159,1372,598,827]" gridcols="11" gridrows="6" pageId="11" pageNumber="32">
<tr id="B59034CBF930DFDAB9A6FDD1FA9EFDF2" box="[159,1372,598,620]" gridrow="0" pageId="11" pageNumber="32" rowspan-0="1">
<th id="F6415DB7F930DFDAB83CFDD1FEE1FDF2" box="[261,291,598,620]" gridcol="1" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB83CFDD1FEE1FDF2" box="[261,291,598,620]" italics="true" pageId="11" pageNumber="32">st1</emphasis>
</th>
<th id="F6415DB7F930DFDAB845FDD1FE5BFDF2" box="[380,409,598,620]" gridcol="2" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB845FDD1FE5BFDF2" box="[380,409,598,620]" italics="true" pageId="11" pageNumber="32">st2</emphasis>
</th>
<th id="F6415DB7F930DFDAB8CBFDD1FDCDFDF2" box="[498,527,598,620]" gridcol="3" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB8CBFDD1FDCDFDF2" box="[498,527,598,620]" italics="true" pageId="11" pageNumber="32">st3</emphasis>
</th>
<th id="F6415DB7F930DFDABB54FDD1FD49FDF2" box="[621,651,598,620]" gridcol="4" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBB54FDD1FD49FDF2" box="[621,651,598,620]" italics="true" pageId="11" pageNumber="32">st4</emphasis>
</th>
<th id="F6415DB7F930DFDABBDDFDD1FCC0FDF2" box="[740,770,598,620]" gridcol="5" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBBDDFDD0FCC0FDF2" box="[740,770,599,620]" italics="true" pageId="11" pageNumber="32">st5</emphasis>
</th>
<th id="F6415DB7F930DFDABA63FDD1FC42FDF2" box="[858,896,598,620]" gridcol="6" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBA63FDD1FC42FDF2" box="[858,896,598,620]" italics="true" pageId="11" pageNumber="32">Zv1</emphasis>
</th>
<th id="F6415DB7F930DFDABAEBFDD1FC35FDF2" box="[978,1015,598,620]" gridcol="7" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBAEBFDD1FC35FDF2" box="[978,1015,598,620]" italics="true" pageId="11" pageNumber="32">Zv2</emphasis>
</th>
<th id="F6415DB7F930DFDABD77FDD1FBB3FDF2" box="[1102,1137,598,620]" gridcol="8" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBD77FDD1FBB3FDF2" box="[1102,1137,598,620]" italics="true" pageId="11" pageNumber="32">Jv1</emphasis>
</th>
<th id="F6415DB7F930DFDABDFAFDD1FB24FDF2" box="[1219,1254,598,620]" gridcol="9" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBDFAFDD1FB24FDF2" box="[1219,1254,598,620]" italics="true" pageId="11" pageNumber="32">Jv2</emphasis>
</th>
<th id="F6415DB7F930DFDABC00FDD1FA9EFDF2" box="[1337,1372,598,620]" gridcol="10" gridrow="0" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBC00FDD1FA9EFDF2" box="[1337,1372,598,620]" italics="true" pageId="11" pageNumber="32">Jv3</emphasis>
</th>
</tr>
<tr id="B59034CBF930DFDAB9A6FD07FA9EFD08" box="[159,1372,640,662]" gridrow="1" pageId="11" pageNumber="32">
<th id="F6415DB7F930DFDAB9A6FD07FF6EFD08" box="[159,172,640,662]" gridcol="0" gridrow="1" pageId="11" pageNumber="32">1</th>
<td id="F6415DB7F930DFDAB83CFD07FEE1FD08" box="[261,291,640,662]" gridcol="1" gridrow="1" pageId="11" pageNumber="32">28</td>
<td id="F6415DB7F930DFDAB845FD07FE5BFD08" box="[380,409,640,662]" gridcol="2" gridrow="1" pageId="11" pageNumber="32">28</td>
<td id="F6415DB7F930DFDAB8CBFD07FDCDFD08" box="[498,527,640,662]" gridcol="3" gridrow="1" pageId="11" pageNumber="32">28</td>
<td id="F6415DB7F930DFDABB54FD07FD49FD08" box="[621,651,640,662]" gridcol="4" gridrow="1" pageId="11" pageNumber="32">23</td>
<td id="F6415DB7F930DFDABBDDFD07FCC0FD08" box="[740,770,640,662]" gridcol="5" gridrow="1" pageId="11" pageNumber="32">-</td>
<td id="F6415DB7F930DFDABA63FD07FC42FD08" box="[858,896,640,662]" gridcol="6" gridrow="1" pageId="11" pageNumber="32">30</td>
<td id="F6415DB7F930DFDABAEBFD07FC35FD08" box="[978,1015,640,662]" gridcol="7" gridrow="1" pageId="11" pageNumber="32">39</td>
<td id="F6415DB7F930DFDABD77FD07FBB3FD08" box="[1102,1137,640,662]" gridcol="8" gridrow="1" pageId="11" pageNumber="32">36</td>
<td id="F6415DB7F930DFDABDFAFD07FB24FD08" box="[1219,1254,640,662]" gridcol="9" gridrow="1" pageId="11" pageNumber="32">39</td>
<td id="F6415DB7F930DFDABC00FD07FA9EFD08" box="[1337,1372,640,662]" gridcol="10" gridrow="1" pageId="11" pageNumber="32">29</td>
</tr>
<tr id="B59034CBF930DFDAB9A6FD2EFA9EFD21" box="[159,1372,681,703]" gridrow="2" pageId="11" pageNumber="32">
<th id="F6415DB7F930DFDAB9A6FD2EFF6EFD21" box="[159,172,681,703]" gridcol="0" gridrow="2" pageId="11" pageNumber="32">2</th>
<td id="F6415DB7F930DFDAB83CFD2EFEE1FD21" box="[261,291,681,703]" gridcol="1" gridrow="2" pageId="11" pageNumber="32">2 8</td>
<td id="F6415DB7F930DFDAB845FD2EFE5BFD21" box="[380,409,681,703]" gridcol="2" gridrow="2" pageId="11" pageNumber="32">2 6</td>
<td id="F6415DB7F930DFDAB8CBFD2EFDCDFD21" box="[498,527,681,703]" gridcol="3" gridrow="2" pageId="11" pageNumber="32">2 8</td>
<td id="F6415DB7F930DFDABB54FD2EFD49FD21" box="[621,651,681,703]" gridcol="4" gridrow="2" pageId="11" pageNumber="32">2 3</td>
<td id="F6415DB7F930DFDABBDDFD2EFCC0FD21" box="[740,770,681,703]" gridcol="5" gridrow="2" pageId="11" pageNumber="32">2 5</td>
<td id="F6415DB7F930DFDABA63FD2EFC42FD21" box="[858,896,681,703]" gridcol="6" gridrow="2" pageId="11" pageNumber="32">1 6</td>
<td id="F6415DB7F930DFDABAEBFD2EFC35FD21" box="[978,1015,681,703]" gridcol="7" gridrow="2" pageId="11" pageNumber="32">4 4</td>
<td id="F6415DB7F930DFDABD77FD2EFBB3FD21" box="[1102,1137,681,703]" gridcol="8" gridrow="2" pageId="11" pageNumber="32">4 0</td>
<td id="F6415DB7F930DFDABDFAFD2EFB24FD21" box="[1219,1254,681,703]" gridcol="9" gridrow="2" pageId="11" pageNumber="32">4 5</td>
<td id="F6415DB7F930DFDABC00FD2EFA9EFD21" box="[1337,1372,681,703]" gridcol="10" gridrow="2" pageId="11" pageNumber="32">3 6</td>
</tr>
<tr id="B59034CBF930DFDAB9A6FD55FA9EFD76" box="[159,1372,722,744]" gridrow="3" pageId="11" pageNumber="32">
<th id="F6415DB7F930DFDAB9A6FD55FF6EFD76" box="[159,172,722,744]" gridcol="0" gridrow="3" pageId="11" pageNumber="32">3</th>
<td id="F6415DB7F930DFDAB83CFD55FEE1FD76" box="[261,291,722,744]" gridcol="1" gridrow="3" pageId="11" pageNumber="32">2 5</td>
<td id="F6415DB7F930DFDAB845FD55FE5BFD76" box="[380,409,722,744]" gridcol="2" gridrow="3" pageId="11" pageNumber="32">2 8</td>
<td id="F6415DB7F930DFDAB8CBFD55FDCDFD76" box="[498,527,722,744]" gridcol="3" gridrow="3" pageId="11" pageNumber="32">2 6</td>
<td id="F6415DB7F930DFDABB54FD55FD49FD76" box="[621,651,722,744]" gridcol="4" gridrow="3" pageId="11" pageNumber="32">2 0</td>
<td id="F6415DB7F930DFDABBDDFD55FCC0FD76" box="[740,770,722,744]" gridcol="5" gridrow="3" pageId="11" pageNumber="32">2 5</td>
<td id="F6415DB7F930DFDABA63FD55FC42FD76" box="[858,896,722,744]" gridcol="6" gridrow="3" pageId="11" pageNumber="32">2 3</td>
<td id="F6415DB7F930DFDABAEBFD55FC35FD76" box="[978,1015,722,744]" gridcol="7" gridrow="3" pageId="11" pageNumber="32">5 1</td>
<td id="F6415DB7F930DFDABD77FD55FBB3FD76" box="[1102,1137,722,744]" gridcol="8" gridrow="3" pageId="11" pageNumber="32">4 5</td>
<td id="F6415DB7F930DFDABDFAFD55FB24FD76" box="[1219,1254,722,744]" gridcol="9" gridrow="3" pageId="11" pageNumber="32">5 0</td>
<td id="F6415DB7F930DFDABC00FD55FA9EFD76" box="[1337,1372,722,744]" gridcol="10" gridrow="3" pageId="11" pageNumber="32">3 8</td>
</tr>
<tr id="B59034CBF930DFDAB9A6FD7BFA9EFC8C" box="[159,1372,764,786]" gridrow="4" pageId="11" pageNumber="32">
<th id="F6415DB7F930DFDAB9A6FD7BFF6EFC8C" box="[159,172,764,786]" gridcol="0" gridrow="4" pageId="11" pageNumber="32">4</th>
<td id="F6415DB7F930DFDAB83CFD7BFEE1FC8C" box="[261,291,764,786]" gridcol="1" gridrow="4" pageId="11" pageNumber="32">-</td>
<td id="F6415DB7F930DFDAB845FD7BFE5BFC8C" box="[380,409,764,786]" gridcol="2" gridrow="4" pageId="11" pageNumber="32">26</td>
<td id="F6415DB7F930DFDAB8CBFD7BFDCDFC8C" box="[498,527,764,786]" gridcol="3" gridrow="4" pageId="11" pageNumber="32">26</td>
