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<document id="7DF3149D6C713A36EE18DA8E4DCAB712" ID-CLB-Dataset="29239" ID-DOI="10.11646/zootaxa.4496.1.7" ID-GBIF-Dataset="3e72ff64-3a86-42fe-9626-dd5bece8188e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1447049" ID-ZooBank="7BBA35FB-8FB5-410E-B5FA-999362AFD204" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538748456993" checkinUser="plazi" docAuthor="Höser, Norbert" docDate="2018" docId="2C55A67B444BFFDAC9D057CBFCE4FD58" docLanguage="en" docName="zootaxa.4496.1.7.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Proctodrilus tuberculatus" docType="treatment" docVersion="8" lastPageNumber="108" masterDocId="D06CDE03444EFFD6C9475225FF95FF94" masterDocTitle="Interrelation of Proctodrilus species (Oligochaeta: Lumbricidae) with lessivage and layering in European soil profiles" masterLastPageNumber="110" masterPageNumber="96" pageNumber="101" updateTime="1698567008538" updateUser="ExternalLinkService">
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<mods:title id="85526F0A4929E9AA068FDA47B2D9A3C3">Interrelation of Proctodrilus species (Oligochaeta: Lumbricidae) with lessivage and layering in European soil profiles</mods:title>
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<mods:namePart id="7DA5FB7471F046E3DB877CCBA4DD4071">Höser, Norbert</mods:namePart>
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<treatment id="2C55A67B444BFFDAC9D057CBFCE4FD58" ID-DOI="http://doi.org/10.5281/zenodo.5964291" ID-GBIF-Taxon="148640270" ID-Zenodo-Dep="5964291" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58" httpUri="http://treatment.plazi.org/id/2C55A67B444BFFDAC9D057CBFCE4FD58" lastPageId="12" lastPageNumber="108" pageId="5" pageNumber="101">
<subSubSection id="ECE644E6444BFFD3C9D057CBFE8CF9DA" pageId="5" pageNumber="101" type="nomenclature">
<paragraph id="A443176D444BFFD3C9D057CBFD50F99C" blockId="5.[151,1436,1518,1614]" box="[151,709,1518,1544]" pageId="5" pageNumber="101">
<heading id="FF0BA001444BFFD3C9D057CBFD50F99C" bold="true" box="[151,709,1518,1544]" fontSize="11" level="1" pageId="5" pageNumber="101" reason="1">
<taxonomicName id="63FC6CEE444BFFD3C9D057CBFD50F99C" authority="(Cernosvitov, 1935)" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[151,709,1518,1544]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="101" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F444BFFD3C9D057CBFD50F99C" bold="true" box="[151,709,1518,1544]" pageId="5" pageNumber="101">
<emphasis id="9688CB7F444BFFD3C9D057CBFE5EF99C" bold="true" box="[151,459,1518,1544]" italics="true" pageId="5" pageNumber="101">Proctodrilus tuberculatus</emphasis>
(
<bibRefCitation id="C06D6A9C444BFFD3C89D57CBFD28F99C" box="[474,701,1518,1544]" pageId="5" pageNumber="101" refId="ref13434" refString="Cernosvitov, L. (1935) Monographie der tschechoslowakischen Lumbriciden. Archiv pro prirodovedecky vyzkum Cech, 19, 1 - 86." type="journal article">Černosvitov, 1935</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A443176D444BFFD3C9D05437FE8CF9DA" blockId="5.[151,1436,1518,1614]" pageId="5" pageNumber="101">In total, 1839 adult individuals were found at 97 localities. They inhabited the finer-grained section of layer boundaries.</paragraph>
</subSubSection>
<subSubSection id="ECE644E6444BFFDFC9D0545BFD94F8D9" lastPageId="9" lastPageNumber="105" pageId="5" pageNumber="101" type="description">
<paragraph id="A443176D444BFFD3C9D0545BFC81F92E" blockId="5.[151,1436,1661,2010]" pageId="5" pageNumber="101">
• Strössendorf (DE), transect Main floodplain. 50°08'N, 11°14'E. Levee deposits. Profile with bandlike layers. 2 ind. at
<quantity id="6304BA88444BFFD3C8045487FE1DF92D" box="[323,392,1698,1722]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="101" unit="cm" value="17.0">17 cm</quantity>
depth, at layer boundary h2 Lu/Ut.
</paragraph>
<paragraph id="A443176D444BFFD3C9D054E3FE5FF896" blockId="5.[151,1436,1661,2010]" pageId="5" pageNumber="101">
• Kleinkorbetha (DE), transect Saale floodplain. 51°15'N, 12°03'E. Floodplain centre. 16 ind. at
<quantity id="6304BA88444BFFD3CC7054E0FA09F949" box="[1335,1436,1733,1758]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" metricValueMax="5.5" metricValueMin="0.9" pageId="5" pageNumber="101" unit="cm" value="32.0" valueMax="55.0" valueMin="9.0">955 cm</quantity>
depth, in h1 Lts/Ls3.
</paragraph>
<paragraph id="A443176D444BFFD3C9D0552BFBE6F8DE" blockId="5.[151,1436,1661,2010]" pageId="5" pageNumber="101">
• Räpitz (DE), Wiesengraben floodplain. 51°16'N, 12°13'E. Floodplain centre. Profile with bandlike layers. 12 ind. at
<quantity id="6304BA88444BFFD3C8695517FE1AF8DD" box="[302,399,1842,1866]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="2.0" metricValueMin="0.4" pageId="5" pageNumber="101" unit="cm" value="12.0" valueMax="20.0" valueMin="4.0">420 cm</quantity>
depth, in slightly stony h2 Ls3, preferentially at layer boundaries.
</paragraph>
<paragraph id="A443176D444BFFD3C9D05573FEA0F806" blockId="5.[151,1436,1661,2010]" pageId="5" pageNumber="101">
• Thronitz (DE), Wiesengraben floodplain. 51°16'N, 12°12'E. Floodplain centre. 1 ind. at
<quantity id="6304BA88444BFFD3CDFA5570FA96F8F9" box="[1213,1283,1877,1902]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="101" unit="cm" value="15.0">15 cm</quantity>
depth, in wet h4 Ls3.
</paragraph>
<paragraph id="A443176D444BFFD3C9D055BBFC2FF84E" blockId="5.[151,1436,1661,2010]" pageId="5" pageNumber="101">
• Döhlen (DE), transect Wiesengraben floodplain. 51°17'N, 12°12'E. Floodplain centre. Profile with bandlike layers. 6 ind. at
<quantity id="6304BA88444BFFD3C8D355E4FD96F84D" box="[404,515,1985,2010]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" metricValueMax="3.5" metricValueMin="1.7" pageId="5" pageNumber="101" unit="cm" value="26.0" valueMax="35.0" valueMin="17.0">1735 cm</quantity>
depth, in h3 Ls3.
<bibRefCitation id="C06D6A9C444BFFD3CB8955E7FC20F84E" box="[718,949,1985,2010]" pageId="5" pageNumber="101" refId="ref14278" refString="Hoser, N. (2015) Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus - Verteilung, in den Auen der Tschernosem- und Parabraunerde-Gebiete beiderseits des Erzgebirges. Opuscula Zoologica Budapest, 46, 133 - 142. https: // doi. org / 10.18348 / opzool. 2015.2.133" type="journal article">
Höser (2015:
<figureCitation id="3CC70BE8444BFFD3CA2F55E4FC3DF84E" box="[872,936,1985,2010]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="5" pageNumber="101">Fig. 1</figureCitation>
)
</bibRefCitation>
.
</paragraph>
<paragraph id="A443176D4448FFD0C9D052BDFEA0FF40" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Quesitz (DE), Wiesengraben floodplain. 51°18'N, 12°11'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884448FFD0CDF852B2FAA5FF3B" box="[1215,1328,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="102" unit="cm" value="27.5" valueMax="30.0" valueMin="25.0">2530 cm</quantity>
depth, in h4 Ls3.
</paragraph>
<paragraph id="A443176D4448FFD0C9D052C5FA13FF6C" blockId="6.[151,1437,151,2013]" box="[151,1414,223,248]" pageId="6" pageNumber="102">
• Kleinlehna (DE), Renne floodplain. 51°18'N, 12°11'E. Floodplain centre. 1 ind. at
<quantity id="6304BA884448FFD0CD3A52C5FB57FF63" box="[1149,1218,224,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="102" unit="cm" value="10.0">10 cm</quantity>
depth, in h3 Ls3.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05320FD95FEAB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Minkwitz (DE), Weisse Elster floodplain. 51°07'N, 12°13'E. Floodplain centre. Profile layered in parts. 6 ind. at
<quantity id="6304BA884448FFD0C9BB530DFED4FEAB" box="[252,321,296,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="102" unit="cm" value="10.0">10 cm</quantity>
depth, in h2 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05368FD16FE13" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Ostrau (DE), Weisse Elster floodplain. 51°06'N, 12°12'E. Floodplain edge. 16 ind. at&gt;
<quantity id="6304BA884448FFD0CDDA5369FB74FEF0" box="[1181,1249,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="102" unit="cm" value="15.0">15 cm</quantity>
depth, in h1 Lts, below a layer inhabited by
<taxonomicName id="63FC6CEE4448FFD0CB565354FDEEFE13" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[529,635,368,392]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="102" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4448FFD0CB565354FDEEFE13" box="[529,635,368,392]" italics="true" pageId="6" pageNumber="102">P. antipai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A443176D4448FFD0C9D053B0FEBAFE5B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Draschwitz DE), Weisse Elster floodplain. 51°06'N, 12°11'E. Floodplain centre. 4 ind. at
<quantity id="6304BA884448FFD0CD8453B1FAA7FE38" box="[1219,1330,404,428]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="102" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">1020 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D053F8FE54FD83" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Profen (DE), Weisse Elster floodplain. 51°08'N, 12°14'E. Floodplain centre. Fossil meander. 6 ind. at
<quantity id="6304BA884448FFD0CC1153F9FE96FD83" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="6" pageNumber="102" unit="cm" value="27.5" valueMax="40.0" valueMin="15.0">1540 cm</quantity>
depth, in h2 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05000FE71FDCB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Zwenkau (DE), transect Alte Elster floodplain. 51°13'N, 12°18'E. Floodplain terrace closest to river. 3 ind. at
<quantity id="6304BA884448FFD0C9A6506DFEB3FDCB" box="[225,294,584,608]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="102" unit="cm" value="12.0">12 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05048FDF4FD3C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Eisdorf (DE), Rützschke/Saugraben bottom land. 51°13'N, 12°12'E. Shallow depression at head of valley. 5 ind. at
<quantity id="6304BA884448FFD0C86950AAFE08FD33" box="[302,413,655,680]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" metricValueMax="4.5" metricValueMin="2.3" pageId="6" pageNumber="102" unit="cm" value="34.0" valueMax="45.0" valueMin="23.0">2345 cm</quantity>
depth, in h3 Ls3.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05090FE21FD64" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Grossschkorlopp (DE), Krebsgraben floodplain. 51°15'N, 12°14'E. Floodplain centre. 6 ind. at
<quantity id="6304BA884448FFD0CC715091FA09FD58" box="[1334,1436,692,716]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="2.2" metricValueMin="0.3" pageId="6" pageNumber="102" unit="cm" value="12.5" valueMax="22.0" valueMin="3.0">322 cm</quantity>
depth, in h2/3 Ls3.
</paragraph>
<paragraph id="A443176D4448FFD0C9D050D8FD77FCAC" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Rehbach (DE), transect Krebsgraben depression. 51°16'N, 12°16'E. Shallow depression at head of valley. 5 ind. at
<quantity id="6304BA884448FFD0C869513AFE08FCA3" box="[302,413,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="6" pageNumber="102" unit="cm" value="18.5" valueMax="22.0" valueMin="15.0">1522 cm</quantity>
depth, in h0/1 Ls3 of the Al.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05160FED9FCEB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Lobas (DE), Lindenberger Schnauder floodplain. 51°00'N, 12°14'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884448FFD0CC5F5161FADAFCC8" box="[1304,1359,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="102" unit="cm" value="8.0">8 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D051A8FEBAFC53" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Kayna (DE), Grosse Schnauder floodplain. 50°59'N, 12°14'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884448FFD0CD8251A9FAA1FC30" box="[1221,1332,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="102" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">1015 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D051F0FEBAFB9B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Oelsen (DE), Grosse Schnauder floodplain. 51°02'N, 12°14'E. Floodplain centre. 5 ind. at
<quantity id="6304BA884448FFD0CD8151F1FAA1FC78" box="[1222,1332,980,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="102" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">1015 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05638FE35FBCC" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Brossen (DE), Grosse Schnauder floodplain. 51°02'N, 12°15'E. Floodplain centre. 3 ind. at
<quantity id="6304BA884448FFD0CDAD5639FAA5FBA0" box="[1258,1328,1052,1076]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="102" unit="cm" value="10.0">10 cm</quantity>
depth, in h2 Lts of the Ah.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05640FE99FB34" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Wintersdorf (DE), Schnauder floodplain. 51°03'N, 12°20'E. Floodplain centre. 1 ind. at&gt;
<quantity id="6304BA884448FFD0CD8D5641FA9AFBE8" box="[1226,1295,1124,1148]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="102" unit="cm" value="30.0">30 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05688FDE1FB7C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Bosengröba (DE), transect Schnauder floodplain. 51°04'N, 12°22'E. Floodplain centre. 22 ind. at
<quantity id="6304BA884448FFD0CC6A5689FA09FB50" box="[1325,1436,1196,1220]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.4" metricValueMin="1.1" pageId="6" pageNumber="102" unit="cm" value="27.5" valueMax="44.0" valueMin="11.0">1144 cm</quantity>
depth, in h2 Lts. Clay skins present.
</paragraph>
<paragraph id="A443176D4448FFD0C9D056D0FD98FAC1" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Ruppersdorf (DE), transect Schnauder floodplain. 51°04'N, 12°22'E. Floodplain centre. Profile with bandlike layers. 47 ind. at 546, concentrated at
<quantity id="6304BA884448FFD0CA5F573DFCF5FAA4" box="[792,864,1304,1328]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="102" unit="cm" value="37.0">37 cm</quantity>
depth, in slightly gravelly h2 Ls3, which is rustmottled from
<quantity id="6304BA884448FFD0C83C5719FE55FAC0" box="[379,448,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="102" unit="cm" value="30.0">30 cm</quantity>
depth.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05744FB22FA08" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
<materialsCitation id="14941D304448FFD0C9D05744FB22FA08" ID-GBIF-Occurrence="1914265956" county="Lt" latitude="50.916668" location="Floodplain" longLatPrecision="1093" longitude="12.433333" municipality="lower Ah" pageId="6" pageNumber="102" specimenCount="1">
• Zehma (DE), transect Pleisse floodplain.
<geoCoordinate id="C1C871AA4448FFD0CB85577AFCB4FAEC" box="[706,801,1375,1400]" direction="north" orientation="latitude" pageId="6" pageNumber="102" precision="925" value="50.916668">50°55'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4448FFD0CA76577AFC1BFAEC" box="[817,910,1375,1400]" direction="east" orientation="longitude" pageId="6" pageNumber="102" precision="925" value="12.433333">12°26'E</geoCoordinate>
.
