202 lines
26 KiB
XML
202 lines
26 KiB
XML
<document id="FF5032E13AD3D5A987B60D7789A89772" ID-CLB-Dataset="35149" ID-DOI="http://dx.doi.org/10.3897/zookeys.536.5982" ID-GBIF-Dataset="f3546db1-d96a-40d8-ac8c-fe806547f201" ID-PMC="PMC4714036" ID-Pensoft-Pub="1313-2970-536-1" ID-PubMed="26798233" ID-ZooBank="A8B92CDD0B8C4384AAC559648BB45AA5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-536-1" ModsDocOrigin="ZooKeys 536" ModsDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" checkinTime="1451243741663" checkinUser="pensoft" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2015" docId="20F714FE5AC070B197AD4980F1FEA4D3" docLanguage="en" docName="ZooKeys 536: 1-92" docOrigin="ZooKeys 536" docSource="http://dx.doi.org/10.3897/zookeys.536.5982" docTitle="Pterostichus (Circinatus) liciniformis Csiki 1930" docType="treatment" docVersion="7" lastPageNumber="38" masterDocId="FFEAFF976442467ED413FFC17531FF81" masterDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="36" updateTime="1731888672137" updateUser="admin">
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<mods:title id="D9D0E0A66FBE3B364D6CD6D310F708A8">The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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<mods:namePart id="2344B133579572DA7ED19DC6B6B37DF6">Shi, Hongliang</mods:namePart>
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<mods:namePart id="6DECDDDFFDB4ADAC3284086727FAA3D8">Liang, Hongbin</mods:namePart>
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<mods:date id="B2CCB52BD77A261CA559CD73759E3C66">2015</mods:date>
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<mods:number id="47A34FAD43DE68E6DF648233168373D9">536</mods:number>
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<mods:classification id="27060A7FB0F65E32CD2BA782FFD0032E">journal article</mods:classification>
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<mods:identifier id="7F78809F48EFA4FA4CD56EAC42ADDE9F" type="DOI">http://dx.doi.org/10.3897/zookeys.536.5982</mods:identifier>
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<treatment id="20F714FE5AC070B197AD4980F1FEA4D3" ID-GBIF-Taxon="152063176" LSID="urn:lsid:plazi:treatment:20F714FE5AC070B197AD4980F1FEA4D3" httpUri="http://treatment.plazi.org/id/20F714FE5AC070B197AD4980F1FEA4D3" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
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<subSubSection id="0FBC729696A50E04D00DFD1A022B9629" pageId="35" pageNumber="36" type="multiple">
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<paragraph id="4DAB038A165DDDE8DBECEEFED6D19E85" pageId="35" pageNumber="36">
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Taxon classification Animalia
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<taxonomicName id="F8AB3046238DD11B399A8B3E75687A1C" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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<taxonomicName id="9444C7FF36FC3993FBC914DB56CB0256" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0D47A260F7802F5238AE91D2A6F89429" pageId="35" pageNumber="36" type="nomenclature">
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<paragraph id="8670C946DDE3EFA27EED6EB948AD110D" pageId="35" pageNumber="36">
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<taxonomicName id="89FAD616DE635435EACE293B0B20B8AB" ID-CoL="4Q3HN" LSID="Chinese common name: 盘胸通缘步甲 (Pán Xīong Tōng Yuán Bù Jiă)" authority="Csiki, 1930" authorityName="Csiki" authorityYear="1930" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) liciniformis" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="liciniformis" subGenus="Circinatus">Pterostichus (Circinatus) liciniformis Csiki, 1930</taxonomicName>
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Figures 16, 42, 52, 64, 80, 102, 122
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</paragraph>
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</subSubSection>
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<subSubSection id="86C7958E6E5598B6866E360879D96187" pageId="36" pageNumber="37" type="reference_group">
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<paragraph id="EE6DFB7A55EBA0017B1805C0F9E339AB" pageId="36" pageNumber="37">