<td id="F6415DB7F930DFDABB54FD7BFD49FC8C" box="[621,651,764,786]" gridcol="4" gridrow="4" pageId="11" pageNumber="32">21</td>
<td id="F6415DB7F930DFDABBDDFD7BFCC0FC8C" box="[740,770,764,786]" gridcol="5" gridrow="4" pageId="11" pageNumber="32">24</td>
<td id="F6415DB7F930DFDABA63FD7BFC42FC8C" box="[858,896,764,786]" gridcol="6" gridrow="4" pageId="11" pageNumber="32">21</td>
<td id="F6415DB7F930DFDABAEBFD7BFC35FC8C" box="[978,1015,764,786]" gridcol="7" gridrow="4" pageId="11" pageNumber="32">31</td>
<td id="F6415DB7F930DFDABD77FD7BFBB3FC8C" box="[1102,1137,764,786]" gridcol="8" gridrow="4" pageId="11" pageNumber="32">30</td>
<td id="F6415DB7F930DFDABDFAFD7BFB24FC8C" box="[1219,1254,764,786]" gridcol="9" gridrow="4" pageId="11" pageNumber="32">34</td>
<td id="F6415DB7F930DFDABC00FD7BFA9EFC8C" box="[1337,1372,764,786]" gridcol="10" gridrow="4" pageId="11" pageNumber="32">33</td>
</tr>
<tr id="B59034CBF930DFDAB9A6FCA2FA9EFCA5" box="[159,1372,805,827]" gridrow="5" pageId="11" pageNumber="32">
<th id="F6415DB7F930DFDAB9A6FCA2FF6EFCA5" box="[159,172,805,827]" gridcol="0" gridrow="5" pageId="11" pageNumber="32">5</th>
<td id="F6415DB7F930DFDAB83CFCA2FEE1FCA5" box="[261,291,805,827]" gridcol="1" gridrow="5" pageId="11" pageNumber="32">2 5</td>
<td id="F6415DB7F930DFDAB845FCA2FE5BFCA5" box="[380,409,805,827]" gridcol="2" gridrow="5" pageId="11" pageNumber="32">2 6</td>
<td id="F6415DB7F930DFDAB8CBFCA2FDCDFCA5" box="[498,527,805,827]" gridcol="3" gridrow="5" pageId="11" pageNumber="32">2 6</td>
<td id="F6415DB7F930DFDABB54FCA2FD49FCA5" box="[621,651,805,827]" gridcol="4" gridrow="5" pageId="11" pageNumber="32">2 1</td>
<td id="F6415DB7F930DFDABBDDFCA2FCC0FCA5" box="[740,770,805,827]" gridcol="5" gridrow="5" pageId="11" pageNumber="32">2 5</td>
<td id="F6415DB7F930DFDABA63FCA2FC42FCA5" box="[858,896,805,827]" gridcol="6" gridrow="5" pageId="11" pageNumber="32">2 9</td>
<td id="F6415DB7F930DFDABAEBFCA2FC35FCA5" box="[978,1015,805,827]" gridcol="7" gridrow="5" pageId="11" pageNumber="32">4 5</td>
<td id="F6415DB7F930DFDABD77FCA2FBB3FCA5" box="[1102,1137,805,827]" gridcol="8" gridrow="5" pageId="11" pageNumber="32">4 5</td>
<td id="F6415DB7F930DFDABDFAFCA2FB24FCA5" box="[1219,1254,805,827]" gridcol="9" gridrow="5" pageId="11" pageNumber="32">5 1</td>
<td id="F6415DB7F930DFDABC00FCA2FA9EFCA5" box="[1337,1372,805,827]" gridcol="10" gridrow="5" pageId="11" pageNumber="32">3 9</td>
</tr>
</table>
</paragraph>
<paragraph id="0B1F3689F930202EB9FEFC07FBD7FBEE" blockId="11.[151,1437,895,2037]" pageId="11" pageNumber="32">
<taxonomicName id="CCA04D0AF930202EB9FEFC07FE94FC06" box="[199,342,896,920]" class="Arachnida" family="Podocinidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF930202EB9FEFC07FE94FC06" box="[199,342,896,920]" italics="true" pageId="11" pageNumber="32">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="939B2A0CF930202EB85CFCF8FE2AFC06" box="[357,488,895,920]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="11" pageNumber="32">Figs 19–21</figureCitation>
). Epistome (
<figureCitation id="939B2A0CF930202EBB4CFCF8FD04FC06" box="[629,710,895,920]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="11" pageNumber="32">Fig. 19</figureCitation>
) and hypostome as in female. Inner lobes of internal malae as in female, outer lobes narrow, serrated, without fimbriae (
<figureCitation id="939B2A0CF930202EBBC4FC23FC92FC22" box="[765,848,932,957]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="11" pageNumber="32">Fig. 21</figureCitation>
). Labrum as in female. Palp chaetotaxy as in female. Movable digit of chelicera with one large tooth, spermatodactyl digitiform, longer than movable digit and with rounded tip; fixed digit multidentate, with two-three distal teeth (subapical tooth clearly larger) and a well separated and oblique more proximal masticatory ridge, composed by three-four small teeth; inner face of chelicera with a narrow scissure arising dorsally from base of fixed digit (without the window of weakly sclerotised integument present in female); other cheliceral structures as in female (
<figureCitation id="939B2A0CF930202EBA8DFBD0FBCAFBEE" box="[948,1032,1111,1136]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="11" pageNumber="32">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="0B1F3689F930202EB9FEFBFAFCB3FB42" blockId="11.[151,1437,895,2037]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB9FEFBFAFF3EFB0A" box="[199,252,1149,1172]" italics="true" pageId="11" pageNumber="32">Legs</emphasis>
(
<figureCitation id="939B2A0CF930202EB835FBFBFEA7FB0A" box="[268,357,1148,1173]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="11" pageNumber="32">Figs 23</figureCitation>
, 
<figureCitation id="939B2A0CF930202EB84AFBFBFE50FB0A" box="[371,402,1148,1172]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="11" pageNumber="32">31</figureCitation>
). Chaetotaxy as in female. Leg II stouter than in female; seta 
<emphasis id="39D4EA9BF930202EBD63FBFAFBB7FB0A" box="[1114,1141,1149,1172]" italics="true" pageId="11" pageNumber="32">av</emphasis>
on femur II inserted on a well-developed tubercle and shorter than in female; seta 
<emphasis id="39D4EA9BF930202EBA35FB26FCE5FB26" box="[780,807,1185,1208]" italics="true" pageId="11" pageNumber="32">av</emphasis>
on genu and tibia II, and setae 
<emphasis id="39D4EA9BF930202EBDB9FB26FB5DFB26" box="[1152,1183,1185,1208]" italics="true" pageId="11" pageNumber="32">mv</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD95FB27FB16FB26" box="[1196,1236,1184,1208]" italics="true" pageId="11" pageNumber="32">pv2</emphasis>
and 
<emphasis id="39D4EA9BF930202EBC30FB27FAF0FB26" box="[1289,1330,1184,1208]" italics="true" pageId="11" pageNumber="32">av2</emphasis>
on tarsus II, thickened, spine-like, and shorter than in female (
<figureCitation id="939B2A0CF930202EBBDBFB43FCFBFB42" box="[738,825,1220,1245]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="11" pageNumber="32">Figs 23</figureCitation>
, 
<figureCitation id="939B2A0CF930202EBA7CFB43FCA6FB42" box="[837,868,1220,1244]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="11" pageNumber="32">31</figureCitation>
).
</paragraph>
<paragraph id="0B1F3689F930202EB9FEFB60FDFEFAD6" blockId="11.[151,1437,895,2037]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB9FEFB60FE9EFA9E" bold="true" box="[199,348,1255,1280]" italics="true" pageId="11" pageNumber="32">Deutonymph</emphasis>
. 
<figureCitation id="939B2A0CF930202EB852FB60FE2EFA9E" box="[363,492,1255,1280]" captionStart-0="FIGURES 25 – 26" captionStart-1="FIGURES 27 – 28" captionStartId-0="12.[151,264,1969,1991]" captionStartId-1="13.[151,264,1437,1459]" captionTargetBox-0="[231,1335,206,1928]" captionTargetBox-1="[183,1388,193,1405]" captionTargetId-0="figure@12.[225,1345,193,1942]" captionTargetId-1="figure@13.[173,1411,181,1417]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." captionText-1="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri-0="https://zenodo.org/record/292766/files/figure.png" httpUri-1="https://zenodo.org/record/292767/files/figure.png" pageId="11" pageNumber="32">Figs 25–28</figureCitation>
, 
<figureCitation id="939B2A0CF930202EB8C0FB6FFD80FA9E" box="[505,578,1256,1280]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="11" pageNumber="32">32–33</figureCitation>
; 
<specimenCount id="1DA6FD00F930202EBB77FB60FD18FA9E" box="[590,730,1255,1280]" pageId="11" pageNumber="32" type="generic">6 specimens</specimenCount>
measured. Exoskeleton soft, depigmented. Idiosoma narrower and with posterior end more acute than in adults, 539 ± 19 (520–570) long and 356 ± 21 (330–390) wide, ratio length/width 1.52 ± 0.04 (1.46–1.58).