<location id="A12341B64448FFD0CAD9577AFB82FAEC" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64448FFD0CAD9577AFB82FAEC" box="[926,1047,1375,1400]" county="Lt" latitude="50.916668" longLatPrecision="1093" longitude="12.433333" municipality="lower Ah" name="Floodplain" pageId="6" pageNumber="102">Floodplain</location>
edge, foot of slope, fossil levee deposits. 8 ind. at
<quantity id="6304BA884448FFD0C8EA57A1FD8EFA08" box="[429,539,1412,1436]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="2.8" metricValueMin="1.1" pageId="6" pageNumber="102" unit="cm" value="19.5" valueMax="28.0" valueMin="11.0">1128 cm</quantity>
depth, in h2
<location id="A12341B64448FFD0CBE957A0FD47FA08" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64448FFD0CBE957A0FD47FA08" box="[686,722,1413,1436]" county="Lt" latitude="50.916668" longLatPrecision="1093" longitude="12.433333" municipality="lower Ah" name="Lts" pageId="6" pageNumber="102">Lts</location>
of the
<collectingMunicipality id="44278D174448FFD0CA6757A1FC1CFA09" box="[800,905,1412,1437]" pageId="6" pageNumber="102">lower Ah</collectingMunicipality>
and h1
<collectingCounty id="4D226FE14448FFD0CAA557A0FC6EFA08" box="[994,1019,1413,1436]" pageId="6" pageNumber="102">Lt</collectingCounty>
of the upper aB.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4448FFD0C9D0578CFD3FFA71" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Selleris (DE), Pleisse floodplain. 50°56'N, 12°25'E. Floodplain centre. Profile with bandlike layers. 2 ind. at
<quantity id="6304BA884448FFD0C9A657E9FE8DFA70" box="[225,280,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="102" unit="cm" value="8.0">8 cm</quantity>
depth, at layer boundary h2 Ls3/Sl.
</paragraph>
<paragraph id="A443176D4448FFD0C9D057D4FA11F99C" blockId="6.[151,1437,151,2013]" box="[151,1412,1519,1544]" pageId="6" pageNumber="102">
• Mockern (DE), Pleisse floodplain. 50°57'N, 12°26'E. Floodplain edge. 5 ind. at
<quantity id="6304BA884448FFD0CD1057D5FB53F99C" box="[1111,1222,1520,1544]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.8" metricValueMin="1.2" pageId="6" pageNumber="102" unit="cm" value="20.0" valueMax="28.0" valueMin="12.0">1228 cm</quantity>
depth, in h2 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05430FA09F9B8" blockId="6.[151,1437,151,2013]" box="[151,1436,1556,1581]" pageId="6" pageNumber="102">
• Ehrenberg (DE), Pleisse floodplain. 50°58'N, 12°27'E. Floodplain centre. 4 ind. at
<quantity id="6304BA884448FFD0CD365431FB75F9B8" box="[1137,1248,1556,1581]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="102" unit="cm" value="17.5" valueMax="25.0" valueMin="10.0">1025 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D0541CFBAEF9E0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Kotteritz (DE), transect Pleisse floodplain. 50°58'N, 12°28'E. Floodplain centre. Profile with bandlike layers. 8 ind. at
<quantity id="6304BA884448FFD0C8045479FE27F9E0" box="[323,434,1628,1652]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="6" pageNumber="102" unit="cm" value="21.0" valueMax="22.0" valueMin="20.0">2022 cm</quantity>
depth, at layer boundary h2 Lt/Ls3, over rust-mottled aB.
</paragraph>
<paragraph id="A443176D4448FFD0C9D054A4FC14F928" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Wilchwitz (DE), transect Pleisse floodplain. 50°59'N, 12°29'E. Second floodplain terrace close to river. 10 ind. at
<quantity id="6304BA884448FFD0C8695481FE08F928" box="[302,413,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="6" pageNumber="102" unit="cm" value="25.0" valueMax="27.0" valueMin="23.0">2327 cm</quantity>
depth, in h1 Lts,&gt;
<quantity id="6304BA884448FFD0CB355481FD3CF928" box="[626,681,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="102" unit="cm" value="2.0">2 cm</quantity>
over clay-poor aB.
</paragraph>
<paragraph id="A443176D4448FFD0C9D054ECFE71F890" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Schelchwitz (DE), transect Pleisse floodplain. 51°00'N, 12°29'E. Floodplain terrace closest to river. 3 ind. at
<quantity id="6304BA884448FFD0C9A654C9FEB3F890" box="[225,294,1772,1796]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="102" unit="cm" value="14.0">14 cm</quantity>
depth, in h2 Lts.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05534FCF6F8D8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Kraschwitz (DE), transect Pleisse floodplain. 51°00'N, 12°29'E. Floodplain centre. 5 ind. at
<quantity id="6304BA884448FFD0CDA6552AFAC5F8BC" box="[1249,1360,1807,1832]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" metricValueMax="5.2" metricValueMin="4.0" pageId="6" pageNumber="102" unit="cm" value="46.0" valueMax="52.0" valueMin="40.0">4052 cm</quantity>
depth, in weakly rust-mottled h0 Lts of the Ba,&gt;
<quantity id="6304BA884448FFD0CBFF5511FD7AF8D8" box="[696,751,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="102" unit="cm" value="6.0">6 cm</quantity>
above Gr.
</paragraph>
<paragraph id="A443176D4448FFD0C9D0557CFB90F800" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="102">
• Poschwitz (DE), transect slope of depression leading to Pleisse floodplain. 51°00'N, 12°28'E. Lower slope. 24 ind. at
<quantity id="6304BA884448FFD0C8165559FE55F800" box="[337,448,1916,1940]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="4.1" metricValueMin="2.0" pageId="6" pageNumber="102" unit="cm" value="30.5" valueMax="41.0" valueMin="20.0">2041 cm</quantity>
depth, in h1 Lts of the Bt+Al.
<bibRefCitation id="C06D6A9C4448FFD0CA5D5558FB95F800" box="[794,1024,1916,1941]" pageId="6" pageNumber="102" refId="ref14220" refString="Hoser, N. (2012) Regenwurmer (Oligochaeta: Lumbricidae) in der Hangcatena. Hercynia, Neue Folge, 45, 193 - 208." type="journal article">Höser (2012: Fig. 3)</bibRefCitation>
.
</paragraph>
<paragraph id="A443176D4448FFD0C9D05584FA08F82D" blockId="6.[151,1437,151,2013]" box="[151,1437,1952,1977]" pageId="6" pageNumber="102">
• Remsa (DE), Pleisse floodplain. 51°00'N, 12°29'E. Floodplain centre. 4 ind. at
<quantity id="6304BA884448FFD0CD0B5585FB04F82C" box="[1100,1169,1952,1977]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="102" unit="cm" value="16.0">16 cm</quantity>
depth, in Lts of the Ah.
</paragraph>
<paragraph id="A443176D4448FFD1C9D055E0FEEFFF40" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="103" pageId="6" pageNumber="102">
• Windischleuba (DE), ten transects Pleisse floodplain. 51°01'N, 12°28'E. Floodplain centre. One profile close to river bandlike, remainder simple layers. 386 ind. at
<quantity id="6304BA884449FFD1CA0C52B2FC25FF3B" box="[843,944,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.9" metricValueMax="7.5" metricValueMin="0.3" pageId="7" pageNumber="103" unit="cm" value="39.0" valueMax="75.0" valueMin="3.0">375 cm</quantity>
, mostly at
<quantity id="6304BA884449FFD1CD6A52BDFB08FF3B" box="[1069,1181,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="7" pageNumber="103" unit="cm" value="25.5" valueMax="30.0" valueMin="21.0">2130 cm</quantity>
depth, in h1 Lts. Clay skins present.
</paragraph>
<paragraph id="A443176D4449FFD1C9D052C5FBB6FE89" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Borgishain (DE), transect Pleisse floodplain. 51°01'N, 12°28'E. Floodplain centre. 23 ind. at
<quantity id="6304BA884449FFD1CDA952FAFADAFF63" box="[1262,1359,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" metricValueMax="4.5" metricValueMin="0.3" pageId="7" pageNumber="103" unit="cm" value="24.0" valueMax="45.0" valueMin="3.0">345 cm</quantity>
depth, in h1/3 Lts of the Ah/aB or weakly rust-mottled Ah, over clay enrichment.
</paragraph>
<paragraph id="A443176D4449FFD1C9D0530DFD3DFEF1" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Fockendorf (DE), transect Pleisse floodplain. 51°03'N, 12°28'E. Several floodplain terraces. 91 ind. at
<quantity id="6304BA884449FFD1CC235302FE96FEF0" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="3.6" metricValueMin="0.3" pageId="7" pageNumber="103" unit="cm" value="19.5" valueMax="36.0" valueMin="3.0">336 cm</quantity>
depth, in h1/2 Lts, in places over Bg.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05355FD1AFE38" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Primmelwitz (DE), transect Pleisse floodplain. 51°03'N, 12°28'E. Floodplain centre. 7 ind. at
<quantity id="6304BA884449FFD1CC6F5355FA09FE13" box="[1320,1436,368,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="7" pageNumber="103" unit="cm" value="22.0" valueMax="24.0" valueMin="20.0">2024 cm</quantity>
depth, in very slightly gravelly h3 Lts.
</paragraph>
<paragraph id="A443176D4449FFD1C9D0539DFBB6FE60" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Treben (DE), transect Pleisse floodplain. 51°03'N, 12°27'E. Floodplain centre. 21 ind. at
<quantity id="6304BA884449FFD1CD8A5392FAA5FE5B" box="[1229,1328,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="4.0" metricValueMin="0.6" pageId="7" pageNumber="103" unit="cm" value="23.0" valueMax="40.0" valueMin="6.0">640 cm</quantity>
depth, in h1/2 Lts, some at layer boundary to Ls3; also at upper boundary of the aB.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05025FB0BFDA8" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Plottendorf (DE), transect Pleisse/Gerstenbach floodplain. 51°03'N, 12°27'E. Floodplain centre. 16 ind. at
<quantity id="6304BA884449FFD1C9A65001FEC5FDA8" box="[225,336,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" metricValueMax="4.5" metricValueMin="2.7" pageId="7" pageNumber="103" unit="cm" value="36.0" valueMax="45.0" valueMin="27.0">2745 cm</quantity>
depth, in h0/1 Lts of the aB. In places
<taxonomicName id="63FC6CEE4449FFD1CA435000FCFBFDA8" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[772,878,549,572]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="103" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4449FFD1CA435000FCFBFDA8" box="[772,878,549,572]" italics="true" pageId="7" pageNumber="103">P. antipai</emphasis>
</taxonomicName>
in overlying layer (
<figureCitation id="3CC70BE84449FFD1CD0C5001FB18FDA8" box="[1099,1165,548,573]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="7" pageNumber="103">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="A443176D4449FFD1C9D0506DFE04FD3C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Serbitz (DE), transect Pleisse floodplain. 51°04'N, 12°27'E. Floodplain centre. 11 ind. mostly at
<quantity id="6304BA884449FFD1CC6C5062FA09FDCB" box="[1323,1436,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.8" metricValueMin="1.6" pageId="7" pageNumber="103" unit="cm" value="22.0" valueMax="28.0" valueMin="16.0">1628 cm</quantity>
depth, in h0/1 Lts, at upper boundary of the aB, singletons at up to
<quantity id="6304BA884449FFD1CABC5049FBD6FD10" box="[1019,1091,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="103" unit="cm" value="50.0">50 cm</quantity>
depth. In places
<taxonomicName id="63FC6CEE4449FFD1CC495048FAE9FD10" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1294,1404,621,644]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="103" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4449FFD1CC495048FAE9FD10" box="[1294,1404,621,644]" italics="true" pageId="7" pageNumber="103">P. antipai</emphasis>
</taxonomicName>
in overlying layer.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05090FC41FD7B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Haselbach (DE), two transects Pleisse floodplain. 51°04'N, 12°27'E. Floodplain centre. Profile with bandlike layers. 21 ind. at
<date id="D04231AD4449FFD1C8E550F2FDDDFD64" box="[418,584,727,752]" pageId="7" pageNumber="103" value="1935-03-05" valueMax="1935-03-25" valueMin="1935-03-05">
35 and
<quantity id="6304BA884449FFD1CB4550F2FDDDFD64" box="[514,584,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="7" pageNumber="103" unit="cm" value="30.0" valueMax="35.0" valueMin="25.0">2535</quantity>
</date>
cm depth, in h2/3 Lts, over the aB.
</paragraph>
<paragraph id="A443176D4449FFD1C9D050D8FDB5FCC9" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Gaulis (DE), transect Pleisse floodplain. 51°11'N, 12°24'E. Fossil levee deposits, floodplain centre. Profile with several layers. 9 ind. at
<quantity id="6304BA884449FFD1CB625105FD00FCA3" box="[549,661,800,824]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" metricValueMax="3.7" metricValueMin="1.2" pageId="7" pageNumber="103" unit="cm" value="24.5" valueMax="37.0" valueMin="12.0">1237 cm</quantity>
depth, mostly at layer boundary in h2 Ls3, over Sl and aB. In places
<taxonomicName id="63FC6CEE4449FFD1C9A65160FEDEFCC8" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[225,331,837,860]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="103" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4449FFD1C9A65160FEDEFCC8" box="[225,331,837,860]" italics="true" pageId="7" pageNumber="103">P. antipai</emphasis>
</taxonomicName>
in overlying layer.
</paragraph>
<paragraph id="A443176D4449FFD1C9D0514DFADDFCEB" blockId="7.[151,1437,151,2013]" box="[151,1352,871,896]" pageId="7" pageNumber="103">
• Rötha (DE), Pleisse floodplain. 51°12'N, 12°24'E. Floodplain centre. 1 ind. at
<quantity id="6304BA884449FFD1CD03514DFB1CFCEB" box="[1092,1161,872,896]" metricMagnitude="-1" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="103" unit="cm" value="28.0">28 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4449FFD1C9D051A8FE99FC53" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Papiermühle (DE), Sprotte floodplain. 50°56'N, 12°24'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884449FFD1CDD851A9FA9BFC30" box="[1183,1294,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="7" pageNumber="103" unit="cm" value="37.5" valueMax="45.0" valueMin="30.0">3045 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4449FFD1C9D051F0FD8DFB84" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">• Nöbden (DE), transect Blaue Flut floodplain. 50°56'N, 12°21'E. Foot of slope. 3 ind. in wet h2 Lts, at layer boundary in inverse profile.</paragraph>
<paragraph id="A443176D4449FFD1C9D05638FB5EFBA0" blockId="7.[151,1437,151,2013]" box="[151,1227,1052,1077]" pageId="7" pageNumber="103">• Gleina (DE), Blaue Flut floodplain. 50°57'N, 12°24'E. Floodplain centre. 4 ind. in h1 Lts.</paragraph>
<paragraph id="A443176D4449FFD1C9D05665FE0DFBE8" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Schlöpitz (DE), transect Blaue Flut floodplain. 50°58'N, 12°23'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884449FFD1CC6E5665FA09FBC3" box="[1321,1436,1088,1112]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="103" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">1030 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4449FFD1C9D056ACFD26FB50" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Rasephas (DE), two transects Blaue Flut floodplain. 51°01'N, 12°27'E. Floodplain centre. 10 ind. at
<quantity id="6304BA884449FFD1CC1156A2FE96FB50" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="7" pageNumber="103" unit="cm" value="22.5" valueMax="35.0" valueMin="10.0">1035 cm</quantity>
depth, in h1/2 Lts. Clay skins present.
