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<mods:title id="5ED55EF9D4F8A40AC7AEF51BCE830B66">A review of the genus Coccoglypta Pilsbry, 1895 (Gastropoda: Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="E00ABC42269AB7EBA286443216E6F252">Chen, Zhe-Yu</mods:namePart>
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<heading id="F36381EAFFF4257FFC7F200ABB44FAEB" box="[961,1307,1333,1360]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="597" reason="2">
Genus
<taxonomicName id="6F944D05FFF4257FFBB0200ABB44FAEB" authority="Pilsbry, 1895" authorityName="Pilsbry" authorityYear="1895" box="[1038,1307,1333,1360]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF4257FFBB0200ABAD8FAF4" box="[1038,1159,1333,1359]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
Pilsbry, 1895
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<paragraph id="A82B3686FFF4257FFC932053BAF1FA3F" blockId="4.[813,1456,1388,1457]" box="[813,1198,1388,1412]" pageId="4" pageNumber="597">
<taxonomicName id="6F944D05FFF4257FFC932053BAF5FA3F" authority="Pilsbry, 1895: 211" authorityName="Pilsbry" authorityPageNumber="211" authorityYear="1895" box="[813,1194,1388,1412]" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="subGenus" subGenus="Coccoglypta">
<emphasis id="9AE0EA94FFF4257FFC932053BD35FA3F" box="[813,874,1388,1412]" italics="true" pageId="4" pageNumber="597">Eulota</emphasis>
(
<emphasis id="9AE0EA94FFF4257FFCC62053BDBAFA3F" box="[888,997,1388,1412]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
) Pilsbry, 1895: 211
</taxonomicName>
.
</paragraph>
<paragraph id="A82B3686FFF4257FFC9320A6BBEFFA0A" blockId="4.[813,1456,1388,1457]" box="[813,1456,1433,1457]" pageId="4" pageNumber="597">
<treatmentCitationGroup id="888411A8FFF4257FFC9320A6BBEFFA0A" box="[813,1456,1433,1457]" pageId="4" pageNumber="597">
<taxonomicName id="6F944D05FFF4257FFC9320A6BDCCFA0A" authorityName="Pilsbry" authorityYear="1895" box="[813,915,1433,1457]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF4257FFC9320A6BDCCFA0A" box="[813,915,1433,1457]" italics="true" pageId="4" pageNumber="597">Coccoglypta</emphasis>
</taxonomicName>
<treatmentCitation id="29351097FFF4257FFC1920A5BA73FA0A" author="YEN T. - C." box="[935,1068,1433,1457]" page="153" pageId="4" pageNumber="597" year="1939">
<bibRefCitation id="CC054B77FFF4257FFC1920A5BA73FA0A" author="YEN T. - C." box="[935,1068,1433,1457]" pageId="4" pageNumber="597" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen 1939: 153</bibRefCitation>
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. —
<treatmentCitation id="29351097FFF4257FFBEF20A6BB0FFA0A" author="SCHILEYKO A. A." box="[1105,1360,1433,1457]" page="1678" pageId="4" pageNumber="597" year="2004">
<bibRefCitation id="CC054B77FFF4257FFBEF20A6BB0FFA0A" author="SCHILEYKO A. A." box="[1105,1360,1433,1457]" pageId="4" pageNumber="597" pagination="1627 - 1763" refId="ref8091" refString="SCHILEYKO A. A. 2004. - Treatise on Recent terrestrial pulmonate molluscs Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1763." type="journal article" year="2004">
Schileyko 2004:
<date id="DC2A1046FFF4257FFB5920A6BB0FFA0A" box="[1255,1360,1433,1457]" pageId="4" pageNumber="597" value="1678" valueMax="1679">1678-1679</date>
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, fig. 2164.