</paragraph>
<paragraph id="0B1F3689F930202EB9FEFAD3FC84F86B" blockId="11.[151,1437,895,2037]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB9FEFAD3FE44FAF2" box="[199,390,1364,1388]" italics="true" pageId="11" pageNumber="32">Dorsal idiosoma</emphasis>
(
<figureCitation id="939B2A0CF930202EB8A0FAD3FDDCFAF3" box="[409,542,1364,1389]" captionStart="FIGURES 25 – 26" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[231,1335,206,1928]" captionTargetId="figure@12.[225,1345,193,1942]" captionTargetPageId="12" captionText="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="11" pageNumber="32">Figs 25–26</figureCitation>
, 
<figureCitation id="939B2A0CF930202EBB14FAD3FD89FAF2" box="[557,587,1364,1388]" captionStart="FIGURES 27 – 28" captionStartId="13.[151,264,1437,1459]" captionTargetBox="[183,1388,193,1405]" captionTargetId="figure@13.[173,1411,181,1417]" captionTargetPageId="13" captionText="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="11" pageNumber="32">28</figureCitation>
, 
<figureCitation id="939B2A0CF930202EBB62FAD3FD64FAF2" box="[603,678,1364,1388]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="11" pageNumber="32">32–33</figureCitation>
). Dorsal shield entire, posteriorly not completely covering dorsal surface, significantly less convex and laterally less expanded than in adults, with a narrow incision along each lateral margin, just anteriorly to 
<emphasis id="39D4EA9BF930202EBB3AFA1BFDDDFA2A" box="[515,543,1436,1460]" italics="true" pageId="11" pageNumber="32">S1</emphasis>
and extending medially about up to level of 
<emphasis id="39D4EA9BF930202EBD23FA1BFBF1FA2A" box="[1050,1075,1436,1460]" italics="true" pageId="11" pageNumber="32">z6</emphasis>
; with 40 pairs of setae, 23 pairs on podonotum (
<emphasis id="39D4EA9BF930202EB877FA47FE45FA46" box="[334,391,1472,1496]" italics="true" pageId="11" pageNumber="32">j1–j6</emphasis>
, 
<emphasis id="39D4EA9BF930202EB8AEFA47FE15FA46" box="[407,471,1472,1496]" italics="true" pageId="11" pageNumber="32">z1–z6</emphasis>
, 
<emphasis id="39D4EA9BF930202EB8DEFA47FDE5FA46" box="[487,551,1472,1496]" italics="true" pageId="11" pageNumber="32">s1–s6</emphasis>
, 
<emphasis id="39D4EA9BF930202EBB0EFA47FDB0FA49" box="[567,626,1472,1496]" italics="true" pageId="11" pageNumber="32">r2-r5</emphasis>
; 
<emphasis id="39D4EA9BF930202EBBBAFA47FD5EFA46" box="[643,668,1472,1496]" italics="true" pageId="11" pageNumber="32">r6</emphasis>
ventrally placed, as in adults) including a marginal supernumerary pair (
<date id="7F1E1049F930202EB9EDFA63FF2CFA62" box="[212,238,1508,1532]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EB9EDFA63FF2CFA62" box="[212,238,1508,1532]" italics="true" pageId="11" pageNumber="32">x1</emphasis>
</date>
) posterior to 
<emphasis id="39D4EA9BF930202EB8B5FA63FE67FA62" box="[396,421,1508,1532]" italics="true" pageId="11" pageNumber="32">s6</emphasis>
setae, and 17 pairs on opisthonotum (
<emphasis id="39D4EA9BF930202EBA65FA63FC5CFA65" box="[860,926,1508,1532]" italics="true" pageId="11" pageNumber="32">J1–J5</emphasis>
, 
<emphasis id="39D4EA9BF930202EBA97FA63FC35FA65" box="[942,1015,1508,1532]" italics="true" pageId="11" pageNumber="32">Z1–Z5</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD3EFA63FB8FFA65" box="[1031,1101,1508,1532]" italics="true" pageId="11" pageNumber="32">S1–S5</emphasis>
) including two pairs of extra setae (
<emphasis id="39D4EA9BF930202EB9E6F98FFE86F9BE" box="[223,324,1544,1568]" italics="true" pageId="11" pageNumber="32">Px2–Px3</emphasis>
) between 
<emphasis id="39D4EA9BF930202EB883F98EFE04F9BE" box="[442,454,1545,1568]" italics="true" pageId="11" pageNumber="32">J</emphasis>
and 
<emphasis id="39D4EA9BF930202EB8C4F98EFDCEF9BE" box="[509,524,1545,1568]" italics="true" pageId="11" pageNumber="32">Z</emphasis>
series (setae 
<date id="7F1E1049F930202EBB9DF98FFD7CF9BE" box="[676,702,1544,1568]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBB9DF98FFD7CF9BE" box="[676,702,1544,1568]" italics="true" pageId="11" pageNumber="32">x2</emphasis>
</date>
, 
<date id="7F1E1049F930202EBBF5F98FFD25F9BE" box="[716,743,1544,1568]" pageId="11" pageNumber="32">
<emphasis id="39D4EA9BF930202EBBF5F98FFD25F9BE" box="[716,743,1544,1568]" italics="true" pageId="11" pageNumber="32">x3</emphasis>
</date>
and 
<emphasis id="39D4EA9BF930202EBA24F98FFC8BF9BE" box="[797,841,1544,1568]" italics="true" pageId="11" pageNumber="32">Px1</emphasis>
absent); with a variable number (10–16) of unpaired setae (
<emphasis id="39D4EA9BF930202EB9D9F9AAFF3BF9DA" box="[224,249,1581,1604]" italics="true" pageId="11" pageNumber="32">Jx</emphasis>
) always present between 
<emphasis id="39D4EA9BF930202EBB24F9AAFDEBF9DA" box="[541,553,1581,1604]" italics="true" pageId="11" pageNumber="32">J</emphasis>
series; disorganized additional setae 
<emphasis id="39D4EA9BF930202EBAF7F9AAFC18F9DA" box="[974,986,1581,1604]" italics="true" pageId="11" pageNumber="32">x</emphasis>
(0–3) may be present between setae 
<emphasis id="39D4EA9BF930202EBC47F9AAFA59F9DA" box="[1406,1435,1581,1604]" italics="true" pageId="11" pageNumber="32">Px</emphasis>
and 
<emphasis id="39D4EA9BF930202EB9F0F9D6FF17F9F6" box="[201,213,1617,1640]" italics="true" pageId="11" pageNumber="32">J</emphasis>
series; in some cases setae 
<emphasis id="39D4EA9BF930202EBB26F9D6FDE8F9F6" box="[543,554,1617,1640]" italics="true" pageId="11" pageNumber="32">r</emphasis>
on unsclerotised integument (i.e. 
<emphasis id="39D4EA9BF930202EBA8FF9D7FC0DF9F6" box="[950,975,1616,1640]" italics="true" pageId="11" pageNumber="32">r3</emphasis>
, 
<figureCitation id="939B2A0CF930202EBAD9F9C8FBF4F9F6" box="[992,1078,1615,1640]" captionStart="FIGURES 25 – 26" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[231,1335,206,1928]" captionTargetId="figure@12.[225,1345,193,1942]" captionTargetPageId="12" captionText="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="11" pageNumber="32">Fig. 25</figureCitation>
; 
<emphasis id="39D4EA9BF930202EBD7CF9D7FB9CF9F6" box="[1093,1118,1616,1640]" italics="true" pageId="11" pageNumber="32">r4</emphasis>
, 
<figureCitation id="939B2A0CF930202EBD56F9C8FB04F9F6" box="[1135,1222,1615,1640]" captionStart="FIGURES 27 – 28" captionStartId="13.[151,264,1437,1459]" captionTargetBox="[183,1388,193,1405]" captionTargetId="figure@13.[173,1411,181,1417]" captionTargetPageId="13" captionText="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="11" pageNumber="32">Fig. 28</figureCitation>
); 
<emphasis id="39D4EA9BF930202EBDE4F9D7FB3AF9F6" box="[1245,1272,1616,1640]" italics="true" pageId="11" pageNumber="32">S3</emphasis>
on the lateral margin of shield or on unsclerotised integument (
<figureCitation id="939B2A0CF930202EBBF3F9F3FCDDF913" box="[714,799,1652,1677]" captionStart="FIGURES 25 – 26" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[231,1335,206,1928]" captionTargetId="figure@12.[225,1345,193,1942]" captionTargetPageId="12" captionText="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="11" pageNumber="32">Fig. 25</figureCitation>
); 
<emphasis id="39D4EA9BF930202EBA0CF9F3FC90F912" box="[821,850,1652,1676]" italics="true" pageId="11" pageNumber="32">Z3</emphasis>
close to lateral margin of shield, but well removed from it; shape of setae as in adults (
<figureCitation id="939B2A0CF930202EBB27F910FDB6F92E" box="[542,628,1687,1712]" captionStart="FIGURES 25 – 26" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[231,1335,206,1928]" captionTargetId="figure@12.[225,1345,193,1942]" captionTargetPageId="12" captionText="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="11" pageNumber="32">Figs 26</figureCitation>
, 
<figureCitation id="939B2A0CF930202EBBB9F91FFD08F92E" box="[640,714,1688,1712]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="11" pageNumber="32">32–33</figureCitation>
); some setae relatively longer than in adults and reaching base of next posterior seta (i.e. 
<emphasis id="39D4EA9BF930202EB895F93BFE04F94A" box="[428,454,1724,1748]" italics="true" pageId="11" pageNumber="32">z3</emphasis>
and 
<emphasis id="39D4EA9BF930202EBB3AF93BFDD9F94D" box="[515,539,1724,1747]" italics="true" pageId="11" pageNumber="32">s5</emphasis>
reaching beyond insertion of 
<emphasis id="39D4EA9BF930202EBA44F93BFC54F94A" box="[893,918,1724,1748]" italics="true" pageId="11" pageNumber="32">z4</emphasis>
and 
<emphasis id="39D4EA9BF930202EBAEDF93BFC2FF94A" box="[980,1005,1724,1748]" italics="true" pageId="11" pageNumber="32">z6</emphasis>
, respectively), 
<emphasis id="39D4EA9BF930202EBD98F93BFB75F94A" box="[1185,1207,1724,1748]" italics="true" pageId="11" pageNumber="32">j1</emphasis>
and 
<emphasis id="39D4EA9BF930202EBDCDF93BFACFF94A" box="[1268,1293,1724,1748]" italics="true" pageId="11" pageNumber="32">z1</emphasis>
the shortest podonotal setae (as in female), opisthonotal setae slightly decreasing in length from anterior to posterior positions with 
<emphasis id="39D4EA9BF930202EB9E9F883FF2CF885" box="[208,238,1796,1819]" italics="true" pageId="11" pageNumber="32">Z5</emphasis>
the shortest (as in female): 
<emphasis id="39D4EA9BF930202EBB10F883FDFCF882" box="[553,574,1796,1820]" italics="true" pageId="11" pageNumber="32">
j 
<date id="7F1E1049F930202EBB0DF883FDFCF882" box="[564,574,1796,1820]" pageId="11" pageNumber="32" value="2016-01-18">1</date>
</emphasis>
16–18, 
<emphasis id="39D4EA9BF930202EBBA2F883FD73F882" box="[667,689,1796,1820]" italics="true" pageId="11" pageNumber="32">j3</emphasis>
38–40, 
<emphasis id="39D4EA9BF930202EBA34F883FCE1F882" box="[781,803,1796,1820]" italics="true" pageId="11" pageNumber="32">j4</emphasis>
35–44, 
<emphasis id="39D4EA9BF930202EBA46F883FC57F885" box="[895,917,1796,1819]" italics="true" pageId="11" pageNumber="32">j5</emphasis>
38–43, 
<emphasis id="39D4EA9BF930202EBACBF883FBC8F882" box="[1010,1034,1796,1820]" italics="true" pageId="11" pageNumber="32">
z 