</paragraph>
<paragraph id="A443176D4449FFD1C9D056F4FAD1FB7C" blockId="7.[151,1437,151,2013]" box="[151,1348,1231,1256]" pageId="7" pageNumber="103">• Altenburg (DE), Deutscher Bach floodplain. 51°00'N, 12°26'E. Floodplain centre. 2 ind. in h1/2 Lts.</paragraph>
<paragraph id="A443176D4449FFD1C9D056D0FE99FAA4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Schlauditz (DE), Gerstenbach floodplain. 50°59'N, 12°22'E. Floodplain centre. 1 ind. at
<quantity id="6304BA884449FFD1CD9556D1FA9EFA98" box="[1234,1291,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="103" unit="cm" value="8.0">8 cm</quantity>
depth, in h1 Lts.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05718FCB1FAEC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Obermolbitz (DE), transect Gerstenbach floodplain. 51°01'N, 12°24'E. Fossil meander. 15 ind. at
<quantity id="6304BA884449FFD1CC6B5719FA09FAC0" box="[1324,1436,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="7" pageNumber="103" unit="cm" value="28.5" valueMax="30.0" valueMin="27.0">2730 cm</quantity>
depth, in h1 Lts, above intensively rust-mottled Go.
</paragraph>
<paragraph id="A443176D4449FFD1C9D057A0FEEFFA54" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">• Untermolbitz (DE), Gerstenbach floodplain. 51°01'N, 12°25'E. Floodplain centre. 3 ind. at h2 Lts. Clay skins present.</paragraph>
<paragraph id="A443176D4449FFD1C9D057E8FE6BF99C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Gerstenberg (DE), transect Gerstenbach floodplain. 51°02'N, 12°26'E. Floodplain centre. 24 ind. at
<quantity id="6304BA884449FFD1CC1157E9FE96F99C" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="3.2" metricValueMin="1.4" pageId="7" pageNumber="103" unit="cm" value="23.0" valueMax="32.0" valueMin="14.0">1432 cm</quantity>
depth, in h2 Lts to Lt.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05430FCA0F9E0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Trebanz (DE), three transects Gerstenbach floodplain. 51°03'N, 12°27'E. Floodplain centre, floodplain edge. Profile partly in bandlike layers. 43 ind. at
<quantity id="6304BA884449FFD1CB8A5412FCABF9C4" box="[717,830,1591,1616]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="4.5" metricValueMin="1.6" pageId="7" pageNumber="103" unit="cm" value="30.5" valueMax="45.0" valueMin="16.0">1645 cm</quantity>
depth, in slightly gravelly h2/3 Lts/Lu of the aB and Go. Near floodplain edge under layer with
<taxonomicName id="63FC6CEE4449FFD1CB845478FCBFF9E0" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[707,810,1629,1652]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="103" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4449FFD1CB845478FCBFF9E0" box="[707,810,1629,1652]" italics="true" pageId="7" pageNumber="103">P. antipai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A443176D4449FFD1C9D054A4FA04F929" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Plottendorf (DE), transect Gerstenbach floodplain. 51°03'N, 12°27'E. Floodplain centre, floodplain edge. 21 ind. at
<quantity id="6304BA884449FFD1C8695481FE08F928" box="[302,413,1700,1725]" metricMagnitude="-1" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="7" pageNumber="103" unit="cm" value="20.5" valueMax="26.0" valueMin="15.0">1526 cm</quantity>
depth, in h1/2 Lts over rust-red aB. Towards floodplain edge
<taxonomicName id="63FC6CEE4449FFD1CD165480FB29F928" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1105,1212,1701,1724]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="103" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4449FFD1CD165480FB29F928" box="[1105,1212,1701,1724]" italics="true" pageId="7" pageNumber="103">P. antipai</emphasis>
</taxonomicName>
in overlying layer.
</paragraph>
<paragraph id="A443176D4449FFD1C9D054ECFAD1F8BC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Leinawald (DE), two transects Teichgraben floodplain. 50°59'N, 12°32'E. Floodplain centre, fossil meander. Profile with bandlike layers. 71 ind. at
<quantity id="6304BA884449FFD1CBDB54C9FD6BF890" box="[668,766,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="7" pageNumber="103" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">410 cm</quantity>
depth, in h0/1 Lts above Ls3, in bandlike clay enrichment, above pale grey, slightly rust-mottled Sw; at
<quantity id="6304BA884449FFD1CB905535FCADF8BC" box="[727,824,1808,1832]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="2.9" metricValueMin="0.4" pageId="7" pageNumber="103" unit="cm" value="16.5" valueMax="29.0" valueMin="4.0">429 cm</quantity>
depth, in h1 Ls3 above Ss. Clay skins present.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05510FD1DF8E4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
• Lohma (DE), transect Spannerbach floodplain. 50°58'N, 12°32'E. Floodplain centre. 6 ind. at
<quantity id="6304BA884449FFD1CC6E5511FA09F8D8" box="[1321,1436,1844,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="103" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">1015 cm</quantity>
depth, in rust-mottled h1 Lts, over Lt.
</paragraph>
<paragraph id="A443176D4449FFD1C9D05558FBC3F848" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="103">
<materialsCitation id="14941D304449FFD1C9D05558FBC3F848" ID-GBIF-Occurrence="1914265965" collectorName="Go. Clay" latitude="51.0" location="Sd" longLatPrecision="1093" longitude="12.5" pageId="7" pageNumber="103" specimenCount="1" stateProvince="Sd">
• Pöppschen (DE), five transects Spannerbach floodplain.
<geoCoordinate id="C1C871AA4449FFD1CA235559FC56F801" box="[868,963,1916,1941]" direction="north" orientation="latitude" pageId="7" pageNumber="103" precision="925" value="51.0">51°00'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4449FFD1CA975559FBB8F800" box="[976,1069,1916,1940]" direction="east" orientation="longitude" pageId="7" pageNumber="103" precision="925" value="12.5">12°30'E</geoCoordinate>
. Lower slope, floodplain centre. 125 ind. at
<quantity id="6304BA884449FFD1C82F5585FE4FF82C" box="[360,474,1952,1977]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="7" pageNumber="103" unit="cm" value="22.5" valueMax="35.0" valueMin="10.0">1035 cm</quantity>
depth, in alluvial fan h1/3 Lts/Ls3, in floodplain centre h1 Lt/Lts; in the Al/Bt, in places reaching
<collectingRegion id="6638D98F4449FFD1C8D255E1FE20F849" box="[405,437,1988,2013]" country="Serbia" name="Sid" pageId="7" pageNumber="103">Sd</collectingRegion>
, Gro,
<collectorName id="090972BB4449FFD1C8BD55E1FDF5F849" box="[506,608,1988,2013]" pageId="7" pageNumber="103">Go. Clay</collectorName>
skins present.
<bibRefCitation id="C06D6A9C4449FFD1CA4155E0FBC6F848" box="[774,1107,1988,2013]" pageId="7" pageNumber="103" refId="ref14220" refString="Hoser, N. (2012) Regenwurmer (Oligochaeta: Lumbricidae) in der Hangcatena. Hercynia, Neue Folge, 45, 193 - 208." type="journal article">Höser (2012: partly in Fig. 2)</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4446FFDEC9D052BDFAA3FF41" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Leinawald (DE), two transects confluence with Wyhra floodplain. 50°58'N, 12°35'E. 10 ind. at
<quantity id="6304BA884446FFDECC7F52B2FA09FF3B" box="[1336,1436,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="8" pageNumber="104" unit="cm" value="8.5" valueMax="15.0" valueMin="2.0">215 cm</quantity>
depth, in wet, rust-mottled h2 Lts with clay skins, over Gro, Gr, rust-coloured Go, with clay below.
</paragraph>
<paragraph id="A443176D4446FFDEC9D052C5FE42FE88" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Langenleuba-Niederhain (DE), transect Wyhra floodplain. 50°58'N, 12°35'E. Floodplain centre. 1 ind. at
<quantity id="6304BA884446FFDECCC752C5FE96FE88" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="104" unit="cm" value="12.0">12 cm</quantity>
depth, in h0/1 Lts.
</paragraph>
<paragraph id="A443176D4446FFDEC9D0530DFD5CFEF0" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Streitwald (DE), two transects Wyhra floodplain. 51°02'N, 12°35'E. Floodplain centre. Profile partly layered. 3 ind. at
<quantity id="6304BA884446FFDEC8E45369FD91FEF0" box="[419,516,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="8" pageNumber="104" unit="cm" value="11.0" valueMax="14.0" valueMin="8.0">814 cm</quantity>
depth, in h1 Ls3.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05355FE56FE44" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Benndorf (DE), transect Wyhra floodplain. 51°03'N, 12°33'E. Fossil meander at floodplain edge. 26 ind. in area of fossil levee deposits at
<quantity id="6304BA884446FFDECB0653B1FD31FE38" box="[577,676,404,428]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="3.8" metricValueMin="0.8" pageId="8" pageNumber="104" unit="cm" value="23.0" valueMax="38.0" valueMin="8.0">838 cm</quantity>
depth, in h1/2 Lts, over a layer inhabited by
<taxonomicName id="63FC6CEE4446FFDECDED53B0FA83FE38" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1194,1302,405,428]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="104" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4446FFDECDED53B0FA83FE38" box="[1194,1302,405,428]" italics="true" pageId="8" pageNumber="104">P. antipai</emphasis>
</taxonomicName>
(
<figureCitation id="3CC70BE84446FFDECC6053B1FAFEFE38" box="[1319,1387,404,429]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="8" pageNumber="104">Fig. 1</figureCitation>
and
<bibRefCitation id="C06D6A9C4446FFDEC9A6539DFEFDFE44" box="[225,360,440,464]" pageId="8" pageNumber="104" refId="ref14083" refString="Hoser, N. (2008) Die Regenwurmer Proctodrilus tuberculatus (CERNOSVITOV, 1935) und Proctodrilus antipai (MICHAELSEN, 1891) als Indikatoren der fossilen Oberflachengliederung der Flussaue (Oligochaeta: Lumbricidae). Hercynia, Neue Folge, 41, 263 - 272." type="journal article">Höser 2008</bibRefCitation>
: Fig. 2).
</paragraph>
<paragraph id="A443176D4446FFDEC9D053F8FCBCFD83" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Beucha (DE), transect Eula floodplain. 51°09'N, 12°35'E. Fossil levee deposits. Profile with several layers. 1 ind. at
<quantity id="6304BA884446FFDEC8695025FEE6FD83" box="[302,371,512,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="104" unit="cm" value="17.0">17 cm</quantity>
depth, in h0/1 Ls3. Clay skins present.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05000FE23FDCB" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Kitzscher (DE), Eula floodplain. 51°09'N, 12°33'E. Floodplain centre. 11 ind. at
<quantity id="6304BA884446FFDECD2F5001FB42FDA8" box="[1128,1239,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.8" metricValueMin="1.2" pageId="8" pageNumber="104" unit="cm" value="25.0" valueMax="38.0" valueMin="12.0">1238 cm</quantity>
depth, in h1 Ls3. Clay skins present.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05048FBE5FD33" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Kötteritzsch (DE), transect Mulde floodplain. 51°10'N, 12°48'E. Two floodplain terraces close to river. Profile layered in places. 12 ind. at
<quantity id="6304BA884446FFDECB2950B5FD5AFD33" box="[622,719,656,680]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="2.0" metricValueMin="0.4" pageId="8" pageNumber="104" unit="cm" value="12.0" valueMax="20.0" valueMin="4.0">420 cm</quantity>
depth, in h2 Ls3. Clay skins present.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05090FA8AFD64" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Kössern (DE), transect Mulde floodplain.
<geoCoordinate id="C1C871AA4446FFDECB855091FCB5FD59" box="[706,800,692,717]" direction="north" orientation="latitude" pageId="8" pageNumber="104" precision="925" value="51.183334">51°11'N</geoCoordinate>
, 12°47'. Floodplain terrace closest to river. Profile with bandlike layers. 70 ind. mostly at
<quantity id="6304BA884446FFDECB1B50F2FD5EFD7B" box="[604,715,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="8" pageNumber="104" unit="cm" value="17.5" valueMax="25.0" valueMin="10.0">1025 cm</quantity>
depth, in h1/2 Ls3, concentrated at layer boundaries.
</paragraph>
<paragraph id="A443176D4446FFDEC9D050D8FE48FCAC" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Nimbschen (DE), Mulde floodplain. 51°13'N, 12°45'E. Floodplain terrace closest to river. 10 ind. at
<quantity id="6304BA884446FFDECC1150D9FE96FCA3" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="20.0" valueMax="25.0" valueMin="15.0">1525 cm</quantity>
depth, in h1/2 Ls3.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05160FE0BFC14" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Grimma (DE), Mulde floodplain. 51°15'N, 12°44'E. Floodplain terrace closest to river. 18 ind. at
<quantity id="6304BA884446FFDECC6B5161FA09FCC8" box="[1324,1436,836,861]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="27.5" valueMax="40.0" valueMin="15.0">1540 cm</quantity>
depth, in h1 Ls3.
</paragraph>
<paragraph id="A443176D4446FFDEC9D051A8FDC3FC5C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Dorna (DE), transect Mulde floodplain. 51°15'N, 12°45'E. Two floodplain terraces close to river. 14 ind. at
<quantity id="6304BA884446FFDEC9A6518AFEC5FC53" box="[225,336,943,968]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="25.0" valueMax="35.0" valueMin="15.0">1535 cm</quantity>
depth, in h1/2 Lts/Ls3.
</paragraph>
<paragraph id="A443176D4446FFDEC9D051F0FD1DFB84" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">• Neichen (DE), Mulde floodplain. 51°17'N, 12°46'E. Floodplain terrace closest to river. 2 ind. in h1 Ls3, underlying substrate slightly gravelly.</paragraph>
<paragraph id="A443176D4446FFDEC9D05638FD73FBCC" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Nučničky (CZ), Labe (Elbe) floodplain. 50°30'N, 14°13'E. Floodplain terrace closest to river. 15 ind. at
<quantity id="6304BA884446FFDECC235639FE96FBC3" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="3.0" metricValueMin="0.2" pageId="8" pageNumber="104" unit="cm" value="16.0" valueMax="30.0" valueMin="2.0">230 cm</quantity>
depth, in slightly gravelly to stony h1 Ls3.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05640FE1FFB50" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Mlékojedy (CZ), Labe (Elbe) floodplain. 50°32'N, 14°08'E. Migratory confluence with the Ohře. Profile with bandlike layers. 10 ind. at
<quantity id="6304BA884446FFDECB0F56ADFD2CFB34" box="[584,697,1160,1184]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="104" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">1030 cm</quantity>
depth, in h2 Ls3. The layer above inhabited by
<taxonomicName id="63FC6CEE4446FFDECD9B56ACFAD2FB0B" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1244,1351,1160,1184]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="104" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4446FFDECD9B56ACFAD2FB0B" box="[1244,1351,1160,1184]" italics="true" pageId="8" pageNumber="104">P. antipai</emphasis>
</taxonomicName>
(
<bibRefCitation id="C06D6A9C4446FFDECC1056ADFEE9FB50" pageId="8" pageNumber="104" refId="ref14278" refString="Hoser, N. (2015) Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus - Verteilung, in den Auen der Tschernosem- und Parabraunerde-Gebiete beiderseits des Erzgebirges. Opuscula Zoologica Budapest, 46, 133 - 142. https: // doi. org / 10.18348 / opzool. 2015.2.133" type="journal article">Höser 2015: Table 3</bibRefCitation>
).