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<paragraph id="A82B3686FFF4257FFC9320F9BBFDFA64" blockId="4.[813,1442,1478,1503]" box="[813,1442,1478,1503]" pageId="4" pageNumber="597">
<typeStatus id="772F8824FFF4257FFC9320F9BD02FA60" box="[813,861,1478,1502]" pageId="4" pageNumber="597">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="6F944D05FFF4257FFC6920F9BB50FA65" authority="Heude, 1886" authorityName="Heude" authorityYear="1886" box="[983,1295,1478,1503]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="597" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF4257FFC6920F9BADAFA65" box="[983,1157,1478,1502]" italics="true" pageId="4" pageNumber="597">Helix pinchoniana</emphasis>
<bibRefCitation id="CC054B77FFF4257FFB3220F8BB50FA65" author="HEUDE R. P. M." box="[1164,1295,1478,1503]" pageId="4" pageNumber="597" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
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, by monotypy.
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<paragraph id="A82B3686FFF4257FFC9320C9BA52F851" blockId="4.[813,1456,1526,2026]" pageId="4" pageNumber="597">DIAGNOSIS. — Shell dextral, depressed-conical, with 5-6 slightly convex whorls; body whorl rounded or slightly keeled; colour yellowish to greenish and light brown; protoconch without notable sculpture, seemingly smooth, or slightly pitted or “hammered”; teleoconch with irregular, rough wrinkles and strong tubercles, occasionally with fine spiral grooves; sculpture on ventral surface sometimes weaker than on dorsal side; aperture rounded to suboval, strongly oblique to shell axis; peristome expanded but not reflected; parietal callus weak, only indicated by fine whitish calcareous layer; umbilicus open, moderately wide to rather narrow, shows all whorls. Penis cylindrical or flattened, moderately long, internally with fine longitudinal pilasters or reticulated sculpture; no epiphallus observed; vas deferens slender, retractor muscle attached on the junction of penis and vas deferens; vagina short, internally with widely-spaced, sometimes converging longitudinal folds; stylophore of moderate size, accessory sac of about same size and shape, each of two alveolar; mucus glands large, distinct or forming a single gland mass, entering accessory sac independently; spermathecal stalk extremely long, bursa oval, small.</paragraph>
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<caption id="FCEB660EFFF5257EFF3A2324BEAEF9FF" ID-DOI="http://doi.org/10.5281/zenodo.3726112" ID-Zenodo-Dep="3726112" httpUri="https://zenodo.org/record/3726112/files/figure.png" pageId="5" pageNumber="598" startId="5.[132,143,1563,1581]" targetBox="[132,1356,220,1516]" targetPageId="5">
<paragraph id="A82B3686FFF5257EFF3A2324BEAEF9FF" blockId="5.[132,1455,1563,1604]" pageId="5" pageNumber="598">FIG. 1. — Phylogenetic tree using Bayesian inference with mitochondrial 16S and CO1 genes. Numbers above or below branches indicate Bayesian posterior probabilities.</paragraph>
</caption>
<subSubSection id="E08E650DFFF5257EFF3A23ECBFFCF856" pageId="5" pageNumber="598" type="discussion">
<paragraph id="A82B3686FFF5257EFF3A23ECBEB7F957" blockId="5.[132,775,1747,2029]" box="[132,232,1747,1773]" pageId="5" pageNumber="598">REMARKS</paragraph>
<paragraph id="A82B3686FFF5257EFF3A23CBBFFCF856" blockId="5.[132,775,1747,2029]" pageId="5" pageNumber="598">
<taxonomicName id="6F944D05FFF5257EFF3A23CBBEA2F8B5" authorityName="Pilsbry" authorityYear="1895" box="[132,253,1780,1806]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF5257EFF3A23CBBEA2F8B5" box="[132,253,1780,1806]" italics="true" pageId="5" pageNumber="598">Coccoglypta</emphasis>
</taxonomicName>
was erected as a section of