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<taxonomicName id="560E19209655E998CFE2B1E0B91BD212" authority="Csiki 1930: 669" authorityName="Csiki" authorityPageNumber="669" authorityYear="1930" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) liciniformis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="liciniformis" subGenus="Circinatus">
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<pageBreakToken id="156431892C5FBC19E97A0225A8FC8408" pageId="36" pageNumber="37" start="start">Pterostichus</pageBreakToken>
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(Circinatus) liciniformis
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<bibRefCitation id="B171DEEB42F8130C58C2DCB5DE125808" author="Csiki, E" editor="Junk, W" journalOrPublisher="W. Junk, Berlin" pageId="59" pageNumber="60" pagination="529 - 737" title="Coleopterorum catalogus. Volumen II. Carabidae II" year="1930">Csiki 1930</bibRefCitation>
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: 669
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</taxonomicName>
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, replacement name for
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<taxonomicName id="354FFF0E8CE00F70000BE0221E942831" authority="Fairmaire, 1888" authorityName="Fairmaire" authorityYear="1888" class="Insecta" family="Carabidae" genus="Steropus" kingdom="Animalia" lsidName="Steropus licinoides" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="licinoides">Steropus licinoides Fairmaire, 1888</taxonomicName>
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;
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<bibRefCitation id="1C7F22856480CD5D5502B1551F5ECD2B" author="Jedlicka, A" journalOrPublisher="Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="59" pageNumber="60" pagination="177 - 346" title="Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera-Carabidae)." volume="26" year="1962">
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<normalizedToken id="084131260B1EB0C5335E9DDECC597D14" originalValue="Jedlička">Jedlicka</normalizedToken>
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||
1962
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||
</bibRefCitation>
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||
: 298 (
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<taxonomicName id="A8358769684AACF3B4512E4C988A6079" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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(incertae sedis));
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<bibRefCitation id="C488563A84DE659A958AB664453761FE" author="Sciaky, R" journalOrPublisher="Annuario del Museo Civico di Storia Naturale di Brescia" pageId="60" pageNumber="61" pagination="217 - 231" title="Circinatus new subgenus and three new species of Pterostichus from China (ColeopteraCarabidae). Natura Bresciana." volume="30" year="1996">Sciaky 1996</bibRefCitation>
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: 224;
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<bibRefCitation id="83D5DA131FFCC5AC4FDF5917FD3BBEBF" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="Apollo Books, Denmark" pageId="59" pageNumber="60" pagination="469 - 521" title="Carabidae: tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1, Archostemata- Myxophaga- Adephaga" year="2003">Bousquet 2003</bibRefCitation>
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: 488;
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<bibRefCitation id="BC8A2B44AF5FE2F30739F3AFDD0431CD" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing, Germany" pageId="59" pageNumber="60" title="Systematic list of extant ground beetles of the world (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz 2005</bibRefCitation>
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: 280;
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<bibRefCitation id="7883311B85DA63249D43316BA4FF239B" author="Allegro, G" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona (Botanica Zoologia)" pageId="59" pageNumber="60" pagination="3 - 18" title="Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae)." volume="34" year="2010">Allegro and Sciaky 2010</bibRefCitation>
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: 7.