</paragraph>
<paragraph id="A443176D4446FFDEC9D056F4FB24FB7C" blockId="8.[151,1437,151,2013]" box="[151,1201,1231,1256]" pageId="8" pageNumber="104">• Dessau (DE), BDF Saalberghau, Elbe floodplain. 51°53'N, 12°12'E. 2 ind. in h1 Us/Ut.</paragraph>
<paragraph id="A443176D4446FFDEC9D056D0FBF2FAA4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Dessau (DE), Saalberghau, transect Elbe floodplain. 51°53'N, 12°12'E. Three floodplain terraces close to river. 15 ind. in h1/2 Us, Lu, Lts. Clay skins present in Lts.
<bibRefCitation id="C06D6A9C4446FFDECA3C573CFBF7FAA4" box="[891,1122,1303,1328]" pageId="8" pageNumber="104" refId="ref14052" refString="Hoser, N. (2005) Regenwurmer im geomorphologischen Relief der Aue des Mittelelbegebietes. Veroffentlichungen der LPR Landschaftsplanung Dr. Reichhoff GmbH, 3, 71 - 76." type="journal article">Höser (2005: Fig. 3)</bibRefCitation>
.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05718FBC6FAC0" blockId="8.[151,1437,151,2013]" box="[151,1107,1340,1365]" pageId="8" pageNumber="104">• Steutz (DE), two BDF, Elbe floodplain. 51°52'N, 12°02'E. 10 ind. in h1 Lts/Lt.</paragraph>
<paragraph id="A443176D4446FFDEC9D05744FD31FA08" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Steutz (DE), transect Elbe floodplain. 51°52'N, 12°02'E. Middle floodplain terrace. 3 ind. in h2 Lu of fossil floodplain channel.
<bibRefCitation id="C06D6A9C4446FFDEC88757A0FD31FA08" box="[448,676,1412,1437]" pageId="8" pageNumber="104" refId="ref14052" refString="Hoser, N. (2005) Regenwurmer im geomorphologischen Relief der Aue des Mittelelbegebietes. Veroffentlichungen der LPR Landschaftsplanung Dr. Reichhoff GmbH, 3, 71 - 76." type="journal article">
Höser (2005:
<figureCitation id="3CC70BE84446FFDECB1D57A1FD0CFA08" box="[602,665,1412,1437]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="8" pageNumber="104">Fig. 1</figureCitation>
)
</bibRefCitation>
</paragraph>
<paragraph id="A443176D4446FFDEC9D0578CFEB1FA70" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Kleneč (CZ), Čepel (Zippel) floodplain. 50°23'N, 14°15'E. Floodplain centre. 4 ind. at
<quantity id="6304BA884446FFDECDF85782FAA5FA54" box="[1215,1328,1447,1472]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="20.0" valueMax="25.0" valueMin="15.0">1525 cm</quantity>
depth, in h2 Lt.
</paragraph>
<paragraph id="A443176D4446FFDEC9D057D4FEC9F9B8" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Söred (HU), transect slope. 47°18'N, 18°18'E. Lower slope, foot of slope. 24 ind. at
<quantity id="6304BA884446FFDECDD057CAFB6DF99C" box="[1175,1272,1519,1544]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="8" pageNumber="104" unit="cm" value="17.5" valueMax="30.0" valueMin="5.0">530 cm</quantity>
depth, in h2/4 Lts and Lt.
</paragraph>
<paragraph id="A443176D4446FFDEC9D0541CFECDF929" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Surduc (RO), transect Someş floodplain. 47°16'N, 23°21'E. Four slightly to medium gravelly floodplain terraces. Profile with several layers. 79 ind. at
<quantity id="6304BA884446FFDECBA35479FCD0F9E0" box="[740,837,1628,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="8" pageNumber="104" unit="cm" value="17.5" valueMax="30.0" valueMin="5.0">530 cm</quantity>
depth, on both terraces close to river in Ls3, in places above ferrous hydroxide (limonite) precipitation (Gso); on both terraces away from river in h0/2 Lts with clay skins.
</paragraph>
<paragraph id="A443176D4446FFDEC9D054ECFE5AF891" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Tinca (RO), Crişul Negru floodplain. 46°46'N, 21°57'E. Floodplain centre. 2 ind. at
<quantity id="6304BA884446FFDECDDB54E2FA99F974" box="[1180,1292,1735,1760]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="18.5" valueMax="22.0" valueMin="15.0">1522 cm</quantity>
depth, in h1 Lts over gravel layer.
</paragraph>
<paragraph id="A443176D4446FFDEC9D05534FB0FF8D9" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Porumbenii Mari (RO), transect Tîrnava Mare floodplain. 46°16'N, 25°08'E. Floodplain terrace close to river. 1 ind. at
<quantity id="6304BA884446FFDEC8C35511FE5CF8D8" box="[388,457,1844,1868]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="104" unit="cm" value="17.0">17 cm</quantity>
depth, in h2 Ls3, at boundary to underlying slightly stony layer.
</paragraph>
<paragraph id="A443176D4446FFDEC9D0557CFC20F849" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="104">
• Beteşti/Betfalva (RO), transect slope on Fâneaţa Îngustă.
<geoCoordinate id="C1C871AA4446FFDECA275572FC2AF8E4" box="[864,959,1879,1904]" direction="north" orientation="latitude" pageId="8" pageNumber="104" precision="925" value="46.3">46°18'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4446FFDECA8E5572FBB3F8E4" box="[969,1062,1879,1904]" direction="east" orientation="longitude" pageId="8" pageNumber="104" precision="925" value="25.116667">25°07'E</geoCoordinate>
. 18 ind. In loess-loam lessive soil on slope plateau, below the Ah, in uppermost zone of the Al and Bt+Al, mostly at
<quantity id="6304BA884446FFDECDC05559FB4EF800" box="[1159,1243,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="8" pageNumber="104" unit="cm" value="6.5" valueMax="7.0" valueMin="6.0">67 cm</quantity>
depth, in h1 Lts/ Lt. Clay skins present. Colluvisol on lower slope, foot of slope, and adjoining basal area with several layers. Species there at
<quantity id="6304BA884446FFDEC8D055E1FD92F848" box="[407,519,1988,2013]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" metricValueMax="3.7" metricValueMin="1.5" pageId="8" pageNumber="104" unit="cm" value="26.0" valueMax="37.0" valueMin="15.0">1537 cm</quantity>
depth, in h1/3 Ls3, Lts.
<bibRefCitation id="C06D6A9C4446FFDECA5E55E0FC24F849" box="[793,945,1988,2013]" pageId="8" pageNumber="104" refId="ref14332" refString="Hoser, N. (2016) Praferenzen der Proctodrilus - Arten (Lumbricidae: Oligochaeta) auf einem Hang des unteren Kokel- Hochlandes in Rumanien. Opuscula Zoologica Budapest, 47, 143 - 149. https: // doi. org / 10.18348 / opzool. 2016.2.143" type="journal article">Höser (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="A443176D4447FFDFC9D052BDFF6AFF6C" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Hoghilag (RO), transect Tîrnava Mare floodplain. 46°14'N, 24°38'E. Floodplain centre. 10 ind. at
<quantity id="6304BA884447FFDFCC6A52B2FA09FF3B" box="[1325,1436,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="9" pageNumber="105" unit="cm" value="22.5" valueMax="35.0" valueMin="10.0">1035 cm</quantity>
depth, in h2 Lts. Clay skins present. In places below a layer containing
<taxonomicName id="63FC6CEE4447FFDFCD425298FB28FF40" authorityName="Zicsi" authorityYear="1985" box="[1029,1213,188,212]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCD425298FB28FF40" box="[1029,1213,188,212]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
(
<bibRefCitation id="C06D6A9C4447FFDFCD8C5299FF64FF6C" pageId="9" pageNumber="105" refId="ref14278" refString="Hoser, N. (2015) Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus - Verteilung, in den Auen der Tschernosem- und Parabraunerde-Gebiete beiderseits des Erzgebirges. Opuscula Zoologica Budapest, 46, 133 - 142. https: // doi. org / 10.18348 / opzool. 2015.2.133" type="journal article">Höser 2015: Table 3</bibRefCitation>
).
</paragraph>
<paragraph id="A443176D4447FFDFC9D05320FBF0FED4" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Săcămaş (RO), transect gorge. 45°54'N, 22°41'E. 6 ind. at
<quantity id="6304BA884447FFDFCA345321FC77FE88" box="[883,994,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="9" pageNumber="105" unit="cm" value="27.5" valueMax="35.0" valueMin="20.0">2035 cm</quantity>
depth, in rust-mottled h1 Ls3. Below a layer with
<taxonomicName id="63FC6CEE4447FFDFC81F530CFD85FED4" authorityName="Zicsi" authorityYear="1985" box="[344,528,296,320]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFC81F530CFD85FED4" box="[344,528,296,320]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
, which itself is below a layer with
<taxonomicName id="63FC6CEE4447FFDFCADF530CFB96FEAB" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[920,1027,296,320]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4447FFDFCADF530CFB96FEAB" box="[920,1027,296,320]" italics="true" pageId="9" pageNumber="105">P. antipai</emphasis>
</taxonomicName>
(
<figureCitation id="3CC70BE84447FFDFCD555302FBC6FED4" box="[1042,1107,295,320]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="9" pageNumber="105">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="A443176D4447FFDFC9D05368FEC5FE39" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05368FEC5FE39" ID-GBIF-Occurrence="1914265963" latitude="45.933334" location="Ilia" longLatPrecision="1127" longitude="22.65" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A75369FE90FEF1" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A75369FE90FEF1" box="[224,261,332,357]" latitude="45.933334" longLatPrecision="1127" longitude="22.65" name="Ilia" pageId="9" pageNumber="105" stateProvince="Mures">Ilia</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFC8F15368FE68FEF0" box="[438,509,333,356]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBCE5369FD7DFEF1" box="[649,744,332,357]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="45.933334">45°56'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCBB25369FCC4FEF0" box="[757,849,332,356]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="22.65">22°39'E</geoCoordinate>
. Two floodplain terraces close to river. Profile with bandlike layers. 51 ind. at
<quantity id="6304BA884447FFDFCB625355FD19FE13" box="[549,652,368,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" metricValueMax="4.0" metricValueMin="0.7" pageId="9" pageNumber="105" unit="cm" value="23.5" valueMax="40.0" valueMin="7.0">740 cm</quantity>
depth, in slightly stony h1/2 Ut above Su, preferentially at layer boundary.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D0539DFE16FE60" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D0539DFE16FE60" ID-GBIF-Occurrence="1914265946" latitude="45.983334" location="Tisa" longLatPrecision="1127" longitude="22.45" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A75392FE9AFE44" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A75392FE9AFE44" box="[224,271,439,464]" latitude="45.983334" longLatPrecision="1127" longitude="22.45" name="Tisa" pageId="9" pageNumber="105" stateProvince="Mures">Tisa</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFC8FE539DFD94FE5B" box="[441,513,440,463]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBCF5392FD72FE44" box="[648,743,439,464]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="45.983334">45°59'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCBB6539DFCDBFE44" box="[753,846,440,464]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="22.45">22°27'E</geoCoordinate>
. Exterior edge of floodplain terrace closest to river. 1 ind. in h0 Ls3.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D05025FAA6FDF4" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05025FAA6FDF4" ID-GBIF-Occurrence="1914265967" latitude="46.0" location="Pojoga" longLatPrecision="1127" longitude="22.35" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A753DAFEBBFD8C" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A753DAFEBBFD8C" box="[224,302,511,536]" latitude="46.0" longLatPrecision="1127" longitude="22.35" name="Pojoga" pageId="9" pageNumber="105" stateProvince="Mures">Pojoga</location>
(RO), two transects
<collectingRegion id="6638D98F4447FFDFCB635025FDF9FD83" box="[548,620,512,535]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBBD53DAFCCCFD8C" box="[762,857,511,536]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="46.0">46°00'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCA2F5025FC50FD8C" box="[872,965,512,536]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="22.35">22°21'E</geoCoordinate>
. Three floodplain terraces close to river. Profile with bandlike layers. 39 ind. at
<quantity id="6304BA884447FFDFCBE05001FC9FFDA8" box="[679,778,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="2.7" metricValueMin="0.1" pageId="9" pageNumber="105" unit="cm" value="14.0" valueMax="27.0" valueMin="1.0">127 cm</quantity>
depth, in h0/1 Ls3, concentrated at boundaries to annual sediment layers. On exterior edge of terrace closest to river
<taxonomicName id="63FC6CEE4447FFDFCA39506CFBA3FDF4" authorityName="Zicsi" authorityYear="1985" box="[894,1078,584,608]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCA39506CFBA3FDF4" box="[894,1078,584,608]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
in the overlying layer.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D05048FDD1FD33" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Vărădia de
<collectingRegion id="6638D98F4447FFDFC8205048FE3AFD10" box="[359,431,621,644]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
(RO), transect
<collectingRegion id="6638D98F4447FFDFCB225048FD39FD10" box="[613,684,621,644]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCA7C5049FC0EFD11" box="[827,923,620,645]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="45.983334">45°59'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCAEE5049FB93FD10" box="[937,1030,620,644]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="22.133333">22°08'E</geoCoordinate>
. Interior edge of floodplain terrace closest to river. 1 ind. in h1 Lts.