<date id="7F1E1049F930202EBAC7F883FBC8F882" box="[1022,1034,1796,1820]" pageId="11" pageNumber="32" value="2013-01-15">1</date>
</emphasis>
13–15, 
<emphasis id="39D4EA9BF930202EBD5EF883FBBDF882" box="[1127,1151,1796,1820]" italics="true" pageId="11" pageNumber="32">z2</emphasis>
43–48, 
<emphasis id="39D4EA9BF930202EBDE5F883FB37F882" box="[1244,1269,1796,1820]" italics="true" pageId="11" pageNumber="32">z4</emphasis>
43–50, 
<emphasis id="39D4EA9BF930202EBC68F883FAA8F885" box="[1361,1386,1796,1819]" italics="true" pageId="11" pageNumber="32">z5</emphasis>
38– 43, 
<geoCoordinate id="6E94504EF930202EB9F8F8AFFF3EF8DE" box="[193,252,1831,1856]" direction="south" orientation="latitude" pageId="11" pageNumber="32" precision="925" value="-4.85">
<emphasis id="39D4EA9BF930202EB9F8F8AFFF1BF8DE" box="[193,217,1832,1856]" italics="true" pageId="11" pageNumber="32">s4</emphasis>
51
</geoCoordinate>
–58, 
<emphasis id="39D4EA9BF930202EB80DF8AFFE8CF8DE" box="[308,334,1832,1856]" italics="true" pageId="11" pageNumber="32">J1</emphasis>
36–40, 
<emphasis id="39D4EA9BF930202EB893F8AFFE06F8DE" box="[426,452,1832,1856]" italics="true" pageId="11" pageNumber="32">
J 
<date id="7F1E1049F930202EB881F8AFFE06F8DE" box="[440,452,1832,1856]" pageId="11" pageNumber="32" value="1935-02-30">2</date>
</emphasis>
30–35, 
<emphasis id="39D4EA9BF930202EBB26F8AFFDF8F8DE" box="[543,570,1832,1856]" italics="true" pageId="11" pageNumber="32">
J 
<date id="7F1E1049F930202EBB14F8AFFDF8F8DE" box="[557,570,1832,1856]" pageId="11" pageNumber="32" value="1934-03-30">3</date>
</emphasis>
30–34, 
<emphasis id="39D4EA9BF930202EBBADF8AFFD6CF8DE" box="[660,686,1832,1856]" italics="true" pageId="11" pageNumber="32">
J 
<date id="7F1E1049F930202EBB98F8AFFD6CF8DE" box="[673,686,1832,1856]" pageId="11" pageNumber="32" value="1925-04-30">4</date>
</emphasis>
25–30, 
<emphasis id="39D4EA9BF930202EBA30F8AEFCE0F8DE" box="[777,802,1833,1856]" italics="true" pageId="11" pageNumber="32">Jx</emphasis>
(about at level of 
<emphasis id="39D4EA9BF930202EBAD7F8AFFBDCF8DE" box="[1006,1054,1832,1856]" italics="true" pageId="11" pageNumber="32">idJ1</emphasis>
) 28–31, 
<emphasis id="39D4EA9BF930202EBDBBF8AEFB59F8DE" box="[1154,1179,1833,1856]" italics="true" pageId="11" pageNumber="32">Jx</emphasis>
(about at level of 
<emphasis id="39D4EA9BF930202EBC5EF8AFFA50F8DE" box="[1383,1426,1832,1856]" italics="true" pageId="11" pageNumber="32">Px3</emphasis>
) 23–28, 
<emphasis id="39D4EA9BF930202EB9D3F8CAFEC1F8FA" box="[234,259,1869,1892]" italics="true" pageId="11" pageNumber="32">Jx</emphasis>
(between 
<emphasis id="39D4EA9BF930202EB84EF8CBFE53F8FA" box="[375,401,1868,1892]" italics="true" pageId="11" pageNumber="32">J4</emphasis>
and 
<emphasis id="39D4EA9BF930202EB8FEF8CBFE23F8FD" box="[455,481,1868,1891]" italics="true" pageId="11" pageNumber="32">J5</emphasis>
) 18–23, 
<emphasis id="39D4EA9BF930202EBB7CF8CBFD9DF8FD" box="[581,607,1868,1891]" italics="true" pageId="11" pageNumber="32">
J 
<date id="7F1E1049F930202EBB6BF8CBFD9DF8FD" box="[594,607,1868,1891]" pageId="11" pageNumber="32" value="1920-05-29">5</date>
</emphasis>
20–29, 
<emphasis id="39D4EA9BF930202EBB80F8CBFD15F8FA" box="[697,727,1868,1892]" italics="true" pageId="11" pageNumber="32">Z1</emphasis>
38–45, 
<emphasis id="39D4EA9BF930202EBA09F8CBFC8FF8FA" box="[816,845,1868,1892]" italics="true" pageId="11" pageNumber="32">Z2</emphasis>
35–43, 
<emphasis id="39D4EA9BF930202EBA91F8CBFC04F8FA" box="[936,966,1868,1892]" italics="true" pageId="11" pageNumber="32">Z4</emphasis>
33–40, 
<emphasis id="39D4EA9BF930202EBD19F8CBFBFCF8FD" box="[1056,1086,1868,1891]" italics="true" pageId="11" pageNumber="32">
Z 
<date id="7F1E1049F930202EBD08F8CBFBFCF8FD" box="[1073,1086,1868,1891]" pageId="11" pageNumber="32" value="1919-05-21">5</date>
</emphasis>
19–21, 
<emphasis id="39D4EA9BF930202EBDAEF8CBFB01F8FA" box="[1175,1219,1868,1892]" italics="true" pageId="11" pageNumber="32">
Px 
<date id="7F1E1049F930202EBD8FF8CBFB01F8FA" box="[1206,1219,1868,1892]" pageId="11" pageNumber="32" value="1933-02-25">2</date>
</emphasis>
25–33, 
<emphasis id="39D4EA9BF930202EBC24F8CBFA8BF8FA" box="[1309,1353,1868,1892]" italics="true" pageId="11" pageNumber="32">
Px 
<date id="7F1E1049F930202EBC02F8CBFA8BF8FA" box="[1339,1353,1868,1892]" pageId="11" pageNumber="32" value="1925-03-30">3</date>
</emphasis>
25–30, 
<date id="7F1E1049F930202EB9AEF8F6FF31F816" box="[151,243,1904,1928]" pageId="11" pageNumber="32" value="1933-10-30">
<emphasis id="39D4EA9BF930202EB9AEF8F6FF61F816" box="[151,163,1905,1928]" italics="true" pageId="11" pageNumber="32">x</emphasis>
30–33
</date>
; with at least 21 pairs of pore-like structures, 10 on podonotum (
<emphasis id="39D4EA9BF930202EBAFDF8F7FC2DF816" box="[964,1007,1904,1928]" italics="true" pageId="11" pageNumber="32">idj1</emphasis>
, 
<emphasis id="39D4EA9BF930202EBAC4F8F7FBEAF816" box="[1021,1064,1904,1928]" italics="true" pageId="11" pageNumber="32">idj3</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD0FF8F7FBA3F816" box="[1078,1121,1904,1928]" italics="true" pageId="11" pageNumber="32">idj6</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD56F8F7FB5CF816" box="[1135,1182,1904,1928]" italics="true" pageId="11" pageNumber="32">idz3</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD92F8F7FB18F816" box="[1195,1242,1904,1928]" italics="true" pageId="11" pageNumber="32">idr5</emphasis>
, 
<emphasis id="39D4EA9BF930202EBDDEF8F7FAD4F816" box="[1255,1302,1904,1928]" italics="true" pageId="11" pageNumber="32">ids6</emphasis>
, 
<emphasis id="39D4EA9BF930202EBC1DF8F7FA97F816" box="[1316,1365,1904,1928]" italics="true" pageId="11" pageNumber="32">gdj2</emphasis>
, 
<emphasis id="39D4EA9BF930202EBC5AF8F7FA56F816" box="[1379,1428,1904,1928]" italics="true" pageId="11" pageNumber="32">gdj4</emphasis>
, 
<emphasis id="39D4EA9BF930202EB9AEF813FF0EF832" box="[151,204,1940,1964]" italics="true" pageId="11" pageNumber="32">gdz5</emphasis>
, 
<emphasis id="39D4EA9BF930202EB9E5F813FED3F832" box="[220,273,1940,1964]" italics="true" pageId="11" pageNumber="32">gdr4</emphasis>
; 
<emphasis id="39D4EA9BF930202EB81AF813FE95F832" box="[291,343,1940,1964]" italics="true" pageId="11" pageNumber="32">gdz6</emphasis>
and 
<emphasis id="39D4EA9BF930202EB8ADF813FE07F832" box="[404,453,1940,1964]" italics="true" pageId="11" pageNumber="32">idx1</emphasis>
apparently absent) and 11 on opisthonotum (
<emphasis id="39D4EA9BF930202EBAEAF813FBC4F832" box="[979,1030,1940,1964]" italics="true" pageId="11" pageNumber="32">idZ1</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD2EF813FB8AF832" box="[1047,1096,1940,1964]" italics="true" pageId="11" pageNumber="32">idJ1</emphasis>
, 
<emphasis id="39D4EA9BF930202EBD61F813FB4BF832" box="[1112,1161,1940,1964]" italics="true" pageId="11" pageNumber="32">idS1</emphasis>
, 
<emphasis id="39D4EA9BF930202EBDA2F813FB0CF832" box="[1179,1230,1940,1964]" italics="true" pageId="11" pageNumber="32">idPx</emphasis>
, 
<emphasis id="39D4EA9BF930202EBDE6F813FAD2F832" box="[1247,1296,1940,1964]" italics="true" pageId="11" pageNumber="32">idJ2</emphasis>
, 
<emphasis id="39D4EA9BF930202EBC19F813FA90F832" box="[1312,1362,1940,1964]" italics="true" pageId="11" pageNumber="32">idS2</emphasis>
, 
<emphasis id="39D4EA9BF930202EBC5AF813FA57F832" box="[1379,1429,1940,1964]" italics="true" pageId="11" pageNumber="32">idS3</emphasis>
, 
<emphasis id="39D4EA9BF930202EB9AEF83FFF05F84E" box="[151,199,1976,2000]" italics="true" pageId="11" pageNumber="32">idJ4</emphasis>
, 
<emphasis id="39D4EA9BF930202EB9E0F83FFECEF84E" box="[217,268,1976,2000]" italics="true" pageId="11" pageNumber="32">idZ4</emphasis>
, 
<emphasis id="39D4EA9BF930202EB824F83FFE8CF84E" box="[285,334,1976,2000]" italics="true" pageId="11" pageNumber="32">idJ5</emphasis>
, 
<emphasis id="39D4EA9BF930202EB866F83FFE51F84E" box="[351,403,1976,2000]" italics="true" pageId="11" pageNumber="32">idZ5</emphasis>
; 
<emphasis id="39D4EA9BF930202EB89CF83FFE1EF84E" box="[421,476,1976,2000]" italics="true" pageId="11" pageNumber="32">gdx2</emphasis>
and 
<emphasis id="39D4EA9BF930202EBB20F83FFD93F84E" box="[537,593,1976,2000]" italics="true" pageId="11" pageNumber="32">gdS3</emphasis>
apparently absent); dorsal surface ornamented throughout with irregular polygonal pattern, less marked in the central region of shield.
</paragraph>
<caption id="5FDF6601F9372029B9AEF836FBDAF87B" httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="12" pageNumber="33" targetBox="[231,1335,206,1928]" targetPageId="12">
<paragraph id="0B1F3689F9372029B9AEF836FBDAF87B" blockId="12.[151,1435,1969,2021]" pageId="12" pageNumber="33">
<emphasis id="39D4EA9BF9372029B9AEF836FE8CF859" bold="true" box="[151,334,1969,1991]" pageId="12" pageNumber="33">FIGURES 25–26</emphasis>
. 
<taxonomicName id="CCA04D0AF9372029B865F836FD95F859" box="[348,599,1969,1991]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF9372029B865F836FD95F859" box="[348,599,1969,1991]" italics="true" pageId="12" pageNumber="33">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; 
<emphasis id="39D4EA9BF9372029BC37F836FAF7F859" box="[1294,1333,1969,1991]" italics="true" pageId="12" pageNumber="33">idj1</emphasis>
, 
<emphasis id="39D4EA9BF9372029BC7AF836FAB2F859" box="[1347,1392,1969,1991]" italics="true" pageId="12" pageNumber="33">gdj2</emphasis>
and 
<emphasis id="39D4EA9BF9372029B9AEF848FF05F87B" box="[151,199,1999,2021]" italics="true" pageId="12" pageNumber="33">gdr4</emphasis>
not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26.
</paragraph>
</caption>
<caption id="5FDF6601F9362028B9AEFA1AFE28FA4C" httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="13" pageNumber="34" targetBox="[183,1388,193,1405]" targetPageId="13">
<paragraph id="0B1F3689F9362028B9AEFA1AFE28FA4C" blockId="13.[151,1436,1437,1490]" pageId="13" pageNumber="34">
<emphasis id="39D4EA9BF9362028B9AEFA1AFE8FFA2D" bold="true" box="[151,333,1437,1459]" pageId="13" pageNumber="34">FIGURES 27–28</emphasis>
. 