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</paragraph>
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<paragraph id="2710C72E968BD5BA8A69C99F311FCBA2" pageId="36" pageNumber="37">
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<taxonomicName id="50FA431AEF554F03B1837E8972400ABA" authority="Synonym" authorityName="Synonym" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) liciniformis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="liciniformis" subGenus="Circinatus">Pterostichus (Circinatus) liciniformis Synonym</taxonomicName>
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:
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<taxonomicName id="D1C34BEDB9BCA225AF7AA1A2D46CEB33" authority="Fairmaire 1888: 10" authorityName="Fairmaire" authorityPageNumber="10" authorityYear="1888" class="Insecta" family="Carabidae" genus="Steropus" kingdom="Animalia" lsidName="Pterostichus licinoides" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="licinoides">
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licinoides
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<bibRefCitation id="41FC433EE6A8E4F61DE323402BBA590B" author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="59" pageNumber="60" pagination="7 - 46" title="Coleopteres de l'interieur de la Chine (suite)." volume="32" year="1888">Fairmaire 1888</bibRefCitation>
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: 10
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</taxonomicName>
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, (Original:
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<taxonomicName id="96DDB2C11783E39B6CDCBBD3EE356B11" class="Insecta" family="Carabidae" genus="Steropus" kingdom="Animalia" lsidName="Steropus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Steropus</taxonomicName>
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; syntype deposited in MNHN?). Junior homonym of
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<taxonomicName id="0D59EE243A57067AE882C545E9949280" authority="Motschulsky, 1866: 261" authorityName="Motschulsky" authorityPageNumber="261" authorityYear="1866" class="Insecta" family="Carabidae" genus="Steropus" kingdom="Animalia" lsidName="Steropus licinoides" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="licinoides">Steropus licinoides Motschulsky, 1866: 261</taxonomicName>
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;
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<bibRefCitation id="5415C05A6516481A7DFD82530C8BA0EA" author="Tschitscherine, T" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="60" pageNumber="61" pagination="260 - 351" title="Materiaux pour servir a l'etude des feroniens. III." volume="30" year="1897">
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<normalizedToken id="79F6CCBD47938B8E8E84C7A5FFB0FA3F" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
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1897
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</bibRefCitation>
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: 294 (
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<taxonomicName id="491D80D26E85B9DEE9025EB2D5DEE91F" class="Insecta" family="Carabidae" genus="Feronia" kingdom="Animalia" lsidName="Feronia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Feronia</taxonomicName>
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(
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<taxonomicName id="8E5235AC84FBEC10D99F134F1B702679" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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)).
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</paragraph>
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</subSubSection>
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<subSubSection id="F9F3B27C27F082A7D4C9F175C62F93A9" pageId="36" pageNumber="37" type="type locality">
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<paragraph id="49A496C2CF632352D7DEDDA29D9F88C7" pageId="36" pageNumber="37">Type locality.</paragraph>
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<paragraph id="6D05F64E97AF78587A0406EC68AF9063" pageId="36" pageNumber="37">
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In the original literature there was no detailed locality indicated, but only
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<normalizedToken id="BE2586F04F2A844E358579549539C35D" originalValue="“Yunnan”">"Yunnan"</normalizedToken>
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was mentioned. According to our knowledge of the type material of the other
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<taxonomicName id="5211719CDF6C9A2B52344489D6F39704" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
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species described in the same paper (see
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<bibRefCitation id="FA0A1085F373033F920F7931C5E703D4" author="Shi, HL" journalOrPublisher="Zootaxa" pageId="60" pageNumber="61" pagination="101 - 135" title="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions." url="10.11646/zootaxa.3664.2.1" volume="3664" year="2013">Shi et al., 2013</bibRefCitation>
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: 118 for a detailed discussion) and of the further specimens of
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<taxonomicName id="2153DBDAC51BB1B89951DD4022296FC7" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus liciniformis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="liciniformis">Pterostichus liciniformis</taxonomicName>
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collected in recent years, we infer that the type locality of this species could be in northeast Dali in Yunnan Province.
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</paragraph>
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</subSubSection>
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<subSubSection id="6E287E702FCCCDC95A8488B305776E64" pageId="36" pageNumber="37" type="notes on types">
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<paragraph id="0FAC63727ACD89D93890613EA756124F" pageId="36" pageNumber="37">Notes on types.</paragraph>
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<paragraph id="B3215353FA4A8A095896492EF2B0B373" pageId="36" pageNumber="37">
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The syntype was in
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<normalizedToken id="EA1745C8110B0B64F9DEEBC0D7E3B114" originalValue="Fairmaire’s">Fairmaire's</normalizedToken>
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collection, and then should have been transferred to the collection of MNHN. But despite a careful examination of the
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<taxonomicName id="D6F2EBCFC9FDA0DC15151815951862EE" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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collection in MNHN, we failed to find the type.