</paragraph>
<paragraph id="A443176D4447FFDFC9D05090FCE3FD7B" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05090FCE3FD7B" ID-GBIF-Occurrence="1914265945" latitude="46.016666" location="Nicolae Balcescu" longLatPrecision="1126" longitude="22.1" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A75091FE30FD59" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A75091FE30FD59" box="[224,421,692,717]" latitude="46.016666" longLatPrecision="1126" longitude="22.1" name="Nicolae Balcescu" pageId="9" pageNumber="105" stateProvince="Mures">Nicolae Bălcescu</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFCB105090FD0AFD58" box="[599,671,693,716]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCA6D5091FC1FFD59" box="[810,906,692,717]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="46.016666">46°01'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCAD05091FC66FD59" box="[919,1011,692,717]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="22.1">22°06'E</geoCoordinate>
. Floodplain terrace closest to river. 4 ind. in h1 Ls3, below humic horizon with
<taxonomicName id="63FC6CEE4447FFDFCBFF50FCFCE5FD64" authorityName="Zicsi" authorityYear="1985" box="[696,880,728,752]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCBFF50FCFCE5FD64" box="[696,880,728,752]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D050D8FC48FC14" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Bata (RO), four transects Mureş floodplain. 46°02'N, 22°03'E. Three floodplain terraces close to river. Profile with bandlike layers. 96 ind. at
<quantity id="6304BA884447FFDFCBE6513AFC96FCA3" box="[673,771,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" metricValueMax="3.5" metricValueMin="0.3" pageId="9" pageNumber="105" unit="cm" value="19.0" valueMax="35.0" valueMin="3.0">335 cm</quantity>
depth, in h0/2 Ls3, concentrated at boundary to Us or Ss; further from the river in Lts. Clay skins present. In places only at
<quantity id="6304BA884447FFDFCA9A5161FBB0FCC8" box="[989,1061,836,860]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="105" unit="cm" value="22.0">22 cm</quantity>
depth and below a layer with
<taxonomicName id="63FC6CEE4447FFDFCCC05160FEE8FC14" authorityName="Zicsi" authorityYear="1985" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCCC05160FEE8FC14" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
(
<figureCitation id="3CC70BE84447FFDFC8CC5142FE5BFC14" box="[395,462,871,896]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="9" pageNumber="105">Fig. 1</figureCitation>
and
<bibRefCitation id="C06D6A9C4447FFDFCB43514DFD1FFC14" box="[516,650,872,896]" pageId="9" pageNumber="105" refId="ref14015" refString="Hoser, N. (2003) Die Verteilung der Regenwurmer in der Aue des Mieresch (Siebenburgen, Banat, Rumanien). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 140, 99 - 116." type="journal article">Höser 2003</bibRefCitation>
: p. 104;
<bibRefCitation id="C06D6A9C4447FFDFCBAD514DFCFAFC14" box="[746,879,871,896]" pageId="9" pageNumber="105" refId="ref14278" refString="Hoser, N. (2015) Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus - Verteilung, in den Auen der Tschernosem- und Parabraunerde-Gebiete beiderseits des Erzgebirges. Opuscula Zoologica Budapest, 46, 133 - 142. https: // doi. org / 10.18348 / opzool. 2015.2.133" type="journal article">Höser 2015</bibRefCitation>
: Table 3).
</paragraph>
<paragraph id="A443176D4447FFDFC9D051A8FE41FB84" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Şoimoş (RO), transect Mureş floodplain. 46°06'N, 21°43'E. Two floodplain terraces close to river. Profile with bandlike layers. 16 ind. at
<quantity id="6304BA884447FFDFCB155195FD23FC53" box="[594,694,944,968]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="2.2" metricValueMin="0.4" pageId="9" pageNumber="105" unit="cm" value="13.0" valueMax="22.0" valueMin="4.0">422 cm</quantity>
depth, in h1 Lts and slightly to very gravelly h1 Ls3, mostly at boundary to Ls4. In places at
<quantity id="6304BA884447FFDFCB7F51F1FD18FC78" box="[568,653,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="9" pageNumber="105" unit="cm" value="7.5" valueMax="9.0" valueMin="6.0">69 cm</quantity>
depth and below a layer with
<taxonomicName id="63FC6CEE4447FFDFCAAB51F0FB33FC78" authorityName="Zicsi" authorityYear="1985" box="[1004,1190,980,1004]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCAAB51F0FB33FC78" box="[1004,1190,980,1004]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
(
<bibRefCitation id="C06D6A9C4447FFDFCDF151F1FAD5FC78" box="[1206,1344,980,1004]" pageId="9" pageNumber="105" refId="ref14015" refString="Hoser, N. (2003) Die Verteilung der Regenwurmer in der Aue des Mieresch (Siebenburgen, Banat, Rumanien). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 140, 99 - 116." type="journal article">Höser 2003</bibRefCitation>
: p. 104;
<bibRefCitation id="C06D6A9C4447FFDFC9A651DDFEF2FB84" box="[225,359,1015,1040]" pageId="9" pageNumber="105" refId="ref14278" refString="Hoser, N. (2015) Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus - Verteilung, in den Auen der Tschernosem- und Parabraunerde-Gebiete beiderseits des Erzgebirges. Opuscula Zoologica Budapest, 46, 133 - 142. https: // doi. org / 10.18348 / opzool. 2015.2.133" type="journal article">Höser 2015</bibRefCitation>
: Table 3).
</paragraph>
<paragraph id="A443176D4447FFDFC9D05638FD40FBCC" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05638FD40FBCC" ID-GBIF-Occurrence="1914265950" latitude="46.1" location="Lipova" longLatPrecision="1125" longitude="21.733334" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A75639FEBAFBA1" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A75639FEBAFBA1" box="[224,303,1052,1077]" latitude="46.1" longLatPrecision="1125" longitude="21.733334" name="Lipova" pageId="9" pageNumber="105" stateProvince="Mures">Lipova</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFC89A5638FDB1FBA0" box="[477,548,1053,1076]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBEA5639FC9EFBA1" box="[685,779,1052,1077]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="46.1">46°06'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCA505639FCE1FBA0" box="[791,884,1052,1076]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="21.733334">21°44'E</geoCoordinate>
. Close to river banks. Profile with several layers. 4 ind. at
<quantity id="6304BA884447FFDFC804561AFE03FBC3" box="[323,406,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="9" pageNumber="105" unit="cm" value="6.5" valueMax="8.0" valueMin="5.0">58 cm</quantity>
depth, in h2 Ls3, over sand.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D05640FDD9FB34" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05640FDD9FB34" ID-GBIF-Occurrence="1914265960" latitude="46.116665" location="Paulis" longLatPrecision="1125" longitude="21.566668" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<location id="A12341B64447FFDFC9A75641FEB1FBE9" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC9A75641FEB1FBE9" box="[224,292,1124,1149]" latitude="46.116665" longLatPrecision="1125" longitude="21.566668" name="Paulis" pageId="9" pageNumber="105" stateProvince="Mures">Păuliş</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFC8975640FD8DFBE8" box="[464,536,1125,1148]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBD85641FD6BFBE9" box="[671,766,1124,1149]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="46.116665">46°07'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCA4D5641FCF2FBE8" box="[778,871,1124,1148]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="21.566668">21°34'E</geoCoordinate>
. Interior edge of floodplain terrace closest to river. 1 ind. at
<quantity id="6304BA884447FFDFC80456ADFE1DFB34" box="[323,392,1160,1184]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="105" unit="cm" value="20.0">20 cm</quantity>
depth, in h1 Ls3.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D05688FE48FA98" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05688FE48FA98" ID-GBIF-Occurrence="1914265966" latitude="46.133335" location="Glogowatz" longLatPrecision="1125" longitude="21.4" municipality="Vladimirescu" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Mures">
<collectingMunicipality id="44278D174447FFDFC9A75689FEEDFB51" box="[224,376,1196,1221]" pageId="9" pageNumber="105">Vladimirescu</collectingMunicipality>
/
<location id="A12341B64447FFDFC8C75689FE6CFB51" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC8C75689FE6CFB51" box="[384,505,1196,1221]" latitude="46.133335" longLatPrecision="1125" longitude="21.4" municipality="Vladimirescu" name="Glogowatz" pageId="9" pageNumber="105" stateProvince="Mures">Glogowatz</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFCBEF5688FD7AFB50" box="[680,751,1197,1220]" country="Romania" name="Mures" pageId="9" pageNumber="105">Mureş</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCA3E5689FC4DFB51" box="[889,984,1196,1221]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="46.133335">46°08'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCAA35689FBD4FB50" box="[996,1089,1196,1220]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="21.4">21°24'E</geoCoordinate>
. Four floodplain terraces close to river. Profile with bandlike layers. 79 ind. at
<quantity id="6304BA884447FFDFCA4B56F5FCE5FB7C" box="[780,880,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" metricValueMax="3.3" metricValueMin="0.4" pageId="9" pageNumber="105" unit="cm" value="18.5" valueMax="33.0" valueMin="4.0">433 cm</quantity>
depth, in h0/2 Lts, Lu, Ls3. Clay skins present.
<bibRefCitation id="C06D6A9C4447FFDFC9A656D0FE4CFA98" box="[225,473,1268,1293]" pageId="9" pageNumber="105" refId="ref14015" refString="Hoser, N. (2003) Die Verteilung der Regenwurmer in der Aue des Mieresch (Siebenburgen, Banat, Rumanien). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 140, 99 - 116." type="journal article">Höser (2003: Table 3)</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D0573CFB40FAC0" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Şomărtin/Martinsberg (RO), transect slope on Pârâul Nou. 45°50'N, 24°38'E. Shallow angle on upper slope. 1 ind. at
<quantity id="6304BA884447FFDFC8045719FE1DFAC0" box="[323,392,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="105" unit="cm" value="14.0">14 cm</quantity>
depth, in h0 Lts of the Al/Bt-Horizon. Under a layer with
<taxonomicName id="63FC6CEE4447FFDFCD505718FB5BFAC0" authorityName="Zicsi" authorityYear="1985" box="[1047,1230,1340,1364]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4447FFDFCD505718FB5BFAC0" box="[1047,1230,1340,1364]" italics="true" pageId="9" pageNumber="105">P. opisthoductus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A443176D4447FFDFC9D05744FD08FA08" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
<materialsCitation id="14941D304447FFDFC9D05744FD08FA08" ID-GBIF-Occurrence="1914265951" latitude="45.783333" location="Kerz" longLatPrecision="1128" longitude="24.583334" municipality="Carta" pageId="9" pageNumber="105" specimenCount="1" stateProvince="Olt">
<collectingMunicipality id="44278D174447FFDFC9A75745FE88FAEC" box="[224,285,1376,1400]" pageId="9" pageNumber="105">Cârţa</collectingMunicipality>
/
<location id="A12341B64447FFDFC8625745FECEFAEC" LSID="urn:lsid:plazi:treatment:2C55A67B444BFFDAC9D057CBFCE4FD58:A12341B64447FFDFC8625745FECEFAEC" box="[293,347,1376,1400]" latitude="45.783333" longLatPrecision="1128" longitude="24.583334" municipality="Carta" name="Kerz" pageId="9" pageNumber="105" stateProvince="Olt">Kerz</location>
(RO), transect
<collectingRegion id="6638D98F4447FFDFCB4C577AFDBAFAEC" box="[523,559,1375,1400]" country="Romania" name="Olt" pageId="9" pageNumber="105">Olt</collectingRegion>
floodplain.
<geoCoordinate id="C1C871AA4447FFDFCBFD577AFC8CFAEC" box="[698,793,1375,1400]" direction="north" orientation="latitude" pageId="9" pageNumber="105" precision="925" value="45.783333">45°47'N</geoCoordinate>
,
<geoCoordinate id="C1C871AA4447FFDFCA62577AFC17FAEC" box="[805,898,1375,1400]" direction="east" orientation="longitude" pageId="9" pageNumber="105" precision="925" value="24.583334">24°35'E</geoCoordinate>
. Floodplain terrace closest to river. 2 ind. at
<quantity id="6304BA884447FFDFCCC75745FE96FA08" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="105" unit="cm" value="17.0">17 cm</quantity>
depth, in h1 Lts. Clay skins present.
</materialsCitation>
</paragraph>
<paragraph id="A443176D4447FFDFC9D0578CFCF7FA70" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Valea Stanciului (RO), transect Jiu floodplain. 43°58'N, 23°53'E. Second floodplain terrace close to river. 1 ind. at
<quantity id="6304BA884447FFDFC86957E9FEE6FA70" box="[302,371,1484,1508]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="105" unit="cm" value="18.0">18 cm</quantity>
depth, in h0/1 Lu, at layer boundary to Ls3.
</paragraph>
<paragraph id="A443176D4447FFDFC9D057D4FE21F9B8" blockId="9.[151,1437,151,1581]" pageId="9" pageNumber="105">
• Zăval (RO), transect Jiu floodplain. 43°50'N, 23°49'E. Floodplain terrace closest to river. 2 ind. at
<quantity id="6304BA884447FFDFCC1357D5FA09F99C" box="[1364,1436,1520,1544]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="105" unit="cm" value="30.0">30 cm</quantity>
depth, in h0/1 Ls3.
</paragraph>
<paragraph id="A443176D4447FFDFC9805479FD94F8D9" blockId="9.[151,1437,1627,2013]" pageId="9" pageNumber="105">
<emphasis id="9688CB7F4447FFDFC9805479FD2FF9E1" bold="true" box="[199,698,1627,1653]" pageId="9" pageNumber="105">
Habitat specifity of
<taxonomicName id="63FC6CEE4447FFDFC882547EFDC0F9E0" box="[453,597,1627,1652]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4447FFDFC882547EFDC0F9E0" bold="true" box="[453,597,1627,1652]" italics="true" pageId="9" pageNumber="105">Proctodrilus</emphasis>
</taxonomicName>
species.
</emphasis>
Our observations suggest that the occurrence of
<taxonomicName id="63FC6CEE4447FFDFCC485479FA0EF9E0" box="[1295,1435,1628,1652]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4447FFDFCC485479FA0EF9E0" box="[1295,1435,1628,1652]" italics="true" pageId="9" pageNumber="105">Proctodrilus</emphasis>
</taxonomicName>
species is not correlated with certain soil
<typeStatus id="7B47A9CF4447FFDFCB3B54A4FD22F90D" box="[636,695,1665,1689]" pageId="9" pageNumber="105">types</typeStatus>
but rather with the structure and the biological state of the soil profiles. Therefore their relation to soil horizons and layers will be considered in the following. The
<taxonomicName id="63FC6CEE4447FFDFCC485481FA0EF928" box="[1295,1435,1700,1724]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4447FFDFCC485481FA0EF928" box="[1295,1435,1700,1724]" italics="true" pageId="9" pageNumber="105">Proctodrilus</emphasis>
</taxonomicName>
species live mainly in sites/habitats that show the influence of sedimentation and erosion, e.g. the soils of floodplains and slopes. They organise themselves into occupying different parts of the soil profile according to the
<typeStatus id="7B47A9CF4447FFDFC9D05534FF52F8BD" box="[151,199,1809,1833]" pageId="9" pageNumber="105">type</typeStatus>
of soil preferred, shaped by factors such as gradient of hydromorphology, or particle size, or in the various layers created by sedimentation.