<taxonomicName id="6F944D05FFF5257EFDA023CBBEE3F895" authority="W. Hartmann, 1841" authorityName="W. Hartmann" authorityYear="1841" class="Gastropoda" family="Bradybaenidae" genus="Eulota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF5257EFDA023CBBC3DF8B5" box="[542,610,1780,1806]" italics="true" pageId="5" pageNumber="598">Eulota</emphasis>
W. Hartmann, 1841
</taxonomicName>
for two species, namely “
<taxonomicName id="6F944D05FFF5257EFE6A222BBC92F895" authority="Heude" authorityName="Heude" box="[468,717,1812,1838]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="dimidiata">
<emphasis id="9AE0EA94FFF5257EFE6A222BBC23F895" box="[468,636,1812,1838]" italics="true" pageId="5" pageNumber="598">Helix dimidiata</emphasis>
Heude
</taxonomicName>
” and “
<taxonomicName id="6F944D05FFF5257EFF2E220CBFF4F8F6" authority="Heude" authorityName="Heude" authorityYear="1886" box="[144,427,1843,1869]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF5257EFF2E220CBF0BF8F6" box="[144,340,1843,1869]" italics="true" pageId="5" pageNumber="598">Helix pinchoniana</emphasis>
Heude
</taxonomicName>
”. Pilsbry (1895) designated the former species as the
<typeStatus id="772F8824FFF5257EFEDD226BBFCEF8D5" box="[355,401,1876,1902]" pageId="5" pageNumber="598">type</typeStatus>
species. Heude, however, has never described a species with the name dimidiata (see
<bibRefCitation id="CC054B77FFF5257EFD13224CBEE1F816" author="JOHNSON R. J." pageId="5" pageNumber="598" refId="ref7571" refString="JOHNSON R. J. 1973. - Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, 111 p. https: // doi. org / 10.5962 / bhl. title. 141074" type="book" year="1973">Johnson 1973</bibRefCitation>
).
<bibRefCitation id="CC054B77FFF5257EFF6922ABBF0AF815" author="YEN T. - C." box="[215,341,1939,1966]" pageId="5" pageNumber="598" refId="ref8670" refString="YEN T. - C. 1939. - Die Chinesischen Land-und Susswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt am Main, 234 p." type="book" year="1939">Yen (1939)</bibRefCitation>
and
<bibRefCitation id="CC054B77FFF5257EFE2F22ACBC46F815" author="ZILCH A." box="[401,537,1939,1966]" pageId="5" pageNumber="598" refId="ref8700" refString="ZILCH A. 1959 - 1960. - Handbuch der Paleozoologie, 6 (2) Euthyneura. Gebruder Borntraeger, Berlin-Nikolassee, 834 p." type="book" year="1959">
Zilch (
<date id="DC2A1046FFF5257EFE6022ACBC46F815" box="[478,537,1939,1966]" pageId="5" pageNumber="598" value="1959" valueMax="1960">1959</date>
</bibRefCitation>
-1960) corrected this mistake and clarified
<typeStatus id="772F8824FFF5257EFEC7228BBFF6F875" box="[377,425,1972,1998]" pageId="5" pageNumber="598">type</typeStatus>
species of
<taxonomicName id="6F944D05FFF5257EFD99228CBCFAF876" authorityName="Pilsbry" authorityYear="1895" box="[551,677,1971,1997]" class="Gastropoda" family="Bradybaenidae" genus="Coccoglypta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="genus">
<emphasis id="9AE0EA94FFF5257EFD99228CBCFAF876" box="[551,677,1971,1997]" italics="true" pageId="5" pageNumber="598">Coccoglypta</emphasis>
</taxonomicName>
as
<taxonomicName id="6F944D05FFF5257EFD72228CBFC1F856" authority="Heude, 1886" authorityName="Heude" authorityYear="1886" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="598" phylum="Mollusca" rank="species" species="pinchoniana">
<emphasis id="9AE0EA94FFF5257EFD72228CBF5AF856" italics="true" pageId="5" pageNumber="598">Helix pinchoniana</emphasis>
<bibRefCitation id="CC054B77FFF5257EFEB222ECBFC1F856" author="HEUDE R. P. M." box="[268,414,2003,2029]" pageId="5" pageNumber="598" pagination="208 - 215" refId="ref7469" refString="HEUDE R. P. M. 1886. - Diagnoses Molluscorum novorum, in Sinis collectorum. Journal de Conchyliologie 34: 208 - 215." type="journal article" year="1886">Heude, 1886</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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