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</paragraph>
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</subSubSection>
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<subSubSection id="20E5611441EF5596CADBBE969372E93B" pageId="36" pageNumber="37" type="type material">
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<paragraph id="B0E524DAF8C018CFD822CDADF7BDEC17" pageId="36" pageNumber="37">Non-type material examined</paragraph>
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<paragraph id="30F4E4CBBF8622A0D3CC795E5F9D78FE" pageId="36" pageNumber="37">
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(total 22 specimens). 1 male (CCCC), "China, Lijiang, Laojunshan, 3500 m, 2009.VI.12, ZHU Xiaoyu lgt."; 3 males, 1 female (IZAS), "China, Yunnan, Lijiang, Yulongxueshan, Yunshanping, in dead log, 2006.11.15, 3200 m, LIU Biao"; 2 males, "China, Yunnan, Lijiang, Yulongxueshan, Yunshanping, in dead log, 2006.11.15, 3200 m, ZHAO Yongshuang"; 2 males (IZAS), "China, Yunnan, Lijiang, Yulongxueshan, Yunshanping, in dead log, 2006.11.15, 3200 m, CHEN Jie"; 2 males (IZAS), "China, Yunnan, Lijiang, Yulongxueshan, Yunshanping, in dead log, 2007.03.14, 3205 m, LIU Biao"; 1 female (IZAS), "Yunnan, Lijiang, Yulongshan, 1979.VIII.10, LIU Guoqing, 2700-3300 m"; 1 male (IZAS), "China, Yunnan Prov., Lijiang county, Laojunshan, grassland,
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<geoCoordinate id="7F244BE343325A38B97BFD972601CA89" direction="north" orientation="latitude" precision="1" value="26.64341">N26.64341</geoCoordinate>
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<geoCoordinate id="EDF42D9DBD1A343BD125744F5B0EB188" direction="east" orientation="longitude" precision="1" value="99.76754">E99.76754</geoCoordinate>
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, 3510 m, 2007.8.19 day, Liang H.B. collector"; 2 males (IZAS), "Yunnan, Lijiang, Yulongxueshan Mt., Maoniuping, 3271 m, 2012.VI.2, pit fall trap, LIU Ye, SHI Hongliang lgt."; 1 male (IZAS), "Yunnan, Lijiang, Yulongxueshan Mt., Maoniuping, 3226, 2012.VI.1, LIU Ye, SHI Hongliang lgt."; 2 males, 1 female (IZAS), "China, Yunnan, Deqen Pref., Zhongdian County, Luoji, pass S. of Jiulong village, 3594 m,
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<geoCoordinate id="F11E669AACFF4F1FEE9E0D303234F991" direction="north" orientation="latitude" precision="1" value="27.67278">N27.67278</geoCoordinate>
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,
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<geoCoordinate id="C51C9784ECC35B95BCE3DFE7112BFE02" direction="east" orientation="longitude" precision="1" value="100.02623">E100.02623</geoCoordinate>
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, 2013.VIII.22, under dead log, SHI Hongliang, LIU Ye & LIU Yizhou lgt."; 1 female (IZAS), "Yunnan, N. of Zhongdian, 3446 m, 2000.VII.23-30, YU Xiaodong lgt."; 1 female (IZAS), "Yunnan, Deqin county, E. slope of Baimaxueshan, 3700 m, 1981.VIII.26, LIAO Subai lgt."; 1 male (NMPC), "China: Yunnan prov., 1.3-2.0, km S of Haba, 17.-20.VI.2007, Haba Xueshan Mts., 2830-, 3000 m, 27°22.1'
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<geoCoordinate id="01DAE75E8E175FD0E0B7E90480A68991" direction="north" orientation="latitude" precision="92" value="100.136665">N 100°08.2'</geoCoordinate>
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E, J.Hajek & J.Ruzicka leg"; "individually collected on soil surface and on plants and shrubs, sparse mixed forest (with dominant pinus); in/ near the brook".