</paragraph>
</subSubSection>
<subSubSection id="ECE644E64447FFDCC980557CFE0BFB84" lastPageId="10" lastPageNumber="106" pageId="9" pageNumber="105" type="reference_group">
<paragraph id="A443176D4447FFDCC980557CFF6CFE44" blockId="9.[151,1437,1627,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="106" pageId="9" pageNumber="105">
<taxonomicName id="63FC6CEE4447FFDFC980557CFEA0F8FB" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[199,309,1880,1904]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="105" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4447FFDFC980557CFEA0F8FB" box="[199,309,1880,1904]" italics="true" pageId="9" pageNumber="105">P. antipai</emphasis>
</taxonomicName>
was exclusively recorded in floodplain soils: cohesive, clay-rich Ah-horizon (
<bibRefCitation id="C06D6A9C4447FFDFCD8E5572FADCF8E4" box="[1225,1353,1879,1904]" pageId="9" pageNumber="105" refId="ref15588" refString="Zicsi, A. (1959) Beitrag zur geographischen Verbreitung und Okologie von Allolobophora antipai (Michaelsen) 1891 (Oligochaeta). Annales Universitatis Scientiarum Budapestinensis de Rolando Eotvos nominatae, Sectio Biologia, 2, 283 - 292." type="journal article">Zicsi 1959</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4447FFDFCC1E557CFF46F801" pageId="9" pageNumber="105" refId="ref13864" refString="Hoser, N. (1986) Die Bindung zweier Unterarten von Allolobophora antipai (Lumbricidae) an Lossboden unterschiedlicher Genese. Pedobiologia, 29, 319 - 326." type="journal article">Höser 1986</bibRefCitation>
), and regularly above or at the upper boundaries of the oxidation horizons of gleys (Go, Gro, Ah-Go). Occasionally the species appears near the surface in wet gleys (Go-Ah). Thus it inhabits the region of fluctuating groundwater levels, including the upper boundary of the capillary water fringe. These
<typeStatus id="7B47A9CF4447FFDFCD2C55E0FB33F849" box="[1131,1190,1989,2013]" pageId="9" pageNumber="105">types</typeStatus>
of soil habitats occur mainly at floodplain edges, though also at the valley bottoms along streams.
<taxonomicName id="63FC6CEE4444FFDCCD4452BCFBFAFF3B" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1027,1135,152,176]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCCD4452BCFBFAFF3B" box="[1027,1135,152,176]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
avoids reduction horizons (Gor, Gr), being found above the upper boundary of clay-rich impervious horizons (
<collectingRegion id="6638D98F4444FFDCCD1C5299FBEEFF41" box="[1115,1147,188,213]" country="Serbia" name="Sid" pageId="10" pageNumber="106">Sd</collectingRegion>
) or of the B-horizons. In floodplains it occupies the exterior edges of the terraces closest to the valley slope, immediately at the edge of the floodplain or in floodplain channels and fossil meanders. Using the moisture classification of
<bibRefCitation id="C06D6A9C4444FFDCCD905321FF76FED4" pageId="10" pageNumber="106" refId="ref15332" refString="Schlichting, E., Blume, H. - P. &amp; Stahr, K. (1995) Bodenkundliches Praktikum. Blackwell Wissenschafts-Verlag, Berlin, Wien, 295 pp." type="book">
Schlichting
<emphasis id="9688CB7F4444FFDCCC265320FA09FE88" box="[1377,1436,260,284]" italics="true" pageId="10" pageNumber="106">et al.</emphasis>
(1995)
</bibRefCitation>
, the species lives in wet to moist habitats (pF 12), i.e., clearly separate from the other three species (see below). In summer their habitat has a closed capillary fringe at 38 dm below surface (= high to moderately high), and small to moderate oscillations of the ground water level. In contrast,
<taxonomicName id="63FC6CEE4444FFDCCA825354FBF9FE1C" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[965,1132,368,392]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCCA825354FBF9FE1C" box="[965,1132,368,392]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
was always found in damp to moist soils (pF 23) with a very low and strongly oscillating capillary fringe during summer (&gt; 15 dm below soil surface).
</paragraph>
<paragraph id="A443176D4444FFDCC98053F8FE2BFD11" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="106">
In the floodplains
<taxonomicName id="63FC6CEE4444FFDCC8D353F8FDD9FE60" authorityName="Zicsi" authorityYear="1985" box="[404,588,476,500]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4444FFDCC8D353F8FDD9FE60" box="[404,588,476,500]" italics="true" pageId="10" pageNumber="106">P. opisthoductus</emphasis>
</taxonomicName>
prefers the fine-grained, humus-rich, rather thin sedimentary layers in soils without gley horizon, and is present in the colluvial topsoils on slopes. It is commonly found in the matted vegetation roots (Aw) of the sediments close to the surface. In floodplains its preferred habitat corresponds roughly to that part of the valley bottom that is regularly inundated by floodwater, mostly excluding the interior edge of the terrace closest to the river.
</paragraph>
<paragraph id="A443176D4444FFDCC98050B5FAE1FC14" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="106">
Most
<taxonomicName id="63FC6CEE4444FFDCC84050B4FE38FD3C" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[263,429,656,680]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCC84050B4FE38FD3C" box="[263,429,656,680]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
individuals were found in areas of clay-poor and clay-skin-bearing horizons (Al, Bt). This species occurs regularly in the clearly layered soils that are well removed from groundwater and exist in floodplains in the vicinity of the river (Fluvisoliment-Bänder,
<bibRefCitation id="C06D6A9C4444FFDCCA3F50F2FBB2FD64" box="[888,1063,727,752]" pageId="10" pageNumber="106" refId="ref15303" refString="Schirmer, W. (1983) Die Talentwicklung an Main und Regnitz seit dem Hochwurm. Geologisches Jahrbuch, Reihe A, 71, 11 - 43." type="journal article">Schirmer 1983</bibRefCitation>
), or were near the river during floodplain creation (
<bibRefCitation id="C06D6A9C4444FFDCC83D50D9FD97FC80" box="[378,514,764,788]" pageId="10" pageNumber="106" refId="ref14083" refString="Hoser, N. (2008) Die Regenwurmer Proctodrilus tuberculatus (CERNOSVITOV, 1935) und Proctodrilus antipai (MICHAELSEN, 1891) als Indikatoren der fossilen Oberflachengliederung der Flussaue (Oligochaeta: Lumbricidae). Hercynia, Neue Folge, 41, 263 - 272." type="journal article">Höser 2008</bibRefCitation>
). Therefore
<taxonomicName id="63FC6CEE4444FFDCCBCD50D8FCA5FC80" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[650,816,764,788]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCCBCD50D8FCA5FC80" box="[650,816,764,788]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
mostly occurs at the interior edge of the terrace closest to the river, where coatings of clay (containing additional silt and humus) are rapidly formed in the sediment during periodic flooding (
<bibRefCitation id="C06D6A9C4444FFDCC82F5161FD91FCC8" box="[360,516,836,861]" pageId="10" pageNumber="106" refId="ref13755" refString="Gerrard, A. J. (1981) Soils and Landforms. An Integration of Geomorphology and Pedology. George Allen &amp; Unwin, London, 219 pp." type="book">Gerrard 1981</bibRefCitation>
). In the slope catena it prefers the Al-horizon on the upper slope, though it is also found in low-gradient slopes and in the inverse profile on the lower part and foot of slopes (
<bibRefCitation id="C06D6A9C4444FFDCCDDD514DFAB4FC14" box="[1178,1313,872,896]" pageId="10" pageNumber="106" refId="ref14220" refString="Hoser, N. (2012) Regenwurmer (Oligochaeta: Lumbricidae) in der Hangcatena. Hercynia, Neue Folge, 45, 193 - 208." type="journal article">Höser 2012</bibRefCitation>
,
<bibRefCitation id="C06D6A9C4444FFDCCC6C5142FAFDFC14" box="[1323,1384,871,896]" pageId="10" pageNumber="106" refId="ref14332" refString="Hoser, N. (2016) Praferenzen der Proctodrilus - Arten (Lumbricidae: Oligochaeta) auf einem Hang des unteren Kokel- Hochlandes in Rumanien. Opuscula Zoologica Budapest, 47, 143 - 149. https: // doi. org / 10.18348 / opzool. 2016.2.143" type="journal article">2016</bibRefCitation>
).
</paragraph>
<paragraph id="A443176D4444FFDCC98051A8FE0BFB84" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="106">
<taxonomicName id="63FC6CEE4444FFDCC98051A8FEB9FC30" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[199,300,908,932]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4444FFDCC98051A8FEB9FC30" box="[199,300,908,932]" italics="true" pageId="10" pageNumber="106">P. thaleri</emphasis>
</taxonomicName>
was found on slopes (Celei) as well as in the floodplain (Copşa Mică). According to the present finds,
<taxonomicName id="63FC6CEE4444FFDCC99A5194FE11FC5C" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[221,388,944,968]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCC99A5194FE11FC5C" box="[221,388,944,968]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4444FFDCC8FC5194FD8BFC5C" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[443,542,944,968]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4444FFDCC8FC5194FD8BFC5C" box="[443,542,944,968]" italics="true" pageId="10" pageNumber="106">P. thaleri</emphasis>
</taxonomicName>
live under geomorphologically similar site conditions, although in fact the two species are mutually exclusive. Therefore we hyothesize that the habitats of the two species differ fundamentally in the degree of lessivage.
</paragraph>
</subSubSection>
<subSubSection id="ECE644E64444FFDAC9805639FCE4FD58" lastPageId="12" lastPageNumber="108" pageId="10" pageNumber="106" type="description">
<paragraph id="A443176D4444FFDCC9805639FCB2F9C4" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="106">
<emphasis id="9688CB7F4444FFDCC9805639FCB5FBA1" bold="true" box="[199,800,1052,1077]" pageId="10" pageNumber="106">Separation of the species in the layered soil profile.</emphasis>
In some parts of the floodplains and slopes, assemblages of
<emphasis id="9688CB7F4444FFDCC9F05664FE7AFBCC" box="[183,495,1088,1112]" italics="true" pageId="10" pageNumber="106">
<taxonomicName id="63FC6CEE4444FFDCC9F05664FEB2FBC3" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[183,295,1088,1112]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">P. antipai</taxonomicName>
,
<taxonomicName id="63FC6CEE4444FFDCC8725664FE7AFBCC" authorityName="Zicsi" authorityYear="1985" box="[309,495,1088,1112]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">P. opisthoductus</taxonomicName>
</emphasis>
, and
<taxonomicName id="63FC6CEE4444FFDCCB765664FD4CFBCC" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[561,729,1088,1112]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCCB765664FD4CFBCC" box="[561,729,1088,1112]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
linked to distinct soil profile layers are conspicuous (
<figureCitation id="3CC70BE84444FFDCCC00561AFA1EFBCC" box="[1351,1419,1087,1112]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="10" pageNumber="106">Fig. 1</figureCitation>
). One profile (Săcămaş) contains these three species in the order shown above, from top to bottom, each in a different layer. There was no observation of any association between the species. Profiles were regularly found in which two species were living, each in a different layer. Seven cases from central
<collectingCountry id="DCEB57FD4444FFDCCD755689FB0FFB50" box="[1074,1178,1196,1220]" name="Germany" pageId="10" pageNumber="106">Germany</collectingCountry>
and northern
<collectingRegion id="6638D98F4444FFDCCC705689FA09FB51" box="[1335,1436,1196,1221]" country="Czech Republic" name="Jihocesky" pageId="10" pageNumber="106">
<collectingCountry id="DCEB57FD4444FFDCCC705689FA09FB51" box="[1335,1436,1196,1221]" name="Czech Republic" pageId="10" pageNumber="106">Bohemia</collectingCountry>
</collectingRegion>
are known in which the layer occupied by
<taxonomicName id="63FC6CEE4444FFDCCBC056F4FD61FB73" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[647,756,1232,1256]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCCBC056F4FD61FB73" box="[647,756,1232,1256]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
lies above that containing
<taxonomicName id="63FC6CEE4444FFDCCD6956F4FB42FB7C" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[1070,1239,1232,1256]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCCD6956F4FB42FB7C" box="[1070,1239,1232,1256]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
(Ostrau, Serbitz, Gaulis, Trebanz, Mlékojedy, two near Plottendorf). In one further profile the relation between the two species is reversed (Benndorf). In this instance the soil profile was of fossil levee deposits that probably were accumulated by a free meander in the course of its cutting through the floodplain edge (
<bibRefCitation id="C06D6A9C4444FFDCCAA55719FBFBFAC0" box="[994,1134,1340,1364]" pageId="10" pageNumber="106" refId="ref14083" refString="Hoser, N. (2008) Die Regenwurmer Proctodrilus tuberculatus (CERNOSVITOV, 1935) und Proctodrilus antipai (MICHAELSEN, 1891) als Indikatoren der fossilen Oberflachengliederung der Flussaue (Oligochaeta: Lumbricidae). Hercynia, Neue Folge, 41, 263 - 272." type="journal article">Höser 2008</bibRefCitation>
). In six cases in
<collectingCountry id="DCEB57FD4444FFDCCC705719FA09FAC1" box="[1335,1436,1340,1365]" name="Romania" pageId="10" pageNumber="106">Romania</collectingCountry>
(Hoghilag, Pojoga, Nicolae Bălcescu, Bata, Şoimoş, Şomărtin)
<taxonomicName id="63FC6CEE4444FFDCCA2E5744FBB6FAEC" authorityName="Zicsi" authorityYear="1985" box="[873,1059,1376,1400]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4444FFDCCA2E5744FBB6FAEC" box="[873,1059,1376,1400]" italics="true" pageId="10" pageNumber="106">P. opisthoductus</emphasis>
</taxonomicName>
occurs above the layer inhabited by
<taxonomicName id="63FC6CEE4444FFDCC9FA57A0FEF3FA08" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[189,358,1412,1436]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCC9FA57A0FEF3FA08" box="[189,358,1412,1436]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
; here the population of
<taxonomicName id="63FC6CEE4444FFDCCBC557A0FCA9FA08" authorityName="Zicsi" authorityYear="1985" box="[642,828,1412,1436]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4444FFDCCBC557A0FCA9FA08" box="[642,828,1412,1436]" italics="true" pageId="10" pageNumber="106">P. opisthoductus</emphasis>
</taxonomicName>
is occasionally concentrated at the lower end of the vegetation-root mat (Aw). The vertical order of species in the stratified soil profile (e.g. the profile in Săcămaş, see
<figureCitation id="3CC70BE84444FFDCC9D057E9FF4CFA70" box="[151,217,1484,1509]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="10" pageNumber="106">Fig. 1</figureCitation>
) demonstrates that
<taxonomicName id="63FC6CEE4444FFDCC8F857E8FDB9FA70" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[447,556,1485,1508]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCC8F857E8FDB9FA70" box="[447,556,1485,1508]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4444FFDCCB2F57E8FCB6FA70" authorityName="Zicsi" authorityYear="1985" box="[616,803,1484,1508]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4444FFDCCB2F57E8FCB6FA70" box="[616,803,1484,1508]" italics="true" pageId="10" pageNumber="106">P. opisthoductus</emphasis>
</taxonomicName>
occur in younger layers and that
<taxonomicName id="63FC6CEE4444FFDCCDEC57E8FA80FA70" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1195,1301,1485,1508]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCCDEC57E8FA80FA70" box="[1195,1301,1485,1508]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
prefers the sediments richest in organic matter. Here different and species-specific ressource requirements and physically induced habitat constraints become evident. Hence in this genus there is, in addition to morphological separation, a similarly clear ecological species separation (
<bibRefCitation id="C06D6A9C4444FFDCCBD5541DFC8EF9C4" box="[658,795,1592,1616]" pageId="10" pageNumber="106" refId="ref13894" refString="Hoser, N. (1997) Standortliche Bindung als Kriterium der Artentrennung bei der Regenwurm-Gattung Proctodrilus Zicsi, 1985. In: Dunger, W. &amp; Voigtlander, K. (Eds.), Bedeutung, Stand und aktuelle Entwicklung der Systematik von Bodentieren, Naturkundemuseum Gorlitz, Gorlitz, pp. 151 - 156." type="book chapter">Höser 1997</bibRefCitation>
).