<taxonomicName id="CCA04D0AF9362028B862FA1AFD97FA2D" box="[347,597,1437,1459]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF9362028B862FA1AFD97FA2D" box="[347,597,1437,1459]" italics="true" pageId="13" pageNumber="34">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="0B1F3689F936202BB9FEFA79FA43FE33" blockId="13.[151,1437,1534,2027]" lastBlockId="14.[151,1437,151,2038]" lastPageId="14" lastPageNumber="35" pageId="13" pageNumber="34">
<emphasis id="39D4EA9BF9362028B9FEFA79FE47F988" box="[199,389,1534,1558]" italics="true" pageId="13" pageNumber="34">Ventral idiosoma</emphasis>
(
<figureCitation id="939B2A0CF9362028B8ADFA79FDD5F989" box="[404,535,1534,1559]" captionStart="FIGURES 27 – 28" captionStartId="13.[151,264,1437,1459]" captionTargetBox="[183,1388,193,1405]" captionTargetId="figure@13.[173,1411,181,1417]" captionTargetPageId="13" captionText="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="13" pageNumber="34">Figs 27–28</figureCitation>
). Tritosternum and presternal area as in adults. Sternal shield weakly sclerotised, with anterior margin and antero-lateral corners not well defined (continuous with presternal lines of sclerotisation), and posterior margin broadly rounded; longer than wide, 220 ± 8 (208–230) long (anterior measuring point at level of first pair of lyrifissures), 100 ± 6 (93–105) wide (widest point just anterior to insertion of 
<emphasis id="39D4EA9BF9362028BDD8F9ECFAC0F91D" box="[1249,1282,1643,1667]" italics="true" pageId="13" pageNumber="34">st3</emphasis>
); shield than narrows sharply behind two pronunced and irregular lobes, arising anteriorly to 
<emphasis id="39D4EA9BF9362028BD29F908FBF2F939" box="[1040,1072,1679,1703]" italics="true" pageId="13" pageNumber="34">st4</emphasis>
; with at least three pairs of setae simple and smooth, subequal in length (
<emphasis id="39D4EA9BF9362028BB66F934FD42F955" box="[607,640,1715,1739]" italics="true" pageId="13" pageNumber="34">
st 
<date id="7F1E1049F9362028BB4DF934FD42F955" box="[628,640,1715,1739]" pageId="13" pageNumber="34" value="1938-01-31">1</date>
</emphasis>
31–38, 
<emphasis id="39D4EA9BF9362028BBE7F934FD3CF955" box="[734,766,1715,1739]" italics="true" pageId="13" pageNumber="34">st2</emphasis>
33–38, 
<emphasis id="39D4EA9BF9362028BA64F934FCBCF955" box="[861,894,1715,1739]" italics="true" pageId="13" pageNumber="34">
st 
<date id="7F1E1049F9362028BA49F934FCBCF955" box="[880,894,1715,1739]" pageId="13" pageNumber="34" value="1938-03-30">3</date>
</emphasis>
30–38), and three pairs of lyrifissures (the first and second pairs oval, respectively behind 
<emphasis id="39D4EA9BF9362028BBA0F950FD78F971" box="[665,698,1751,1775]" italics="true" pageId="13" pageNumber="34">st1</emphasis>
and between 
<emphasis id="39D4EA9BF9362028BA5DF950FCB5F970" box="[868,887,1751,1774]" italics="true" pageId="13" pageNumber="34">st</emphasis>
2 and 
<emphasis id="39D4EA9BF9362028BAFFF950FC1BF970" box="[966,985,1751,1774]" italics="true" pageId="13" pageNumber="34">st</emphasis>
3, the third pair the smallest, circular, posterior to 
<emphasis id="39D4EA9BF9362028B824F97CFEFDF88D" box="[285,319,1787,1811]" italics="true" pageId="13" pageNumber="34">st3</emphasis>
and on lateral margins of shield); 
<emphasis id="39D4EA9BF9362028BB85F97CFD1FF88D" box="[700,733,1787,1811]" italics="true" pageId="13" pageNumber="34">st4</emphasis>
shorter than other sternal setae (18–21), generally on the tip of sternal lobes or out of shield, on soft integument (
<figureCitation id="939B2A0CF9362028BBEBF899FCEAF8A9" box="[722,808,1822,1847]" captionStart="FIGURES 27 – 28" captionStartId="13.[151,264,1437,1459]" captionTargetBox="[183,1388,193,1405]" captionTargetId="figure@13.[173,1411,181,1417]" captionTargetPageId="13" captionText="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="13" pageNumber="34">Fig. 27</figureCitation>
); ornamentation of shield not well discernible. Distinct endopodal platelets absent. Peritrematal shields absent, only with two small weakly sclerotised portions, the proximal one bearing a gland pore 
<emphasis id="39D4EA9BF9362028BB22F8E0FDF5F8E0" box="[539,567,1895,1918]" italics="true" pageId="13" pageNumber="34">gp</emphasis>
; peritreme extending anteriorly beyond coxae I, up to level between 
<emphasis id="39D4EA9BF9362028BC0CF8E0FA8CF8E1" box="[1333,1358,1895,1919]" italics="true" pageId="13" pageNumber="34">s1</emphasis>
and 
<emphasis id="39D4EA9BF9362028BCBAF8E0FA5EF8E1" box="[1411,1436,1895,1919]" italics="true" pageId="13" pageNumber="34">z1</emphasis>
(as in female), and posteriorly not behind coxa IV; a gland pore 
<emphasis id="39D4EA9BF9362028BA97F80CFC08F83C" box="[942,970,1931,1954]" italics="true" pageId="13" pageNumber="34">gp</emphasis>
posteriorly to stigma, on unsclerotised integument (other post-stigmatic pore-like structures not discernible). Setae 
<emphasis id="39D4EA9BF9362028BD35F828FBE7F859" box="[1036,1061,1967,1991]" italics="true" pageId="13" pageNumber="34">r6</emphasis>
as in adults. Exopodal platelets absent; glands 
<emphasis id="39D4EA9BF9362028B805F854FEA6F875" box="[316,356,2003,2027]" italics="true" pageId="13" pageNumber="34">gv2</emphasis>
not discernible. Anal shield subtriangular, longer than wide, 93 ± 2 (89–96) long, 77 ± 4 (73–83) wide (at level of pores 
<emphasis id="39D4EA9BF935202BB8ACFF1FFE7FFF2E" box="[405,445,152,176]" italics="true" pageId="14" pageNumber="35">gv3</emphasis>
); anal opening situated about at mid-level of shield; para-anal setae longer than unpaired post-anal seta (as in adults); pores 
<emphasis id="39D4EA9BF935202BBB15FF3BFD91FF4A" box="[556,595,188,212]" italics="true" pageId="14" pageNumber="35">gv3</emphasis>
not well discernible (often circumscribed by integument sclerotised in a spiral fashion), on lateral margins of anal shield, at level of anal opening as in adults; cribrum and sculptural lines as in female. Opisthogastric unsclerotised cuticle with 16 setae (including 
<emphasis id="39D4EA9BF935202BBAE5FE83FC3EFE85" box="[988,1020,260,283]" italics="true" pageId="14" pageNumber="35">st5</emphasis>
), in addition to a variable number of setae 
<emphasis id="39D4EA9BF935202BB9E0FEAEFF3CFEDE" box="[217,254,297,320]" italics="true" pageId="14" pageNumber="35">UR</emphasis>
, and 5 pairs of discernible circular lyrifissures 
<emphasis id="39D4EA9BF935202BBA1CFEAFFC85FEA1" box="[805,839,296,319]" italics="true" pageId="14" pageNumber="35">ivp</emphasis>
; position of 
<emphasis id="39D4EA9BF935202BBAE6FEAFFBC7FEDE" box="[991,1029,296,320]" italics="true" pageId="14" pageNumber="35">Jv1</emphasis>
and 
<emphasis id="39D4EA9BF935202BBD7BFEAFFBABFEDE" box="[1090,1129,296,320]" italics="true" pageId="14" pageNumber="35">Jv2</emphasis>
setae as in male; all setae smooth and needle-like, subequal in length (
<emphasis id="39D4EA9BF935202BBBB6FECBFD72FEFD" box="[655,688,332,355]" italics="true" pageId="14" pageNumber="35">
st 
<date id="7F1E1049F935202BBB9BFECBFD72FEFD" box="[674,688,332,355]" pageId="14" pageNumber="35" value="1919-05-23">5</date>
</emphasis>
19–23, 
<emphasis id="39D4EA9BF935202BBA37FECBFCF6FEFA" box="[782,820,332,356]" italics="true" pageId="14" pageNumber="35">
Jv 
<date id="7F1E1049F935202BBA10FECBFCF6FEFA" box="[809,820,332,356]" pageId="14" pageNumber="35" value="2014-01-18">1</date>
</emphasis>
14–18, 
<emphasis id="39D4EA9BF935202BBAABFECBFC7BFEFA" box="[914,953,332,356]" italics="true" pageId="14" pageNumber="35">
Jv 
<date id="7F1E1049F935202BBA95FECBFC7BFEFA" box="[940,953,332,356]" pageId="14" pageNumber="35" value="1918-02-20">2</date>
</emphasis>
18–20, 
<emphasis id="39D4EA9BF935202BBD2EFECBFBFCFEFA" box="[1047,1086,332,356]" italics="true" pageId="14" pageNumber="35">
Jv 
<date id="7F1E1049F935202BBD08FECBFBFCFEFA" box="[1073,1086,332,356]" pageId="14" pageNumber="35" value="1918-03-19">3</date>
</emphasis>
18–19, 
<emphasis id="39D4EA9BF935202BBDA2FECBFB01FEFA" box="[1179,1219,332,356]" italics="true" pageId="14" pageNumber="35">
Jv 
<date id="7F1E1049F935202BBD8CFECBFB01FEFA" box="[1205,1219,332,356]" pageId="14" pageNumber="35" value="2014-04-16">4</date>
</emphasis>
14–16, 
<emphasis id="39D4EA9BF935202BBC19FECBFA85FEFD" box="[1312,1351,332,355]" italics="true" pageId="14" pageNumber="35">
Jv 
<date id="7F1E1049F935202BBC00FECBFA85FEFD" box="[1337,1351,332,355]" pageId="14" pageNumber="35" value="2013-05-15">5</date>
</emphasis>
13–15, 
<emphasis id="39D4EA9BF935202BB9AEFEF7FF03FE16" box="[151,193,368,392]" italics="true" pageId="14" pageNumber="35">
Zv 
<date id="7F1E1049F935202BB98FFEF7FF03FE16" box="[182,193,368,392]" pageId="14" pageNumber="35" value="2013-01-15">1</date>
</emphasis>
13–15, 
<emphasis id="39D4EA9BF935202BB825FEF7FE85FE16" box="[284,327,368,392]" italics="true" pageId="14" pageNumber="35">
Zv 
<date id="7F1E1049F935202BB803FEF7FE85FE16" box="[314,327,368,392]" pageId="14" pageNumber="35" value="2013-02-16">2</date>
</emphasis>
13–16, 
<emphasis id="39D4EA9BF935202BB89BFEF7FE0EFE16" box="[418,460,368,392]" italics="true" pageId="14" pageNumber="35">
Zv 
<date id="7F1E1049F935202BB886FEF7FE0EFE16" box="[447,460,368,392]" pageId="14" pageNumber="35" value="2013-03-16">3</date>
</emphasis>
13–16, 
<emphasis id="39D4EA9BF935202BBB1EFEF7FD93FE16" box="[551,593,368,392]" italics="true" pageId="14" pageNumber="35">
Zv 
<date id="7F1E1049F935202BBB7DFEF7FD93FE16" box="[580,593,368,392]" pageId="14" pageNumber="35" value="2013-04-14">4</date>
</emphasis>
13–14, 
<emphasis id="39D4EA9BF935202BBB94FEF7FD1AFE19" box="[685,728,368,391]" italics="true" pageId="14" pageNumber="35">
Zv 
<date id="7F1E1049F935202BBBF3FEF7FD1AFE19" box="[714,728,368,391]" pageId="14" pageNumber="35" value="2014-11-05">5</date>
</emphasis>
11–14, 
<emphasis id="39D4EA9BF935202BBA0BFEF7FCBCFE16" box="[818,894,368,392]" italics="true" pageId="14" pageNumber="35">
R1–R 
<date id="7F1E1049F935202BBA49FEF7FCBCFE16" box="[880,894,368,392]" pageId="14" pageNumber="35" value="2016-10-05">5</date>
</emphasis>
10–16, 
<emphasis id="39D4EA9BF935202BBAE3FEF6FC3DFE16" box="[986,1023,369,392]" italics="true" pageId="14" pageNumber="35">UR</emphasis>
11–14); metapodal platelets present, proportionally smaller than in female, elongate and oriented at 45°; parapodal elements or other platelets absent.