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</paragraph>
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</subSubSection>
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<subSubSection id="C21487BFF67CDA13424E85944BF37867" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="2F80490372B12691355B352A3D8CC247" pageId="36" pageNumber="37">Diagnosis.</paragraph>
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<paragraph id="3CD5A42402AFA69E96C76B68F7AF36AB" pageId="36" pageNumber="37">Pronotum with single mid-lateral seta; posterior seta distant from hind angle, hind angle completely rounded; elytral shoulder angle completely rounded, without humeral tooth; male and female with similar faint linear elytral microsculpture; fifth tarsomeres glabrous beneath; males with terminal sternum depressed in middle.</paragraph>
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<paragraph id="76E4F65614FA6E137FDAEDD1440AC215" pageId="36" pageNumber="37">
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This species differs from all other
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<taxonomicName id="B0A4A4FFDF04B2B53A51AC84F3B5E915" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
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||
species by: body form relatively wide; elytral humeral angle completely rounded; elytral lateral expansion wide and reflexed.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="927CF90A6FE88EBDFC27134C5058F2D9" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="supplemental description">
|
||
<paragraph id="E8B764C3A3DBA1CE43C1A456BBE55638" pageId="36" pageNumber="37">Supplemental description.</paragraph>
|
||
<paragraph id="57BA7B165B3E289663FE621CEEA2EFEB" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||
Male sternum. Penultimate sternum without special structures; terminal sternum depressed in middle, depression occupying apical half of sternum, finely rugose inside depression, a faint vertical ridge present in middle of depression, frontal- lateral corners of depression forming two indistinct tubercles (Fig. 122). Endophallus (Fig. 42D, E, F) bent to ventral side across left side of aedeagus, major parts of endophallus located on ventral side of aedeagus (in lateral view); gonopore (gp) located at level near aedeagal base, pointing to ventral face, gonopore lobe (gpl) bent in apical direction. Five distinct lobes recognized: basal lobe (bl) close to base of apical lamella, small and rounded; ventral lobe (vl) with a long and strongly chitinized piece, rounded membranous lobe borne on ventral-apical part of chitinized piece; right lobe (rl) rounded and large, shallowly constricted in middle, decorated
|
||
<pageBreakToken id="DB2FDB562C32008911B6DB1154216288" pageId="37" pageNumber="38" start="start">with</pageBreakToken>
|
||
fine scales; left lobe I (lf-I) close to base of vl, apex enlarged; left lobe II (lf-II) narrower than lf-I, pointing to left side of aedeagus. Female genitalia. Spermatheca with seminal canal approx 3.5 times as long as receptaculum; receptaculum clavate (Fig. 64), gradually expanded to apex; seminal canal inserted at base of common oviduct, base of seminal canal sclerotized. Stylomere II with two ensiform setae at basal half of outer margin, and one near middle of inner margin; two short nematiform setae located in a furrow near apex. Female sternum VIII (Fig. 80B) with fine setae on posterior margin; posterior margin almost straight, slightly notched in middle; anterior region more weakly chitinized than posterior region, without denser pigmentation, deeply notched in middle; middle transparent region diamond-shaped, with two very narrow oblique extensions adjacent to anterior and posterior notches in middle. Female tergum VIII (Fig. 80A) very short, mainly semi-chitinized with sparse and irregular spots; lateral-anterior region with two chitinized patches; anterior margin not notched in middle, posterior margin evenly arcuate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9364596870458E996E910BE13B7B1D3F" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="822C38E4CD049993439F6FDABF41D622" pageId="37" pageNumber="38">Distribution.</paragraph>
|
||