</paragraph>
<paragraph id="A443176D4444FFDDC9805479FC8BFF6C" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,248]" lastPageId="11" lastPageNumber="107" pageId="10" pageNumber="106">
<emphasis id="9688CB7F4444FFDCC9805479FBAAF9E1" bold="true" box="[199,1087,1628,1653]" pageId="10" pageNumber="106">Preference for layer boundaries as a taxonomic characteristic of the genus</emphasis>
. The four
<taxonomicName id="63FC6CEE4444FFDCCDF05479FAD6F9E0" box="[1207,1347,1628,1652]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4444FFDCCDF05479FAD6F9E0" box="[1207,1347,1628,1652]" italics="true" pageId="10" pageNumber="106">Proctodrilus</emphasis>
</taxonomicName>
species prefer proximity to the layer boundaries in the soil profile, where they are met with on the side where the substrate particles are finer (
<figureCitation id="3CC70BE84444FFDCC8295481FE25F928" box="[366,432,1700,1725]" captionStart="FIGURE 1" captionStartId="11.[151,250,1397,1419]" captionTargetBox="[158,1428,328,1363]" captionTargetId="figure@11.[151,1436,301,1376]" captionTargetPageId="11" captionText="FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size." httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="10" pageNumber="106">Fig. 1</figureCitation>
). This is most striking in
<taxonomicName id="63FC6CEE4444FFDCCB9E5480FC14F928" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[729,897,1700,1724]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCCB9E5480FC14F928" box="[729,897,1700,1724]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
, and well seen in more than 13 out of 97 study sites. Such assemblages were also recorded for
<taxonomicName id="63FC6CEE4444FFDCCB8054ECFCA4F974" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[711,817,1736,1760]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4444FFDCCB8054ECFCA4F974" box="[711,817,1736,1760]" italics="true" pageId="10" pageNumber="106">P. thaleri</emphasis>
</taxonomicName>
(near Copşa Mică) and
<taxonomicName id="63FC6CEE4444FFDCCD1454ECFA9AF974" authorityName="Zicsi" authorityYear="1985" box="[1107,1295,1736,1760]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4444FFDCCD1454ECFA9AF974" box="[1107,1295,1736,1760]" italics="true" pageId="10" pageNumber="106">P. opisthoductus</emphasis>
</taxonomicName>
(near Valea Lungă, Spini, Dobra, Bata).
<taxonomicName id="63FC6CEE4444FFDCC8AB54C8FDC8F890" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[492,605,1773,1796]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCC8AB54C8FDC8F890" box="[492,605,1773,1796]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
appears in higher numbers not only at layer boundaries (Immelborn, Kotteritz, Deutsches Holz, Břežany nad Ohři) but also in thin water-conducting horizons (Sw). The four species mostly occur above fluctuating groundwater levels, where temporary minima of soil moisture occur, for example for
<taxonomicName id="63FC6CEE4444FFDCC986557CFEFCF8E4" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[193,361,1880,1904]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4444FFDCC986557CFEFCF8E4" box="[193,361,1880,1904]" italics="true" pageId="10" pageNumber="106">P. tuberculatus</emphasis>
</taxonomicName>
at the interior, generally clearly layered edges of floodplain terraces close to the river (
<bibRefCitation id="C06D6A9C4444FFDCCC10557DFEA6F800" pageId="10" pageNumber="106" refId="ref14015" refString="Hoser, N. (2003) Die Verteilung der Regenwurmer in der Aue des Mieresch (Siebenburgen, Banat, Rumanien). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 140, 99 - 116." type="journal article">Höser 2003: Table 3</bibRefCitation>
). In such locations the capillary water of the layer boundaries will be a sufficient water supply. This water forms in the lower part of the fine-grained substrate that overlies a coarser-grained one (“hängendes Kapillarwasser”:
<bibRefCitation id="C06D6A9C4444FFDCC81A55E1FDD8F848" box="[349,589,1988,2013]" pageId="10" pageNumber="106" refId="ref14770" refString="Muckenhausen, E. (1993) Die Bodenkunde und ihre geologischen, geomorphologischen, mineralogischen und petrologischen Grundlagen. DLG-Verlag, Frankfurt am Main, 579 pp., att. 24 pls." type="book">Mückenhausen 1993</bibRefCitation>
). In addition, the clay-rich habitat of
<taxonomicName id="63FC6CEE4444FFDCCD4255E0FBE6F848" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1029,1139,1989,2012]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="106" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4444FFDCCD4255E0FBE6F848" box="[1029,1139,1989,2012]" italics="true" pageId="10" pageNumber="106">P. antipai</emphasis>
</taxonomicName>
is mostly reached by the capillary limit of the groundwater. The four
<taxonomicName id="63FC6CEE4445FFDDCBC152BDFC87FF24" box="[646,786,152,176]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4445FFDDCBC152BDFC87FF24" box="[646,786,152,176]" italics="true" pageId="11" pageNumber="107">Proctodrilus</emphasis>
</taxonomicName>
species are adapted to temporary minima of soil moisture by their enteronephric excretory system (
<bibRefCitation id="C06D6A9C4445FFDDCB3D5299FD6FFF41" box="[634,762,188,213]" pageId="11" pageNumber="107" refId="ref15274" refString="Rosa, D. (1906) Sui nefridii con sbocco intestinale commune del l' Allolobophora antipae MICH. Archivio zoologico italiano, 3, 73 - 97." type="journal article">Rosa 1906</bibRefCitation>
,
<bibRefCitation id="C06D6A9C4445FFDDCA4E5299FC1DFF41" box="[777,904,188,213]" pageId="11" pageNumber="107" refId="ref15630" refString="Zicsi, A. (1985) Uber die Gattungen Helodrilus Hoffmeister, 1845 und Proctodrilus gen. n. (Oligochaeta: Lumbricidae). Acta Zoologica Hungarica, 31, 275 - 289." type="journal article">Zicsi 1985</bibRefCitation>
), by which they are able to retain water and reabsorb it through the intestine (
<bibRefCitation id="C06D6A9C4445FFDDCB4D52FAFD17FF6C" box="[522,642,223,248]" pageId="11" pageNumber="107" refId="ref13326" refString="Bahl, K. N. (1947) Excretion in the Oligochaeta. Biological Reviews of the Cambridge Philosophical Society, 22, 109 - 147. https: // doi. org / 10.1111 / j. 1469 - 185 X. 1947. tb 00325. x" type="journal article">Bahl 1947</bibRefCitation>
,
<bibRefCitation id="C06D6A9C4445FFDDCBCA52FAFC87FF6C" box="[653,786,223,248]" pageId="11" pageNumber="107" refId="ref15487" refString="Stolte, H. - A. (1969) Oligochaeta. 8. Lieferung. In: Stolte, H. - A. (Ed.), Dr. H. G. Bronns Klassen und Ordnungen des Tierreichs. Vierter Band, III. Abteilung, Annelides, 3. Buch, Akademische Verlagsgesellschaft Geest &amp; Portig, Leipzig, pp. 1287 - 1453." type="book chapter">Stolte 1969</bibRefCitation>
).
</paragraph>
<caption id="F08347E54445FFDDC9D05750FBA4FA5C" httpUri="https://zenodo.org/record/1447051/files/figure.png" pageId="11" pageNumber="107" startId="11.[151,250,1397,1419]" targetBox="[158,1428,328,1363]" targetPageId="11">
<paragraph id="A443176D4445FFDDC9D05750FBA4FA5C" blockId="11.[151,1436,1397,1480]" pageId="11" pageNumber="107">
<emphasis id="9688CB7F4445FFDDC9D05750FE86FA1E" bold="true" box="[151,275,1397,1419]" pageId="11" pageNumber="107">FIGURE 1.</emphasis>
Ecological species separation in the genus
<taxonomicName id="63FC6CEE4445FFDDCB8C5750FC14FA1F" authority=": Soil" authorityName="Soil" box="[715,897,1397,1419]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4445FFDDCB8C5750FCDFFA1F" box="[715,842,1397,1419]" italics="true" pageId="11" pageNumber="107">Proctodrilus</emphasis>
: Soil
</taxonomicName>
profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of
<taxonomicName id="63FC6CEE4445FFDDCC5A57B1FA09FA3E" box="[1309,1436,1428,1450]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4445FFDDCC5A57B1FA09FA3E" box="[1309,1436,1428,1450]" italics="true" pageId="11" pageNumber="107">Proctodrilus</emphasis>
</taxonomicName>
species in the soil profile: close to layer boundaries and in the layer with finer particle size.
</paragraph>
</caption>
<paragraph id="A443176D4445FFDDC98057D0FAE8F8BA" blockId="11.[151,1437,1525,2018]" pageId="11" pageNumber="107">
<emphasis id="9688CB7F4445FFDDC98057D0FDE9F99A" bold="true" box="[199,636,1525,1550]" pageId="11" pageNumber="107">Distributional range of the species.</emphasis>
The species records presented here lie within the currently known distributional ranges (
<bibRefCitation id="C06D6A9C4445FFDDC8C8543CFDBEF9A5" box="[399,555,1561,1586]" pageId="11" pageNumber="107" refId="ref13376" refString="Bouche, M. B. (1972) Lombriciens de France. Ecologie et systematique. Annales de Zoologie-Ecologie animale, Numero Hors- Serie, 72 (2), 1 - 671." type="journal article">Bouché 1972</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCB7F543CFCBEF9A5" box="[568,811,1561,1586]" pageId="11" pageNumber="107" refId="ref13458" refString="Csuzdi, Cs. &amp; Zicsi, A. (2003) Earthworms of Hungary (Annelida: Oligochaeta, Lumbricidae). Hungarian Natural History Museum, Budapest, 271 pp." type="book">Csuzdi &amp; Zicsi 2003</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCA7F543CFB9BF9A6" box="[824,1038,1561,1586]" pageId="11" pageNumber="107" refId="ref13491" refString="Csuzdi, Cs., Zicsi, A. &amp; Misirlioglu, M. (2006) An annotated checklist of the earthworm fauna of Turkey (Oligochaeta: Lumbricidae). Zootaxa, 1175, 1 - 29." type="journal article">
Csuzdi
<emphasis id="9688CB7F4445FFDDCAC8543FFC57F9A5" box="[911,962,1561,1585]" italics="true" pageId="11" pageNumber="107">et al</emphasis>
. 2006
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Lehmitz
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;
<bibRefCitation id="C06D6A9C4445FFDDCD90541BFA02F9C2" box="[1239,1431,1597,1622]" pageId="11" pageNumber="107" refId="ref14912" refString="Pop, V. V., Pop, A. A. &amp; Csuzdi, Cs. (2012) An annotated checklist of the Romanian earthworm fauna (Oligochaeta, Lumbricidae). In: Pavlicek, T., Cardet, P., Csuzdi, Cs., Le Bayon, R. - C. &amp; Gobat, J. - M. (Eds.), Advances in Earthworm Taxonomy V (Annelida: Oligochaeta), Kasparek Verlag, Heidelberg, pp. 59 - 70." type="book chapter">
Pop
<emphasis id="9688CB7F4445FFDDCC57541BFADBF9C1" box="[1296,1358,1597,1621]" italics="true" pageId="11" pageNumber="107">et al.</emphasis>
2012
</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDC9D05444FD93F9ED" box="[151,518,1633,1658]" pageId="11" pageNumber="107" refId="ref15365" refString="Stojanovic, M. &amp; Milutinovic, T. (2014) The earthworms (Oligochaeta: Lumbricidae) of the Pannonian region of Serbia, Vojvodina Province: Zoogeography and Diversity. North-Western Journal of Zoology, 10, 305 - 313." type="journal article">Stojanović &amp; Milutinović 2014</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCB515444FC89F9ED" box="[534,796,1633,1658]" pageId="11" pageNumber="107" refId="ref15409" refString="Stojanovic, M., Tsekova, R., Pesic, S., Milanovic, J. &amp; Milutinovic, T. (2013) Diversity and a biogeographical review of the earthworms (Oligochaeta: Lumbricidae) of the Balkan Mountains (Stara Planina Mountains) in Serbia and Bulgaria. Turkish Journal of Zoology, 37, 635 - 642. https: // doi. org / 10.3906 / zoo- 1301 - 33" type="journal article">
Stojanović
<emphasis id="9688CB7F4445FFDDCBDE5447FD5BF9ED" box="[665,718,1633,1657]" italics="true" pageId="11" pageNumber="107">et al</emphasis>
. 2013
</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCA6B5444FC7AF9ED" box="[812,1007,1633,1658]" pageId="11" pageNumber="107" refId="ref15557" refString="Valchovski, H. I. (2012) Checklist of earthworms (Oligochaeta: Lumbricidae) from Bulgaria - a review. Zootaxa, 3458, 86 - 102." type="journal article">Valchovski 2012</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCAB85444FBEAF9ED" box="[1023,1151,1633,1658]" pageId="11" pageNumber="107" refId="ref15666" refString="Zicsi, A. (1994) Die Regenwurmer Osterreichs (Oligochaeta: Lumbricidae) mit Bestimmungstabellen der Arten. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Osterreich, 131, 37 - 74." type="journal article">Zicsi 1994</bibRefCitation>
). These authors and our records make clear that among the four
<taxonomicName id="63FC6CEE4445FFDDCB1354A0FD75F909" box="[596,736,1669,1693]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4445FFDDCB1354A0FD75F909" box="[596,736,1669,1693]" italics="true" pageId="11" pageNumber="107">Proctodrilus</emphasis>
</taxonomicName>
species
<taxonomicName id="63FC6CEE4445FFDDCA7854A3FC70F909" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[831,997,1669,1693]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4445FFDDCA7854A3FC70F909" box="[831,997,1669,1693]" italics="true" pageId="11" pageNumber="107">P. tuberculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4445FFDDCD5C54A3FB13F909" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[1051,1158,1670,1693]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4445FFDDCD5C54A3FB13F909" box="[1051,1158,1670,1693]" italics="true" pageId="11" pageNumber="107">P. antipai</emphasis>
</taxonomicName>
occupy the widest range and extend farthest to the north (to northern central Europe), while
<taxonomicName id="63FC6CEE4445FFDDCAC7548FFBA2F955" authorityName="Zicsi" authorityYear="1985" box="[896,1079,1705,1729]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4445FFDDCAC7548FFBA2F955" box="[896,1079,1705,1729]" italics="true" pageId="11" pageNumber="107">P. opisthoductus</emphasis>
</taxonomicName>
does not reach the border of the German uplands.
<taxonomicName id="63FC6CEE4445FFDDC82454EBFE5DF971" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[355,456,1741,1765]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4445FFDDC82454EBFE5DF971" box="[355,456,1741,1765]" italics="true" pageId="11" pageNumber="107">P. thaleri</emphasis>
</taxonomicName>
has the narrowest distribution of a Carpathian endemic, as established by
<bibRefCitation id="C06D6A9C4445FFDDCC6B54EBFF70F89D" pageId="11" pageNumber="107" refId="ref14912" refString="Pop, V. V., Pop, A. A. &amp; Csuzdi, Cs. (2012) An annotated checklist of the Romanian earthworm fauna (Oligochaeta, Lumbricidae). In: Pavlicek, T., Cardet, P., Csuzdi, Cs., Le Bayon, R. - C. &amp; Gobat, J. - M. (Eds.), Advances in Earthworm Taxonomy V (Annelida: Oligochaeta), Kasparek Verlag, Heidelberg, pp. 59 - 70." type="book chapter">
Pop
<emphasis id="9688CB7F4445FFDDCC2654EBFA09F971" box="[1377,1436,1741,1765]" italics="true" pageId="11" pageNumber="107">et al.</emphasis>
(2012)
</bibRefCitation>
. As regards the zoogeographical categorisation in chorological
<typeStatus id="7B47A9CF4445FFDDCA9C54D7FB8CF89E" box="[987,1049,1778,1802]" pageId="11" pageNumber="107">types</typeStatus>
,
<taxonomicName id="63FC6CEE4445FFDDCD6D54D7FB42F89D" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[1066,1239,1777,1801]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4445FFDDCD6D54D7FB42F89D" box="[1066,1239,1777,1801]" italics="true" pageId="11" pageNumber="107">P. tuberculatus</emphasis>
</taxonomicName>
is a transaegian species while
<taxonomicName id="63FC6CEE4445FFDDC8735533FE0BF8B9" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[308,414,1814,1837]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4445FFDDC8735533FE0BF8B9" box="[308,414,1814,1837]" italics="true" pageId="11" pageNumber="107">P. antipai</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4445FFDDC8925533FD18F8B9" authorityName="Zicsi" authorityYear="1985" box="[469,653,1813,1837]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4445FFDDC8925533FD18F8B9" box="[469,653,1813,1837]" italics="true" pageId="11" pageNumber="107">P. opisthoductus</emphasis>
</taxonomicName>
are Central European species (
<bibRefCitation id="C06D6A9C4445FFDDCAAF5530FB22F8BA" box="[1000,1207,1813,1838]" pageId="11" pageNumber="107" refId="ref13531" refString="Csuzdi, Cs., Pop, V. V. &amp; Pop, A. A. (2011) The earthworm fauna of the Carpathian Basin with new records and description of three new species (Oligochaeta: Lumbricidae). Zoologischer Anzeiger, 250, 2 - 18. https: // doi. org / 10.1016 / j. jcz. 2010.10.001" type="journal article">
Csuzdi
<emphasis id="9688CB7F4445FFDDCD795533FBE3F8B9" box="[1086,1142,1813,1837]" italics="true" pageId="11" pageNumber="107">et al.</emphasis>
2011
</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCD855533FAE4F8BA" box="[1218,1393,1813,1838]" pageId="11" pageNumber="107" refId="ref14912" refString="Pop, V. V., Pop, A. A. &amp; Csuzdi, Cs. (2012) An annotated checklist of the Romanian earthworm fauna (Oligochaeta, Lumbricidae). In: Pavlicek, T., Cardet, P., Csuzdi, Cs., Le Bayon, R. - C. &amp; Gobat, J. - M. (Eds.), Advances in Earthworm Taxonomy V (Annelida: Oligochaeta), Kasparek Verlag, Heidelberg, pp. 59 - 70." type="book chapter">
Pop
<emphasis id="9688CB7F4445FFDDCDB25533FABBF8B9" box="[1269,1326,1813,1837]" italics="true" pageId="11" pageNumber="107">et al.</emphasis>
2012
</bibRefCitation>
).
</paragraph>
<paragraph id="A443176D4445FFDAC980551CFBF6FF41" blockId="11.[151,1437,1525,2018]" lastBlockId="12.[151,1437,151,717]" lastPageId="12" lastPageNumber="108" pageId="11" pageNumber="107">
The distribution center of the four
<taxonomicName id="63FC6CEE4445FFDDCB0D551CFD43F8C5" box="[586,726,1849,1873]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4445FFDDCB0D551CFD43F8C5" box="[586,726,1849,1873]" italics="true" pageId="11" pageNumber="107">Proctodrilus</emphasis>
</taxonomicName>
species is the European region of terrestrial soils influenced by lessivage (
<bibRefCitation id="C06D6A9C4445FFDDC84B557BFDAEF8E2" box="[268,571,1885,1910]" pageId="11" pageNumber="107" refId="ref13699" refString="Ganssen, R. &amp; Gracanin, Z. (1972) Bodengeographie mit besonderer Berucksichtigung der Boden Mitteleuropas. K. F. Koehler Verlag, Stuttgart, 325 pp." type="book">Ganssen &amp; Gračanin 1972</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4445FFDDCB00557BFCF2F8E2" box="[583,871,1885,1910]" pageId="11" pageNumber="107" refId="ref13731" refString="Ganssen, R. &amp; Hadrich, F. (1965) Atlas zur Bodenkunde. Bibliographisches Institut, Mannheim, 85 pp." type="book">Ganssen &amp; Hädrich 1965</bibRefCitation>
). Furthermore, floodplains also offer habitats with similar properties (
<bibRefCitation id="C06D6A9C4445FFDDC82B55A4FE65F80E" box="[364,496,1921,1946]" pageId="11" pageNumber="107" refId="ref14419" refString="Kopp, E. (1964) Zur Genese der Boden aus Hochflutlehm auf der Niederterrasse im Raume Bonn-Koln-Krefeld. Eiszeitalter und Gegenwart, 15, 81 - 91." type="journal article">Kopp 1964</bibRefCitation>
). The lessivage processes result in the leaching of clay, analogous to the process in floodplain soils where sedimentation and floodplain dynamics result in clay migration (
<bibRefCitation id="C06D6A9C4445FFDDCD9A5583FAFFF82A" box="[1245,1386,1957,1982]" pageId="11" pageNumber="107" refId="ref14419" refString="Kopp, E. (1964) Zur Genese der Boden aus Hochflutlehm auf der Niederterrasse im Raume Bonn-Koln-Krefeld. Eiszeitalter und Gegenwart, 15, 81 - 91." type="journal article">Kopp 1964</bibRefCitation>
).
<taxonomicName id="63FC6CEE4445FFDDCCC05583FEB7F875" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="107" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4445FFDDCCC05583FEB7F875" italics="true" pageId="11" pageNumber="107">P. tuberculatus</emphasis>
</taxonomicName>
(in Bt+Al) clearly prefers such clay-poor horizons of the stratified soil profiles on slopes and the interior edges of floodplain terraces, while
<taxonomicName id="63FC6CEE4442FFDACB3F52BCFD77FF3B" baseAuthorityName="Michaelsen" baseAuthorityYear="1891" box="[632,738,152,176]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="antipai">
<emphasis id="9688CB7F4442FFDACB3F52BCFD77FF3B" box="[632,738,152,176]" italics="true" pageId="12" pageNumber="108">P. antipai</emphasis>
</taxonomicName>
(in Ah-Sw) inhabits favourable bands in the sedimented strata of the clay-rich floodplain edge. In this way both species occupy a Europe-wide range.
</paragraph>
<paragraph id="A443176D4442FFDAC98052C5FCE4FD58" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="108">
In north- central Europe, where lessivage took place in several phases during the late glacial period and the Holocene (
<bibRefCitation id="C06D6A9C4442FFDAC8575321FE25FE89" box="[272,432,260,285]" pageId="12" pageNumber="108" refId="ref14513" refString="Kundler, P. (1961) Lessives (Parabraunerden, Fahlerden) aus Geschiebemergel der Wurm-Eiszeit im norddeutschen Tiefland. Zeitschrift fur Pflanzenernahrung, Dungung und Bodenkunde, 95, 97 - 110. https: // doi. org / 10.1002 / jpln. 19610950202" type="journal article">Kundler 1961</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4442FFDAC8FC5321FDFEFE88" box="[443,619,260,285]" pageId="12" pageNumber="108" refId="ref14656" refString="Lieberoth, I. (1982) Bodenkunde. VEB Deutscher Landwirtschaftsverlag, Berlin, 432 pp." type="book">Lieberoth 1982</bibRefCitation>
), the clay-leached horizons occurring in the terrestrial domain are mostly relicts (
<bibRefCitation id="C06D6A9C4442FFDAC9AB530DFEE8FED4" box="[236,381,296,320]" pageId="12" pageNumber="108" refId="ref15172" refString="Reuter, G. (2001) Durch holozane Klimaanderung induzierte disharmonische Bodentyp-Interferenzen im mittleren Europa. Erganzung III zu: Nova Acta Leopoldina, Neue Folge, 88 (Nr. 331), 1 - 5. [http: // www 1. uni-giessen. de / bodenkunde / akpp / artikel / reuter 01. php]" type="journal article">Reuter 2001</bibRefCitation>
;
<bibRefCitation id="C06D6A9C4442FFDAC8CE5302FD98FED4" box="[393,525,295,320]" pageId="12" pageNumber="108" refId="ref14483" refString="Kuhn, P. (2003) Spatglaziale und holozane Lessivegenese auf jungweichselzeitlichen Sedimenten Deutschlands. Greifswalder Geographische Arbeiten, 28, 1 - 167 + 1 - 37." type="journal article">Kühn 2003</bibRefCitation>
). Characteristics indicating recent lessivage that derive from the earlier stage of brown loam (terra fusca) are lacking there (
<bibRefCitation id="C06D6A9C4442FFDACBD45369FCA6FEF1" box="[659,819,332,357]" pageId="12" pageNumber="108" refId="ref15007" refString="Reuter, G. (1964 a) Vergleichende Untersuchungen an lessivierten Boden in verschiedenen Klimagebieten. Proceedings of the 8 th International Congress of Soil Science, Bucharest, 5, 723 - 732." type="journal article">Reuter 1964a</bibRefCitation>
).
<taxonomicName id="63FC6CEE4442FFDACA0E5368FC66FEF0" baseAuthorityName="Cernosvitov" baseAuthorityYear="1935" box="[841,1011,332,356]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="tuberculatus">
<emphasis id="9688CB7F4442FFDACA0E5368FC66FEF0" box="[841,1011,332,356]" italics="true" pageId="12" pageNumber="108">P. tuberculatus</emphasis>
</taxonomicName>
apparently prefers such soils, where lessivage is advanced, almost complete, or a relict of previous pedogenetic processes. Clearly recent lessivage is evident under warm climatic conditons, e.g. in southeast Europe (
<bibRefCitation id="C06D6A9C4442FFDACA3353B1FB85FE39" box="[884,1040,404,429]" pageId="12" pageNumber="108" refId="ref15043" refString="Reuter, G. (1964 b) Zur Mikromorphologie lessivierter Boden in verschiedenen Klimagebieten. In: Jongerius, A. (Ed.), Soil Micromorphology, Elsevier, Amsterdam, pp. 213 - 218." type="book chapter">Reuter 1964b</bibRefCitation>
,
<bibRefCitation id="C06D6A9C4442FFDACD5C53B1FBC2FE38" box="[1051,1111,404,428]" pageId="12" pageNumber="108" refId="ref15147" refString="Reuter, G. (1999) Statement zur Lessive-Problematik in Mitteleuropa. Mitteilungen der Deutschen Bodenkundlichen Gesellschaft, 91, 1065 - 1068." type="journal article">1999</bibRefCitation>
), where
<taxonomicName id="63FC6CEE4442FFDACDF253B0FAF8FE38" authorityName="Zicsi" authorityYear="1985" box="[1205,1389,404,428]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4442FFDACDF253B0FAF8FE38" box="[1205,1389,404,428]" italics="true" pageId="12" pageNumber="108">P. opisthoductus</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4442FFDAC9D0539CFF6BFE44" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[151,254,440,464]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4442FFDAC9D0539CFF6BFE44" box="[151,254,440,464]" italics="true" pageId="12" pageNumber="108">P. thaleri</emphasis>
</taxonomicName>
are found. Apparently
<taxonomicName id="63FC6CEE4442FFDACB43539CFD29FE44" authorityName="Zicsi" authorityYear="1985" box="[516,700,440,464]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="opisthoductus">
<emphasis id="9688CB7F4442FFDACB43539CFD29FE44" box="[516,700,440,464]" italics="true" pageId="12" pageNumber="108">P. opisthoductus</emphasis>
</taxonomicName>
and
<taxonomicName id="63FC6CEE4442FFDACBB4539CFCCFFE44" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[755,858,440,464]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4442FFDACBB4539CFCCFFE44" box="[755,858,440,464]" italics="true" pageId="12" pageNumber="108">P. thaleri</emphasis>
</taxonomicName>
are tied to horizons in different layers in which the drainage of the clay-complexes of a brown loam character (
<bibRefCitation id="C06D6A9C4442FFDACA1053F9FC69FE61" box="[855,1020,476,501]" pageId="12" pageNumber="108" refId="ref14449" refString="Kubiena, W. L. (1956) Zur Mikromorphologie, Systematik und Entwicklung der rezenten und fossilen Lossboden. Eiszeitalter und Gegenwart, 7, 102 - 112, 4 pls." type="journal article">Kubiena 1956</bibRefCitation>
) is still ongoing, in other words in horizons of recent lessivage in the warm-humid climate of southeast Europe. This explains the geographic location of their distribution areal and leads to the hypthesis that
<taxonomicName id="63FC6CEE4442FFDACA4B5000FCE7FDA8" authorityName="Hoser &amp; Zicsi" authorityYear="2009" box="[780,882,548,572]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="species" species="thaleri">
<emphasis id="9688CB7F4442FFDACA4B5000FCE7FDA8" box="[780,882,548,572]" italics="true" pageId="12" pageNumber="108">P. thaleri</emphasis>
</taxonomicName>
occurs only in those parts of Europe where—due to ongoing (recent) lessivage in soil profiles—active flow-plasma is present in highest concentrations. Pathways in the soil structure created by lessivage (
<bibRefCitation id="C06D6A9C4442FFDACB145049FD6DFD11" box="[595,760,620,645]" pageId="12" pageNumber="108" refId="ref14449" refString="Kubiena, W. L. (1956) Zur Mikromorphologie, Systematik und Entwicklung der rezenten und fossilen Lossboden. Eiszeitalter und Gegenwart, 7, 102 - 112, 4 pls." type="journal article">Kubiena 1956</bibRefCitation>
), and also by other similarly clay-filled inhomogeneities of lessived floodplain soils such as fractures of the soil aggregate apparently favour these four small
<taxonomicName id="63FC6CEE4442FFDACC4850B5FA0EFD3C" box="[1295,1435,656,680]" class="Clitellata" family="Lumbricidae" genus="Proctodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="12" pageNumber="108" phylum="Annelida" rank="genus">
<emphasis id="9688CB7F4442FFDACC4850B5FA0EFD3C" box="[1295,1435,656,680]" italics="true" pageId="12" pageNumber="108">Proctodrilus</emphasis>
</taxonomicName>
species in their movement through the soil column (
<bibRefCitation id="C06D6A9C4442FFDACB9B5091FCF0FD58" box="[732,869,692,716]" pageId="12" pageNumber="108" refId="ref14083" refString="Hoser, N. (2008) Die Regenwurmer Proctodrilus tuberculatus (CERNOSVITOV, 1935) und Proctodrilus antipai (MICHAELSEN, 1891) als Indikatoren der fossilen Oberflachengliederung der Flussaue (Oligochaeta: Lumbricidae). Hercynia, Neue Folge, 41, 263 - 272." type="journal article">Höser 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>