</paragraph>
<paragraph id="0B1F3689F935202BB9FEFE3FFF29FD86" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
<taxonomicName id="CCA04D0AF935202BB9FEFE3FFE94FE4E" box="[199,342,440,464]" class="Arachnida" family="Podocinidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF935202BB9FEFE3FFE94FE4E" box="[199,342,440,464]" italics="true" pageId="14" pageNumber="35">Gnathosoma</emphasis>
</taxonomicName>
. All gnathosomal structures as in female, except for palpal trochanter and femur, distinctly shorter than in female; epistome generally not discernible; all morphological details of the cheliceral dentition exactly as in female.
</paragraph>
<paragraph id="0B1F3689F935202BB9FEFDA2FC22FDA3" blockId="14.[151,1437,151,2038]" box="[199,992,548,573]" pageId="14" pageNumber="35">
<emphasis id="39D4EA9BF935202BB9FEFDA2FF3EFDA2" box="[199,252,549,572]" italics="true" pageId="14" pageNumber="35">Legs</emphasis>
. Leg II stouter than others (as in adults). Chaetotaxy as in female.
</paragraph>
</subSubSection>
<subSubSection id="43BA6502F9352035B9FEFDC0FCB8FD36" lastPageId="16" lastPageNumber="37" pageId="14" pageNumber="35" type="discussion">
<paragraph id="0B1F3689F935202BB9FEFDC0FF3CFCC3" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
<emphasis id="39D4EA9BF935202BB9FEFDC0FEF9FDFE" bold="true" box="[199,315,583,608]" pageId="14" pageNumber="35">Remarks.</emphasis>
In adults of 
<taxonomicName id="CCA04D0AF935202BB8F0FDCFFD9AFDFE" box="[457,600,584,608]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BB8F0FDCFFE1EFDC1" box="[457,476,584,607]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB8D3FDCFFD9AFDFE" box="[490,600,584,608]" italics="true" pageId="14" pageNumber="35">lignicolus</emphasis>
</taxonomicName>
the dorsal shield is highly convex and laterally expanded, often extending onto the ventral surface in mounted specimens. This prevents the observation of marginal and submarginal morphological details, which in this species include some setae and pore-like structures very close to lateral margins of shield (
<figureCitation id="939B2A0CF935202BB848FD33FE7BFD52" box="[369,441,692,717]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1778,1800]" captionTargetBox="[164,1424,214,1739]" captionTargetId="figure@5.[151,1436,193,1756]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4 – 7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292760/files/figure.png" pageId="14" pageNumber="35">Fig. 3</figureCitation>
). In deutonymphs, the dorsal shield is significantly less convex and expanded than in adults (as shown by the position of the setae 
<emphasis id="39D4EA9BF935202BBBB4FD5EFD5AFD6E" box="[653,664,729,752]" italics="true" pageId="14" pageNumber="35">r</emphasis>
and 
<emphasis id="39D4EA9BF935202BBBF7FD5FFD1EFD71" box="[718,732,728,751]" italics="true" pageId="14" pageNumber="35">S</emphasis>
, on the lateral margin of the shield or on unsclerotised cuticle) and very marginal structures of the adults are absent (with the exception of the setae 
<date id="7F1E1049F935202BBDB2FD7BFB66FC8A" box="[1163,1188,764,788]" pageId="14" pageNumber="35">
<emphasis id="39D4EA9BF935202BBDB2FD7BFB66FC8A" box="[1163,1188,764,788]" italics="true" pageId="14" pageNumber="35">x1</emphasis>
</date>
; 
<figureCitation id="939B2A0CF935202BBD81FD7BFAD0FC8B" box="[1208,1298,764,789]" captionStart="FIGURES 25 – 26" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[231,1335,206,1928]" captionTargetId="figure@12.[225,1345,193,1942]" captionTargetPageId="12" captionText="FIGURES 25 – 26. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph. 25. Dorsal shield (partial view; idj 1, gdj 2 and gdr 4 not illustrated); 26. Dorsal setae. Scale bars: 100 µm for Fig. 25; 50 µm for Fig. 26." httpUri="https://zenodo.org/record/292766/files/figure.png" pageId="14" pageNumber="35">Figs 25</figureCitation>
, 
<figureCitation id="939B2A0CF935202BBC1BFD7BFA80FC8A" box="[1314,1346,764,788]" captionStart="FIGURES 27 – 28" captionStartId="13.[151,264,1437,1459]" captionTargetBox="[183,1388,193,1405]" captionTargetId="figure@13.[173,1411,181,1417]" captionTargetPageId="13" captionText="FIGURES 27 – 28. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), deutonymph, idiosoma (partial views). 27. Ventral view; 28. Lateral view. Scale bars: 100 µm." httpUri="https://zenodo.org/record/292767/files/figure.png" pageId="14" pageNumber="35">28</figureCitation>
). These observations suggest that the origin of these structures might be correlated with the lateral expansion of the shield in adults.
</paragraph>
<paragraph id="0B1F3689F935202BB9FEFCE0FC14FBC6" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
Based on our observations mostly in adult females, and in contrast to what has been reported in literature for other laelapid mites, in 
<taxonomicName id="CCA04D0AF935202BB8A4FC0BFDE9FC3A" box="[413,555,908,932]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BB8A4FC0BFE72FC3D" box="[413,432,908,931]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB884FC0BFDE9FC3A" box="[445,555,908,932]" italics="true" pageId="14" pageNumber="35">lignicolus</emphasis>
</taxonomicName>
the correct chaetotaxy for femur I seems be 2 3/1 2/3 2 (three setae 
<emphasis id="39D4EA9BF935202BBC19FC0BFAFEFC3A" box="[1312,1340,908,932]" italics="true" pageId="14" pageNumber="35">ad</emphasis>
and two setae 
<emphasis id="39D4EA9BF935202BB9EEFC37FF31FC56" box="[215,243,944,968]" italics="true" pageId="14" pageNumber="35">pd</emphasis>
). In fact, in this species the most proximal dorsal seta of femur I is inserted more or less in the middle of the segment, but in most specimens it appears shifted more toward the anterior (2 3/1 2/3 2) than the posterior margin (2 2/1 3/3 2) of the segment. Likewise, in 
<taxonomicName id="CCA04D0AF935202BBBFDFC7FFC90FB8E" box="[708,850,1016,1040]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BBBFDFC7FFD15FB91" box="[708,727,1016,1039]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BBBDDFC7FFC90FB8E" box="[740,850,1016,1040]" italics="true" pageId="14" pageNumber="35">lignicolus</emphasis>
</taxonomicName>
the more proximal ventral seta of trochanter II (1 0/ 1 0/2 1, one seta 
<emphasis id="39D4EA9BF935202BB868FB9AFEA9FBAA" box="[337,363,1053,1076]" italics="true" pageId="14" pageNumber="35">av</emphasis>
and two setae 
<emphasis id="39D4EA9BF935202BBB29FB9AFDE9FBAA" box="[528,555,1053,1076]" italics="true" pageId="14" pageNumber="35">pv</emphasis>
) and the only ventral seta of femur IV (1 2/0 1/1 1, one seta 
<emphasis id="39D4EA9BF935202BBDF0FB9BFB27FBAA" box="[1225,1253,1052,1076]" italics="true" pageId="14" pageNumber="35">pd</emphasis>
; 
<figureCitation id="939B2A0CF935202BBDCAFB9BFA80FBAA" box="[1267,1346,1052,1077]" captionStart="FIGURES 22 – 24" captionStartId="10.[151,264,1580,1602]" captionTargetBox="[174,1419,211,1540]" captionTargetId="figure@10.[151,1436,193,1558]" captionTargetPageId="10" captionText="FIGURES 22 – 24. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 22. Female, leg IV, femur, genu and tibia (dorsal view); 23. Male, leg II, femur, genu, tibia and tarsus (lateral view); 24. Female, leg II, femur, genu, tibia and tarsus (lateral view). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292765/files/figure.png" pageId="14" pageNumber="35">Fig. 22</figureCitation>
) appear closer to the posterior than the anterior margin of the relative leg segment.
</paragraph>
<paragraph id="0B1F3689F935202BB9FEFBE3FE16F90E" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
Males and females of 
<taxonomicName id="CCA04D0AF935202BB8F0FBE3FD98FBE2" box="[457,602,1124,1148]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BB8F0FBE3FE1EFBE5" box="[457,476,1124,1147]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB8D5FBE3FD98FBE2" box="[492,602,1124,1148]" italics="true" pageId="14" pageNumber="35">lignicolus</emphasis>
</taxonomicName>
show differences at species-level for some secondary sexual characters. Among others (i.e. different shape and chaetotaxy of leg II), of special interest are some features of internal malae and opisthonotal setation. Based on descriptions from literature and direct examination of some European species [i.e. representatives of the 
<taxonomicName id="CCA04D0AF935202BB8FDFB48FD83FB78" box="[452,577,1230,1254]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="cuneifer">
<emphasis id="39D4EA9BF935202BB8FDFB48FE15FB78" box="[452,471,1231,1254]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB8DFFB49FD83FB78" box="[486,577,1230,1254]" italics="true" pageId="14" pageNumber="35">cuneifer</emphasis>
</taxonomicName>
-group and 
<taxonomicName id="CCA04D0AF935202BBBFCFB48FBE0FB79" authority="Masan, 1992" authorityName="Masan" authorityYear="1992" box="[709,1058,1230,1255]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="39D4EA9BF935202BBBFCFB48FD1AFB78" box="[709,728,1231,1254]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BBBDEFB49FCB0FB78" box="[743,882,1230,1254]" italics="true" pageId="14" pageNumber="35">tuberculatus</emphasis>
(
<bibRefCitation id="6F314B78F935202BBAB8FB49FBD9FB79" author="Masan" box="[897,1051,1230,1255]" pageId="14" pageNumber="35" refString="Masan, P. (1992) A new species of the gamasid mite from the genus Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes) from Czechoslovakia. Biologia (Bratislava), 47, 383 - 388." type="journal article" year="1992">Mašán, 1992</bibRefCitation>
)
</taxonomicName>
], in 
<taxonomicName id="CCA04D0AF935202BBD6DFB49FB31FB78" box="[1108,1267,1230,1254]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="39D4EA9BF935202BBD6DFB49FB31FB78" box="[1108,1267,1230,1254]" italics="true" pageId="14" pageNumber="35">Cosmolaelaps</emphasis>
</taxonomicName>
species whose females display external lobes of the internal malae with well-developed fimbriae, these structures are generally equally developed also in males. Interestingly, the females of 
<taxonomicName id="CCA04D0AF935202BBA75FA91FC19FAB3" box="[844,987,1301,1325]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BBA75FA91FC9DFAB3" box="[844,863,1302,1325]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BBA54FA92FC19FAB3" box="[877,987,1301,1325]" italics="true" pageId="14" pageNumber="35">lignicolus</emphasis>
</taxonomicName>
have external lobes with fimbriae welldeveloped, whereas the males of the same species have external lobes merely fringed, with true fimbriation absent (
<figureCitation id="939B2A0CF935202BB9A6FADBFEDDFAEB" box="[159,287,1372,1397]" captionStart="FIGURES 16 – 21" captionStartId="9.[151,264,1586,1608]" captionTargetBox="[164,1418,210,1548]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 16 – 21. Cosmolaelaps lignicolus (G. &amp; R. Canestrini). 16. Female, epistome, four different specimens; 17. Female, chelicera; 18. Female, gnathosoma (ventral view; labrum not illustrated); 19. Male, epistome; 20. Male, chelicera; 21. Male, corniculi and internal malae (dorsal view; labrum not illustrated). Scale bars: 100 µm for Fig. 17; 50 µm for Figs 16, 18 – 21." httpUri="https://zenodo.org/record/292881/files/figure.png" pageId="14" pageNumber="35">Figs 18, 21</figureCitation>
). We have also observed fimbriae well-developed and fimbriae absent, respectively in female and male specimens of the 
<taxonomicName id="CCA04D0AF935202BB860FA07FE0AFA06" box="[345,456,1408,1432]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="vacuus">
<emphasis id="39D4EA9BF935202BB860FA07FEAEFA09" box="[345,364,1408,1431]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB843FA06FE0AFA06" box="[378,456,1409,1432]" italics="true" pageId="14" pageNumber="35">vacuus</emphasis>
</taxonomicName>
-group collected from 
<collectingCountry id="73B77619F935202BBBFBFAF8FD37FA06" box="[706,757,1407,1432]" name="Italy" pageId="14" pageNumber="35">Italy</collectingCountry>
[see also figures 200-HW and 200-HM in 
<bibRefCitation id="6F314B78F935202BBDECFAF8FF23FA22" pageId="14" pageNumber="35" refString="Hirschmann, W., Bernhard, F., Greim, E. &amp; Gotz, H. (1969) Gangsystematik der Parasitiformes, Teile 75. Zwanzig neue Hypoaspis - Arten. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 12, 133 - 141." type="journal article">
Hirschmann 
<emphasis id="39D4EA9BF935202BBC5DFA06FA57FA06" box="[1380,1429,1408,1432]" italics="true" pageId="14" pageNumber="35">et al</emphasis>
. (1969)
</bibRefCitation>
for 
<taxonomicName id="CCA04D0AF935202BB82BFA24FD91FA22" authority="Bernhard" authorityName="Bernhard" box="[274,595,1443,1468]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="serratosimilis">
<emphasis id="39D4EA9BF935202BB82BFA24FEE7FA24" box="[274,293,1443,1466]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BB80AFA24FE0FFA25" box="[307,461,1443,1467]" italics="true" pageId="14" pageNumber="35">serratosimilis</emphasis>
(Bernhard)
</taxonomicName>
, a junior synonym of 
<taxonomicName id="CCA04D0AF935202BBA72FA24FC78FA25" box="[843,954,1443,1467]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="vacuus">
<emphasis id="39D4EA9BF935202BBA72FA24FC9CFA24" box="[843,862,1443,1466]" italics="true" pageId="14" pageNumber="35">C</emphasis>
. 
<emphasis id="39D4EA9BF935202BBA54FA23FC78FA25" box="[877,954,1444,1467]" italics="true" pageId="14" pageNumber="35">vacuus</emphasis>
</taxonomicName>
, syn. by 
<bibRefCitation id="6F314B78F935202BBD1AFA23FB2CFA25" author="Bregetova" box="[1059,1262,1443,1468]" pageId="14" pageNumber="35" refString="Bregetova, N. G. (1977) Laelaptidae. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.), Key to the Soil - Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 483 - 554. [in Russian]" type="book chapter" year="1977">Bregetova (1977)</bibRefCitation>
]. The existence of this secondary sexual dimorphism might be due to specific differences in the biology of the sexes but, in the absence of further information, it is difficult to propose a hypothesis. In regard to the chaetotaxy of the opisthonotum, in the females setae 
<emphasis id="39D4EA9BF935202BBB1EF989FDA8F9B8" box="[551,618,1550,1574]" italics="true" pageId="14" pageNumber="35">J2–J4</emphasis>
seem be unstable in position and presence (
<figureCitation id="939B2A0CF935202BBD66F98AFB64F9B8" box="[1119,1190,1549,1574]" captionStart="FIGURES 1 – 2" captionStartId="4.[151,264,1939,1961]" captionTargetBox="[155,1421,421,1907]" captionTargetId="figure@4.[150,1436,409,1918]" captionTargetPageId="4" captionText="FIGURES 1 – 2. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), adult female. 1. Habitus of the dorsal shield (slightly pressed specimen; setae x 2 and x 3, and most postero-marginal pore-like structures of the dorsal shield not visible in this specimen); 2. Detail of seta Z 5 and lyrifissure idZ 5 (ventral view). Scale bars: 100 µm for Fig. 1; 50 µm for Fig. 2." httpUri="https://zenodo.org/record/292759/files/figure.png" pageId="14" pageNumber="35">Figs 1</figureCitation>
, 
<figureCitation id="939B2A0CF935202BBD8FF989FB21F9B8" box="[1206,1251,1550,1574]" captionStart="FIGURES 3 – 7" captionStartId="5.[151,264,1778,1800]" captionTargetBox="[164,1424,214,1739]" captionTargetId="figure@5.[151,1436,193,1756]" captionTargetPageId="5" captionText="FIGURES 3 – 7. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), female. 3. Section of dorsal shield; 4 – 7. Opisthonotum, four different specimens (marginal setae and pore-like structures not illustrated). Scale bars: 100 µm." httpUri="https://zenodo.org/record/292760/files/figure.png" pageId="14" pageNumber="35">4–7</figureCitation>
), whereas in the males, a slightly narrower idiosoma and a reduction of the distance between the setae and pore-like structures of some series (i.e. clearly discernible in the case of 
<emphasis id="39D4EA9BF935202BBBF3F9D2FCC9F9F3" box="[714,779,1621,1645]" italics="true" pageId="14" pageNumber="35">idPx2</emphasis>
, 
<emphasis id="39D4EA9BF935202BBA23F9D2FCF5F9F3" box="[794,823,1621,1645]" italics="true" pageId="14" pageNumber="35">Z4</emphasis>
and 
<emphasis id="39D4EA9BF935202BBA56F9D2FC61F9F3" box="[879,931,1621,1645]" italics="true" pageId="14" pageNumber="35">idZ4</emphasis>
) are associated with the apparent absence of the setae 
<emphasis id="39D4EA9BF935202BB9C6F9FFFEA1F90E" box="[255,355,1656,1680]" italics="true" pageId="14" pageNumber="35">Px1–Px3</emphasis>
(
<figureCitation id="939B2A0CF935202BB84AF9F0FE04F90E" box="[371,454,1655,1680]" captionStart="FIGURE 13 – 15" captionStartId="8.[151,250,1851,1873]" captionTargetBox="[161,1420,221,1818]" captionTargetId="figure@8.[151,1436,193,1829]" captionTargetPageId="8" captionText="FIGURE 13 – 15. Cosmolaelaps lignicolus (G. &amp; R. Canestrini), male. 13. Opisthonotum (slightly pressed specimen; marginal setae and pore-like structures not illustrated); 14. Section of dorsal and peritrematal shields; 15. Ventral idiosoma (partial view). Scale bars = 100 µm." httpUri="https://zenodo.org/record/292763/files/figure.png" pageId="14" pageNumber="35">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="0B1F3689F935202BB9FEF91CFD57F8DC" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
Finally, it is interesting to note that in deutonymphal stage described here, the fimbriation of internal malae is developed as in females, and the opisthonotum is slightly hypertrichous (with a greater number of setae 
<emphasis id="39D4EA9BF935202BBC17F947FA85F949" box="[1326,1351,1728,1751]" italics="true" pageId="14" pageNumber="35">Jx</emphasis>
than in adults and the presence of supernumerary setae 
<emphasis id="39D4EA9BF935202BBB88F964FD7FF964" box="[689,701,1763,1786]" italics="true" pageId="14" pageNumber="35">x</emphasis>
). In one deutonymph has been observed a dorsal seta with a well distinct bulging base (
<figureCitation id="939B2A0CF935202BB8A1F882FE2CF880" box="[408,494,1797,1822]" captionStart="FIGURES 29 – 34" captionStartId="15.[151,264,1915,1937]" captionTargetBox="[251,1331,223,1864]" captionTargetId="figure@15.[234,1351,205,1881]" captionTargetPageId="15" captionText="FIGURES 29 – 34. Cosmolaelaps lignicolus (G. &amp; R. Canestrini) and C. vacuus (Michael). 29 – 30. C. lignicolus, female, dorsal setae; 31. C. lignicolus, male, leg II, femur and genu (lateral view); 32 – 33. C. lignicolus, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. C. vacuus, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29 – 30 and 32 – 33 not to scale (see text for length of setae)." httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="14" pageNumber="35">Fig. 33</figureCitation>
). However, it is not possible conclude whether this seta should be considered an aberrant seta or a rare attribute of the species.
</paragraph>
<paragraph id="0B1F3689F935202BB9FEF8CBFC5FF86B" blockId="14.[151,1437,151,2038]" pageId="14" pageNumber="35">
<emphasis id="39D4EA9BF935202BB9FEF8CBFD94F8FB" bold="true" box="[199,598,1868,1893]" pageId="14" pageNumber="35">Ecological notes and distribution.</emphasis>
The previous records of 
<taxonomicName id="CCA04D0AF935202BBA57F8CAFB40F8FB" box="[878,1154,1869,1893]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF935202BBA57F8CAFB40F8FB" box="[878,1154,1869,1893]" italics="true" pageId="14" pageNumber="35">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
documented its presence in three localities of peninsular 
<collectingCountry id="73B77619F935202BBB3FF8F7FDF8F817" box="[518,570,1904,1929]" name="Italy" pageId="14" pageNumber="35">Italy</collectingCountry>
, where this species was respectively collected from a soaked tree (G. &amp; R. Canestrini 1882), under stones, from decomposing organic substrate (
<bibRefCitation id="6F314B78F935202BBAF4F813FB4BF833" author="Canestrini" box="[973,1161,1940,1965]" pageId="14" pageNumber="35" refString="Canestrini, G. (1885) Prospetto dell'Acarofauna Italiana, Volume 1, Famiglie: Oribatini, Gamasini, Hoplopini, Trombidini, Rhyncholophini. Proserpini, Padova, 163 pp., pls. I - IX." type="book" year="1885">Canestrini 1885</bibRefCitation>
), and from unspecified habitat (this latter record is reported in 
<bibRefCitation id="6F314B78F935202BBB72F83FFC5FF84F" author="Castagnoli" box="[587,925,1976,2001]" pageId="14" pageNumber="35" refString="Castagnoli, M. &amp; Pegazzano, F. (1985) Catalogue of the Berlese Acaroteca. Istituto Sperimentale per la Zoologia Agraria, Firenze, 490 pp." type="book" year="1985">Castagnoli &amp; Pegazzano 1985</bibRefCitation>
and it is relative to some specimens collected by the Italian entomologist Agostino Dodero near the city of Genoa).
</paragraph>
<caption id="5FDF6601F934202AB9AEF8FCFCAEF873" httpUri="https://zenodo.org/record/292768/files/figure.png" pageId="15" pageNumber="36" targetBox="[251,1331,223,1864]" targetPageId="15">
<paragraph id="0B1F3689F934202AB9AEF8FCFCAEF873" blockId="15.[151,1436,1915,2029]" pageId="15" pageNumber="36">
<emphasis id="39D4EA9BF934202AB9AEF8FCFE91F80F" bold="true" box="[151,339,1915,1937]" pageId="15" pageNumber="36">FIGURES 29–34.</emphasis>
<taxonomicName id="CCA04D0AF934202AB863F8FCFD91F80F" box="[346,595,1915,1937]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF934202AB863F8FCFD91F80F" box="[346,595,1915,1937]" italics="true" pageId="15" pageNumber="36">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
(G. &amp; R. Canestrini) and 
<taxonomicName id="CCA04D0AF934202ABA6EF8FBFBEAF80F" authority="Michael" authorityName="Michael" box="[855,1064,1915,1937]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="vacuus">
<emphasis id="39D4EA9BF934202ABA6EF8FBFCAAF80F" box="[855,872,1916,1937]" italics="true" pageId="15" pageNumber="36">C</emphasis>
. 
<emphasis id="39D4EA9BF934202ABA4CF8FBFC79F80F" box="[885,955,1916,1937]" italics="true" pageId="15" pageNumber="36">vacuus</emphasis>
(Michael)
</taxonomicName>
. 29–30. 
<taxonomicName id="CCA04D0AF934202ABD47F8FBFAC2F80F" box="[1150,1280,1915,1937]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF934202ABD47F8FBFB4DF80F" box="[1150,1167,1916,1937]" italics="true" pageId="15" pageNumber="36">C</emphasis>
. 
<emphasis id="39D4EA9BF934202ABDA5F8FCFAC2F80F" box="[1180,1280,1915,1937]" italics="true" pageId="15" pageNumber="36">lignicolus</emphasis>
</taxonomicName>
, female, dorsal setae; 31. 
<taxonomicName id="CCA04D0AF934202AB83BF81CFE44F82E" box="[258,390,1946,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF934202AB83BF81CFED1F82E" box="[258,275,1947,1968]" italics="true" pageId="15" pageNumber="36">C</emphasis>
. 
<emphasis id="39D4EA9BF934202AB81BF81DFE44F82E" box="[290,390,1946,1968]" italics="true" pageId="15" pageNumber="36">lignicolus</emphasis>
</taxonomicName>
, male, leg II, femur and genu (lateral view); 32–33. 
<taxonomicName id="CCA04D0AF934202ABA8DF81CFBFBF82E" box="[948,1081,1946,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF934202ABA8DF81CFC07F82E" box="[948,965,1947,1968]" italics="true" pageId="15" pageNumber="36">C</emphasis>
. 
<emphasis id="39D4EA9BF934202ABAECF81DFBFBF82E" box="[981,1081,1946,1968]" italics="true" pageId="15" pageNumber="36">lignicolus</emphasis>
</taxonomicName>
, deutonymph, dorsal setae (arrow indicates a seta with a bulging base); 34. 
<taxonomicName id="CCA04D0AF934202ABB05F83DFD63F851" box="[572,673,1978,1999]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="vacuus">
<emphasis id="39D4EA9BF934202ABB05F83DFD8FF851" box="[572,589,1978,1999]" italics="true" pageId="15" pageNumber="36">C</emphasis>
. 
<emphasis id="39D4EA9BF934202ABB62F83DFD63F851" box="[603,673,1978,1999]" italics="true" pageId="15" pageNumber="36">vacuus</emphasis>
</taxonomicName>
, male, leg II, femur and genu (lateral view). Scale bars: 50 µm for Figs 31 and 34; Figs 29–30 and 32–33 not to scale (see text for length of setae).
</paragraph>
</caption>
<paragraph id="0B1F3689F92B2035B9FEFF10FCB8FD36" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="37">
All the new specimens of 
<taxonomicName id="CCA04D0AF92B2035B8CAFF1FFD41FF2E" box="[499,643,152,176]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF92B2035B8CAFF1FFDC4FF31" box="[499,518,152,175]" italics="true" pageId="16" pageNumber="37">C</emphasis>
. 
<emphasis id="39D4EA9BF92B2035BB2CFF1FFD41FF2E" box="[533,643,152,176]" italics="true" pageId="16" pageNumber="37">lignicolus</emphasis>
</taxonomicName>
have been exclusively collected from anthills of 
<taxonomicName id="CCA04D0AF92B2035BD81FF1FFA57FF31" box="[1208,1429,152,175]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="emarginatus">
<emphasis id="39D4EA9BF92B2035BD81FF1FFA57FF31" box="[1208,1429,152,175]" italics="true" pageId="16" pageNumber="37">Lasius emarginatus</emphasis>
</taxonomicName>
. This ant is a very thermophilous Euro-Caucasian, Mediterranean and sub-Mediterranean species, inhabiting lowland and submontane areas. Its nests can be found in warm and rocky natural habitats, as well as in rural and urban disturbed areas (
<bibRefCitation id="6F314B78F92B2035B8A7FE83FDF0FE82" author="Seifert" box="[414,562,260,285]" pageId="16" pageNumber="37" refString="Seifert, B. (2007) Die Ameisen Mittel - und Nordeuropas. Iutra Verlags- und Vertriebsgesellschaft, Tauer, 368 pp." type="book" year="2007">Seifert 2007</bibRefCitation>
). The senior author has observed 
<taxonomicName id="CCA04D0AF92B2035BA84FE83FB17FE82" box="[957,1237,260,284]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF92B2035BA84FE83FB17FE82" box="[957,1237,260,284]" italics="true" pageId="16" pageNumber="37">Cosmolaelaps lignicolus</emphasis>
</taxonomicName>
in anthills under stones or most often within logs and stumps, mainly in various forest habitats, where the most abundant samples were also obtained. The species was also collected from nests in open areas, albeit with series of fewer specimens. The relationship between this laelapid mite and its ant host is demonstrated by further observations: (1), 
<taxonomicName id="CCA04D0AF92B2035BCBBFEF7FEC4FE32" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF92B2035BCBBFEF7FA57FE19" box="[1410,1429,368,391]" italics="true" pageId="16" pageNumber="37">C</emphasis>
. 
<emphasis id="39D4EA9BF92B2035B9AEFE13FEC4FE32" box="[151,262,404,428]" italics="true" pageId="16" pageNumber="37">lignicolus</emphasis>
</taxonomicName>
has never been found by the senior author in many samples obtained from rotting logs or other substrates not colonised by ant species; (2), most specimens have been observed in deeper sections of the anthills, where the cocoons of the ant host were present; in cases where the nests were not dissected in depth, individuals of 
<taxonomicName id="CCA04D0AF92B2035BCBBFE5BFEC4FD86" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF92B2035BCBBFE5BFA57FE6D" box="[1410,1429,476,499]" italics="true" pageId="16" pageNumber="37">C</emphasis>
. 
<emphasis id="39D4EA9BF92B2035B9AEFD87FEC4FD86" box="[151,262,512,536]" italics="true" pageId="16" pageNumber="37">lignicolus</emphasis>
</taxonomicName>
have been observed to come out occasionally and slowly from the crevices of inner and more humid parts of the anthill; (3), deutonymphs have also been found in anthills, suggesting that a large part of the life cycle is accomplished in this habitat; (4), 
<taxonomicName id="CCA04D0AF92B2035BB03FDCFFD09FDFE" box="[570,715,584,608]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lignicolus">
<emphasis id="39D4EA9BF92B2035BB03FDCFFD8FFDC1" box="[570,589,584,607]" italics="true" pageId="16" pageNumber="37">C</emphasis>
. 
<emphasis id="39D4EA9BF92B2035BB64FDCFFD09FDFE" box="[605,715,584,608]" italics="true" pageId="16" pageNumber="37">lignicolus</emphasis>
</taxonomicName>
has not been found in anthills of some syntopic species of 
<taxonomicName id="CCA04D0AF92B2035BCBCFDCEFEE0FD1A" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="emarginatus">
<emphasis id="39D4EA9BF92B2035BCBCFDCEFA56FDFE" box="[1413,1428,585,608]" italics="true" pageId="16" pageNumber="37">L</emphasis>
. 
<emphasis id="39D4EA9BF92B2035B9AEFDEAFEE0FD1A" box="[151,290,621,644]" italics="true" pageId="16" pageNumber="37">emarginatus</emphasis>
</taxonomicName>
and with very similar ecology [i.e. 
<taxonomicName id="CCA04D0AF92B2035BB8BFDEAFC58FD1A" authority="Emery" authorityName="Emery" box="[690,922,620,644]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="lasioides">
<emphasis id="39D4EA9BF92B2035BB8BFDEAFD03FD1A" box="[690,705,621,644]" italics="true" pageId="16" pageNumber="37">L</emphasis>
. 
<emphasis id="39D4EA9BF92B2035BBF6FDEBFCF1FD1A" box="[719,819,620,644]" italics="true" pageId="16" pageNumber="37">lasioides</emphasis>
(Emery)
</taxonomicName>
, 
<taxonomicName id="CCA04D0AF92B2035BA9CFDEAFC3CFD1A" box="[933,1022,621,644]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="39D4EA9BF92B2035BA9CFDEAFC76FD1A" box="[933,948,621,644]" italics="true" pageId="16" pageNumber="37">L</emphasis>
. 
<emphasis id="39D4EA9BF92B2035BAFAFDEAFC3CFD1A" box="[963,1022,621,644]" italics="true" pageId="16" pageNumber="37">niger</emphasis>
</taxonomicName>
(L.) and 
<taxonomicName id="CCA04D0AF92B2035BD5EFDEAFA98FD1B" authority="Seifert" authorityName="Seifert" box="[1127,1370,620,645]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="platythorax">
<emphasis id="39D4EA9BF92B2035BD5EFDEAFBB4FD1A" box="[1127,1142,621,644]" italics="true" pageId="16" pageNumber="37">L</emphasis>
. 
<emphasis id="39D4EA9BF92B2035BDBDFDEBFAC7FD1A" box="[1156,1285,620,644]" italics="true" pageId="16" pageNumber="37">platythorax</emphasis>
Seifert
</taxonomicName>
] or in those of other formicid species, suggesting a high host specificity.
</paragraph>
</subSubSection>
</treatment>
</document>