<paragraph id="8075B20D4C85B790572941AEC7DA6600" pageId="37" pageNumber="38">This species is relatively widely distributed in northwest Yunnan. So far it has been recorded from several localities in Lijiang, Shangri-La (=Zhongdian), and Deqin (Map 1).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="018FDD94EAE53B7C12B1A984AD5F92D4" pageId="37" pageNumber="38" type="affinities">
|
||
<paragraph id="79276439BDB8CFAEEE2AD55F3DE595C9" pageId="37" pageNumber="38">Affinities.</paragraph>
|
||
<paragraph id="78FB8F3D1A6F2A9A556038D72D3C69D5" pageId="37" pageNumber="38">
|
||
As indicated by
|
||
<bibRefCitation id="DB3C96A160B28A8DFC99712A33232FE1" author="Allegro, G" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona (Botanica Zoologia)" pageId="59" pageNumber="60" pagination="3 - 18" title="Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae)." volume="34" year="2010">Allegro and Sciaky (2010</bibRefCitation>
|
||
: 7),
|
||
<taxonomicName id="8EA44A464DACE587D9668AA15CEE358A" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus liciniformis" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="liciniformis">Pterostichus liciniformis</taxonomicName>
|
||
appears to be isolated in the species group. This species could be close to the other two species from north Yunnan,
|
||
<taxonomicName id="B8FA958D1F556AC12F4144638225D4D1" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus wangjiani" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="wangjiani" status="sp. nov.">Pterostichus wangjiani</taxonomicName>
|
||
<taxonomicNameLabel id="D0ED0655BB12B65DCBFBA0F782E9A3A7" rank="species">sp. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="33D3AFB0514BE2FB42D55BBDC7A68E55" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus dimorphus" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="dimorphus" status="sp. nov.">Pterostichus dimorphus</taxonomicName>
|
||
<taxonomicNameLabel id="0CF59353E2A42758C97BCB99322EDD20" rank="species">sp. n.</taxonomicNameLabel>
|
||
Despite differences in the pronotal hind angle, elytral shoulder angle and microsculpture, these species are very similar in the depressed male terminal sternum and aedeagus. Nevertheless,
|
||
<taxonomicName id="81A887E9CE1BC0AB64727CDAF926E11F" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus liciniformis" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="liciniformis">Pterostichus liciniformis</taxonomicName>
|
||
could represent an early branch in the
|
||
<taxonomicName id="74E03D4008EA4AFA9BC97BE2447FEEFC" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="baenningeri">baenningeri</taxonomicName>
|
||
-group. A detailed discussion is provided below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41BE33EE4F2440F65F6EF8BAD616B02E" pageId="37" pageNumber="38" type="geographical variation">
|
||
<paragraph id="27786B6C7DF91D940F22EF22D4A21F6B" pageId="37" pageNumber="38">Geographical variation.</paragraph>
|
||
<paragraph id="DAFB7B1BB8ED33FA0F0F9E4E6190C403" pageId="37" pageNumber="38">
|
||
We studied males from two localities, Lijiang (Yulongxueshan Mt.;
|
||
<geoCoordinate id="5BAD06270CF0D5181ED9D55D9DA33F51" direction="north" orientation="latitude" precision="555" value="27.2">N27.20°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DC03504740EF9F7DBC6EB4CAB4268C4C" direction="east" orientation="longitude" precision="555" value="100.27">E100.27°</geoCoordinate>
|
||
) and Shangri-La (= Zhongdian; pass S. of Jiulong village;
|
||
<geoCoordinate id="E6CDEFDAB2B58C960EA11138486C6EA7" direction="north" orientation="latitude" precision="555" value="27.67">N27.67°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A19018BE3A3146586E73270513959F6D" direction="east" orientation="longitude" precision="555" value="100.03">E100.03°</geoCoordinate>
|
||
). The specimens from these two localities have no differences in the external or aedeagal (endophallus excluded) characters. However, for the endophallus, we found one difference: in a specimen from Lijiang, the apex of lf-II is distinctly enlarged in dorsal view; in the specimen from Shangri-La, lf-II is smaller than that from Lijiang, and the apex less enlarged in ventral view. (Figs 42, 52)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |